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<document id="1E616F960C6A6391CCF4B381FF9AA0AF" ID-DOI="10.5281/zenodo.5303571" ID-ISSN="0374-1036" ID-Zenodo-Dep="5303571" ID-ZooBank="0360453D-E09E-4CC4-9BB0-981AEC7BED66S" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630104360660" checkinUser="marcus" docAuthor="Tsai, Jing-Fu &amp; Rédei, Dávid" docDate="2015" docId="03B63F659F5D1C40FE2BFB49FDB8FDA7" docLanguage="en" docName="ActaEntMusNatPra.2015.55.2.625-664.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 55 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Acanthosoma asahinai Ishihara 1943" docType="treatment" docVersion="3" lastPageNumber="634" masterDocId="FF8F471D9F5A1C49FFADFFB2FFA0FFF4" masterDocTitle="The genus Acanthosoma in Taiwan (Hemiptera: Heteroptera: Acanthosomatidae)" masterLastPageNumber="664" masterPageNumber="625" pageNumber="632" updateTime="1699153447944" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="4EDE3EEFD760161CA5B4DECB001C01D4">The genus Acanthosoma in Taiwan (Hemiptera: Heteroptera: Acanthosomatidae)</mods:title>
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<mods:affiliation id="2AB60C7687D63D978183AA064AB9D271">Systematic Entomology, Graduate School of Agriculture, Hokkaido University, Sapporo, 060 - 8589 Japan; e-mail: jingfu. tsai @ gmail. com</mods:affiliation>
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<mods:affiliation id="4AFA3C1A15E971123BAD9EF94802BCB4">Institute of Entomology, College of Life Sciences, Nankai University, Weijin Road 94, 300071 Tianjin, China; e-mail: david. redei @ gmail. com &amp; Department of Zoology, Hungarian Natural History Museum, H- 1088 Budapest, Baross u. 13, Hungary</mods:affiliation>
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<treatment id="03B63F659F5D1C40FE2BFB49FDB8FDA7" LSID="urn:lsid:plazi:treatment:03B63F659F5D1C40FE2BFB49FDB8FDA7" httpUri="http://treatment.plazi.org/id/03B63F659F5D1C40FE2BFB49FDB8FDA7" lastPageId="9" lastPageNumber="634" pageId="7" pageNumber="632">
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<heading id="D0E8391F9F5D1C4EFE2BFB49FC8CFAE7" bold="true" box="[390,812,1275,1299]" centered="true" fontSize="8" level="2" pageId="7" pageNumber="632" reason="3">
<taxonomicName id="4C1FF5F09F5D1C4EFE2BFB49FC8CFAE7" ID-CoL="7NYCZ" authority="Ishihara, 1943" authorityName="Ishihara" authorityYear="1943" box="[390,812,1275,1299]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="632" phylum="Arthropoda" rank="species" species="asahinai">
<emphasis id="B96B52619F5D1C4EFE2BFB49FC8CFAE7" bold="true" box="[390,812,1275,1299]" pageId="7" pageNumber="632">
<emphasis id="B96B52619F5D1C4EFE2BFB49FD23FAE7" bold="true" box="[390,643,1275,1299]" italics="true" pageId="7" pageNumber="632">Acanthosoma asahinai</emphasis>
<bibRefCitation id="EF8EF3829F5D1C4EFD24FB4EFC8CFAE7" author="ISHIHARA T." box="[649,812,1276,1299]" pageId="7" pageNumber="632" pagination="495 - 497" refId="ref19649" refString="ISHIHARA T. 1943: Zwei Acanthosoma-arten der Insel Formosa (Hemiptera: Pentatomidae). Transactions of the Natural History Society of Taiwan 33: 495 - 497." type="journal article" year="1943">Ishihara, 1943</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="C305DDF89F5D1C4EFE60FAA8FD45FAD9" box="[461,741,1306,1325]" pageId="7" pageNumber="632" type="description">
<paragraph id="8BA08E739F5D1C4EFE60FAA8FD45FAD9" blockId="7.[113,1089,1306,1614]" box="[461,741,1306,1325]" pageId="7" pageNumber="632">
(
<figureCitation id="132492F69F5D1C4EFE7EFAA8FD80FAD9" box="[467,544,1306,1325]" captionStart="Figs 110" captionStartId="10.[151,188,1450,1469]" captionTargetBox="[154,1125,221,1407]" captionTargetId="figure-10@10.[151,1126,221,1420]" captionTargetPageId="10" captionText="Figs 110.Acanthosoma spp., males. 1 A.atayal sp.nov., posterior part of abdomen, ventral view; 2 same, genital capsule and exposed part of segment VIII, ventral view; 3 same, genital capsule, lateral view; 4 A. haemorrhoidale formosanum subsp. nov., holotype, genital capsule, posterior view; 5 A.haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM), genital capsule, posterior view; 6 A. asahinai Ishihara, 1943, posterior part of abdomen, ventral view; 7 same, genital capsule, ventral view; 8 A. fallax sp. nov., posterior part of abdomen, ventral view; 9 same, genital capsule, ventral view; 10 A. pugnax sp. nov., genital capsule and exposed part of segment VIII, ventral view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303573" httpUri="https://zenodo.org/record/5303573/files/figure.png" pageId="7" pageNumber="632">Figs 67</figureCitation>
,
<figureCitation id="132492F69F5D1C4EFD84FAA8FDC0FAD9" box="[553,608,1306,1325]" captionStart="Figs 2044" captionStartId="14.[151,188,1196,1215]" captionTargetBox="[151,1126,231,1174]" captionTargetId="figure-84@14.[151,1126,231,1178]" captionTargetPageId="14" captionText="Figs 2044. Acanthosoma spp., right parameres in different views. 2021 A. atayal sp. nov.; 22 A. haemorrhoidale formosanum subsp. nov., holotype; 23 A. haemorrhoidale ouchii Ishihara, 1950 (China: Sichuan, Mt. Emei, USNM); 24 A. haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM); 2528 A. asahinai Ishihara, 1943; 2932 A. fallax sp. nov.; 3336 A. pugnax sp. nov.; 3739 A. axicia sp. nov.; 4041 A. laevicorne Dallas, 1851; 4244 A. expansum Horváth, 1905. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303577" httpUri="https://zenodo.org/record/5303577/files/figure.png" pageId="7" pageNumber="632">2528</figureCitation>
,
<figureCitation id="132492F69F5D1C4EFDC4FAA8FD00FAD9" box="[617,672,1306,1325]" captionStart="Figs 4551" captionStartId="17.[113,150,1468,1487]" captionTargetBox="[115,1086,506,1453]" captionTargetId="figure-12@17.[113,1088,506,1462]" captionTargetPageId="17" captionText="Figs 4551. Acanthosoma spp. 45 A. atayal sp. nov., female terminalia, posteroventral view; 46 A. haemorrhoidale formosanum subsp. nov., paratype, female terminalia, posteroventral view; 47 A. asahinai Ishihara, 1943, hind tibia of male; 48 same, female terminalia, dorsal view; 49 same, posteroventral view; 50 A. fallax sp. nov., female terminalia, dorsal view; 51 same, posteroventral view. PO6, PO7 = Pendergrasts organs of ventrites VI and VII, respectively. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303579" httpUri="https://zenodo.org/record/5303579/files/figure.png" pageId="7" pageNumber="632">4749</figureCitation>
,
<figureCitation id="132492F69F5D1C4EFD04FAA8FD7FFAD9" box="[681,735,1306,1325]" captionStart="Figs 6467" captionStartId="24.[151,188,1541,1560]" captionTargetBox="[182,1088,227,1493]" captionTargetId="figure-10@24.[151,1126,212,1512]" captionTargetPageId="24" captionText="Figs 6467. Acanthosoma asahinai Ishihara, 1943. 6465 male; 6667 female. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303587" httpUri="https://zenodo.org/record/5303587/files/figure.png" pageId="7" pageNumber="632">6467</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C305DDF89F5D1C4EFFDCFA8EFB8DFA6B" pageId="7" pageNumber="632" type="reference_group">
<paragraph id="8BA08E739F5D1C4EFFDCFA8EFDBDFA9E" blockId="7.[113,1089,1306,1614]" pageId="7" pageNumber="632">
<treatmentCitationGroup id="AB0FA95D9F5D1C4EFFDCFA8EFDBDFA9E" pageId="7" pageNumber="632">
<treatmentCitation id="0ABEA8629F5D1C4EFFDCFA8EFE4CFABB" author="ISHIHARA T." box="[113,492,1340,1359]" page="495" pageId="7" pageNumber="632" year="1943">
<taxonomicName id="4C1FF5F09F5D1C4EFFDCFA8EFE4CFABB" ID-CoL="7NYCZ" authority="Ishihara, 1943: 495" authorityName="Ishihara" authorityPageNumber="495" authorityYear="1943" box="[113,492,1340,1359]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="632" phylum="Arthropoda" rank="species" species="asahinai">
<emphasis id="B96B52619F5D1C4EFFDCFA8EFE96FABB" box="[113,310,1340,1359]" italics="true" pageId="7" pageNumber="632">Acanthosoma asahinai</emphasis>
<materialsCitation id="3B77842E9F5D1C4EFE92FA8EFE4CFABB" box="[319,492,1340,1359]" collectionCode="ELKU" country="Taiwan" location="Mareppa-Oiwake" municipality="Formosa" pageId="7" pageNumber="632" specimenCount="1" stateProvince="Nantou" typeStatus="holotype">
<bibRefCitation id="EF8EF3829F5D1C4EFE92FA8EFE4CFABB" author="ISHIHARA T." box="[319,492,1340,1359]" pageId="7" pageNumber="632" pagination="495 - 497" refId="ref19649" refString="ISHIHARA T. 1943: Zwei Acanthosoma-arten der Insel Formosa (Hemiptera: Pentatomidae). Transactions of the Natural History Society of Taiwan 33: 495 - 497." type="journal article" year="1943">Ishihara, 1943: 495</bibRefCitation>
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.
<typeStatus id="54A430D19F5D1C4EFE5AFA8EFDEAFABB" box="[503,586,1340,1359]" pageId="7" pageNumber="632" type="holotype">Holotype</typeStatus>
: J,
<collectingMunicipality id="6BC414099F5D1C4EFDDCFA8EFD1CFABB" box="[625,700,1340,1359]" pageId="7" pageNumber="632">Formosa</collectingMunicipality>
[=
<collectingCountry id="F308CEE39F5D1C4EFD73FA8EFC81FABB" box="[734,801,1340,1359]" name="Taiwan" pageId="7" pageNumber="632">Taiwan</collectingCountry>
], Taityū- syū [=
<collectingRegion id="49DB40919F5D1C4EFC02FA8EFB98FABB" box="[943,1080,1340,1359]" country="Taiwan" name="Nantou" pageId="7" pageNumber="632">Nantou County</collectingRegion>
],
<location id="8EC0D8A89F5D1C4EFF3DFAE5FE85FA9E" LSID="urn:lsid:plazi:treatment:03B63F659F5D1C40FE2BFB49FDB8FDA7:8EC0D8A89F5D1C4EFF3DFAE5FE85FA9E" box="[144,293,1367,1386]" country="Taiwan" municipality="Formosa" name="Mareppa-Oiwake" pageId="7" pageNumber="632" stateProvince="Nantou">Mareppa-Oiwake</location>
[= Renai, Tsuifeng]; ELKU!
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA08E739F5D1C4EFFDCFAC3FB8DFA6B" blockId="7.[113,1089,1306,1614]" pageId="7" pageNumber="632">
<treatmentCitationGroup id="AB0FA95D9F5D1C4EFFDCFAC3FB8DFA6B" pageId="7" pageNumber="632">
<treatmentCitation id="0ABEA8629F5D1C4EFFDCFAC3FDB0FA70" author="HSIAO T. Y. &amp; LIU S. L." box="[113,528,1393,1412]" page=")" pageId="7" pageNumber="632" year="1977">
<taxonomicName id="4C1FF5F09F5D1C4EFFDCFAC3FDB0FA70" ID-CoL="7NYCZ" authority=": HSIAO &amp; LIU (1977: 175)" authorityName=": HSIAO &amp; LIU" authorityPageNumber="175" authorityYear="1977" box="[113,528,1393,1412]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="632" phylum="Arthropoda" rank="species" species="asahinai">
<emphasis id="B96B52619F5D1C4EFFDCFAC3FE93FA70" box="[113,307,1393,1412]" italics="true" pageId="7" pageNumber="632">Acanthosoma asahinai</emphasis>
:
<bibRefCitation id="EF8EF3829F5D1C4EFE93FAC3FDB0FA70" author="HSIAO T. Y. &amp; LIU S. L." box="[318,528,1393,1412]" pageId="7" pageNumber="632" pagination="300 - 302" refId="ref19524" refString="HSIAO T. Y. &amp; LIU S. L. 1977: Family Acanthosomatidae. Pp. 159 - 180, 300 - 302, plates 25 - 29. In: HSIAO T. Y., REN S. Z., ZHENG L. Y., JING H. L. &amp; LIU S. L.: A handbook for the determination of the Chinese Hemiptera-Heteroptera. Volume I. Science Press, Beijing, iii + 330 pp, 52 plates (in Chinese, English summary)." type="book chapter" year="1977">HSIAO &amp; LIU (1977: 175)</bibRefCitation>
</taxonomicName>
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(listed, distribution),
<treatmentCitation id="0ABEA8629F5D1C4EFD65FAC3FCF5FA70" author="HUA L. Z." box="[712,853,1393,1412]" page="166" pageId="7" pageNumber="632" year="2000">
<bibRefCitation id="EF8EF3829F5D1C4EFD65FAC3FCF5FA70" author="HUA L. Z." box="[712,853,1393,1412]" pageId="7" pageNumber="632" refId="ref19620" refString="HUA L. Z. 2000: List of Chinese insects. Vol. I. Zhongshan University Press, Guangzhou, 2 + 7 + 448 pp." type="book" year="2000">HUA (2000: 166)</bibRefCitation>
</treatmentCitation>
(listed, distribution), GÖLL- NER- SCHEIDING (2006: 166) (catalogue, distribution),
<treatmentCitation id="0ABEA8629F5D1C4EFDE2FA3EFCBCFA6B" author="ZHENG S. Z. &amp; LIN Y. X." box="[591,796,1420,1439]" page="92" pageId="7" pageNumber="632" year="2013">
<bibRefCitation id="EF8EF3829F5D1C4EFDE2FA3EFCBCFA6B" author="ZHENG S. Z. &amp; LIN Y. X." box="[591,796,1420,1439]" pageId="7" pageNumber="632" refId="ref21903" refString="ZHENG S. Z. &amp; LIN Y. X. 2013: Stinkbug. In: XU H. Y. (ed.): [Taiwan Nature Identification Series]. Vol. 29. Morning Star Publishing, Taipei, 381 pp (in Chinese)." type="book" year="2013">ZHENG &amp; LIN (2013: 92)</bibRefCitation>
</treatmentCitation>
(partim) (redescription, photo).
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</paragraph>
</subSubSection>
<subSubSection id="C305DDF89F5D1C41FFDCFA04FC6FFDEE" lastPageId="8" lastPageNumber="633" pageId="7" pageNumber="632" type="materials_examined">
<paragraph id="8BA08E739F5D1C4EFFDCFA04FBE1FA3D" blockId="7.[113,1089,1306,1614]" box="[113,1089,1457,1482]" pageId="7" pageNumber="632">
<emphasis id="B96B52619F5D1C4EFFDCFA04FEF5FA3D" bold="true" box="[113,341,1462,1481]" pageId="7" pageNumber="632">Type material examined.</emphasis>
<typeStatus id="54A430D19F5D1C4EFEF6FA04FE0EFA33" box="[347,430,1462,1481]" pageId="7" pageNumber="632" type="holotype">HOLOTYPE</typeStatus>
: J,
<date id="FFA1A8B39F5D1C4EFE74FA04FDEEFA3D" box="[473,590,1462,1481]" pageId="7" pageNumber="632" value="1936-08-20">20 VIII 1936</date>
\ MAREPPA \ ---OIWAKE [pink, printed], S. Asahina \
</paragraph>
<paragraph id="8BA08E739F5D1C4EFFDCFA63FBE1FA10" blockId="7.[113,1089,1306,1614]" box="[113,1089,1489,1508]" pageId="7" pageNumber="632">
&lt;collected&gt; [in Japanese script] [handwritten], No. 464 [yellow, printed],
<typeStatus id="54A430D19F5D1C4EFD59FA63FCC9FA10" box="[756,873,1489,1508]" pageId="7" pageNumber="632" type="holotype">HOLOTYPE</typeStatus>
\
<taxonomicName id="4C1FF5F09F5D1C4EFCDAFA63FC4CFA10" authorityName="Curtis" authorityYear="1824" box="[887,1004,1489,1508]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="632" phylum="Arthropoda" rank="genus">Acanthosoma</taxonomicName>
\ asahinai
</paragraph>
<paragraph id="8BA08E739F5D1C4EFFDCFA59FE75F9ED" blockId="7.[113,1089,1306,1614]" pageId="7" pageNumber="632">
\ Ishihara [orange, handwritten]; pinned, segments IIIIV of left and IV of right antenna, right mid leg, tarsus of left hind leg lacking; deposited in
<collectionCode id="ED0E16B69F5D1C4EFE38F9B4FE71F9ED" box="[405,465,1542,1561]" pageId="7" pageNumber="632">ELKU</collectionCode>
.
</paragraph>
<paragraph id="8BA08E739F5D1C4EFFDCF993FBE0F9C0" blockId="7.[113,1089,1306,1614]" box="[113,1088,1563,1588]" pageId="7" pageNumber="632">
<emphasis id="B96B52619F5D1C4EFFDCF993FE85F9C0" bold="true" box="[113,293,1569,1588]" pageId="7" pageNumber="632">Material examined.</emphasis>
<materialsCitation id="3B77842E9F5D1C4EFE86F993FC42F9C0" box="[299,994,1563,1588]" collectingDate="2009-03-18" collectionCode="NCHU" collectorName="S. W. Hou" country="Taiwan" location="Dong Ao Ling" pageId="7" pageNumber="632" specimenCount="1" stateProvince="Yilan">
<emphasis id="B96B52619F5D1C4EFE86F993FE51F9C0" bold="true" box="[299,497,1569,1588]" pageId="7" pageNumber="632">
<collectingCountry id="F308CEE39F5D1C4EFE86F993FE25F9C0" box="[299,389,1569,1588]" name="Taiwan" pageId="7" pageNumber="632">TAIWAN</collectingCountry>
:
<collectingRegion id="49DB40919F5D1C4EFE22F993FE63F9C6" box="[399,451,1569,1588]" country="Taiwan" name="Yilan" pageId="7" pageNumber="632">YILAN</collectingRegion>
CO.:
</emphasis>
<location id="8EC0D8A89F5D1C4EFE55F993FDD8F9C0" LSID="urn:lsid:plazi:treatment:03B63F659F5D1C40FE2BFB49FDB8FDA7:8EC0D8A89F5D1C4EFE55F993FDD8F9C0" box="[504,632,1569,1588]" country="Taiwan" name="Dong Ao Ling" pageId="7" pageNumber="632" stateProvince="Yilan">Dong Ao Ling</location>
,
<date id="FFA1A8B39F5D1C4EFD2EF993FD41F9C0" box="[643,737,1569,1588]" pageId="7" pageNumber="632" value="2009-03-18">
<collectingDate id="EFE5515B9F5D1C4EFD2EF993FD41F9C0" box="[643,737,1569,1588]" pageId="7" pageNumber="632" value="2009-03-18">18.iii.2009</collectingDate>
</date>
, leg.
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(1 J
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)
</materialsCitation>
; Yuanyang
</paragraph>
<paragraph id="8BA08E739F5D1C4EFFDCF989FBE1F9BA" blockId="7.[113,1089,1306,1614]" box="[113,1089,1589,1614]" pageId="7" pageNumber="632">
Lake 100, logging Rd. 16K,
<date id="FFA1A8B39F5D1C4EFEC7F989FE69F9BA" box="[362,457,1595,1614]" pageId="7" pageNumber="632" value="2010-03-29">29.iii.2010</date>
, leg. S.F. Huang &amp; S.J. Huang (
<specimenCount id="9D1945FA9F5D1C4EFD4FF989FCA3F9BA" box="[738,771,1589,1614]" pageId="7" pageNumber="632" type="female">1 ♀</specimenCount>
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).
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<collectingRegion id="49DB40919F5D1C4EFCFAF989FC0DF9B8" box="[855,941,1595,1614]" country="Taiwan" name="Taichung" pageId="7" pageNumber="632">TAICHUNG</collectingRegion>
CO.:
</emphasis>
Anmashan,
</paragraph>
<paragraph id="8BA08E739F521C41FF3AFF50FC6FFDEE" blockId="8.[151,1128,219,538]" pageId="8" pageNumber="633">
<date id="FFA1A8B39F521C41FF3AFF50FF5AFF01" box="[151,250,226,245]" pageId="8" pageNumber="633" value="2002-12-13">13.xii.2002</date>
, leg. C.S. Lin, sweeping net,
<collectionCode id="ED0E16B69F521C41FE55FF50FD96FF01" box="[504,566,226,245]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="8" pageNumber="633">NMNS</collectionCode>
ENT 3973942 (
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).
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<collectingRegion id="49DB40919F521C41FCEFFF50FC28FF07" box="[834,904,226,245]" country="Taiwan" name="Nantou" pageId="8" pageNumber="633">NANTOU</collectingRegion>
CO.:
</emphasis>
Tunyuan,
<date id="FFA1A8B39F521C41FBBDFF50FBC3FF01" box="[1040,1123,226,245]" pageId="8" pageNumber="633" value="2003-06-05">5.vi.2003</date>
, leg. J.F. Tsai (1 J [dissected], 1 J
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<collectionCode id="ED0E16B69F521C41FE42FF4EFD8FFEFB" box="[495,559,252,271]" country="Taiwan" httpUri="http://grbio.org/cool/e7yf-mhkb" name="National Chung Hsing University" pageId="8" pageNumber="633">NCHU</collectionCode>
); Sungk[u]ang,
<date id="FFA1A8B39F521C41FD14FF4EFCBCFEFB" box="[697,796,252,271]" pageId="8" pageNumber="633" value="1985-07-29">29.vii.1985</date>
, leg. T. Hattori, NSMT-I-He 34906 (
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); Huisun Forest Station Emergency rd. to Huisun
<collectingRegion id="49DB40919F521C41FD38FEA5FD62FEDE" box="[661,706,279,298]" country="United Kingdom" name="Isle of Anglesey" pageId="8" pageNumber="633">Mon</collectingRegion>
.,
<date id="FFA1A8B39F521C41FD60FEA5FC8BFEDE" box="[717,811,279,298]" pageId="8" pageNumber="633" value="2010-09-10">10.ix.2010</date>
,
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, pine forest,
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acc. Ti-25a Colls, ex UV light sheet, leg. C. Skinner &amp; W. Hunting (
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); Huisun Experimental Forest Station,
<date id="FFA1A8B39F521C41FEB7FEFEFED7FEAB" box="[282,375,332,351]" pageId="8" pageNumber="633" value="2010-11-12">12.xi.2010</date>
, ex
<taxonomicName id="4C1FF5F09F521C41FE37FEFEFDB3FEAB" box="[410,531,332,351]" class="Magnoliopsida" family="Anacardiaceae" genus="Rhus" kingdom="Plantae" order="Sapindales" pageId="8" pageNumber="633" phylum="Tracheophyta" rank="species" species="javanica">
<emphasis id="B96B52619F521C41FE37FEFEFDB3FEAB" box="[410,531,332,351]" italics="true" pageId="8" pageNumber="633">Rhus javanica</emphasis>
</taxonomicName>
L.
<taxonomicName id="4C1FF5F09F521C41FD9CFEFEFC11FEAB" authority="(DC.) Rehder &amp; E. H. Wilson" authorityName="Rehder &amp; E. H. Wilson" baseAuthorityName="DC." box="[561,945,332,351]" pageId="8" pageNumber="633" rank="variety" variety="roxburghii">
var.
<emphasis id="B96B52619F521C41FDFBFEFEFD10FEAB" box="[598,688,332,351]" italics="true" pageId="8" pageNumber="633">roxburghii</emphasis>
(DC.) Rehder &amp; E.H. Wilson
</taxonomicName>
, leg. Z.Y. Wang (
<specimenCount id="9D1945FA9F521C41FBE5FEFEFBC7FEAB" box="[1096,1127,326,351]" pageId="8" pageNumber="633" type="female">1 ♀</specimenCount>
<collectionCode id="ED0E16B69F521C41FF3AFED5FF76FE8E" box="[151,214,359,378]" country="Taiwan" httpUri="http://grbio.org/cool/e7yf-mhkb" name="National Chung Hsing University" pageId="8" pageNumber="633">NCHU</collectionCode>
, 1 J
<specimenCount id="9D1945FA9F521C41FEAFFED5FE81FE8E" box="[258,289,353,378]" pageId="8" pageNumber="633" type="female">1 ♀</specimenCount>
<collectionCode id="ED0E16B69F521C41FE8BFED5FEC2FE8E" box="[294,354,359,378]" country="Taiwan" httpUri="http://grbio.org/cool/e7yf-mhkb" name="National Chung Hsing University" pageId="8" pageNumber="633">NCHU</collectionCode>
&gt;
<collectionCode id="ED0E16B69F521C41FED4FED5FE14FE8E" box="[377,436,359,378]" pageId="8" pageNumber="633">SEHU</collectionCode>
); Jenai, Chunyang,
<date id="FFA1A8B39F521C41FDF7FED5FD10FE8E" box="[602,688,359,378]" pageId="8" pageNumber="633" value="1995-05-31">31.v.1995</date>
, UV light, leg. M.L. Chan &amp; C.S. Lin, 2005141 (
<specimenCount id="9D1945FA9F521C41FBF0FED5FF07FE60" pageId="8" pageNumber="633" type="female">1 ♀</specimenCount>
<collectionCode id="ED0E16B69F521C41FF00FE30FF4EFE61" box="[173,238,386,405]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="8" pageNumber="633">NMNS</collectionCode>
); Meifeng,
<date id="FFA1A8B39F521C41FEFFFE30FE12FE61" box="[338,434,386,405]" pageId="8" pageNumber="633" value="2001-12-18">18.xii.2001</date>
<date id="FFA1A8B39F521C41FE12FE30FDB6FE61" box="[447,534,386,405]" pageId="8" pageNumber="633" value="2002-02-19">19.ii.2002</date>
, Malaise trap, leg. C.S. Lin &amp; W.T. Yang,
<collectionCode id="ED0E16B69F521C41FC2BFE30FC64FE61" box="[902,964,386,405]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="8" pageNumber="633">NMNS</collectionCode>
ENT 44672 (
<specimenCount id="9D1945FA9F521C41FBEAFE30FBC8FE60" box="[1095,1128,379,405]" pageId="8" pageNumber="633" type="female">1 ♀</specimenCount>
<collectionCode id="ED0E16B69F521C41FF3AFE2EFF78FE5B" box="[151,216,412,431]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="8" pageNumber="633">NMNS</collectionCode>
); Nanshanxi,
<date id="FFA1A8B39F521C41FEE1FE2EFE3DFE5B" box="[332,413,412,431]" pageId="8" pageNumber="633" value="1994-09-06">6.ix.1994</date>
(1 J
<collectionCode id="ED0E16B69F521C41FE60FE2EFDABFE5B" box="[461,523,412,431]" pageId="8" pageNumber="633">OMHJ</collectionCode>
); Shizitou,
<date id="FFA1A8B39F521C41FDC7FE2EFD1FFE5B" box="[618,703,412,431]" pageId="8" pageNumber="633" value="1994-10-18">18.x.1994</date>
(1 J
<specimenCount id="9D1945FA9F521C41FD42FE2EFCAEFE5B" box="[751,782,406,431]" pageId="8" pageNumber="633" type="female">1 ♀</specimenCount>
<collectionCode id="ED0E16B69F521C41FCBEFE2EFCF1FE5B" box="[787,849,412,431]" pageId="8" pageNumber="633">OMHJ</collectionCode>
).
<emphasis id="B96B52619F521C41FCF2FE2EFC6BFE5B" bold="true" box="[863,971,412,431]" pageId="8" pageNumber="633">
<collectingRegion id="49DB40919F521C41FCF2FE2EFC3FFE5A" box="[863,927,412,431]" country="Taiwan" name="Miaoli" pageId="8" pageNumber="633">MIAOLI</collectingRegion>
CO.:
</emphasis>
Simaxian logging Rd.,
<date id="FFA1A8B39F521C41FF12FE05FEB2FE3E" box="[191,274,439,458]" pageId="8" pageNumber="633" value="2010-09-08">8.ix.2010</date>
, leg. Y.C. Liao (1 J
<collectionCode id="ED0E16B69F521C41FE10FE05FE5DFE3E" box="[445,509,439,458]" country="Taiwan" httpUri="http://grbio.org/cool/e7yf-mhkb" name="National Chung Hsing University" pageId="8" pageNumber="633">NCHU</collectionCode>
); Nanzhuang, Luhu,
<date id="FFA1A8B39F521C41FD01FE05FD5FFE3E" box="[684,767,439,458]" pageId="8" pageNumber="633" value="2010-09-09">9.ix.2010</date>
, ex
<taxonomicName id="4C1FF5F09F521C41FCB2FE05FC37FE3E" box="[799,919,439,458]" class="Magnoliopsida" family="Anacardiaceae" genus="Rhus" kingdom="Plantae" order="Sapindales" pageId="8" pageNumber="633" phylum="Tracheophyta" rank="species" species="javanica">
<emphasis id="B96B52619F521C41FCB2FE05FC37FE3E" box="[799,919,439,458]" italics="true" pageId="8" pageNumber="633">Rhus javanica</emphasis>
</taxonomicName>
L.
<taxonomicName id="4C1FF5F09F521C41FC1EFE05FEC1FE11" authority="(DC.) Rehder &amp; E. H. Wilson" authorityName="Rehder &amp; E. H. Wilson" baseAuthorityName="DC." pageId="8" pageNumber="633" rank="variety" variety="roxburghii">
var.
<emphasis id="B96B52619F521C41FC7AFE05FB91FE3E" box="[983,1073,439,458]" italics="true" pageId="8" pageNumber="633">roxburghii</emphasis>
(DC.) Rehder &amp; E.H. Wilson
</taxonomicName>
, leg. Y.C. Liao (1 J
<collectionCode id="ED0E16B69F521C41FDB1FE60FDC1FE11" box="[540,609,466,485]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="8" pageNumber="633">HNHM</collectionCode>
).
<emphasis id="B96B52619F521C41FDDCFE60FCA1FE11" bold="true" box="[625,769,466,485]" pageId="8" pageNumber="633">
<collectingRegion id="49DB40919F521C41FDDCFE60FD73FE17" box="[625,723,466,485]" country="Taiwan" name="Kaohsiung" pageId="8" pageNumber="633">KAOHSIUNG</collectingRegion>
CO.:
</emphasis>
Yushan Nat. Pk.
<date id="FFA1A8B39F521C41FC30FE60FBB4FE11" box="[925,1044,466,485]" pageId="8" pageNumber="633" value="1990-10-24" valueMax="1990-10-28" valueMin="1990-10-24">2428.x.1990</date>
,
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<quantity id="4CE723969F521C41FF40FE5EFEFFFE0B" box="[237,351,492,511]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="8" pageNumber="633" unit="m" value="1850.0" valueMax="1900.0" valueMin="1800.0">18001900m</quantity>
, leg. C.K. Starr &amp; S.S. Lu,
<collectionCode id="ED0E16B69F521C41FDE6FE5EFD29FE0B" box="[587,649,492,511]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="8" pageNumber="633">NMNS</collectionCode>
ENT 76468 (1 J
<collectionCode id="ED0E16B69F521C41FC82FE5EFCD0FE0B" box="[815,880,492,511]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="8" pageNumber="633">NMNS</collectionCode>
).
<emphasis id="B96B52619F521C41FCD2FE5EFBA0FE0B" bold="true" box="[895,1024,492,511]" pageId="8" pageNumber="633">
<collectingRegion id="49DB40919F521C41FCD2FE5EFC73FE0A" box="[895,979,492,511]" country="Taiwan" name="Pingtung" pageId="8" pageNumber="633">PINGTUNG</collectingRegion>
CO.:
</emphasis>
Mt. Dahan, ex
<taxonomicName id="4C1FF5F09F521C41FF1CFDB5FE8AFDEE" box="[177,298,519,538]" class="Magnoliopsida" family="Anacardiaceae" genus="Rhus" kingdom="Plantae" order="Sapindales" pageId="8" pageNumber="633" phylum="Tracheophyta" rank="species" species="javanica">
<emphasis id="B96B52619F521C41FF1CFDB5FE8AFDEE" box="[177,298,519,538]" italics="true" pageId="8" pageNumber="633">Rhus javanica</emphasis>
</taxonomicName>
L.
<taxonomicName id="4C1FF5F09F521C41FEEAFDB5FD65FDEE" authority="(DC.) Rehder &amp; E. H. Wilson" authorityName="Rehder &amp; E. H. Wilson" baseAuthorityName="DC." box="[327,709,519,538]" pageId="8" pageNumber="633" rank="variety" variety="roxburghii">
var.
<emphasis id="B96B52619F521C41FEC1FDB5FE66FDEE" box="[364,454,519,538]" italics="true" pageId="8" pageNumber="633">roxburghii</emphasis>
(DC.) Rehder &amp; E.H. Wilson
</taxonomicName>
, leg. Y.X. Shieh (
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<collectionCode id="ED0E16B69F521C41FC2DFDB5FC65FDEE" box="[896,965,519,538]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="8" pageNumber="633">HNHM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C305DDF89F521C41FF3AFD81FC44FCDE" pageId="8" pageNumber="633" type="diagnosis">
<paragraph id="8BA08E739F521C41FF3AFD81FC44FCDE" blockId="8.[151,1127,563,1610]" pageId="8" pageNumber="633">
<emphasis id="B96B52619F521C41FF3AFD81FEA9FDBE" bold="true" box="[151,265,563,586]" pageId="8" pageNumber="633">Diagnosis.</emphasis>
This species is readily recognized based on the horizontally directed, acute humeral processes of both sexes, the conspicuous subbasal dilation of the hind tibia of the male, and the elongate, posteriorly directed projections of the male genital capsule. The detailed original description (
<bibRefCitation id="EF8EF3829F521C41FEB0FD21FE18FD5E" author="ISHIHARA T." box="[285,440,659,682]" pageId="8" pageNumber="633" pagination="495 - 497" refId="ref19649" refString="ISHIHARA T. 1943: Zwei Acanthosoma-arten der Insel Formosa (Hemiptera: Pentatomidae). Transactions of the Natural History Society of Taiwan 33: 495 - 497." type="journal article" year="1943">ISHIHARA 1943</bibRefCitation>
) allows unambiguous recognition of the species; as an addition, the male (
<figureCitation id="132492F69F521C41FEAEFD01FEC4FD3E" box="[259,356,691,714]" captionStart="Figs 110" captionStartId="10.[151,188,1450,1469]" captionTargetBox="[154,1125,221,1407]" captionTargetId="figure-10@10.[151,1126,221,1420]" captionTargetPageId="10" captionText="Figs 110.Acanthosoma spp., males. 1 A.atayal sp.nov., posterior part of abdomen, ventral view; 2 same, genital capsule and exposed part of segment VIII, ventral view; 3 same, genital capsule, lateral view; 4 A. haemorrhoidale formosanum subsp. nov., holotype, genital capsule, posterior view; 5 A.haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM), genital capsule, posterior view; 6 A. asahinai Ishihara, 1943, posterior part of abdomen, ventral view; 7 same, genital capsule, ventral view; 8 A. fallax sp. nov., posterior part of abdomen, ventral view; 9 same, genital capsule, ventral view; 10 A. pugnax sp. nov., genital capsule and exposed part of segment VIII, ventral view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303573" httpUri="https://zenodo.org/record/5303573/files/figure.png" pageId="8" pageNumber="633">Figs 67</figureCitation>
,
<figureCitation id="132492F69F521C41FEDCFD01FE15FD3E" box="[369,437,691,714]" captionStart="Figs 2044" captionStartId="14.[151,188,1196,1215]" captionTargetBox="[151,1126,231,1174]" captionTargetId="figure-84@14.[151,1126,231,1178]" captionTargetPageId="14" captionText="Figs 2044. Acanthosoma spp., right parameres in different views. 2021 A. atayal sp. nov.; 22 A. haemorrhoidale formosanum subsp. nov., holotype; 23 A. haemorrhoidale ouchii Ishihara, 1950 (China: Sichuan, Mt. Emei, USNM); 24 A. haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM); 2528 A. asahinai Ishihara, 1943; 2932 A. fallax sp. nov.; 3336 A. pugnax sp. nov.; 3739 A. axicia sp. nov.; 4041 A. laevicorne Dallas, 1851; 4244 A. expansum Horváth, 1905. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303577" httpUri="https://zenodo.org/record/5303577/files/figure.png" pageId="8" pageNumber="633">2528</figureCitation>
) and female (
<figureCitation id="132492F69F521C41FDE6FD01FD66FD3E" box="[587,710,691,714]" captionStart="Figs 4551" captionStartId="17.[113,150,1468,1487]" captionTargetBox="[115,1086,506,1453]" captionTargetId="figure-12@17.[113,1088,506,1462]" captionTargetPageId="17" captionText="Figs 4551. Acanthosoma spp. 45 A. atayal sp. nov., female terminalia, posteroventral view; 46 A. haemorrhoidale formosanum subsp. nov., paratype, female terminalia, posteroventral view; 47 A. asahinai Ishihara, 1943, hind tibia of male; 48 same, female terminalia, dorsal view; 49 same, posteroventral view; 50 A. fallax sp. nov., female terminalia, dorsal view; 51 same, posteroventral view. PO6, PO7 = Pendergrasts organs of ventrites VI and VII, respectively. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303579" httpUri="https://zenodo.org/record/5303579/files/figure.png" pageId="8" pageNumber="633">Figs 4849</figureCitation>
) genitalia are described and figured in the present paper. The female of this species is similar in size, colour, and in the presence of sharp humeral processes to that of
<taxonomicName id="4C1FF5F09F521C41FD84FD41FD24FCFE" authorityName="Tsai &amp; Rédei" authorityYear="2015" box="[553,644,755,778]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="633" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="B96B52619F521C41FD84FD41FD24FCFE" box="[553,644,755,778]" italics="true" pageId="8" pageNumber="633">A. fallax</emphasis>
</taxonomicName>
, but the broadly rounded posterior margin of laterotergites VIII sharply differs from the condition found in the latter species.
</paragraph>
</subSubSection>
<subSubSection id="C305DDF89F521C41FF3AFC81FD5EFABE" pageId="8" pageNumber="633" type="description">
<paragraph id="8BA08E739F521C41FF3AFC81FE30FB5E" blockId="8.[151,1127,563,1610]" pageId="8" pageNumber="633">
<emphasis id="B96B52619F521C41FF3AFC81FD22FCBE" bold="true" box="[151,642,819,842]" pageId="8" pageNumber="633">Description of male and female terminalia.</emphasis>
<emphasis id="B96B52619F521C41FD24FC81FD60FCBE" box="[649,704,819,842]" italics="true" pageId="8" pageNumber="633">Male</emphasis>
. Genital capsule (
<figureCitation id="132492F69F521C41FC2EFC81FC43FCBE" box="[899,995,819,842]" captionStart="Figs 110" captionStartId="10.[151,188,1450,1469]" captionTargetBox="[154,1125,221,1407]" captionTargetId="figure-10@10.[151,1126,221,1420]" captionTargetPageId="10" captionText="Figs 110.Acanthosoma spp., males. 1 A.atayal sp.nov., posterior part of abdomen, ventral view; 2 same, genital capsule and exposed part of segment VIII, ventral view; 3 same, genital capsule, lateral view; 4 A. haemorrhoidale formosanum subsp. nov., holotype, genital capsule, posterior view; 5 A.haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM), genital capsule, posterior view; 6 A. asahinai Ishihara, 1943, posterior part of abdomen, ventral view; 7 same, genital capsule, ventral view; 8 A. fallax sp. nov., posterior part of abdomen, ventral view; 9 same, genital capsule, ventral view; 10 A. pugnax sp. nov., genital capsule and exposed part of segment VIII, ventral view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303573" httpUri="https://zenodo.org/record/5303573/files/figure.png" pageId="8" pageNumber="633">Figs 67</figureCitation>
) with a pair of elongate posterolateral projections directed posteriad, running subparallel, slightly curved mesad subapically, apically with a tuft of setae; dorsal rim with a deep, obtuse concavity at midline, provided with several fine setae; ventral infolding with a pair of fine setal tufts and a pair of heavily sclerotized and pigmented denticles laterad of paramere sockets. Paramere (
<figureCitation id="132492F69F521C41FF32FC61FEB6FC1E" box="[159,278,979,1002]" captionStart="Figs 2044" captionStartId="14.[151,188,1196,1215]" captionTargetBox="[151,1126,231,1174]" captionTargetId="figure-84@14.[151,1126,231,1178]" captionTargetPageId="14" captionText="Figs 2044. Acanthosoma spp., right parameres in different views. 2021 A. atayal sp. nov.; 22 A. haemorrhoidale formosanum subsp. nov., holotype; 23 A. haemorrhoidale ouchii Ishihara, 1950 (China: Sichuan, Mt. Emei, USNM); 24 A. haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM); 2528 A. asahinai Ishihara, 1943; 2932 A. fallax sp. nov.; 3336 A. pugnax sp. nov.; 3739 A. axicia sp. nov.; 4041 A. laevicorne Dallas, 1851; 4244 A. expansum Horváth, 1905. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303577" httpUri="https://zenodo.org/record/5303577/files/figure.png" pageId="8" pageNumber="633">Figs 2528</figureCitation>
) T-shaped, dorsal arm with blunt apex, without distinct subapical denticle, ventral arm bird-head-shaped apically.
<emphasis id="B96B52619F521C41FE48FC41FD95FBFE" box="[485,565,1011,1034]" italics="true" pageId="8" pageNumber="633">Female</emphasis>
(
<figureCitation id="132492F69F521C41FDEFFC41FD1FFBFE" box="[578,703,1011,1034]" captionStart="Figs 4551" captionStartId="17.[113,150,1468,1487]" captionTargetBox="[115,1086,506,1453]" captionTargetId="figure-12@17.[113,1088,506,1462]" captionTargetPageId="17" captionText="Figs 4551. Acanthosoma spp. 45 A. atayal sp. nov., female terminalia, posteroventral view; 46 A. haemorrhoidale formosanum subsp. nov., paratype, female terminalia, posteroventral view; 47 A. asahinai Ishihara, 1943, hind tibia of male; 48 same, female terminalia, dorsal view; 49 same, posteroventral view; 50 A. fallax sp. nov., female terminalia, dorsal view; 51 same, posteroventral view. PO6, PO7 = Pendergrasts organs of ventrites VI and VII, respectively. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303579" httpUri="https://zenodo.org/record/5303579/files/figure.png" pageId="8" pageNumber="633">Figs 4849</figureCitation>
). Posterior margin of ventrite VII with a deep, broadly U-shaped incision surrounding valvifers VIII; tergite VIII with dark markings proximally and distally as in
<figureCitation id="132492F69F521C41FE6AFB81FDB5FBBE" box="[455,533,1075,1098]" captionStart="Figs 4551" captionStartId="17.[113,150,1468,1487]" captionTargetBox="[115,1086,506,1453]" captionTargetId="figure-12@17.[113,1088,506,1462]" captionTargetPageId="17" captionText="Figs 4551. Acanthosoma spp. 45 A. atayal sp. nov., female terminalia, posteroventral view; 46 A. haemorrhoidale formosanum subsp. nov., paratype, female terminalia, posteroventral view; 47 A. asahinai Ishihara, 1943, hind tibia of male; 48 same, female terminalia, dorsal view; 49 same, posteroventral view; 50 A. fallax sp. nov., female terminalia, dorsal view; 51 same, posteroventral view. PO6, PO7 = Pendergrasts organs of ventrites VI and VII, respectively. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303579" httpUri="https://zenodo.org/record/5303579/files/figure.png" pageId="8" pageNumber="633">Fig. 48</figureCitation>
; posterior margin of laterotergites VIII broadly rounded, far surpassing posterolateral angles of abdominal ventrite VII, mesal portion (surrounding postgenital segments) with dark marking; Pendergrasts organs of ventrites VI and VII subelliptical, of equal size.
</paragraph>
<paragraph id="8BA08E739F521C41FF3AFB01FBACFB1E" blockId="8.[151,1127,563,1610]" pageId="8" pageNumber="633">
<emphasis id="B96B52619F521C41FF3AFB01FE9EFB3E" bold="true" box="[151,318,1203,1226]" pageId="8" pageNumber="633">Measurements</emphasis>
(in mm). Body length to apex of membrane 14.315.9 (J) / 15.917.2 (
<specimenCount id="9D1945FA9F521C41FBEEFB1EFBF6FB3F" box="[1091,1110,1196,1227]" pageId="8" pageNumber="633" type="female"></specimenCount>
); to imaginary line connecting apices of projections of genital capsule 15.217.1 (J)
</paragraph>
<paragraph id="8BA08E739F521C41FF3AFB41FD5EFABE" blockId="8.[151,1127,563,1610]" pageId="8" pageNumber="633">
<emphasis id="B96B52619F521C41FF3AFB41FEB5FAFE" bold="true" box="[151,277,1267,1290]" pageId="8" pageNumber="633">Bionomics.</emphasis>
Adults, larvae and eggs of this species are frequently observed on
<taxonomicName id="4C1FF5F09F521C41FC7DFB41FBC7FAFE" box="[976,1127,1267,1290]" class="Magnoliopsida" family="Anacardiaceae" genus="Rhus" kingdom="Plantae" order="Sapindales" pageId="8" pageNumber="633" phylum="Tracheophyta" rank="species" species="javanica">
<emphasis id="B96B52619F521C41FC7DFB41FBC7FAFE" box="[976,1127,1267,1290]" italics="true" pageId="8" pageNumber="633">Rhus javanica</emphasis>
</taxonomicName>
L.
<taxonomicName id="4C1FF5F09F521C41FF19FAA1FC9FFADE" authority="(DC.) Rehder &amp; E. H. Wilson" authorityName="Rehder &amp; E. H. Wilson" baseAuthorityName="DC." box="[180,831,1299,1322]" pageId="8" pageNumber="633" rank="variety" variety="roxburghii">
var.
<emphasis id="B96B52619F521C41FF4CFAA1FEF1FADE" box="[225,337,1299,1322]" italics="true" pageId="8" pageNumber="633">roxburghii</emphasis>
(DC.) Rehder &amp; E.H. Wilson (Anacardiaceae)
</taxonomicName>
which is apparently its host plant. The breeding season is from September to October.
</paragraph>
</subSubSection>
<subSubSection id="C305DDF89F521C41FF3AFAE1FE86FA5E" pageId="8" pageNumber="633" type="distribution">
<paragraph id="8BA08E739F521C41FF3AFAE1FE86FA5E" blockId="8.[151,1127,563,1610]" pageId="8" pageNumber="633">
<emphasis id="B96B52619F521C41FF3AFAE1FE8AFA9E" bold="true" box="[151,298,1363,1386]" pageId="8" pageNumber="633">Distribution.</emphasis>
The species is apparently endemic to
<collectingCountry id="F308CEE39F521C41FD61FAE1FCBAFA9E" box="[716,794,1363,1386]" name="Taiwan" pageId="8" pageNumber="633">Taiwan</collectingCountry>
and it is relatively frequent in the
<emphasis id="B96B52619F521C41FF13FAC1FE0CFA7E" box="[190,428,1395,1418]" italics="true" pageId="8" pageNumber="633">
<taxonomicName id="4C1FF5F09F521C41FF13FAC1FE81FA7E" box="[190,289,1395,1418]" class="Magnoliopsida" family="Lauraceae" genus="Machilus" kingdom="Plantae" order="Laurales" pageId="8" pageNumber="633" phylum="Tracheophyta" rank="genus">Machilus</taxonomicName>
-
<taxonomicName id="4C1FF5F09F521C41FE87FAC1FE0CFA7E" box="[298,428,1395,1418]" class="Magnoliopsida" family="Fagaceae" genus="Castanopsis" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="633" phylum="Tracheophyta" rank="genus">Castanopsis</taxonomicName>
</emphasis>
forest zone of the moutainous regions ranging between
<quantity id="4CE723969F521C41FC51FAC1FF0CFA5E" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.8" metricValueMin="0.5" pageId="8" pageNumber="633" unit="m" value="1150.0" valueMax="1800.0" valueMin="500.0">5001800 m</quantity>
in altitude.
</paragraph>
</subSubSection>
<subSubSection id="C305DDF89F521C40FF3AFA01FDB8FDA7" lastPageId="9" lastPageNumber="634" pageId="8" pageNumber="633" type="discussion">
<paragraph id="8BA08E739F521C41FF3AFA01FD45F9FE" blockId="8.[151,1127,563,1610]" pageId="8" pageNumber="633">
<emphasis id="B96B52619F521C41FF3AFA01FEA8FA3E" bold="true" box="[151,264,1459,1482]" pageId="8" pageNumber="633">Remarks.</emphasis>
<taxonomicName id="4C1FF5F09F521C41FEBFFA01FDAEFA3E" authorityName=": HSIAO &amp; LIU" authorityYear="1977" box="[274,526,1459,1482]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="633" phylum="Arthropoda" rank="species" species="asahinai">
<emphasis id="B96B52619F521C41FEBFFA01FDAEFA3E" box="[274,526,1459,1482]" italics="true" pageId="8" pageNumber="633">Acanthosoma asahinai</emphasis>
</taxonomicName>
was described based on a single male from Mareppa-Oiwake (now Renai, Tsuifeng) in
<collectingCountry id="F308CEE39F521C41FD86FA61FDDDFA1E" box="[555,637,1491,1514]" name="Taiwan" pageId="8" pageNumber="633">Taiwan</collectingCountry>
. The identity of the species was confirmed by reexamination of the
<typeStatus id="54A430D19F521C41FED6FA41FE78F9FE" box="[379,472,1523,1546]" pageId="8" pageNumber="633" type="holotype">holotype</typeStatus>
during the present study.
</paragraph>
<paragraph id="8BA08E739F521C40FF18F9A1FC0CFEE7" blockId="8.[151,1127,563,1610]" lastBlockId="9.[113,1089,220,595]" lastPageId="9" lastPageNumber="634" pageId="8" pageNumber="633">
Except of
<bibRefCitation id="EF8EF3829F521C41FEB2F9A1FE51F9DE" author="ZHENG S. Z. &amp; LIN Y. X." box="[287,497,1555,1578]" pageId="8" pageNumber="633" refId="ref21903" refString="ZHENG S. Z. &amp; LIN Y. X. 2013: Stinkbug. In: XU H. Y. (ed.): [Taiwan Nature Identification Series]. Vol. 29. Morning Star Publishing, Taipei, 381 pp (in Chinese)." type="book" year="2013">ZHENG &amp; LIN (2013)</bibRefCitation>
no subsequent authors provided original data on the species. The top left photo in p. 92 of
<bibRefCitation id="EF8EF3829F521C41FE7DF981FD08F9BE" author="ZHENG S. Z. &amp; LIN Y. X." box="[464,680,1587,1610]" pageId="8" pageNumber="633" refId="ref21903" refString="ZHENG S. Z. &amp; LIN Y. X. 2013: Stinkbug. In: XU H. Y. (ed.): [Taiwan Nature Identification Series]. Vol. 29. Morning Star Publishing, Taipei, 381 pp (in Chinese)." type="book" year="2013">ZHENG &amp; LIN (2013)</bibRefCitation>
shows
<taxonomicName id="4C1FF5F09F521C41FD5AF981FCD0F9BE" authorityName=": HSIAO &amp; LIU" authorityYear="1977" box="[759,880,1587,1610]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="633" phylum="Arthropoda" rank="species" species="asahinai">
<emphasis id="B96B52619F521C41FD5AF981FCD0F9BE" box="[759,880,1587,1610]" italics="true" pageId="8" pageNumber="633">A. asahinai</emphasis>
</taxonomicName>
but the top right one is
<taxonomicName id="4C1FF5F09F531C40FFDCFF6EFF6BFF07" authority="Tsai &amp; Rédei, 2015" authorityName="Tsai &amp; Rédei" authorityYear="2015" box="[113,203,220,243]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="634" phylum="Arthropoda" rank="species" species="fallax" status="sp. nov.">
<emphasis id="B96B52619F531C40FFDCFF6EFF6BFF07" box="[113,203,220,243]" italics="true" pageId="9" pageNumber="634">A. fallax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A258EF1A9F531C40FF7DFF6EFE84FF07" box="[208,292,220,243]" pageId="9" pageNumber="634" rank="species">sp. nov.</taxonomicNameLabel>
; their redescription of
<taxonomicName id="4C1FF5F09F531C40FDB4FF6EFD32FF07" authorityName=": HSIAO &amp; LIU" authorityYear="1977" box="[537,658,220,243]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="634" phylum="Arthropoda" rank="species" species="asahinai">
<emphasis id="B96B52619F531C40FDB4FF6EFD32FF07" box="[537,658,220,243]" italics="true" pageId="9" pageNumber="634">A. asahinai</emphasis>
</taxonomicName>
includes characters of both species. The male is redescribed and the female is described for the first time in this paper.
</paragraph>
<paragraph id="8BA08E739F531C40FF3DFEAEFDB8FDA7" blockId="9.[113,1089,220,595]" pageId="9" pageNumber="634">
The new species belongs to the
<taxonomicName id="4C1FF5F09F531C40FE4AFEAEFD1DFEC7" authorityName="Dallas" authorityYear="1851" box="[487,701,284,307]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="634" phylum="Arthropoda" rank="species" species="forfex">
<emphasis id="B96B52619F531C40FE4AFEAEFD1DFEC7" box="[487,701,284,307]" italics="true" pageId="9" pageNumber="634">Acanthosoma forfex</emphasis>
</taxonomicName>
species group hereby defined in the following way: genital capsule with a pair of greatly elongate posterolateral projections; paramere T-shaped, with a long dorsal and a somewhat shorter ventral arm. The group comprises the following species:
<taxonomicName id="4C1FF5F09F531C40FE75FECEFDF2FE67" authorityName=": HSIAO &amp; LIU" authorityYear="1977" box="[472,594,380,403]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="634" phylum="Arthropoda" rank="species" species="asahinai">
<emphasis id="B96B52619F531C40FE75FECEFDF2FE67" box="[472,594,380,403]" italics="true" pageId="9" pageNumber="634">A. asahinai</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1FF5F09F531C40FDCDFECEFC26FE67" authority="Reuter, 1881" authorityName="Reuter" authorityYear="1881" box="[608,902,380,403]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="634" phylum="Arthropoda" rank="species" species="forcipatum">
<emphasis id="B96B52619F531C40FDCDFECEFD53FE67" box="[608,755,380,403]" italics="true" pageId="9" pageNumber="634">A. forcipatum</emphasis>
<bibRefCitation id="EF8EF3829F531C40FD57FECEFC26FE67" author="REUTER O. M." box="[762,902,380,403]" pageId="9" pageNumber="634" pagination="67 - 86" refId="ref20954" refString="REUTER O. M. 1881: Acanthosomina et Urolabidina nova et minus cognita. Berliner Entomologische Zeitschrift 25: 67 - 86." type="journal article" year="1881">Reuter, 1881</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1FF5F09F531C40FC3FFECEFF09FE47" authority="Dallas, 1851" authorityName="Dallas" authorityYear="1851" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="634" phylum="Arthropoda" rank="species" species="forfex">
<emphasis id="B96B52619F531C40FC3FFECEFC4EFE67" box="[914,1006,380,403]" italics="true" pageId="9" pageNumber="634">A. forfex</emphasis>
<bibRefCitation id="EF8EF3829F531C40FC5BFECEFF09FE47" author="DALLAS W. S." pageId="9" pageNumber="634" refId="ref18627" refString="DALLAS W. S. 1851: List of the specimens of hemipterous insects in the collection of the British Museum. Part 1. Trustees of the British Museum, London, 368 pp, XI plates." type="book" year="1851">Dallas, 1851</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1FF5F09F531C40FF19FE2EFE74FE47" authority="Jakovlev, 1880" authorityName="Jakovlev" authorityYear="1880" box="[180,468,412,435]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="634" phylum="Arthropoda" rank="species" species="forficula">
<emphasis id="B96B52619F531C40FF19FE2EFE8CFE47" box="[180,300,412,435]" italics="true" pageId="9" pageNumber="634">A. forficula</emphasis>
<bibRefCitation id="EF8EF3829F531C40FE9FFE2EFE74FE47" author="JAKOVLEV V. E." box="[306,468,412,435]" pageId="9" pageNumber="634" pagination="385 - 398" refId="ref19702" refString="JAKOVLEV V. E. 1880: Contributions to the fauna of bugs of Russia and the neighbouring countries. IV. Bulletin de la Societe Imperiale des Naturalistes de Moscou 55 (2): 385 - 398 (in Russian and German)." type="journal article" year="1880">Jakovlev, 1880</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1FF5F09F531C40FE72FE2EFC33FE47" authority="Yamamoto &amp; Hayashi, 2011" authorityName="Yamamoto &amp; Hayashi" authorityYear="2011" box="[479,915,412,435]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="634" phylum="Arthropoda" rank="species" species="ishiharai">
<emphasis id="B96B52619F531C40FE72FE2EFDFCFE47" box="[479,604,412,435]" italics="true" pageId="9" pageNumber="634">A. ishiharai</emphasis>
<bibRefCitation id="EF8EF3829F531C40FDCFFE2EFC33FE47" author="YAMAMOTO A. &amp; HAYASHI M." box="[610,915,412,435]" pageId="9" pageNumber="634" pagination="145 - 152" refId="ref21476" refString="YAMAMOTO A. &amp; HAYASHI M. 2011: A new species of the genus Acanthosoma from Japan (Heteroptera, Acanthosomatidae). Japanese Journal of Systematic Entomology 17: 145 - 152." type="journal article" year="2011">Yamamoto &amp; Hayashi, 2011</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1FF5F09F531C40FC30FE2EFEB5FE27" authority="Jakovlev, 1880" authorityName="Jakovlev" authorityYear="1880" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="634" phylum="Arthropoda" rank="species" species="labiduroides">
<emphasis id="B96B52619F531C40FC30FE2EFBE1FE47" box="[925,1089,412,435]" italics="true" pageId="9" pageNumber="634">A. labiduroides</emphasis>
<bibRefCitation id="EF8EF3829F531C40FFDCFE0EFEB5FE27" author="JAKOVLEV V. E." box="[113,277,444,467]" pageId="9" pageNumber="634" pagination="385 - 398" refId="ref19702" refString="JAKOVLEV V. E. 1880: Contributions to the fauna of bugs of Russia and the neighbouring countries. IV. Bulletin de la Societe Imperiale des Naturalistes de Moscou 55 (2): 385 - 398 (in Russian and German)." type="journal article" year="1880">Jakovlev, 1880</bibRefCitation>
</taxonomicName>
, the newly described
<taxonomicName id="4C1FF5F09F531C40FDAEFE0EFDFFFE27" authority="Tsai &amp; Rédei, 2015" authorityName="Tsai &amp; Rédei" authorityYear="2015" box="[515,607,444,467]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="634" phylum="Arthropoda" rank="species" species="fallax" status="sp. nov.">
<emphasis id="B96B52619F531C40FDAEFE0EFDFFFE27" box="[515,607,444,467]" italics="true" pageId="9" pageNumber="634">A. fallax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A258EF1A9F531C40FDCAFE0EFD1EFE27" box="[615,702,444,467]" pageId="9" pageNumber="634" rank="species">sp. nov.</taxonomicNameLabel>
, and perhaps
<taxonomicName id="4C1FF5F09F531C40FCF9FE0EFF10FE07" authority="(Distant, 1900)" baseAuthorityName="Distant" baseAuthorityYear="1900" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="634" phylum="Arthropoda" rank="species" species="hampsoni">
<emphasis id="B96B52619F531C40FCF9FE0EFC7CFE27" box="[852,988,444,467]" italics="true" pageId="9" pageNumber="634">A. hampsoni</emphasis>
(
<bibRefCitation id="EF8EF3829F531C40FC46FE0EFF08FE07" author="DISTANT W. L." pageId="9" pageNumber="634" pagination="220 - 234" refId="ref18825" refString="DISTANT W. L. 1900: Rhynchotal notes. - VI. Heteroptera: Dinidorinae, Phyllocephalinae, Urolabidinae, and Acanthosominae. Annals and Magazine of Natural History, Series 7 6: 220 - 234." type="journal article" year="1900">Distant, 1900</bibRefCitation>
)
</taxonomicName>
(the latter one doubtfully associated).
<taxonomicName id="4C1FF5F09F531C40FDF5FE6EFCEEFE07" authorityName=": HSIAO &amp; LIU" authorityYear="1977" box="[600,846,476,499]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="634" phylum="Arthropoda" rank="species" species="asahinai">
<emphasis id="B96B52619F531C40FDF5FE6EFCEEFE07" box="[600,846,476,499]" italics="true" pageId="9" pageNumber="634">Acanthosoma asahinai</emphasis>
</taxonomicName>
is apparently sister to
<taxonomicName id="4C1FF5F09F531C40FFDCFE4EFEB5FDE7" authorityName="Jakovlev" authorityYear="1880" box="[113,277,508,531]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="634" phylum="Arthropoda" rank="species" species="labiduroides">
<emphasis id="B96B52619F531C40FFDCFE4EFEB5FDE7" box="[113,277,508,531]" italics="true" pageId="9" pageNumber="634">A. labiduroides</emphasis>
</taxonomicName>
based on the high similarity of the genital capsule and female terminalia. A study on the molecular phylogeny of the family also supports the sister relationship of the two species (TSAI et al., in preparation).
</paragraph>
</subSubSection>
</treatment>
</document>