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<document id="8FC71AD92D97DC0E309288AA1A642701" ID-CLB-Dataset="25319" ID-DOI="10.11646/zootaxa.4679.3.12" ID-GBIF-Dataset="44e80db5-8f5d-4bed-a189-baed3b1020c8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3471116" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1570099173655" checkinUser="plazi" docAuthor="Montreuil, Olivier &amp; Frolov, Andrey V." docDate="2019" docId="03B6CB587A46B969FF18F870A3F55E6F" docLanguage="en" docName="zootaxa.4679.3.12.pdf" docOrigin="Zootaxa 4679 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trigonochilus oedipus" docType="treatment" docVersion="7" lastPageNumber="598" masterDocId="FF8FB3207A46B96BFF8FFFD3A7795C0D" masterDocTitle="A review of the Malagasy species of the genus Trigonochilus Brenske (Coleoptera Scarabaeidae: Rutelinae: Adoretini) with description of a new species" masterLastPageNumber="600" masterPageNumber="596" pageNumber="596" updateTime="1698757505706" updateUser="ExternalLinkService">
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<mods:title id="3B8F8EAFB01E431D62E2F576C14BEF99">A review of the Malagasy species of the genus Trigonochilus Brenske (Coleoptera Scarabaeidae: Rutelinae: Adoretini) with description of a new species</mods:title>
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<mods:namePart id="9E184C6D203128DB64364FB706FAEA8E">Montreuil, Olivier</mods:namePart>
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<mods:namePart id="5760C80D7C8B2941C9148E847D6A8593">Frolov, Andrey V.</mods:namePart>
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<mods:title id="E1790E7D3A06D9E01DD314D134D378A9">Zootaxa</mods:title>
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<mods:date id="6349494E31AEC1F772A4C5B4FFD0F7A1">2019</mods:date>
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<mods:number id="CD6A06374FFFD9380702E118F9D4EFB3">2019-10-03</mods:number>
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<mods:number id="C94ED6A3DB8B6DF5416BD998BC673E49">4679</mods:number>
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<mods:identifier id="FBF52ACC4A361C2762EDA0622F742F2F" type="DOI">10.11646/zootaxa.4679.3.12</mods:identifier>
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<treatment id="03B6CB587A46B969FF18F870A3F55E6F" ID-GBIF-Taxon="159628562" LSID="urn:lsid:plazi:treatment:03B6CB587A46B969FF18F870A3F55E6F" httpUri="http://treatment.plazi.org/id/03B6CB587A46B969FF18F870A3F55E6F" lastPageId="2" lastPageNumber="598" pageId="0" pageNumber="596">
<subSubSection id="C30529C57A46B96BFF18F870A5F15BB0" box="[151,648,1955,1982]" pageId="0" pageNumber="596" type="nomenclature">
<paragraph id="8BA07A4E7A46B96BFF18F870A5F15BB0" blockId="0.[151,648,1955,2017]" box="[151,648,1955,1982]" pageId="0" pageNumber="596">
<heading id="D0E8CD227A46B96BFF18F870A5F15BB0" bold="true" box="[151,648,1955,1982]" fontSize="11" level="1" pageId="0" pageNumber="596" reason="1">
<taxonomicName id="4C1F01CD7A46B96BFF18F870A5F15BB0" ID-CoL="7D3YY" authority="(Fairmaire, 1903)" baseAuthorityName="Fairmaire" baseAuthorityYear="1903" box="[151,648,1955,1982]" class="Insecta" family="Scarabaeidae" genus="Trigonochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="596" phylum="Arthropoda" rank="species" species="oedipus">
<emphasis id="B96BA65C7A46B96BFF18F870A5F15BB0" bold="true" box="[151,648,1955,1982]" pageId="0" pageNumber="596">
<emphasis id="B96BA65C7A46B96BFF18F870A6DF5BB3" bold="true" box="[151,422,1955,1982]" italics="true" pageId="0" pageNumber="596">Trigonochilus oedipus</emphasis>
(
<bibRefCitation id="EF8E07BF7A46B96BFE3AF870A5F95BB0" author="Fairmaire, L." box="[437,640,1955,1981]" pageId="0" pageNumber="596" pagination="9 - 11" refId="ref2390" refString="Fairmaire, L. (1903) Description de Coleopteres nouveaux de Madagascar. Le Naturaliste, 25, 9 - 11." type="journal article" year="1903">Fairmaire, 1903</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30529C57A46B96BFF18F81AA6685BEC" box="[151,273,1993,2017]" pageId="0" pageNumber="596" type="description">
<paragraph id="8BA07A4E7A46B96BFF18F81AA6685BEC" blockId="0.[151,648,1955,2017]" box="[151,273,1993,2017]" pageId="0" pageNumber="596">
<figureCitation id="132466CB7A46B96BFF18F81AA7A35BEC" box="[151,218,1993,2017]" captionStart="FIGURES 1" captionStartId="1.[151,264,1895,1920]" captionTargetBox="[187,1410,347,1860]" captionTargetId="figure@1.[178,1410,336,1868]" captionTargetPageId="1" captionText="FIGURES 1A–F. Trigonochilus oedipus (Fairmaire, 1903) (A–C), T. macrotarsus Montreuil &amp; Frolov, new species (D–F) and locality map (G). A, male, dorsal view; B, aedeagus, lateral and dorsal view; C, female, dorsal view; D, male, holotype, dorsal view; E, aedeagus, lateral and dosal view; F, female, paratype, dorsal view; G, locality map (black circle – T. oedipus, white circle – T. macrotarsus). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3471118" httpUri="https://zenodo.org/record/3471118/files/figure.png" pageId="0" pageNumber="596">Figs. 1</figureCitation>
A–E
</paragraph>
</subSubSection>
<subSubSection id="C30529C57A47B96AFF18FF4AA5DF5CC2" pageId="1" pageNumber="597" type="reference_group">
<paragraph id="8BA07A4E7A47B96AFF18FF4AA6805CBC" blockId="1.[151,678,152,208]" box="[151,505,152,177]" pageId="1" pageNumber="597">
<treatmentCitationGroup id="AB0F5D607A47B96AFF18FF4AA6805CBC" box="[151,505,152,177]" pageId="1" pageNumber="597">
<taxonomicName id="4C1F01CD7A47B96AFF18FF4AA68F5CBC" authority="Fairmaire, 1903: 9" authorityName="Fairmaire" authorityPageNumber="9" authorityYear="1903" box="[151,502,152,177]" class="Insecta" family="Rutelidae" genus="Adoretus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="597" phylum="Arthropoda" rank="species" species="oedipus">
<emphasis id="B96BA65C7A47B96AFF18FF4AA6455CBC" box="[151,316,153,177]" italics="true" pageId="1" pageNumber="597">Adoretus oedipus</emphasis>
<treatmentCitation id="0ABE5C5F7A47B96AFECDFF4BA68F5CBC" author="Fairmaire, L." box="[322,502,152,177]" page="9" pageId="1" pageNumber="597" year="1903">
<bibRefCitation id="EF8E07BF7A47B96AFECDFF4BA68F5CBC" author="Fairmaire, L." box="[322,502,152,177]" pageId="1" pageNumber="597" pagination="9 - 11" refId="ref2390" refString="Fairmaire, L. (1903) Description de Coleopteres nouveaux de Madagascar. Le Naturaliste, 25, 9 - 11." type="journal article" year="1903">Fairmaire, 1903: 9</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA07A4E7A47B96AFF18FF64A5DF5CC2" blockId="1.[151,678,152,208]" box="[151,678,183,208]" pageId="1" pageNumber="597">
<treatmentCitationGroup id="AB0F5D607A47B96AFF18FF64A5DF5CC2" box="[151,678,183,208]" pageId="1" pageNumber="597">
<taxonomicName id="4C1F01CD7A47B96AFF18FF64A5DB5CC2" ID-CoL="7D3YY" authority=": Ohaus, 1912: 416" authorityName="Ohaus" authorityPageNumber="416" authorityYear="1912" box="[151,674,183,208]" class="Insecta" family="Scarabaeidae" genus="Trigonochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="597" phylum="Arthropoda" rank="species" species="oedipus">
<emphasis id="B96BA65C7A47B96AFF18FF64A6125CC2" box="[151,363,183,207]" italics="true" pageId="1" pageNumber="597">Trigonochilus oedipus</emphasis>
:
<treatmentCitation id="0ABE5C5F7A47B96AFEF6FF64A5565CDD" author="Ohaus, F." box="[377,559,183,208]" page="416" pageId="1" pageNumber="597" year="1912">
<bibRefCitation id="EF8E07BF7A47B96AFEF6FF64A5565CDD" author="Ohaus, F." box="[377,559,183,208]" pageId="1" pageNumber="597" pagination="141 - 156" refId="ref2807" refString="Ohaus, F. (1912) Revision der Adoretini (Col. lamell. Rutelin.). Deutsche Entomologische Zeitschrift, 1912, 141 - 156. https: // doi. org / 10.1002 / mmnd. 48019120203" type="journal article" year="1912">Ohaus, 1912: 416</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0ABE5C5F7A47B96AFDB6FF64A5DB5CC2" author="Ohaus, F." box="[569,674,183,207]" page="197" pageId="1" pageNumber="597" year="1918">
<bibRefCitation id="EF8E07BF7A47B96AFDB6FF64A5DB5CC2" author="Ohaus, F." box="[569,674,183,207]" pageId="1" pageNumber="597" pagination="1 - 241" refId="ref2848" refString="Ohaus, F. (1918) Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum catalogus. Vol. XX. Pars 66. W. Junk, Berlin, pp. 1 - 241." type="book chapter" year="1918">1918: 197</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30529C57A47B969FF18FED3A41F5E33" lastPageId="2" lastPageNumber="598" pageId="1" pageNumber="597" type="materials_examined">
<paragraph id="8BA07A4E7A47B96AFF18FED3A68C5D31" blockId="1.[151,1437,255,316]" pageId="1" pageNumber="597">
<emphasis id="B96BA65C7A47B96AFF18FED3A6D05D1A" bold="true" box="[151,425,255,280]" pageId="1" pageNumber="597">Type material examined</emphasis>
.
<materialsCitation id="3B7770137A47B96AFE3BFF2CA68C5D31" ID-GBIF-Occurrence="2423812411" collectionCode="MNHN" collectorName="Coll. Leon Fairmaire" country="Madagascar" location="Madagascar" pageId="1" pageNumber="597" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54A4C4EC7A47B96AFE3BFF2CA56E5D1A" box="[436,535,255,279]" pageId="1" pageNumber="597" type="holotype">Holotype</typeStatus>
<specimenCount id="9D19B1C77A47B96AFD92FED3A5485D1A" box="[541,561,256,279]" pageId="1" pageNumber="597" type="male">♂</specimenCount>
(
<figureCitation id="132466CB7A47B96AFDB0FF2CA5EB5D15" box="[575,658,255,280]" captionStart="FIGURES 1" captionStartId="1.[151,264,1895,1920]" captionTargetBox="[187,1410,347,1860]" captionTargetId="figure@1.[178,1410,336,1868]" captionTargetPageId="1" captionText="FIGURES 1A–F. Trigonochilus oedipus (Fairmaire, 1903) (A–C), T. macrotarsus Montreuil &amp; Frolov, new species (D–F) and locality map (G). A, male, dorsal view; B, aedeagus, lateral and dorsal view; C, female, dorsal view; D, male, holotype, dorsal view; E, aedeagus, lateral and dosal view; F, female, paratype, dorsal view; G, locality map (black circle – T. oedipus, white circle – T. macrotarsus). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3471118" httpUri="https://zenodo.org/record/3471118/files/figure.png" pageId="1" pageNumber="597">Fig. 1D</figureCitation>
), “
<taxonomicName id="4C1F01CD7A47B96AFD20FF2CA4125D1A" authorityName="Fairmaire" authorityYear="1903" box="[687,875,255,279]" class="Insecta" family="Rutelidae" genus="Adoretus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="597" phylum="Arthropoda" rank="species" species="oedipus">Adoretus oedipus</taxonomicName>
m. Madag / TYPE / Museum Paris
<collectingCountry id="F3083ADE7A47B96AFB6EFF2CA21B5D1A" box="[1249,1378,255,279]" name="Madagascar" pageId="1" pageNumber="597">Madagascar</collectingCountry>
<collectorName id="26EA1F987A47B96AFAE7FF2CA6445D36" pageId="1" pageNumber="597">Coll. Leon Fairmaire</collectorName>
1906” (
<collectionCode id="ED0EE28B7A47B96AFE1DFEF7A6935D31" box="[402,490,292,316]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="597">MNHN</collectionCode>
).
</materialsCitation>
</paragraph>
<caption id="DF602AC67A47B96AFF18F8B4A61D5BE6" ID-DOI="http://doi.org/10.5281/zenodo.3471118" ID-Zenodo-Dep="3471118" httpUri="https://zenodo.org/record/3471118/files/figure.png" pageId="1" pageNumber="597" startId="1.[151,264,1895,1920]" targetBox="[187,1410,347,1860]" targetPageId="1">
<paragraph id="8BA07A4E7A47B96AFF18F8B4A61D5BE6" blockId="1.[151,1437,1895,2028]" pageId="1" pageNumber="597">
<emphasis id="B96BA65C7A47B96AFF18F8B4A6325B8D" bold="true" box="[151,331,1895,1920]" pageId="1" pageNumber="597">FIGURES 1A–F.</emphasis>
<taxonomicName id="4C1F01CD7A47B96AFEDEF8BBA5AB5B8D" authority="(Fairmaire, 1903)" authorityName="Fairmaire" authorityYear="1903" baseAuthorityName="Fairmaire" baseAuthorityYear="1903" box="[337,722,1895,1920]" class="Insecta" family="Scarabaeidae" genus="Trigonochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="597" phylum="Arthropoda" rank="species" species="oedipus">
<emphasis id="B96BA65C7A47B96AFEDEF8BBA55C5B8D" box="[337,549,1896,1920]" italics="true" pageId="1" pageNumber="597">Trigonochilus oedipus</emphasis>
(
<bibRefCitation id="EF8E07BF7A47B96AFDBFF8B4A5B25B8D" author="Fairmaire, L." box="[560,715,1895,1920]" pageId="1" pageNumber="597" pagination="9 - 11" refId="ref2390" refString="Fairmaire, L. (1903) Description de Coleopteres nouveaux de Madagascar. Le Naturaliste, 25, 9 - 11." type="journal article" year="1903">Fairmaire, 1903</bibRefCitation>
)
</taxonomicName>
(A–C),
<taxonomicName id="4C1F01CD7A47B96AFC91F8BBA3145B8D" authority="Montreuil &amp; Frolov" authorityName="Montreuil &amp; Frolov" authorityYear="2019" box="[798,1133,1895,1920]" class="Insecta" family="Scarabaeidae" genus="Trigonochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="597" phylum="Arthropoda" rank="species" species="macrotarsus" status="sp. nov.">
<emphasis id="B96BA65C7A47B96AFC91F8BBA4D55B72" box="[798,940,1896,1919]" italics="true" pageId="1" pageNumber="597">T. macrotarsus</emphasis>
Montreuil &amp; Frolov
</taxonomicName>
,
<taxonomicNameLabel id="A2581B277A47B96AFBF9F8BBA3915B8D" box="[1142,1256,1895,1920]" pageId="1" pageNumber="597" rank="species">new species</taxonomicNameLabel>
(D–F) and locality map (G). A, male, dorsal view; B, aedeagus, lateral and dorsal view; C, female, dorsal view; D, male, holotype, dorsal view; E, aedeagus, lateral and dosal view; F, female, paratype, dorsal view; G, locality map (black circle –
<taxonomicName id="4C1F01CD7A47B96AFB8AF863A31E5BC5" baseAuthorityName="Fairmaire" baseAuthorityYear="1903" box="[1029,1127,1968,1992]" class="Insecta" family="Scarabaeidae" genus="Trigonochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="597" phylum="Arthropoda" rank="species" species="oedipus">
<emphasis id="B96BA65C7A47B96AFB8AF863A31E5BC5" box="[1029,1127,1968,1992]" italics="true" pageId="1" pageNumber="597">T. oedipus</emphasis>
</taxonomicName>
, white circle –
<taxonomicName id="4C1F01CD7A47B96AFA8EF863A2F75BCA" authorityName="Montreuil &amp; Frolov" authorityYear="2019" box="[1281,1422,1968,1991]" class="Insecta" family="Scarabaeidae" genus="Trigonochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="597" phylum="Arthropoda" rank="species" species="macrotarsus">
<emphasis id="B96BA65C7A47B96AFA8EF863A2F75BCA" box="[1281,1422,1968,1991]" italics="true" pageId="1" pageNumber="597">T. macrotarsus</emphasis>
</taxonomicName>
). Scale bar = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BA07A4E7A44B969FF48FF4AA32F5CDB" blockId="2.[151,1437,153,610]" pageId="2" pageNumber="598">
<emphasis id="B96BA65C7A44B969FF48FF4AA6C95CBC" bold="true" box="[199,432,153,177]" pageId="2" pageNumber="598">Additional material.</emphasis>
<materialsCitation id="3B7770137A44B969FE37FF49A4C95CBF" ID-GBIF-Occurrence="2423812417" box="[440,944,154,178]" collectionCode="MNHN" location="Diego-Suarez" pageId="2" pageNumber="598" specimenCount="1" specimenCount-male="1" stateProvince="Antsiranana">
<specimenCount id="9D19B1C77A44B969FE37FF49A69D5CBC" box="[440,484,154,178]" pageId="2" pageNumber="598" type="male">1 ♂</specimenCount>
, “
<location id="8EC02C957A44B969FE74FF49A5E85CBF" LSID="urn:lsid:plazi:treatment:03B6CB587A46B969FF18F870A3F55E6F:8EC02C957A44B969FE74FF49A5E85CBF" box="[507,657,154,178]" name="Diego-Suarez" pageId="2" pageNumber="598" stateProvince="Antsiranana">Diego-Suarez</location>
? [
<collectingRegion id="49DBB4AC7A44B969FD26FF49A44B5CBF" box="[681,818,154,178]" country="Madagascar" name="Antsiranana" pageId="2" pageNumber="598">Antsiranana</collectingRegion>
]” (
<collectionCode id="ED0EE28B7A44B969FCDEFF49A4D05CBF" box="[849,937,154,178]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="598">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7770137A44B969FC32FF4AA6AA5CDB" ID-GBIF-Occurrence="2423812416" collectionCode="MNHN" country="Madagascar" location="Amber" municipality="Mt. d'Ambre" pageId="2" pageNumber="598" specimenCount="7" specimenCount-female="2" specimenCount-male="5">
<specimenCount id="9D19B1C77A44B969FC32FF4AA4845CBC" box="[957,1021,153,177]" pageId="2" pageNumber="598" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="9D19B1C77A44B969FB85FF49A3315CBC" box="[1034,1096,154,178]" pageId="2" pageNumber="598" type="female">2 ♀♀</specimenCount>
, “
<collectingMunicipality id="6BC4E0347A44B969FBEEFF49A3895CBF" box="[1121,1264,154,178]" pageId="2" pageNumber="598">Mt. d’Ambre</collectingMunicipality>
[
<location id="8EC02C957A44B969FB71FF49A2325CBF" LSID="urn:lsid:plazi:treatment:03B6CB587A46B969FF18F870A3F55E6F:8EC02C957A44B969FB71FF49A2325CBF" box="[1278,1355,154,178]" country="Madagascar" municipality="Mt. d'Ambre" name="Amber" pageId="2" pageNumber="598">Amber</location>
Moun- tains]
<collectingCountry id="F3083ADE7A44B969FF56FF6DA6255CDB" box="[217,348,190,214]" name="Madagascar" pageId="2" pageNumber="598">Madagascar</collectingCountry>
” (
<collectionCode id="ED0EE28B7A44B969FEFBFF6DA6B55CDB" box="[372,460,190,214]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="598">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7770137A44B969FE51FF6DA32B5CDB" ID-GBIF-Occurrence="2423812413" box="[478,1106,190,214]" collectionCode="MNHN" country="Madagascar" location="Montagne d'Ambre" municipality="Mt. d'Ambre" pageId="2" pageNumber="598" specimenCount="2" specimenCount-male="2">
<specimenCount id="9D19B1C77A44B969FE51FF6DA5655CD8" box="[478,540,190,214]" pageId="2" pageNumber="598" type="male">2 ♂♂</specimenCount>
, “
<location id="8EC02C957A44B969FDBEFF6DA47C5CDB" LSID="urn:lsid:plazi:treatment:03B6CB587A46B969FF18F870A3F55E6F:8EC02C957A44B969FDBEFF6DA47C5CDB" box="[561,773,190,214]" country="Madagascar" name="Montagne d'Ambre" pageId="2" pageNumber="598">Montagne d’Ambre</location>
[
<emphasis id="B96BA65C7A44B969FC9AFF6DA4305CDB" box="[789,841,190,214]" italics="true" pageId="2" pageNumber="598">idem</emphasis>
]
<collectingCountry id="F3083ADE7A44B969FCD7FF6DA4A25CDB" box="[856,987,190,214]" name="Madagascar" pageId="2" pageNumber="598">Madagascar</collectingCountry>
” (
<collectionCode id="ED0EE28B7A44B969FC7CFF6DA3325CDB" box="[1011,1099,190,214]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="598">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA07A4E7A44B969FF48FF31A6795D6B" blockId="2.[151,1437,153,610]" pageId="2" pageNumber="598">
<emphasis id="B96BA65C7A44B969FF48FF31A6635CF7" bold="true" box="[199,282,226,250]" pageId="2" pageNumber="598">Female</emphasis>
(
<figureCitation id="132466CB7A44B969FEA7FF31A6025CF7" box="[296,379,226,250]" captionStart="FIGURES 1" captionStartId="1.[151,264,1895,1920]" captionTargetBox="[187,1410,347,1860]" captionTargetId="figure@1.[178,1410,336,1868]" captionTargetPageId="1" captionText="FIGURES 1A–F. Trigonochilus oedipus (Fairmaire, 1903) (A–C), T. macrotarsus Montreuil &amp; Frolov, new species (D–F) and locality map (G). A, male, dorsal view; B, aedeagus, lateral and dorsal view; C, female, dorsal view; D, male, holotype, dorsal view; E, aedeagus, lateral and dosal view; F, female, paratype, dorsal view; G, locality map (black circle – T. oedipus, white circle – T. macrotarsus). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3471118" httpUri="https://zenodo.org/record/3471118/files/figure.png" pageId="2" pageNumber="598">Fig. 1C</figureCitation>
). Similar to the male in colour and setation, but have a little wider elytra, obtuse posterior angles of pronotum (hind angles are somewhat protruding in males), shorter antennae, relatively slender metatibiae and metatarsi, well developed smaller claw of metatarsi (this claw is completely reduced in males), and rounded apical sternite of the abdomen.
</paragraph>
<paragraph id="8BA07A4E7A44B969FF48FEA2A41F5E33" blockId="2.[151,1437,153,610]" pageId="2" pageNumber="598">
<emphasis id="B96BA65C7A44B969FF48FEA2A6BB5D84" bold="true" box="[199,450,369,393]" pageId="2" pageNumber="598">Differential diagnosis.</emphasis>
<taxonomicName id="4C1F01CD7A44B969FE47FEA1A5D15D87" baseAuthorityName="Fairmaire" baseAuthorityYear="1903" box="[456,680,370,394]" class="Insecta" family="Scarabaeidae" genus="Trigonochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="598" phylum="Arthropoda" rank="species" species="oedipus">
<emphasis id="B96BA65C7A44B969FE47FEA1A5D15D87" box="[456,680,370,394]" italics="true" pageId="2" pageNumber="598">Trigonochilus oedipus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C1F01CD7A44B969FCA3FEA1A4B15D84" authority="Montreuil &amp; Frolov, 2019" authorityName="Montreuil &amp; Frolov" authorityYear="2019" box="[812,968,370,393]" class="Insecta" family="Scarabaeidae" genus="Trigonochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="598" phylum="Arthropoda" rank="species" species="macrotarsus" status="sp. nov.">
<emphasis id="B96BA65C7A44B969FCA3FEA1A4B15D84" box="[812,968,370,393]" italics="true" pageId="2" pageNumber="598">T. macrotarsus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2581B277A44B969FC5FFEA1A3295D87" box="[976,1104,370,394]" pageId="2" pageNumber="598" rank="species">new species</taxonomicNameLabel>
in a number of characters. The coloration of the head and pronotum is uniformly light brown. The elytra are yellow with dark brown sutural margins, base and humeri. The elytra are also more ovoid in this species (rather parallel-sided in
<taxonomicName id="4C1F01CD7A44B969FBB3FE69A3A15DDC" authority="Montreuil &amp; Frolov, 2019" authorityName="Montreuil &amp; Frolov" authorityYear="2019" box="[1084,1240,442,465]" class="Insecta" family="Scarabaeidae" genus="Trigonochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="598" phylum="Arthropoda" rank="species" species="macrotarsus" status="sp. nov.">
<emphasis id="B96BA65C7A44B969FBB3FE69A3A15DDC" box="[1084,1240,442,465]" italics="true" pageId="2" pageNumber="598">T. macrotarsus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2581B277A44B969FB51FE69A21B5DDF" box="[1246,1378,442,466]" pageId="2" pageNumber="598" rank="species">new species</taxonomicNameLabel>
). The legs are brown similar to the head and pronotum except for tarsi which are blackish brown. Apical margin of labrum with a short, pointed process medially, as opposed to a long, delated process in
<taxonomicName id="4C1F01CD7A44B969FC21FDD1A3325E14" authority="Montreuil &amp; Frolov, 2019" authorityName="Montreuil &amp; Frolov" authorityYear="2019" box="[942,1099,514,537]" class="Insecta" family="Scarabaeidae" genus="Trigonochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="598" phylum="Arthropoda" rank="species" species="macrotarsus" status="sp. nov.">
<emphasis id="B96BA65C7A44B969FC21FDD1A3325E14" box="[942,1099,514,537]" italics="true" pageId="2" pageNumber="598">T. macrotarsus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2581B277A44B969FBDDFDD1A3AB5E17" box="[1106,1234,514,538]" pageId="2" pageNumber="598" rank="species">new species</taxonomicNameLabel>
Metatarsi with only one claw in males. Aedeagus with parameres split at the apex (
<figureCitation id="132466CB7A44B969FC84FDF5A4215E33" box="[779,856,550,574]" captionStart="FIGURES 1" captionStartId="1.[151,264,1895,1920]" captionTargetBox="[187,1410,347,1860]" captionTargetId="figure@1.[178,1410,336,1868]" captionTargetPageId="1" captionText="FIGURES 1A–F. Trigonochilus oedipus (Fairmaire, 1903) (A–C), T. macrotarsus Montreuil &amp; Frolov, new species (D–F) and locality map (G). A, male, dorsal view; B, aedeagus, lateral and dorsal view; C, female, dorsal view; D, male, holotype, dorsal view; E, aedeagus, lateral and dosal view; F, female, paratype, dorsal view; G, locality map (black circle – T. oedipus, white circle – T. macrotarsus). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3471118" httpUri="https://zenodo.org/record/3471118/files/figure.png" pageId="2" pageNumber="598">Fig. 1B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30529C57A44B969FF18FD9AA3F55E6F" box="[151,1164,585,610]" pageId="2" pageNumber="598" type="distribution">
<paragraph id="8BA07A4E7A44B969FF18FD9AA3F55E6F" blockId="2.[151,1437,153,610]" box="[151,1164,585,610]" pageId="2" pageNumber="598">
<emphasis id="B96BA65C7A44B969FF18FD9AA65B5E6C" bold="true" box="[151,290,585,609]" pageId="2" pageNumber="598">Distribution</emphasis>
. Northernmost part of
<collectingCountry id="F3083ADE7A44B969FD97FD99A5E45E6F" box="[536,669,586,610]" name="Madagascar" pageId="2" pageNumber="598">Madagascar</collectingCountry>
:
<collectingRegion id="49DBB4AC7A44B969FD2AFD99A45F5E6F" box="[677,806,586,610]" country="Madagascar" name="Antsiranana" pageId="2" pageNumber="598">Antsiranana</collectingRegion>
and Amber Mountains (
<figureCitation id="132466CB7A44B969FBA5FD99A3065E6F" box="[1066,1151,586,610]" captionStart="FIGURES 1" captionStartId="1.[151,264,1895,1920]" captionTargetBox="[187,1410,347,1860]" captionTargetId="figure@1.[178,1410,336,1868]" captionTargetPageId="1" captionText="FIGURES 1A–F. Trigonochilus oedipus (Fairmaire, 1903) (A–C), T. macrotarsus Montreuil &amp; Frolov, new species (D–F) and locality map (G). A, male, dorsal view; B, aedeagus, lateral and dorsal view; C, female, dorsal view; D, male, holotype, dorsal view; E, aedeagus, lateral and dosal view; F, female, paratype, dorsal view; G, locality map (black circle – T. oedipus, white circle – T. macrotarsus). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3471118" httpUri="https://zenodo.org/record/3471118/files/figure.png" pageId="2" pageNumber="598">Fig. 1G</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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