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<document id="1F266DE69B6FC96EB027339FB432560D" ID-DOI="10.5281/zenodo.5303571" ID-ISSN="0374-1036" ID-Zenodo-Dep="5303571" ID-ZooBank="0360453D-E09E-4CC4-9BB0-981AEC7BED66S" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630104360660" checkinUser="marcus" docAuthor="Tsai, Jing-Fu &amp; Rédei, Dávid" docDate="2015" docId="03B63F659F5E1C4FFE4CFBA9FD2BFD27" docLanguage="en" docName="ActaEntMusNatPra.2015.55.2.625-664.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 55 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Acanthosoma atayal Tsai &amp; Rédei 2015, sp. nov." docType="treatment" docVersion="3" lastPageNumber="631" masterDocId="FF8F471D9F5A1C49FFADFFB2FFA0FFF4" masterDocTitle="The genus Acanthosoma in Taiwan (Hemiptera: Heteroptera: Acanthosomatidae)" masterLastPageNumber="664" masterPageNumber="625" pageNumber="629" updateTime="1699153447944" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="A6CA0BCA166B9A32B0C72CFA0A1FB974" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E8C55FFCCB56C4E7574322F713A9F27E">
<mods:title id="76DD184B4C03917352B7D68636D9C090">The genus Acanthosoma in Taiwan (Hemiptera: Heteroptera: Acanthosomatidae)</mods:title>
</mods:titleInfo>
<mods:name id="D71749D2A58DECB7EB0202BADADCD932" type="personal">
<mods:role id="D6B23DB5BCAB9668E8FB99A77E57986B">
<mods:roleTerm id="9C7067A18B0B6DE0BF5BEDE392D704CE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5F1F79EA2E901690AB006AEF630822BF">Tsai, Jing-Fu</mods:namePart>
<mods:affiliation id="5B64968CC58E3A2FCC404E0C566AEC04">Systematic Entomology, Graduate School of Agriculture, Hokkaido University, Sapporo, 060 - 8589 Japan; e-mail: jingfu. tsai @ gmail. com</mods:affiliation>
</mods:name>
<mods:name id="3B23828C9C609EFB061ADB01C289F531" type="personal">
<mods:role id="5FF72F7CA1E7009263959ABDBEE4C68A">
<mods:roleTerm id="699FE52C71DD1A41DE924BFF1E0C3260">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E04FD3BF850A99D6A6D8505CC3A67F0C">Rédei, Dávid</mods:namePart>
<mods:affiliation id="2244DCA940B6C242732D8A4B0C570F01">Institute of Entomology, College of Life Sciences, Nankai University, Weijin Road 94, 300071 Tianjin, China; e-mail: david. redei @ gmail. com &amp; Department of Zoology, Hungarian Natural History Museum, H- 1088 Budapest, Baross u. 13, Hungary</mods:affiliation>
</mods:name>
<mods:typeOfResource id="931AEF5BAFF95DAAF9BDD310E1C84DA9">text</mods:typeOfResource>
<mods:relatedItem id="92B46CA4AABB3EAA682069BA624E0BF8" type="host">
<mods:titleInfo id="F0B2A1F50F81F8E396C0FE7CB8D323EF">
<mods:title id="68DB94267E65A932BE7B4822DF1C2E78">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="33E4FF66B4CB9E02AAAC6363898D535A">
<mods:date id="DF78A86342B170B3BD0C964F68EBBF27">2015</mods:date>
<mods:detail id="6F7BD8F1D3BDA9039C11742C43DF1760" type="pubDate">
<mods:number id="41DC28A0AF4D53ED0FAFC85F272A5944">2015-12-31</mods:number>
</mods:detail>
<mods:detail id="4D76801C643B0F6D63B4215510C6EBF2" type="volume">
<mods:number id="1BD89C40CE1DED1DA2164E7B077252D6">55</mods:number>
</mods:detail>
<mods:detail id="4CF13430C32935A51AF0A6B88560D992" type="issue">
<mods:number id="8AEC4BF5C30D8756FB0576FB79FAC6D8">2</mods:number>
</mods:detail>
<mods:extent id="93CE2B5AF056028A25098AC8EC091FDF" unit="page">
<mods:start id="6599E714774536941D025AE9729CCA6E">625</mods:start>
<mods:end id="A8FF6870462A4A4E987B109E508CE7AC">664</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="2DF3660CA49E6BA2317435EDFB73961E">journal article</mods:classification>
<mods:identifier id="AC97CE5DC832E999CD2E9DFC3C6BE59A" type="DOI">10.5281/zenodo.5303571</mods:identifier>
<mods:identifier id="0463BB02FE7A7CBB8342CA95E607AC08" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="E27F222C657C29EB34B2D62A50361637" type="Zenodo-Dep">5303571</mods:identifier>
<mods:identifier id="0F9F219274ECDBED8DD7B40E68184429" type="ZooBank">0360453D-E09E-4CC4-9BB0-981AEC7BED66S</mods:identifier>
</mods:mods>
<treatment id="03B63F659F5E1C4FFE4CFBA9FD2BFD27" LSID="urn:lsid:plazi:treatment:03B63F659F5E1C4FFE4CFBA9FD2BFD27" httpUri="http://treatment.plazi.org/id/03B63F659F5E1C4FFE4CFBA9FD2BFD27" lastPageId="6" lastPageNumber="631" pageId="4" pageNumber="629">
<subSubSection id="C305DDF89F5E1C4DFE4CFBA9FCBDFBC7" box="[481,797,1051,1075]" pageId="4" pageNumber="629" type="nomenclature">
<paragraph id="8BA08E739F5E1C4DFE4CFBA9FCBDFBC7" blockId="4.[481,797,1051,1075]" box="[481,797,1051,1075]" pageId="4" pageNumber="629">
<heading id="D0E8391F9F5E1C4DFE4CFBA9FCBDFBC7" bold="true" box="[481,797,1051,1075]" centered="true" fontSize="8" level="2" pageId="4" pageNumber="629" reason="3">
<emphasis id="B96B52619F5E1C4DFE4CFBA9FCBDFBC7" bold="true" box="[481,797,1051,1075]" pageId="4" pageNumber="629">
<taxonomicName id="4C1FF5F09F5E1C4DFE4CFBA9FD61FBC7" authority="Tsai &amp; Rédei, 2015" authorityName="Tsai &amp; Rédei" authorityYear="2015" box="[481,705,1051,1075]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="629" phylum="Arthropoda" rank="species" species="atayal" status="sp. nov.">
<emphasis id="B96B52619F5E1C4DFE4CFBA9FD61FBC7" bold="true" box="[481,705,1051,1075]" italics="true" pageId="4" pageNumber="629">Acanthosoma atayal</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A258EF1A9F5E1C4DFD65FBAEFCBDFBC7" box="[712,797,1052,1075]" pageId="4" pageNumber="629" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C305DDF89F5E1C4DFDAEFB88FD5BFBB9" box="[515,763,1082,1101]" pageId="4" pageNumber="629" type="description">
<paragraph id="8BA08E739F5E1C4DFDAEFB88FD5BFBB9" blockId="4.[151,1128,1082,1410]" box="[515,763,1082,1101]" pageId="4" pageNumber="629">
(
<figureCitation id="132492F69F5E1C4DFDA4FB88FDF5FBB9" box="[521,597,1082,1101]" captionStart="Figs 1–10" captionStartId="10.[151,188,1450,1469]" captionTargetBox="[154,1125,221,1407]" captionTargetId="figure-10@10.[151,1126,221,1420]" captionTargetPageId="10" captionText="Figs 1–10.Acanthosoma spp., males. 1 – A.atayal sp.nov., posterior part of abdomen, ventral view; 2 – same, genital capsule and exposed part of segment VIII, ventral view; 3 – same, genital capsule, lateral view; 4 – A. haemorrhoidale formosanum subsp. nov., holotype, genital capsule, posterior view; 5 – A.haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM), genital capsule, posterior view; 6 – A. asahinai Ishihara, 1943, posterior part of abdomen, ventral view; 7 – same, genital capsule, ventral view; 8 – A. fallax sp. nov., posterior part of abdomen, ventral view; 9 – same, genital capsule, ventral view; 10 – A. pugnax sp. nov., genital capsule and exposed part of segment VIII, ventral view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303573" httpUri="https://zenodo.org/record/5303573/files/figure.png" pageId="4" pageNumber="629">Figs 1–3</figureCitation>
,
<figureCitation id="132492F69F5E1C4DFDF2FB88FD36FBB9" box="[607,662,1082,1101]" captionStart="Figs 20–44" captionStartId="14.[151,188,1196,1215]" captionTargetBox="[151,1126,231,1174]" captionTargetId="figure-84@14.[151,1126,231,1178]" captionTargetPageId="14" captionText="Figs 20–44. Acanthosoma spp., right parameres in different views. 20–21 – A. atayal sp. nov.; 22 – A. haemorrhoidale formosanum subsp. nov., holotype; 23 – A. haemorrhoidale ouchii Ishihara, 1950 (China: Sichuan, Mt. Emei, USNM); 24 – A. haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM); 25–28 – A. asahinai Ishihara, 1943; 29–32 – A. fallax sp. nov.; 33–36 – A. pugnax sp. nov.; 37–39 – A. axicia sp. nov.; 40–41 – A. laevicorne Dallas, 1851; 42–44 – A. expansum Horváth, 1905. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303577" httpUri="https://zenodo.org/record/5303577/files/figure.png" pageId="4" pageNumber="629">20–21</figureCitation>
,
<figureCitation id="132492F69F5E1C4DFD32FB88FD16FBB9" box="[671,694,1082,1101]" captionStart="Figs 45–51" captionStartId="17.[113,150,1468,1487]" captionTargetBox="[115,1086,506,1453]" captionTargetId="figure-12@17.[113,1088,506,1462]" captionTargetPageId="17" captionText="Figs 45–51. Acanthosoma spp. 45 – A. atayal sp. nov., female terminalia, posteroventral view; 46 – A. haemorrhoidale formosanum subsp. nov., paratype, female terminalia, posteroventral view; 47 – A. asahinai Ishihara, 1943, hind tibia of male; 48 – same, female terminalia, dorsal view; 49 – same, posteroventral view; 50 – A. fallax sp. nov., female terminalia, dorsal view; 51 – same, posteroventral view. PO6, PO7 = Pendergrast’s organs of ventrites VI and VII, respectively. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303579" httpUri="https://zenodo.org/record/5303579/files/figure.png" pageId="4" pageNumber="629">45</figureCitation>
,
<figureCitation id="132492F69F5E1C4DFD12FB88FD55FBB9" box="[703,757,1082,1101]" captionStart="Figs 56–59" captionStartId="22.[151,188,1539,1558]" captionTargetBox="[191,1126,235,1500]" captionTargetId="figure-10@22.[151,1126,220,1520]" captionTargetPageId="22" captionText="Figs 56–59. Acanthosoma atayal sp. nov. 56–57 – male; 58–59 – female. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303583" httpUri="https://zenodo.org/record/5303583/files/figure.png" pageId="4" pageNumber="629">56–59</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C305DDF89F5E1C4DFF3AFBD6FC34FA76" pageId="4" pageNumber="629" type="materials_examined">
<paragraph id="8BA08E739F5E1C4DFF3AFBD6FC34FA76" blockId="4.[151,1128,1082,1410]" pageId="4" pageNumber="629">
<emphasis id="B96B52619F5E1C4DFF3AFBD6FEBDFB83" bold="true" box="[151,285,1124,1143]" pageId="4" pageNumber="629">Type material.</emphasis>
<materialsCitation id="3B77842E9F5E1C4DFE89FBD6FC34FA76" collectingDate="1980-09-12" collectingDateMax="2010-10-15" collectingDateMin="1980-09-12" collectionCode="NMNS, NTU, NCHU, HNHM, NMPC, TFRI, ZJPC, SEHU, NSMT, TARI" collectorName="Provincial Rd. No. &amp; S. W. Hou &amp; C. B. Sun &amp; C. L. Chen &amp; W. C. Chuang &amp; Yuchih &amp; Puli &amp; J. Dalihod &amp; Tseifeng &amp; H. Takizawa &amp; Alishan &amp; K. Nagami &amp; Tayuling &amp; K. S. Lin &amp; C. W. Wang" country="Taiwan" county="Yuanfeng" elevation="2560" location="Rd. No." municipality="Provincial Rd." pageId="4" pageNumber="629" specimenCode="ENT 639-546" specimenCount="9" specimenCount-female="9" stateProvince="Nantou" typeStatus="holotype">
<typeStatus id="54A430D19F5E1C4DFE89FBD6FED8FB81" box="[292,376,1124,1143]" pageId="4" pageNumber="629" type="holotype">HOLOTYPE</typeStatus>
: J,
<emphasis id="B96B52619F5E1C4DFE0DFBD6FDD8FB83" bold="true" box="[416,632,1124,1143]" pageId="4" pageNumber="629">
<collectingCountry id="F308CEE39F5E1C4DFE0DFBD6FE5AFB83" box="[416,506,1124,1143]" name="Taiwan" pageId="4" pageNumber="629">TAIWAN</collectingCountry>
:
<collectingRegion id="49DB40919F5E1C4DFDA8FBD6FDEAFB81" box="[517,586,1124,1143]" country="Taiwan" name="Nantou" pageId="4" pageNumber="629">NANTOU</collectingRegion>
CO.:
</emphasis>
<collectingCounty id="62C1F6FF9F5E1C4DFDD3FBD6FD72FB83" box="[638,722,1124,1143]" pageId="4" pageNumber="629">Yuanfeng</collectingCounty>
,
<collectorName id="26EAEBA59F5E1C4DFD71FBD6FC20FB83" box="[732,896,1124,1143]" pageId="4" pageNumber="629">
<collectingMunicipality id="6BC414099F5E1C4DFD71FBD6FCFBFB83" box="[732,859,1124,1143]" pageId="4" pageNumber="629">
Provincial
<location id="8EC0D8A89F5E1C4DFC96FBD6FCFBFB83" LSID="urn:lsid:plazi:treatment:03B63F659F5E1C4FFE4CFBA9FD2BFD27:8EC0D8A89F5E1C4DFC96FBD6FCFBFB83" box="[827,859,1124,1143]" country="Taiwan" county="Yuanfeng" municipality="Provincial Rd." name="Rd. No." pageId="4" pageNumber="629" stateProvince="Nantou">Rd.</location>
</collectingMunicipality>
No.
</collectorName>
14A 22.5K,
<date id="FFA1A8B39F5E1C4DFC46FBD6FBE2FB83" box="[1003,1090,1124,1143]" pageId="4" pageNumber="629" value="2010-10-15">
<collectingDate id="EFE5515B9F5E1C4DFC46FBD6FBE2FB83" box="[1003,1090,1124,1143]" pageId="4" pageNumber="629" value="2010-10-15">15.x.2010</collectingDate>
</date>
, ex.
<taxonomicName id="4C1FF5F09F5E1C4DFF3AFBCDFE13FB66" authority="Hayata" authorityName="Hayata" box="[151,435,1151,1170]" class="Magnoliopsida" family="Rosaceae" genus="Photinia" kingdom="Plantae" order="Rosales" pageId="4" pageNumber="629" phylum="Tracheophyta" rank="species" species="niitakayamensis">
<emphasis id="B96B52619F5E1C4DFF3AFBCDFECFFB66" box="[151,367,1151,1170]" italics="true" pageId="4" pageNumber="629">Photinia niitakayamensis</emphasis>
Hayata
</taxonomicName>
, leg.
<collectorName id="26EAEBA59F5E1C4DFE72FBCDFD93FB66" box="[479,563,1151,1170]" pageId="4" pageNumber="629">S.W. Hou</collectorName>
; mounted on card, intact, deposited in
<collectionCode id="ED0E16B69F5E1C4DFCD7FBCDFC18FB66" box="[890,952,1151,1170]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="4" pageNumber="629">NMNS</collectionCode>
(
<figureCitation id="132492F69F5E1C4DFC6EFBCDFB86FB66" box="[963,1062,1151,1170]" captionStart="Figs 56–59" captionStartId="22.[151,188,1539,1558]" captionTargetBox="[191,1126,235,1500]" captionTargetId="figure-10@22.[151,1126,220,1520]" captionTargetPageId="22" captionText="Figs 56–59. Acanthosoma atayal sp. nov. 56–57 – male; 58–59 – female. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303583" httpUri="https://zenodo.org/record/5303583/files/figure.png" pageId="4" pageNumber="629">Figs 56–57</figureCitation>
). PARA-
<subScript id="179B8C369F5E1C4DFF3AFB2CFF65FB5F" attach="none" box="[151,197,1182,1195]" fontSize="6" pageId="4" pageNumber="629">TYPES</subScript>
:
<emphasis id="B96B52619F5E1C4DFF7CFB2BFE39FB58" bold="true" box="[209,409,1177,1196]" pageId="4" pageNumber="629">
<collectingCountry id="F308CEE39F5E1C4DFF7CFB2BFE8BFB58" box="[209,299,1177,1196]" name="Taiwan" pageId="4" pageNumber="629">TAIWAN</collectingCountry>
:
<collectingRegion id="49DB40919F5E1C4DFE99FB2BFECBFB5F" box="[308,363,1177,1196]" country="Taiwan" name="Taipei" pageId="4" pageNumber="629">TAIPEI</collectingRegion>
CO.:
</emphasis>
<location id="8EC0D8A89F5E1C4DFE32FB2BFE74FB58" LSID="urn:lsid:plazi:treatment:03B63F659F5E1C4FFE4CFBA9FD2BFD27:8EC0D8A89F5E1C4DFE32FB2BFE74FB58" box="[415,468,1177,1196]" country="Taiwan" county="Yuanfeng" municipality="Provincial Rd." name="Wulai" pageId="4" pageNumber="629" stateProvince="Nantou">Wulai</location>
,
<date id="FFA1A8B39F5E1C4DFE73FB2BFD95FB58" box="[478,565,1177,1196]" pageId="4" pageNumber="629" value="2008-10-15">
<collectingDate id="EFE5515B9F5E1C4DFE73FB2BFD95FB58" box="[478,565,1177,1196]" pageId="4" pageNumber="629" value="2008-10-15">15.x.2008</collectingDate>
</date>
, leg.
<collectorName id="26EAEBA59F5E1C4DFDC9FB2BFD13FB58" box="[612,691,1177,1196]" pageId="4" pageNumber="629">C.B. Sun</collectorName>
(1 J [dissected]
<collectionCode id="ED0E16B69F5E1C4DFCE4FB2BFCD9FB58" box="[841,889,1177,1196]" pageId="4" pageNumber="629">NTU</collectionCode>
).
<emphasis id="B96B52619F5E1C4DFC25FB2BFBABFB58" bold="true" box="[904,1035,1177,1196]" pageId="4" pageNumber="629">
<collectingRegion id="49DB40919F5E1C4DFC25FB2BFC7EFB5F" box="[904,990,1177,1196]" country="Taiwan" name="Taichung" pageId="4" pageNumber="629">TAICHUNG</collectingRegion>
CO.:
</emphasis>
<location id="8EC0D8A89F5E1C4DFBBDFB2BFBC3FB58" LSID="urn:lsid:plazi:treatment:03B63F659F5E1C4FFE4CFBA9FD2BFD27:8EC0D8A89F5E1C4DFBBDFB2BFBC3FB58" box="[1040,1123,1177,1196]" country="Taiwan" county="Yuanfeng" municipality="Provincial Rd." name="Anmasan" pageId="4" pageNumber="629" stateProvince="Nantou">Anmasan</location>
,
<date id="FFA1A8B39F5E1C4DFF3AFB06FF5BFB33" box="[151,251,1204,1223]" pageId="4" pageNumber="629" value="1983-07-18">
<collectingDate id="EFE5515B9F5E1C4DFF3AFB06FF5BFB33" box="[151,251,1204,1223]" pageId="4" pageNumber="629" value="1983-07-18">18.vii.1983</collectingDate>
</date>
, leg.
<collectorName id="26EAEBA59F5E1C4DFE86FB06FE25FB33" box="[299,389,1204,1223]" pageId="4" pageNumber="629">C.L. Chen</collectorName>
(
<specimenCount id="9D1945FA9F5E1C4DFE39FB06FE16FB33" box="[404,438,1198,1223]" pageId="4" pageNumber="629" type="female">1 ♀</specimenCount>
<collectionCode id="ED0E16B69F5E1C4DFE16FB06FE5BFB33" box="[443,507,1204,1223]" country="Taiwan" httpUri="http://grbio.org/cool/e7yf-mhkb" name="National Chung Hsing University" pageId="4" pageNumber="629">NCHU</collectionCode>
).
<emphasis id="B96B52619F5E1C4DFDA6FB06FDDEFB33" bold="true" box="[523,638,1204,1223]" pageId="4" pageNumber="629">
<collectingRegion id="49DB40919F5E1C4DFDA6FB06FDF0FB31" box="[523,592,1204,1223]" country="Taiwan" name="Nantou" pageId="4" pageNumber="629">NANTOU</collectingRegion>
CO.:
</emphasis>
same as
<typeStatus id="54A430D19F5E1C4DFD7CFB06FCBCFB33" box="[721,796,1204,1223]" pageId="4" pageNumber="629" type="holotype">holotype</typeStatus>
(1 J
<specimenCount id="9D1945FA9F5E1C4DFCFCFB06FCD5FB33" box="[849,885,1198,1223]" pageId="4" pageNumber="629" type="female">1 ♀</specimenCount>
<collectionCode id="ED0E16B69F5E1C4DFCD4FB06FC1AFB33" box="[889,954,1204,1223]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="4" pageNumber="629">NMNS</collectionCode>
); same as
<typeStatus id="54A430D19F5E1C4DFBBBFB06FBC3FB33" box="[1046,1123,1204,1223]" pageId="4" pageNumber="629" type="holotype">holotype</typeStatus>
, ex
<taxonomicName id="4C1FF5F09F5E1C4DFF1EFB7DFC87FB16" authority="(Hayata) Kudo &amp; Masamune ex Kudo" authorityName="Kudo" baseAuthorityName="Hayata" box="[179,807,1231,1250]" class="Magnoliopsida" family="Fagaceae" genus="Cyclobalanopsis" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="629" phylum="Tracheophyta" rank="species" species="stenophylloides">
<emphasis id="B96B52619F5E1C4DFF1EFB7DFE6EFB16" box="[179,462,1231,1250]" italics="true" pageId="4" pageNumber="629">Cyclobalanopsis stenophylloides</emphasis>
(Hayata) Kudo &amp; Masamune ex Kudo
</taxonomicName>
(1 J
<collectionCode id="ED0E16B69F5E1C4DFCCDFB7DFC00FB16" box="[864,928,1231,1250]" country="Taiwan" httpUri="http://grbio.org/cool/e7yf-mhkb" name="National Chung Hsing University" pageId="4" pageNumber="629">NCHU</collectionCode>
);
<location id="8EC0D8A89F5E1C4DFC1DFB7DFBA4FB16" LSID="urn:lsid:plazi:treatment:03B63F659F5E1C4FFE4CFBA9FD2BFD27:8EC0D8A89F5E1C4DFC1DFB7DFBA4FB16" box="[944,1028,1231,1250]" country="Taiwan" county="Yuanfeng" municipality="Provincial Rd." name="Yuanfeng" pageId="4" pageNumber="629" stateProvince="Nantou">Yuanfeng</location>
,
<collectorName id="26EAEBA59F5E1C4DFBA2FB7DFF7DFB08" pageId="4" pageNumber="629">
<location id="8EC0D8A89F5E1C4DFBA2FB7DFF17FB08" LSID="urn:lsid:plazi:treatment:03B63F659F5E1C4FFE4CFBA9FD2BFD27:8EC0D8A89F5E1C4DFBA2FB7DFF17FB08" country="Taiwan" county="Yuanfeng" municipality="Provincial Rd." name="Provincial Rd." pageId="4" pageNumber="629" stateProvince="Nantou">
Provincial
<location id="8EC0D8A89F5E1C4DFF3AFB5BFF17FB08" LSID="urn:lsid:plazi:treatment:03B63F659F5E1C4FFE4CFBA9FD2BFD27:8EC0D8A89F5E1C4DFF3AFB5BFF17FB08" box="[151,183,1257,1276]" country="Taiwan" county="Yuanfeng" municipality="Provincial Rd." name="Rd. No." pageId="4" pageNumber="629" stateProvince="Nantou">Rd.</location>
</location>
No.
</collectorName>
14A, 24.5K,
<date id="FFA1A8B39F5E1C4DFEE0FB5BFE3AFB08" box="[333,410,1257,1276]" pageId="4" pageNumber="629" value="2010-10-01">
<collectingDate id="EFE5515B9F5E1C4DFEE0FB5BFE3AFB08" box="[333,410,1257,1276]" pageId="4" pageNumber="629" value="2010-10-01">1.x.2010</collectingDate>
</date>
, ex
<taxonomicName id="4C1FF5F09F5E1C4DFE12FB5BFD7EFB08" authority="Hayata" authorityName="Hayata" box="[447,734,1257,1276]" class="Magnoliopsida" family="Rosaceae" genus="Photinia" kingdom="Plantae" order="Rosales" pageId="4" pageNumber="629" phylum="Tracheophyta" rank="species" species="niitakayamensis">
<emphasis id="B96B52619F5E1C4DFE12FB5BFD38FB08" box="[447,664,1257,1276]" italics="true" pageId="4" pageNumber="629">Photinia niitakayamensis</emphasis>
Hayata
</taxonomicName>
, leg.
<collectorName id="26EAEBA59F5E1C4DFCA3FB5BFCC3FB08" box="[782,867,1257,1276]" pageId="4" pageNumber="629">S.W. Hou</collectorName>
(1 J
<specimenCount id="9D1945FA9F5E1C4DFC35FB5BFC1BFB08" box="[920,955,1251,1276]" pageId="4" pageNumber="629" type="female">1 ♀</specimenCount>
<collectionCode id="ED0E16B69F5E1C4DFC6DFB5BFBA0FB08" box="[960,1024,1257,1276]" country="Taiwan" httpUri="http://grbio.org/cool/e7yf-mhkb" name="National Chung Hsing University" pageId="4" pageNumber="629">NCHU</collectionCode>
);
<location id="8EC0D8A89F5E1C4DFBA2FB5BFBC3FB08" LSID="urn:lsid:plazi:treatment:03B63F659F5E1C4FFE4CFBA9FD2BFD27:8EC0D8A89F5E1C4DFBA2FB5BFBC3FB08" box="[1039,1123,1257,1276]" country="Taiwan" county="Yuanfeng" municipality="Provincial Rd." name="Yuanfeng" pageId="4" pageNumber="629" stateProvince="Nantou">Yuanfeng</location>
,
<collectorName id="26EAEBA59F5E1C4DFF3AFAB6FE9AFAE3" box="[151,314,1284,1303]" pageId="4" pageNumber="629">
<location id="8EC0D8A89F5E1C4DFF3AFAB6FEB4FAE3" LSID="urn:lsid:plazi:treatment:03B63F659F5E1C4FFE4CFBA9FD2BFD27:8EC0D8A89F5E1C4DFF3AFAB6FEB4FAE3" box="[151,276,1284,1303]" country="Taiwan" county="Yuanfeng" municipality="Provincial Rd." name="Provincial Rd." pageId="4" pageNumber="629" stateProvince="Nantou">
Provincial
<location id="8EC0D8A89F5E1C4DFF58FAB6FEB4FAE3" LSID="urn:lsid:plazi:treatment:03B63F659F5E1C4FFE4CFBA9FD2BFD27:8EC0D8A89F5E1C4DFF58FAB6FEB4FAE3" box="[245,276,1284,1303]" country="Taiwan" county="Yuanfeng" municipality="Provincial Rd." name="Rd. No." pageId="4" pageNumber="629" stateProvince="Nantou">Rd.</location>
</location>
No.
</collectorName>
14A, 24.5K,
<date id="FFA1A8B39F5E1C4DFE05FAB6FE54FAE3" box="[424,500,1284,1303]" pageId="4" pageNumber="629" value="2010-10-01">
<collectingDate id="EFE5515B9F5E1C4DFE05FAB6FE54FAE3" box="[424,500,1284,1303]" pageId="4" pageNumber="629" value="2010-10-01">1.x.2010</collectingDate>
</date>
, ex
<taxonomicName id="4C1FF5F09F5E1C4DFDB5FAB6FC95FAE3" authority="Hayata" authorityName="Hayata" box="[536,821,1284,1303]" class="Magnoliopsida" family="Rosaceae" genus="Photinia" kingdom="Plantae" order="Rosales" pageId="4" pageNumber="629" phylum="Tracheophyta" rank="species" species="niitakayamensis">
<emphasis id="B96B52619F5E1C4DFDB5FAB6FD50FAE3" box="[536,752,1284,1303]" italics="true" pageId="4" pageNumber="629">Photinia niitakayamensis</emphasis>
Hayata
</taxonomicName>
, leg.
<collectorName id="26EAEBA59F5E1C4DFCCEFAB6FC17FAE3" box="[867,951,1284,1303]" pageId="4" pageNumber="629">S.W. Hou</collectorName>
(1 J
<specimenCount id="9D1945FA9F5E1C4DFC44FAB6FBABFAE3" box="[1001,1035,1278,1303]" pageId="4" pageNumber="629" type="female">1 ♀</specimenCount>
<collectionCode id="ED0E16B69F5E1C4DFBA2FAB6FBF2FAE3" box="[1039,1106,1284,1303]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="4" pageNumber="629">HNHM</collectionCode>
, 1 J
<specimenCount id="9D1945FA9F5E1C4DFF00FAADFF6FFAC5" box="[173,207,1304,1330]" pageId="4" pageNumber="629" type="female">1 ♀</specimenCount>
<collectionCode id="ED0E16B69F5E1C4DFF79FAADFEB3FAC6" box="[212,275,1311,1330]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="4" pageNumber="629">NMPC</collectionCode>
, 1 J
<collectionCode id="ED0E16B69F5E1C4DFEEEFAADFED5FAC6" box="[323,373,1311,1330]" country="Taiwan" httpUri="http://grbio.org/cool/fitg-90ys" name="Taiwan Fisheries Research Institute" pageId="4" pageNumber="629">TFRI</collectionCode>
);
<location id="8EC0D8A89F5E1C4DFE28FAADFDA6FAC6" LSID="urn:lsid:plazi:treatment:03B63F659F5E1C4FFE4CFBA9FD2BFD27:8EC0D8A89F5E1C4DFE28FAADFDA6FAC6" box="[389,518,1311,1330]" country="Taiwan" county="Yuanfeng" municipality="Provincial Rd." name="Ho Huan Shan" pageId="4" pageNumber="629" stateProvince="Nantou">Ho Huan Shan</location>
,
<date id="FFA1A8B39F5E1C4DFDBDFAADFDD3FAC6" box="[528,627,1311,1330]" pageId="4" pageNumber="629" value="1990-07-26">
<collectingDate id="EFE5515B9F5E1C4DFDBDFAADFDD3FAC6" box="[528,627,1311,1330]" pageId="4" pageNumber="629" value="1990-07-26">26.vii.1990</collectingDate>
</date>
, sweeping, leg.
<collectorName id="26EAEBA59F5E1C4DFD53FAADFCD5FAC6" box="[766,885,1311,1330]" pageId="4" pageNumber="629">W.C. Chuang</collectorName>
,
<collectionCode id="ED0E16B69F5E1C4DFCD3FAADFC1CFAC6" box="[894,956,1311,1330]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="4" pageNumber="629">NMNS</collectionCode>
<specimenCode id="DBB926089F5E1C4DFC6FFAADFB98FAC6" box="[962,1080,1311,1330]" collectionCode="NMNS" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="4" pageNumber="629">ENT 639-546</specimenCode>
(
<specimenCount id="9D1945FA9F5E1C4DFBEAFAADFBC8FAC5" box="[1095,1128,1304,1330]" pageId="4" pageNumber="629" type="female">1 ♀</specimenCount>
<collectionCode id="ED0E16B69F5E1C4DFF3AFA8BFF78FAB8" box="[151,216,1337,1356]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="4" pageNumber="629">NMNS</collectionCode>
);
<location id="8EC0D8A89F5E1C4DFF4AFA8BFED3FAB8" LSID="urn:lsid:plazi:treatment:03B63F659F5E1C4FFE4CFBA9FD2BFD27:8EC0D8A89F5E1C4DFF4AFA8BFED3FAB8" box="[231,371,1337,1356]" country="Taiwan" county="Yuanfeng" municipality="Provincial Rd." name="Sun Moon Lake" pageId="4" pageNumber="629" stateProvince="Nantou">Sun Moon Lake</location>
,
<collectorName id="26EAEBA59F5E1C4DFED6FA8BFE17FAB8" box="[379,439,1337,1356]" pageId="4" pageNumber="629">Yuchih</collectorName>
env.,
<collectorName id="26EAEBA59F5E1C4DFE47FA8BFDAEFAB8" box="[490,526,1337,1356]" pageId="4" pageNumber="629">Puli</collectorName>
,
<date id="FFA1A8B39F5E1C4DFDBAFA8BFD34FAB8" box="[535,660,1337,1356]" pageId="4" pageNumber="629" value="1995-06-12" valueMax="1995-06-16" valueMin="1995-06-12">
<collectingDate id="EFE5515B9F5E1C4DFDBAFA8BFD34FAB8" box="[535,660,1337,1356]" pageId="4" pageNumber="629" value="1995-06-12" valueMax="1995-06-16" valueMin="1995-06-12">12–16.vi.1995</collectingDate>
</date>
, leg.
<collectorName id="26EAEBA59F5E1C4DFD6FFA8BFCB9FAB8" box="[706,793,1337,1356]" pageId="4" pageNumber="629">J. Dalihod</collectorName>
(1 J
<collectionCode id="ED0E16B69F5E1C4DFCE6FA8BFCDEFAB8" box="[843,894,1337,1356]" pageId="4" pageNumber="629">ZJPC</collectionCode>
);
<collectorName id="26EAEBA59F5E1C4DFC21FA8BFC77FAB8" box="[908,983,1337,1356]" pageId="4" pageNumber="629">Tseifeng</collectorName>
,
<date id="FFA1A8B39F5E1C4DFC4DFA8BFB9EFAB8" box="[992,1086,1337,1356]" pageId="4" pageNumber="629" value="1985-08-01">
<collectingDate id="EFE5515B9F5E1C4DFC4DFA8BFB9EFAB8" box="[992,1086,1337,1356]" pageId="4" pageNumber="629" value="1985-08-01">1.viii.1985</collectingDate>
</date>
, leg.
<collectorName id="26EAEBA59F5E1C4DFF3AFAE6FEA3FA93" box="[151,259,1364,1383]" pageId="4" pageNumber="629">H. Takizawa</collectorName>
(
<specimenCount id="9D1945FA9F5E1C4DFEBDFAE6FEE1FA93" box="[272,321,1358,1383]" pageId="4" pageNumber="629" type="female">2 ♀♀</specimenCount>
<collectionCode id="ED0E16B69F5E1C4DFEEAFAE6FE22FA93" box="[327,386,1364,1383]" pageId="4" pageNumber="629">SEHU</collectionCode>
).
<emphasis id="B96B52619F5E1C4DFE3FFAE6FE5AFA93" bold="true" box="[402,506,1364,1383]" pageId="4" pageNumber="629">
<collectingRegion id="49DB40919F5E1C4DFE3FFAE6FE6CFA91" box="[402,460,1364,1383]" country="Taiwan" name="Chiayi" pageId="4" pageNumber="629">CHIAYI</collectingRegion>
CO.:
</emphasis>
<collectorName id="26EAEBA59F5E1C4DFE52FAE6FDE4FA93" box="[511,580,1364,1383]" pageId="4" pageNumber="629">Alishan</collectorName>
,
<date id="FFA1A8B39F5E1C4DFDE0FAE6FD0AFA93" box="[589,682,1364,1383]" pageId="4" pageNumber="629" value="1982-06-20">
<collectingDate id="EFE5515B9F5E1C4DFDE0FAE6FD0AFA93" box="[589,682,1364,1383]" pageId="4" pageNumber="629" value="1982-06-20">20.vi.1982</collectingDate>
</date>
, leg.
<collectorName id="26EAEBA59F5E1C4DFD77FAE6FC9BFA93" box="[730,827,1364,1383]" pageId="4" pageNumber="629">K. Nagami</collectorName>
, NMNT-I-He 34905 (
<specimenCount id="9D1945FA9F5E1C4DFC54FAE6FBBAFA93" box="[1017,1050,1358,1383]" pageId="4" pageNumber="629" type="female">1 ♀</specimenCount>
<collectionCode id="ED0E16B69F5E1C4DFBB2FAE6FBFEFA93" box="[1055,1118,1364,1383]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="4" pageNumber="629">NSMT</collectionCode>
).
<emphasis id="B96B52619F5E1C4DFF3AFADDFEAEFA76" bold="true" box="[151,270,1391,1410]" pageId="4" pageNumber="629">
<collectingRegion id="49DB40919F5E1C4DFF3AFADDFF42FA74" box="[151,226,1391,1410]" country="Taiwan" name="Hualien" pageId="4" pageNumber="629">HUALIEN</collectingRegion>
CO.:
</emphasis>
<collectorName id="26EAEBA59F5E1C4DFEB9FADDFEC2FA76" box="[276,354,1391,1410]" pageId="4" pageNumber="629">Tayuling</collectorName>
,
<quantity id="4CE723969F5E1C4DFEC7FADDFE0DFA76" box="[362,429,1391,1410]" metricMagnitude="3" metricUnit="m" metricValue="2.56" pageId="4" pageNumber="629" unit="m" value="2560.0">
<elevation id="003269409F5E1C4DFEC7FADDFE0DFA76" box="[362,429,1391,1410]" metricMagnitude="3" metricUnit="m" metricValue="2.56" pageId="4" pageNumber="629" unit="m" value="2560.0">2560 m</elevation>
</quantity>
,
<date id="FFA1A8B39F5E1C4DFE18FADDFD93FA76" box="[437,563,1391,1410]" pageId="4" pageNumber="629" value="1980-09-12" valueMax="1980-09-15" valueMin="1980-09-12">
<collectingDate id="EFE5515B9F5E1C4DFE18FADDFD93FA76" box="[437,563,1391,1410]" pageId="4" pageNumber="629" value="1980-09-12" valueMax="1980-09-15" valueMin="1980-09-12">12–15.ix.1980</collectingDate>
</date>
, leg.
<collectorName id="26EAEBA59F5E1C4DFDCDFADDFD09FA76" box="[608,681,1391,1410]" pageId="4" pageNumber="629">K.S. Lin</collectorName>
&amp;
<collectorName id="26EAEBA59F5E1C4DFD69FADDFC85FA76" box="[708,805,1391,1410]" pageId="4" pageNumber="629">C.W. Wang</collectorName>
(1 J
<collectionCode id="ED0E16B69F5E1C4DFCFBFADDFC2AFA76" box="[854,906,1391,1410]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="4" pageNumber="629">TARI</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C305DDF89F5E1C4DFF3AFA28FC62FA05" pageId="4" pageNumber="629" type="diagnosis">
<paragraph id="8BA08E739F5E1C4DFF3AFA28FC62FA05" blockId="4.[151,1127,1434,1585]" pageId="4" pageNumber="629">
<emphasis id="B96B52619F5E1C4DFF3AFA28FEA8FA45" bold="true" box="[151,264,1434,1457]" pageId="4" pageNumber="629">Diagnosis.</emphasis>
The species can be recognized by its humeri being distinctly produced but relatively short and broad, apically obtuse, directed slightly posterolaterad, punctured, and tinged with reddish. The male (
<figureCitation id="132492F69F5E1C4DFEC8FA68FE63FA05" box="[357,451,1498,1521]" captionStart="Figs 1–10" captionStartId="10.[151,188,1450,1469]" captionTargetBox="[154,1125,221,1407]" captionTargetId="figure-10@10.[151,1126,221,1420]" captionTargetPageId="10" captionText="Figs 1–10.Acanthosoma spp., males. 1 – A.atayal sp.nov., posterior part of abdomen, ventral view; 2 – same, genital capsule and exposed part of segment VIII, ventral view; 3 – same, genital capsule, lateral view; 4 – A. haemorrhoidale formosanum subsp. nov., holotype, genital capsule, posterior view; 5 – A.haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM), genital capsule, posterior view; 6 – A. asahinai Ishihara, 1943, posterior part of abdomen, ventral view; 7 – same, genital capsule, ventral view; 8 – A. fallax sp. nov., posterior part of abdomen, ventral view; 9 – same, genital capsule, ventral view; 10 – A. pugnax sp. nov., genital capsule and exposed part of segment VIII, ventral view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303573" httpUri="https://zenodo.org/record/5303573/files/figure.png" pageId="4" pageNumber="629">Figs 1–3</figureCitation>
) and female (
<figureCitation id="132492F69F5E1C4DFDFAFA68FD04FA05" box="[599,676,1498,1521]" captionStart="Figs 45–51" captionStartId="17.[113,150,1468,1487]" captionTargetBox="[115,1086,506,1453]" captionTargetId="figure-12@17.[113,1088,506,1462]" captionTargetPageId="17" captionText="Figs 45–51. Acanthosoma spp. 45 – A. atayal sp. nov., female terminalia, posteroventral view; 46 – A. haemorrhoidale formosanum subsp. nov., paratype, female terminalia, posteroventral view; 47 – A. asahinai Ishihara, 1943, hind tibia of male; 48 – same, female terminalia, dorsal view; 49 – same, posteroventral view; 50 – A. fallax sp. nov., female terminalia, dorsal view; 51 – same, posteroventral view. PO6, PO7 = Pendergrast’s organs of ventrites VI and VII, respectively. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303579" httpUri="https://zenodo.org/record/5303579/files/figure.png" pageId="4" pageNumber="629">Fig. 45</figureCitation>
) terminalia are diagnostic.
</paragraph>
</subSubSection>
<subSubSection id="C305DDF89F5E1C4CFF3AFA48FC31FA87" lastPageId="5" lastPageNumber="630" pageId="4" pageNumber="629" type="description">
<paragraph id="8BA08E739F5E1C4CFF3AFA48FC80FE87" blockId="4.[151,1127,1434,1585]" lastBlockId="5.[113,1089,220,1619]" lastPageId="5" lastPageNumber="630" pageId="4" pageNumber="629">
<emphasis id="B96B52619F5E1C4DFF3AFA48FED4F9E5" bold="true" box="[151,372,1529,1553]" pageId="4" pageNumber="629">
Description.
<emphasis id="B96B52619F5E1C4DFE8EFA4BFED4F9E5" bold="true" box="[291,372,1529,1553]" italics="true" pageId="4" pageNumber="629">Colour.</emphasis>
</emphasis>
Ground colour of dorsum green (turning to yellowish in dead specimens), suffused with reddish brown to various extent; pronotum broadly suffused with red on calli and along its posterior margin, humeral processes reddish brown to red with more or less distinct dark suffusion; endocorium of fore wings usually with strong ferruginous suffusion; abdomen pale yellow, with distinct red suffusion on exposed parts of segments VII (J,
<specimenCount id="9D1945FA9F5F1C4CFBB3FEA7FB91FEC0" box="[1054,1073,277,308]" pageId="5" pageNumber="630" type="female">♀</specimenCount>
), VIII (
<specimenCount id="9D1945FA9F5F1C4CFF1FFE87FF65FEA0" box="[178,197,309,340]" pageId="5" pageNumber="630" type="female">♀</specimenCount>
), and dorsal surface of genital capsule (J); connexival plates and corresponding ventrites of abdominal segments II–VII without black markings.
</paragraph>
<paragraph id="8BA08E739F5F1C4CFF3DFECEFE80FD67" blockId="5.[113,1089,220,1619]" pageId="5" pageNumber="630">
<emphasis id="B96B52619F5F1C4CFF3DFECEFE0DFE67" bold="true" box="[144,429,379,403]" pageId="5" pageNumber="630">
<emphasis id="B96B52619F5F1C4CFF3DFECEFE06FE67" bold="true" box="[144,422,379,403]" italics="true" pageId="5" pageNumber="630">Integument and vestiture</emphasis>
.
</emphasis>
Body glabrous except of fine, scattered hairs on appendages and terminalia; mandibular plates with a few scattered black punctures, interocular area with a pair of irregular longitudinal rows of black punctures; pronotum, scutellum and sclerotized parts of fore wings with rather uniform black punctation except on calli, distal portions of humeral processes and costal margin of fore wing, punctation of exocorium and neighbouring part of endocorium not conspicuously different; ventral surface of body virtually unpunctured except of scattered punctures on ventral surface of humeral process and faint, superficial punctures along posterior margin of abdominal ventrite VIII and on ventral surface of genital capsule of male.
</paragraph>
<paragraph id="8BA08E739F5F1C4CFF3DFD29FDA9FC27" blockId="5.[113,1089,220,1619]" pageId="5" pageNumber="630">
<emphasis id="B96B52619F5F1C4CFF3DFD29FE9CFD47" box="[144,316,667,691]" italics="true" pageId="5" pageNumber="630">
<emphasis id="B96B52619F5F1C4CFF3DFD29FF5EFD47" bold="true" box="[144,254,667,691]" italics="true" pageId="5" pageNumber="630">Structure.</emphasis>
Body
</emphasis>
elongate oval with distinctly produced humeri, distance between tips of humeral processes about 1.2–1.25 (J,
<specimenCount id="9D1945FA9F5F1C4CFDA8FD07FDB8FD20" box="[517,536,693,724]" pageId="5" pageNumber="630" type="female">♀</specimenCount>
) times as long as greatest width of body posteriad of humeri.
<emphasis id="B96B52619F5F1C4CFF66FD6EFEA5FD07" box="[203,261,732,755]" italics="true" pageId="5" pageNumber="630">Head</emphasis>
1.2–1.25 (J,
<specimenCount id="9D1945FA9F5F1C4CFE37FD67FE0DFD00" box="[410,429,725,756]" pageId="5" pageNumber="630" type="female">♀</specimenCount>
) times as broad as its median length, about 1.55–1.6 times as broad as interocular distance, finely transversely rugose, anterior portion of clypeus depressed, with a broad median longitudinal groove.
<emphasis id="B96B52619F5F1C4CFD9EFCAEFD24FCC7" box="[563,644,796,819]" italics="true" pageId="5" pageNumber="630">Labium</emphasis>
slightly surpassing mid coxae.
<emphasis id="B96B52619F5F1C4CFC7AFCAEFBE1FCC7" box="[983,1089,796,819]" italics="true" pageId="5" pageNumber="630">Pronotum</emphasis>
with anterolateral margin weakly concave, continued in humeral processes without conspicuous break in its outline; humeri short and broad, subtriangular, directed subhorizontally, apically obtuse.
<emphasis id="B96B52619F5F1C4CFEB1FCCEFDFBFC67" box="[284,603,892,915]" italics="true" pageId="5" pageNumber="630">Thoracic pleuron and sternum</emphasis>
. Mesosternal carina distinctly surpassing base of head, highly elevated, rather broadly rounded anteriorly; metathoracic scent gland ostiole with a long, slightly arched peritreme.
</paragraph>
<paragraph id="8BA08E739F5F1C4CFF3DFC6EFBE1FB47" blockId="5.[113,1089,220,1619]" pageId="5" pageNumber="630">
<emphasis id="B96B52619F5F1C4CFF3DFC6EFE57FC07" box="[144,503,988,1011]" italics="true" pageId="5" pageNumber="630">Male and female terminalia. Male</emphasis>
. Genital capsule (
<figureCitation id="132492F69F5F1C4CFD19FC6EFCB0FC07" box="[692,784,988,1011]" captionStart="Figs 1–10" captionStartId="10.[151,188,1450,1469]" captionTargetBox="[154,1125,221,1407]" captionTargetId="figure-10@10.[151,1126,221,1420]" captionTargetPageId="10" captionText="Figs 1–10.Acanthosoma spp., males. 1 – A.atayal sp.nov., posterior part of abdomen, ventral view; 2 – same, genital capsule and exposed part of segment VIII, ventral view; 3 – same, genital capsule, lateral view; 4 – A. haemorrhoidale formosanum subsp. nov., holotype, genital capsule, posterior view; 5 – A.haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM), genital capsule, posterior view; 6 – A. asahinai Ishihara, 1943, posterior part of abdomen, ventral view; 7 – same, genital capsule, ventral view; 8 – A. fallax sp. nov., posterior part of abdomen, ventral view; 9 – same, genital capsule, ventral view; 10 – A. pugnax sp. nov., genital capsule and exposed part of segment VIII, ventral view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303573" httpUri="https://zenodo.org/record/5303573/files/figure.png" pageId="5" pageNumber="630">Figs 1–3</figureCitation>
) with a pair of short posterolateral projections each with a dense tuft of setae apically; dorsal rim broadly U-shaped; ventral rim with a tuft of setae medially, ventral infolding densely pilose. Paramere (
<figureCitation id="132492F69F5F1C4CFC10FBAEFB99FBC7" box="[957,1081,1052,1075]" captionStart="Figs 20–44" captionStartId="14.[151,188,1196,1215]" captionTargetBox="[151,1126,231,1174]" captionTargetId="figure-84@14.[151,1126,231,1178]" captionTargetPageId="14" captionText="Figs 20–44. Acanthosoma spp., right parameres in different views. 20–21 – A. atayal sp. nov.; 22 – A. haemorrhoidale formosanum subsp. nov., holotype; 23 – A. haemorrhoidale ouchii Ishihara, 1950 (China: Sichuan, Mt. Emei, USNM); 24 – A. haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM); 25–28 – A. asahinai Ishihara, 1943; 29–32 – A. fallax sp. nov.; 33–36 – A. pugnax sp. nov.; 37–39 – A. axicia sp. nov.; 40–41 – A. laevicorne Dallas, 1851; 42–44 – A. expansum Horváth, 1905. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303577" httpUri="https://zenodo.org/record/5303577/files/figure.png" pageId="5" pageNumber="630">Figs 20–21</figureCitation>
) L-shaped, apex finger-shaped.
<emphasis id="B96B52619F5F1C4CFE18FB8EFDA5FBA7" box="[437,517,1084,1107]" italics="true" pageId="5" pageNumber="630">Female</emphasis>
(
<figureCitation id="132492F69F5F1C4CFDBEFB8EFDC1FBA7" box="[531,609,1084,1107]" captionStart="Figs 45–51" captionStartId="17.[113,150,1468,1487]" captionTargetBox="[115,1086,506,1453]" captionTargetId="figure-12@17.[113,1088,506,1462]" captionTargetPageId="17" captionText="Figs 45–51. Acanthosoma spp. 45 – A. atayal sp. nov., female terminalia, posteroventral view; 46 – A. haemorrhoidale formosanum subsp. nov., paratype, female terminalia, posteroventral view; 47 – A. asahinai Ishihara, 1943, hind tibia of male; 48 – same, female terminalia, dorsal view; 49 – same, posteroventral view; 50 – A. fallax sp. nov., female terminalia, dorsal view; 51 – same, posteroventral view. PO6, PO7 = Pendergrast’s organs of ventrites VI and VII, respectively. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303579" httpUri="https://zenodo.org/record/5303579/files/figure.png" pageId="5" pageNumber="630">Fig. 45</figureCitation>
). Posterior margin with a deep, shield-shaped median incision surrounding valvifers VIII; posterior margin of laterotergites VIII prominent, far surpassing posterolateral angle of abdominal segment VII. Pendergrast’s organs on ventrites VI and VII broadly separeted, those on ventrite VII much smaller than those on ventrite VI.
</paragraph>
<paragraph id="8BA08E739F5F1C4CFF3DFB0EFC31FA87" blockId="5.[113,1089,220,1619]" pageId="5" pageNumber="630">
<emphasis id="B96B52619F5F1C4CFF3DFB0EFE91FB27" bold="true" box="[144,305,1212,1235]" italics="true" pageId="5" pageNumber="630">Measurements</emphasis>
(in mm). Body length from apex of head to apex of membrane 11.9–12.2 (J) / 13.7–14.9 (
<specimenCount id="9D1945FA9F5F1C4CFE8AFB67FE9AFB00" box="[295,314,1237,1268]" pageId="5" pageNumber="630" type="female">♀</specimenCount>
); greatest width of body posteriad of humeral processes 5.06–8.34; median length of head 1.75–1.95, width across eyes 2.15–2.40, interocular distance 1.35–1.55; length of antennal segments (I) 1.50–1.55: (IIa) 1.65–1.80: (IIb) 1.25–1.40: (III) 1.85–2.00: (IV) 1.80–1.85; median length of pronotum 2.00–2.75, greatest width (across tips of humeri) 6.41–8.49; median length of scutellum 3.45–4.45, width at base 2.85–3.70.
</paragraph>
</subSubSection>
<subSubSection id="C305DDF89F5F1C4CFFDCFACEFEEAF9A7" pageId="5" pageNumber="630" type="etymology">
<paragraph id="8BA08E739F5F1C4CFFDCFACEFC02FA27" blockId="5.[113,1089,220,1619]" pageId="5" pageNumber="630">
<emphasis id="B96B52619F5F1C4CFFDCFACEFF52FA67" bold="true" box="[113,242,1404,1427]" pageId="5" pageNumber="630">Etymology.</emphasis>
The specific epithet is derived from the name of the Atayal people, one of Taiwan’s indigenous tribes residing in the mountainous regions of northern and central
<collectingCountry id="F308CEE39F5F1C4CFC5EFA2EFBE1FA47" box="[1011,1089,1436,1459]" name="Taiwan" pageId="5" pageNumber="630">Taiwan</collectingCountry>
where the new species occurs; noun in apposition, ending not to be changed.
</paragraph>
<paragraph id="8BA08E739F5F1C4CFFDCFA6EFEEAF9A7" blockId="5.[113,1089,220,1619]" pageId="5" pageNumber="630">
<emphasis id="B96B52619F5F1C4CFFDCFA6EFF4FFA07" bold="true" box="[113,239,1500,1523]" pageId="5" pageNumber="630">Bionomics.</emphasis>
Several specimens were collected in multiple occasions from
<taxonomicName id="4C1FF5F09F5F1C4CFC2DFA6EFE2CF9E7" authority="Hayata" authorityName="Hayata" authorityYear="1911" class="Magnoliopsida" family="Rosaceae" genus="Photinia" kingdom="Plantae" order="Rosales" pageId="5" pageNumber="630" phylum="Tracheophyta" rank="species" species="niitakyamensis">
<emphasis id="B96B52619F5F1C4CFC2DFA6EFF1AF9E7" italics="true" pageId="5" pageNumber="630">Photinia niitakyamensis</emphasis>
Hayata (Rosaceae)
</taxonomicName>
during its fruiting period. The breeding season of the species is around October. A few specimens were found on
<taxonomicName id="4C1FF5F09F5F1C4CFD2EF9AEFEE5F9A7" authority="Kudo &amp; Masam." authorityName="Kudo &amp; Masam." class="Magnoliopsida" family="Fagaceae" genus="Cyclobalanopsis" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="630" phylum="Tracheophyta" rank="species" species="stenophylloides">
<emphasis id="B96B52619F5F1C4CFD2EF9AEFC42F9C7" box="[643,994,1564,1587]" italics="true" pageId="5" pageNumber="630">Cyclobalanopsis stenophylloides</emphasis>
Kudô &amp; Masam. (Fagaceae)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C305DDF89F5C1C4FFF3AFF6EFDECFEE7" pageId="6" pageNumber="631" type="distribution">
<paragraph id="8BA08E739F5C1C4FFF3AFF6EFDECFEE7" blockId="6.[151,1127,220,723]" pageId="6" pageNumber="631">
<emphasis id="B96B52619F5C1C4FFF3AFF6EFE89FF07" bold="true" box="[151,297,220,243]" pageId="6" pageNumber="631">Distribution.</emphasis>
The species is endemic to
<collectingCountry id="F308CEE39F5C1C4FFD96FF6EFD2CFF07" box="[571,652,220,243]" name="Taiwan" pageId="6" pageNumber="631">Taiwan</collectingCountry>
, apparently restricted to mountainous regions ranging from
<quantity id="4CE723969F5C1C4FFE87FF4EFE72FEE7" box="[298,466,252,275]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="6" pageNumber="631" unit="m" value="2000.0" valueMax="2500.0" valueMin="1500.0">1500 to 2500 m</quantity>
in altitude.
</paragraph>
</subSubSection>
<subSubSection id="C305DDF89F5C1C4FFF3AFEAEFD2BFD27" pageId="6" pageNumber="631" type="discussion">
<paragraph id="8BA08E739F5C1C4FFF3AFEAEFD2BFD27" blockId="6.[151,1127,220,723]" pageId="6" pageNumber="631">
<emphasis id="B96B52619F5C1C4FFF3AFEAEFEA6FEC7" bold="true" box="[151,262,284,307]" pageId="6" pageNumber="631">Remarks.</emphasis>
This species belongs to the
<taxonomicName id="4C1FF5F09F5C1C4FFD95FEAEFCDBFEC7" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[568,891,284,307]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="631" phylum="Arthropoda" rank="species" species="haemorrhoidale">
<emphasis id="B96B52619F5C1C4FFD95FEAEFCDBFEC7" box="[568,891,284,307]" italics="true" pageId="6" pageNumber="631">Acanthosoma haemorrhoidale</emphasis>
</taxonomicName>
species group hereby defined in the following way: posterolateral angles of genital capsule produced into a pair of short and broad projections but never into conspicuously elongate processes, of trapezoid shape in posterior view, ventral infolding perpendicular, facing posteriad in rest; paramere simple, rod-like, apex either rounded or hooked but never bifurcate. The group currently contains
<taxonomicName id="4C1FF5F09F5C1C4FFBE2FE2EFDA5FE27" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="631" phylum="Arthropoda" rank="species" species="haemorrhoidale">
<emphasis id="B96B52619F5C1C4FFBE2FE2EFEE2FE27" italics="true" pageId="6" pageNumber="631">A. haemorrhoidale</emphasis>
(Linnaeus, 1758)
</taxonomicName>
,
<taxonomicName id="4C1FF5F09F5C1C4FFDBCFE0EFCA5FE27" authority="Liu, 1980" authorityName="Liu" authorityYear="1980" box="[529,773,444,467]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="631" phylum="Arthropoda" rank="species" species="emeiense">
<emphasis id="B96B52619F5C1C4FFDBCFE0EFD32FE27" box="[529,658,444,467]" italics="true" pageId="6" pageNumber="631">A. emeiense</emphasis>
<bibRefCitation id="EF8EF3829F5C1C4FFD34FE0EFCA5FE27" author="LIU S. L." box="[665,773,444,467]" pageId="6" pageNumber="631" pagination="233 - 242" refId="ref20554" refString="LIU S. L. 1980: Six new species of Acanthosomatidae from China (Hemiptera: Heteroptera). Zoological Research 1: 233 - 242 (in Chinese, English summary)." type="journal article" year="1980">Liu, 1980</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1FF5F09F5C1C4FFCBCFE0EFBE3FE27" authority="Jakovlev, 1880" authorityName="Jakovlev" authorityYear="1880" box="[785,1091,444,467]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="631" phylum="Arthropoda" rank="species" species="spinicolle">
<emphasis id="B96B52619F5C1C4FFCBCFE0EFC38FE27" box="[785,920,444,467]" italics="true" pageId="6" pageNumber="631">A. spinicolle</emphasis>
<bibRefCitation id="EF8EF3829F5C1C4FFC0DFE0EFBE3FE27" author="JAKOVLEV V. E." box="[928,1091,444,467]" pageId="6" pageNumber="631" pagination="385 - 398" refId="ref19702" refString="JAKOVLEV V. E. 1880: Contributions to the fauna of bugs of Russia and the neighbouring countries. IV. Bulletin de la Societe Imperiale des Naturalistes de Moscou 55 (2): 385 - 398 (in Russian and German)." type="journal article" year="1880">Jakovlev, 1880</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1FF5F09F5C1C4FFBE2FE0EFE58FE07" authority="Hsiao &amp; Liu, 1977" authorityName="Hsiao &amp; Liu" authorityYear="1977" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="631" phylum="Arthropoda" rank="species" species="nigrodorsum">
<emphasis id="B96B52619F5C1C4FFBE2FE0EFE80FE07" italics="true" pageId="6" pageNumber="631">A. nigrodorsum</emphasis>
<bibRefCitation id="EF8EF3829F5C1C4FFE85FE6EFE58FE07" author="HSIAO T. Y. &amp; LIU S. L." box="[296,504,476,499]" pageId="6" pageNumber="631" pagination="300 - 302" refId="ref19524" refString="HSIAO T. Y. &amp; LIU S. L. 1977: Family Acanthosomatidae. Pp. 159 - 180, 300 - 302, plates 25 - 29. In: HSIAO T. Y., REN S. Z., ZHENG L. Y., JING H. L. &amp; LIU S. L.: A handbook for the determination of the Chinese Hemiptera-Heteroptera. Volume I. Science Press, Beijing, iii + 330 pp, 52 plates (in Chinese, English summary)." type="book chapter" year="1977">Hsiao &amp; Liu, 1977</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1FF5F09F5C1C4FFDA9FE6EFCF7FE07" authority="(Distant, 1900)" baseAuthorityName="Distant" baseAuthorityYear="1900" box="[516,855,476,499]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="631" phylum="Arthropoda" rank="species" species="murreeanum">
<emphasis id="B96B52619F5C1C4FFDA9FE6EFD0BFE07" box="[516,683,476,499]" italics="true" pageId="6" pageNumber="631">A. murreeanum</emphasis>
(
<bibRefCitation id="EF8EF3829F5C1C4FFD16FE6EFCEFFE07" author="DISTANT W. L." box="[699,847,476,499]" pageId="6" pageNumber="631" pagination="220 - 234" refId="ref18825" refString="DISTANT W. L. 1900: Rhynchotal notes. - VI. Heteroptera: Dinidorinae, Phyllocephalinae, Urolabidinae, and Acanthosominae. Annals and Magazine of Natural History, Series 7 6: 220 - 234." type="journal article" year="1900">Distant, 1900</bibRefCitation>
)
</taxonomicName>
and the newly described
<taxonomicName id="4C1FF5F09F5C1C4FFF3AFE4EFF59FDE7" authority="Tsai &amp; Rédei, 2015" authorityName="Tsai &amp; Rédei" authorityYear="2015" box="[151,249,508,531]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="631" phylum="Arthropoda" rank="species" species="atayal" status="sp. nov.">
<emphasis id="B96B52619F5C1C4FFF3AFE4EFF59FDE7" box="[151,249,508,531]" italics="true" pageId="6" pageNumber="631">A. atayal</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A258EF1A9F5C1C4FFEADFE4EFEF3FDE7" box="[256,339,508,531]" pageId="6" pageNumber="631" rank="species">sp. nov.</taxonomicNameLabel>
The new species differs from all of the previously described members of the species group in its relatively smaller and more slender body, the shape of its humeral processes, and the reduced Pendergrast’s organs on ventrite VII. It is particularly similar to
<taxonomicName id="4C1FF5F09F5C1C4FFF3AFDEEFEC1FD87" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[151,353,604,627]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="631" phylum="Arthropoda" rank="species" species="haemorrhoidale">
<emphasis id="B96B52619F5C1C4FFF3AFDEEFEC1FD87" box="[151,353,604,627]" italics="true" pageId="6" pageNumber="631">A. haemorrhoidale</emphasis>
</taxonomicName>
; besides of the above characters, it differs from the latter species in the angularly broken apical portion of the paramere (
<figureCitation id="132492F69F5C1C4FFD06FDCEFD5AFD67" box="[683,762,636,659]" captionStart="Figs 20–44" captionStartId="14.[151,188,1196,1215]" captionTargetBox="[151,1126,231,1174]" captionTargetId="figure-84@14.[151,1126,231,1178]" captionTargetPageId="14" captionText="Figs 20–44. Acanthosoma spp., right parameres in different views. 20–21 – A. atayal sp. nov.; 22 – A. haemorrhoidale formosanum subsp. nov., holotype; 23 – A. haemorrhoidale ouchii Ishihara, 1950 (China: Sichuan, Mt. Emei, USNM); 24 – A. haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM); 25–28 – A. asahinai Ishihara, 1943; 29–32 – A. fallax sp. nov.; 33–36 – A. pugnax sp. nov.; 37–39 – A. axicia sp. nov.; 40–41 – A. laevicorne Dallas, 1851; 42–44 – A. expansum Horváth, 1905. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303577" httpUri="https://zenodo.org/record/5303577/files/figure.png" pageId="6" pageNumber="631">Fig. 20</figureCitation>
) (strongly hooked in
<taxonomicName id="4C1FF5F09F5C1C4FFC49FDCEFF47FD47" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="631" phylum="Arthropoda" rank="species" species="haemorrhoidale">
<emphasis id="B96B52619F5C1C4FFC49FDCEFF47FD47" italics="true" pageId="6" pageNumber="631">A. haemorrhoidale</emphasis>
</taxonomicName>
, cf.
<figureCitation id="132492F69F5C1C4FFEB7FD2EFE34FD47" box="[282,404,668,691]" captionStart="Figs 20–44" captionStartId="14.[151,188,1196,1215]" captionTargetBox="[151,1126,231,1174]" captionTargetId="figure-84@14.[151,1126,231,1178]" captionTargetPageId="14" captionText="Figs 20–44. Acanthosoma spp., right parameres in different views. 20–21 – A. atayal sp. nov.; 22 – A. haemorrhoidale formosanum subsp. nov., holotype; 23 – A. haemorrhoidale ouchii Ishihara, 1950 (China: Sichuan, Mt. Emei, USNM); 24 – A. haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM); 25–28 – A. asahinai Ishihara, 1943; 29–32 – A. fallax sp. nov.; 33–36 – A. pugnax sp. nov.; 37–39 – A. axicia sp. nov.; 40–41 – A. laevicorne Dallas, 1851; 42–44 – A. expansum Horváth, 1905. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303577" httpUri="https://zenodo.org/record/5303577/files/figure.png" pageId="6" pageNumber="631">Figs 22–24</figureCitation>
) and the strongly convex posterior margin of laterotergites VIII of female (weakly convex in
<taxonomicName id="4C1FF5F09F5C1C4FFE1FFD0EFDDBFD27" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[434,635,700,723]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="631" phylum="Arthropoda" rank="species" species="haemorrhoidale">
<emphasis id="B96B52619F5C1C4FFE1FFD0EFDDBFD27" box="[434,635,700,723]" italics="true" pageId="6" pageNumber="631">A. haemorrhoidale</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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