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<document id="5AEDC6D00B4E42BC2F95C71883AAC5C1" ID-DOI="10.5281/zenodo.5303571" ID-ISSN="0374-1036" ID-Zenodo-Dep="5303571" ID-ZooBank="0360453D-E09E-4CC4-9BB0-981AEC7BED66S" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630104360660" checkinUser="marcus" docAuthor="Tsai, Jing-Fu &amp; Rédei, Dávid" docDate="2015" docId="03B63F659F571C46FE1EFE89FD09FC6D" docLanguage="en" docName="ActaEntMusNatPra.2015.55.2.625-664.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 55 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Acanthosoma pugnax Tsai &amp; Rédei 2015, sp. nov." docType="treatment" docVersion="4" lastPageNumber="640" masterDocId="FF8F471D9F5A1C49FFADFFB2FFA0FFF4" masterDocTitle="The genus Acanthosoma in Taiwan (Hemiptera: Heteroptera: Acanthosomatidae)" masterLastPageNumber="664" masterPageNumber="625" pageNumber="638" updateTime="1699153447944" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="AF4CA66DF1EEF98553D7027EA4A96CA8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="49D3739A8F7F0BF1C2D4A5B4B0A482F0">
<mods:title id="82932F341E06B8DA30EE77C202B8C472">The genus Acanthosoma in Taiwan (Hemiptera: Heteroptera: Acanthosomatidae)</mods:title>
</mods:titleInfo>
<mods:name id="CEE39B97A29FB726096D882D2D35C192" type="personal">
<mods:role id="D92B54144161FAAF6CC66EFBD9A94B66">
<mods:roleTerm id="3C7CFF86522FFA0F039403D4166F20FF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="56955E9B94F51908777A57297FFCE8AD">Tsai, Jing-Fu</mods:namePart>
<mods:affiliation id="AA188D045D40FB359E45728881606412">Systematic Entomology, Graduate School of Agriculture, Hokkaido University, Sapporo, 060 - 8589 Japan; e-mail: jingfu. tsai @ gmail. com</mods:affiliation>
</mods:name>
<mods:name id="F9F2EAA1EE436B2DC71027761CC4FA43" type="personal">
<mods:role id="9E47C84D899C32612445FE7049CC276A">
<mods:roleTerm id="927321FAE486B64C009E2A01FD1E1D97">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="62D46620D03D6FC148AB3C06598FE6AE">Rédei, Dávid</mods:namePart>
<mods:affiliation id="708288CCE3B8596CF5633D7D60AAA3B4">Institute of Entomology, College of Life Sciences, Nankai University, Weijin Road 94, 300071 Tianjin, China; e-mail: david. redei @ gmail. com &amp; Department of Zoology, Hungarian Natural History Museum, H- 1088 Budapest, Baross u. 13, Hungary</mods:affiliation>
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<mods:typeOfResource id="F82766C5DE1C42DC0FCEB395B58319D4">text</mods:typeOfResource>
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<mods:title id="7CC23F93EF6F536654A00B3521188FF7">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="1E6DFD8D02AAB4FF20D3417248F488EC">
<mods:date id="C1F2076FAC62E6875FA10D24B18884C0">2015</mods:date>
<mods:detail id="686225389F2E3EDF88916C2417846E0F" type="pubDate">
<mods:number id="55E40BB9C9F7D0EFFCC902F856D6A283">2015-12-31</mods:number>
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<mods:detail id="064E20D3AE85B06285D72843314D0C20" type="volume">
<mods:number id="7ACB39AA6B25F60046FD12501E9CB38D">55</mods:number>
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<mods:number id="2DB0B84654222F106EE9F7FEF0AC787D">2</mods:number>
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<mods:classification id="A0D02A0F8953BC4F7FFF5FF85162A827">journal article</mods:classification>
<mods:identifier id="F07FC2E5BB011E3F8D57E615C4D66F7C" type="DOI">10.5281/zenodo.5303571</mods:identifier>
<mods:identifier id="E4577BCE8CAEB75D830ED8680597B50F" type="ISSN">0374-1036</mods:identifier>
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<treatment id="03B63F659F571C46FE1EFE89FD09FC6D" ID-DOI="http://doi.org/10.5281/zenodo.5331609" ID-Zenodo-Dep="5331609" LSID="urn:lsid:plazi:treatment:03B63F659F571C46FE1EFE89FD09FC6D" httpUri="http://treatment.plazi.org/id/03B63F659F571C46FE1EFE89FD09FC6D" lastPageId="15" lastPageNumber="640" pageId="13" pageNumber="638">
<subSubSection id="C305DDF89F571C44FE1EFE89FCA0FEA7" box="[435,768,315,339]" pageId="13" pageNumber="638" type="nomenclature">
<paragraph id="8BA08E739F571C44FE1EFE89FCA0FEA7" blockId="13.[435,768,315,339]" box="[435,768,315,339]" pageId="13" pageNumber="638">
<heading id="D0E8391F9F571C44FE1EFE89FCA0FEA7" bold="true" box="[435,768,315,339]" centered="true" fontSize="8" level="2" pageId="13" pageNumber="638" reason="3">
<emphasis id="B96B52619F571C44FE1EFE89FCA0FEA7" bold="true" box="[435,768,315,339]" pageId="13" pageNumber="638">
<taxonomicName id="4C1FF5F09F571C44FE1EFE89FD03FEA7" authority="Tsai &amp; Rédei, 2015" authorityName="Tsai &amp; Rédei" authorityYear="2015" box="[435,675,315,339]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="638" phylum="Arthropoda" rank="species" species="pugnax" status="sp. nov.">
<emphasis id="B96B52619F571C44FE1EFE89FD03FEA7" bold="true" box="[435,675,315,339]" italics="true" pageId="13" pageNumber="638">Acanthosoma pugnax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A258EF1A9F571C44FD07FE8EFCA0FEA7" box="[682,768,316,339]" pageId="13" pageNumber="638" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C305DDF89F571C44FE4FFEE8FB8BFE38" pageId="13" pageNumber="638" type="description">
<paragraph id="8BA08E739F571C44FE4FFEE8FB8BFE38" blockId="13.[113,1090,346,460]" pageId="13" pageNumber="638">
(
<figureCitation id="132492F69F571C44FE45FEE8FD8AFE99" box="[488,554,346,365]" captionStart="Figs 1–10" captionStartId="10.[151,188,1450,1469]" captionTargetBox="[154,1125,221,1407]" captionTargetId="figure-10@10.[151,1126,221,1420]" captionTargetPageId="10" captionText="Figs 1–10.Acanthosoma spp., males. 1 – A.atayal sp.nov., posterior part of abdomen, ventral view; 2 – same, genital capsule and exposed part of segment VIII, ventral view; 3 – same, genital capsule, lateral view; 4 – A. haemorrhoidale formosanum subsp. nov., holotype, genital capsule, posterior view; 5 – A.haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM), genital capsule, posterior view; 6 – A. asahinai Ishihara, 1943, posterior part of abdomen, ventral view; 7 – same, genital capsule, ventral view; 8 – A. fallax sp. nov., posterior part of abdomen, ventral view; 9 – same, genital capsule, ventral view; 10 – A. pugnax sp. nov., genital capsule and exposed part of segment VIII, ventral view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303573" httpUri="https://zenodo.org/record/5303573/files/figure.png" pageId="13" pageNumber="638">Figs 10</figureCitation>
,
<figureCitation id="132492F69F571C44FD9EFEE8FDCBFE99" box="[563,619,346,365]" captionStart="Figs 20–44" captionStartId="14.[151,188,1196,1215]" captionTargetBox="[151,1126,231,1174]" captionTargetId="figure-84@14.[151,1126,231,1178]" captionTargetPageId="14" captionText="Figs 20–44. Acanthosoma spp., right parameres in different views. 20–21 – A. atayal sp. nov.; 22 – A. haemorrhoidale formosanum subsp. nov., holotype; 23 – A. haemorrhoidale ouchii Ishihara, 1950 (China: Sichuan, Mt. Emei, USNM); 24 – A. haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM); 25–28 – A. asahinai Ishihara, 1943; 29–32 – A. fallax sp. nov.; 33–36 – A. pugnax sp. nov.; 37–39 – A. axicia sp. nov.; 40–41 – A. laevicorne Dallas, 1851; 42–44 – A. expansum Horváth, 1905. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303577" httpUri="https://zenodo.org/record/5303577/files/figure.png" pageId="13" pageNumber="638">33–36</figureCitation>
,
<figureCitation id="132492F69F571C44FDDEFEE8FD2AFE99" box="[627,650,346,365]" captionStart="Figs 52–55" captionStartId="18.[151,188,1522,1541]" captionTargetBox="[151,1124,840,1491]" captionTargetId="figure-10@18.[151,1126,839,1496]" captionTargetPageId="18" captionText="Figs 52–55. Acanthosoma spp., female terminalia, posteroventral view. 52 – A. axicia sp. nov.; 53 – A. pugnax sp. nov.; 54 – A. laevicorne Dallas, 1851; 55 – A. expansum Horváth, 1905. PO6, PO7 = Pendergrast’s organs of ventrites VI and VII, respectively. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303581" httpUri="https://zenodo.org/record/5303581/files/figure.png" pageId="13" pageNumber="638">53</figureCitation>
,
<figureCitation id="132492F69F571C44FD3EFEE8FD6AFE99" box="[659,714,346,365]" captionStart="Figs 72–75" captionStartId="26.[151,188,1547,1566]" captionTargetBox="[194,1071,246,1496]" captionTargetId="figure-10@26.[151,1126,217,1516]" captionTargetPageId="26" captionText="Figs 72–75. Acanthosoma pugnax sp. nov. 72–73 – male (holotype); 74–75 – female (paratype). Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303591" httpUri="https://zenodo.org/record/5303591/files/figure.png" pageId="13" pageNumber="638">72–75</figureCitation>
)
<materialsCitation id="3B77842E9F571C44FFDCFE36FBAEFE63" box="[113,1038,383,408]" collectingDate="2013-04-15" collectorName="S. Wu" country="Taiwan" location="Wuling Hostel" pageId="13" pageNumber="638" specimenCount="1" stateProvince="Taichung" typeStatus="holotype">
<emphasis id="B96B52619F571C44FFDCFE36FF57FE63" bold="true" box="[113,247,388,407]" pageId="13" pageNumber="638">Type material.</emphasis>
<typeStatus id="54A430D19F571C44FF50FE36FEF0FE61" box="[253,336,388,407]" pageId="13" pageNumber="638" type="holotype">HOLOTYPE</typeStatus>
: J,
<emphasis id="B96B52619F571C44FEDAFE36FDFEFE63" bold="true" box="[375,606,388,407]" pageId="13" pageNumber="638">
<collectingCountry id="F308CEE39F571C44FEDAFE36FE71FE63" box="[375,465,388,407]" name="Taiwan" pageId="13" pageNumber="638">TAIWAN</collectingCountry>
:
<collectingRegion id="49DB40919F571C44FE76FE36FD91FE61" box="[475,561,388,407]" country="Taiwan" name="Taichung" pageId="13" pageNumber="638">TAICHUNG</collectingRegion>
CO.:
</emphasis>
<location id="8EC0D8A89F571C44FDC9FE36FD44FE63" LSID="urn:lsid:plazi:treatment:03B63F659F571C46FE1EFE89FD09FC6D:8EC0D8A89F571C44FDC9FE36FD44FE63" box="[612,740,388,407]" country="Taiwan" name="Wuling Hostel" pageId="13" pageNumber="638" stateProvince="Taichung">Wuling Hostel</location>
,
<date id="FFA1A8B39F571C44FD40FE36FCE8FE63" box="[749,840,388,407]" pageId="13" pageNumber="638" value="2013-04-15">
<collectingDate id="EFE5515B9F571C44FD40FE36FCE8FE63" box="[749,840,388,407]" pageId="13" pageNumber="638" value="2013-04-15">15.iv.2013</collectingDate>
</date>
, MV lamp, leg.
<collectorName id="26EAEBA59F571C44FC7BFE36FBAEFE63" box="[982,1038,388,407]" pageId="13" pageNumber="638">S. Wu</collectorName>
</materialsCitation>
; male genitalia dissected, preserved in glass vial with glycerol, pinned with the specimen;
<materialsCitation id="3B77842E9F571C44FC8BFE2DFB87FE38" collectingDate="1979-07-06" collectingDateMax="1979-07-09" collectingDateMin="1979-07-06" collectionCode="NMNS, TARI" collectorName="W. J. Wu" country="Taiwan" elevation="2275" location="Anmashan" pageId="13" pageNumber="638" specimenCount="1" specimenCount-female="1" stateProvince="Taichung" typeStatus="paratype">
deposited in
<collectionCode id="ED0E16B69F571C44FC22FE2DFC6CFE46" box="[911,972,415,434]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="13" pageNumber="638">NMNS</collectionCode>
(
<figureCitation id="132492F69F571C44FC7BFE2DFB98FE46" box="[982,1080,415,434]" captionStart="Figs 72–75" captionStartId="26.[151,188,1547,1566]" captionTargetBox="[194,1071,246,1496]" captionTargetId="figure-10@26.[151,1126,217,1516]" captionTargetPageId="26" captionText="Figs 72–75. Acanthosoma pugnax sp. nov. 72–73 – male (holotype); 74–75 – female (paratype). Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303591" httpUri="https://zenodo.org/record/5303591/files/figure.png" pageId="13" pageNumber="638">Figs 72–73</figureCitation>
).
<typeStatus id="54A430D19F571C44FFDCFE0BFF60FE3F" box="[113,192,441,460]" pageId="13" pageNumber="638" type="paratype">PARATYPE</typeStatus>
:
<emphasis id="B96B52619F571C44FF61FE0BFE11FE38" bold="true" box="[204,433,441,460]" pageId="13" pageNumber="638">
<collectingCountry id="F308CEE39F571C44FF61FE0BFE86FE38" box="[204,294,441,460]" name="Taiwan" pageId="13" pageNumber="638">TAIWAN</collectingCountry>
:
<collectingRegion id="49DB40919F571C44FE83FE0BFE24FE3F" box="[302,388,441,460]" country="Taiwan" name="Taichung" pageId="13" pageNumber="638">TAICHUNG</collectingRegion>
CO.:
</emphasis>
<location id="8EC0D8A89F571C44FE1BFE0BFDB3FE38" LSID="urn:lsid:plazi:treatment:03B63F659F571C46FE1EFE89FD09FC6D:8EC0D8A89F571C44FE1BFE0BFDB3FE38" box="[438,531,441,460]" country="Taiwan" name="Anmashan" pageId="13" pageNumber="638" stateProvince="Taichung">Anmashan</location>
,
<quantity id="4CE723969F571C44FDB1FE0BFDFFFE38" box="[540,607,441,460]" metricMagnitude="3" metricUnit="m" metricValue="2.275" pageId="13" pageNumber="638" unit="m" value="2275.0">
<elevation id="003269409F571C44FDB1FE0BFDFFFE38" box="[540,607,441,460]" metricMagnitude="3" metricUnit="m" metricValue="2.275" pageId="13" pageNumber="638" unit="m" value="2275.0">2275 m</elevation>
</quantity>
,
<date id="FFA1A8B39F571C44FDCAFE0BFD75FE38" box="[615,725,441,460]" pageId="13" pageNumber="638" value="1979-07-06" valueMax="1979-07-09" valueMin="1979-07-06">
<collectingDate id="EFE5515B9F571C44FDCAFE0BFD75FE38" box="[615,725,441,460]" pageId="13" pageNumber="638" value="1979-07-06" valueMax="1979-07-09" valueMin="1979-07-06">6–9.vii.1979</collectingDate>
</date>
, leg.
<collectorName id="26EAEBA59F571C44FCAFFE0BFCEAFE38" box="[770,842,441,460]" pageId="13" pageNumber="638">W.J. Wu</collectorName>
(
<specimenCount id="9D1945FA9F571C44FCFAFE0BFCD6FE38" box="[855,886,435,460]" pageId="13" pageNumber="638" type="female">1 ♀</specimenCount>
<collectionCode id="ED0E16B69F571C44FCD6FE0BFC0DFE38" box="[891,941,441,460]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="13" pageNumber="638">TARI</collectionCode>
) (
<figureCitation id="132492F69F571C44FC10FE0BFB81FE38" box="[957,1057,441,460]" captionStart="Figs 72–75" captionStartId="26.[151,188,1547,1566]" captionTargetBox="[194,1071,246,1496]" captionTargetId="figure-10@26.[151,1126,217,1516]" captionTargetPageId="26" captionText="Figs 72–75. Acanthosoma pugnax sp. nov. 72–73 – male (holotype); 74–75 – female (paratype). Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303591" httpUri="https://zenodo.org/record/5303591/files/figure.png" pageId="13" pageNumber="638">Figs 74–75</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C305DDF89F571C44FFDCFE57FBE2FDA8" pageId="13" pageNumber="638" type="diagnosis">
<paragraph id="8BA08E739F571C44FFDCFE57FBE2FDA8" blockId="13.[113,1090,485,1596]" pageId="13" pageNumber="638">
<emphasis id="B96B52619F571C44FFDCFE57FF46FE08" bold="true" box="[113,230,485,508]" pageId="13" pageNumber="638">Diagnosis.</emphasis>
Recognized by the following combination of characters: humeral process dark, greatly elongate, straight, apically sharp, directed anterolaterad; mesosternal carina short, slightly surpassing base of head; male genital capsule with a sclerotized ridge along margin of dorsal rim (
<figureCitation id="132492F69F571C44FF45FDF7FE94FDA8" box="[232,308,581,604]" captionStart="Figs 1–10" captionStartId="10.[151,188,1450,1469]" captionTargetBox="[154,1125,221,1407]" captionTargetId="figure-10@10.[151,1126,221,1420]" captionTargetPageId="10" captionText="Figs 1–10.Acanthosoma spp., males. 1 – A.atayal sp.nov., posterior part of abdomen, ventral view; 2 – same, genital capsule and exposed part of segment VIII, ventral view; 3 – same, genital capsule, lateral view; 4 – A. haemorrhoidale formosanum subsp. nov., holotype, genital capsule, posterior view; 5 – A.haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM), genital capsule, posterior view; 6 – A. asahinai Ishihara, 1943, posterior part of abdomen, ventral view; 7 – same, genital capsule, ventral view; 8 – A. fallax sp. nov., posterior part of abdomen, ventral view; 9 – same, genital capsule, ventral view; 10 – A. pugnax sp. nov., genital capsule and exposed part of segment VIII, ventral view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303573" httpUri="https://zenodo.org/record/5303573/files/figure.png" pageId="13" pageNumber="638">Fig. 10</figureCitation>
). The paramere (
<figureCitation id="132492F69F571C44FE4CFDF7FDF7FDA8" box="[481,599,581,604]" captionStart="Figs 20–44" captionStartId="14.[151,188,1196,1215]" captionTargetBox="[151,1126,231,1174]" captionTargetId="figure-84@14.[151,1126,231,1178]" captionTargetPageId="14" captionText="Figs 20–44. Acanthosoma spp., right parameres in different views. 20–21 – A. atayal sp. nov.; 22 – A. haemorrhoidale formosanum subsp. nov., holotype; 23 – A. haemorrhoidale ouchii Ishihara, 1950 (China: Sichuan, Mt. Emei, USNM); 24 – A. haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM); 25–28 – A. asahinai Ishihara, 1943; 29–32 – A. fallax sp. nov.; 33–36 – A. pugnax sp. nov.; 37–39 – A. axicia sp. nov.; 40–41 – A. laevicorne Dallas, 1851; 42–44 – A. expansum Horváth, 1905. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303577" httpUri="https://zenodo.org/record/5303577/files/figure.png" pageId="13" pageNumber="638">Figs 33–36</figureCitation>
) and female terminalia (
<figureCitation id="132492F69F571C44FCFFFDF7FC3CFDA8" box="[850,924,581,604]" captionStart="Figs 52–55" captionStartId="18.[151,188,1522,1541]" captionTargetBox="[151,1124,840,1491]" captionTargetId="figure-10@18.[151,1126,839,1496]" captionTargetPageId="18" captionText="Figs 52–55. Acanthosoma spp., female terminalia, posteroventral view. 52 – A. axicia sp. nov.; 53 – A. pugnax sp. nov.; 54 – A. laevicorne Dallas, 1851; 55 – A. expansum Horváth, 1905. PO6, PO7 = Pendergrast’s organs of ventrites VI and VII, respectively. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303581" httpUri="https://zenodo.org/record/5303581/files/figure.png" pageId="13" pageNumber="638">Fig. 53</figureCitation>
) are diagnostic.
</paragraph>
</subSubSection>
<subSubSection id="C305DDF89F571C46FFDCFDD7FBE1FE6D" lastPageId="15" lastPageNumber="640" pageId="13" pageNumber="638" type="description">
<paragraph id="8BA08E739F571C44FFDCFDD7FF7CFCC8" blockId="13.[113,1090,485,1596]" pageId="13" pageNumber="638">
<emphasis id="B96B52619F571C44FFDCFDD7FEF6FD88" bold="true" box="[113,342,612,636]" pageId="13" pageNumber="638">
Description.
<emphasis id="B96B52619F571C44FEAEFDD6FEF6FD88" bold="true" box="[259,342,612,636]" italics="true" pageId="13" pageNumber="638">Colour.</emphasis>
</emphasis>
Ground colour of dorsum green suffused with ferruginous to various extent, occasionally completely reddish brown; humeral processes suffused with black except their anterior margin; endocorium of fore wings with strong ferruginous suffusion; abdomen pale yellow, with distinct red suffusion on exposed parts of segments VII (J,
<specimenCount id="9D1945FA9F571C44FC1AFD0CFC6AFD29" box="[951,970,702,733]" pageId="13" pageNumber="638" type="female">♀</specimenCount>
), VIII (
<specimenCount id="9D1945FA9F571C44FBB3FD0CFB91FD29" box="[1054,1073,702,733]" pageId="13" pageNumber="638" type="female">♀</specimenCount>
), and dorsal surface of genital capsule (J); minute denticles at posterolateral angles of ventrites III–VI black but otherwise lateral margin of pregenital abdomen without conspicuous black markings.
</paragraph>
<paragraph id="8BA08E739F571C44FF3DFCF7FF69FB88" blockId="13.[113,1090,485,1596]" pageId="13" pageNumber="638">
<emphasis id="B96B52619F571C44FF3DFCF7FE0DFCA8" bold="true" box="[144,429,836,860]" pageId="13" pageNumber="638">
<emphasis id="B96B52619F571C44FF3DFCF7FE06FCA8" bold="true" box="[144,422,836,860]" italics="true" pageId="13" pageNumber="638">Integument and vestiture</emphasis>
.
</emphasis>
Body glabrous except of fine, scattered hairs on appendages and terminalia; mandibular plates with a few scattered black punctures, interocular area with a pair of irregular longitudinal rows of black punctures; maxillary plate with an obtuse tubercle anteriad and mesad of antennal insertion; pronotum, scutellum and sclerotized parts of fore wings with rather uniform black punctation except on calli, distal portions of humeral processes and costal margin of fore wing, punctation of exocorium and neighbouring part of endocorium not conspicuously different; ventral surface of body virtually unpunctured except of scattered punctures on ventral surface of humeral process and faint, superficial punctures along posterior margin of abdominal ventrite VIII and on ventral surface of genital capsule of male.
</paragraph>
<paragraph id="8BA08E739F571C44FF3DFB36FE24FA48" blockId="13.[113,1090,485,1596]" pageId="13" pageNumber="638">
<emphasis id="B96B52619F571C44FF3DFB36FE9CFB68" box="[144,316,1156,1180]" italics="true" pageId="13" pageNumber="638">
<emphasis id="B96B52619F571C44FF3DFB36FF5EFB68" bold="true" box="[144,254,1156,1180]" italics="true" pageId="13" pageNumber="638">Structure.</emphasis>
Body
</emphasis>
elongate oval with distinctly produced humeri, distance between tips of humeral processes about 1.65 (
<specimenCount id="9D1945FA9F571C44FE6CFB2CFE74FB49" box="[449,468,1182,1213]" pageId="13" pageNumber="638" type="female">♀</specimenCount>
) times as long as greatest width of body posteriad of humeri.
<emphasis id="B96B52619F571C44FF02FB77FF49FB28" box="[175,233,1221,1244]" italics="true" pageId="13" pageNumber="638">Head</emphasis>
1.18 (J) / 1.24 (
<specimenCount id="9D1945FA9F571C44FE34FB0CFE0CFB29" box="[409,428,1214,1245]" pageId="13" pageNumber="638" type="female">♀</specimenCount>
) times as broad as its median length, about 1.58 (J) / 1.53 (
<specimenCount id="9D1945FA9F571C44FB88FB0CFB98FB29" box="[1061,1080,1214,1245]" pageId="13" pageNumber="638" type="female">♀</specimenCount>
) times as broad as interocular distance, finely transversely rugose, anterior portion of clypeus with a broad median longitudinal groove.
<emphasis id="B96B52619F571C44FD9FFAB7FD23FAE8" box="[562,643,1285,1308]" italics="true" pageId="13" pageNumber="638">Labium</emphasis>
projecting between or slightly surpassing mid coxae.
<emphasis id="B96B52619F571C44FF46FA97FEF5FAC8" box="[235,341,1317,1340]" italics="true" pageId="13" pageNumber="638">Pronotum</emphasis>
with anterolateral margin weakly concave, gradually transitioning into humeral process which is long, straight, spiniform, directed subhorizontally, apically sharp.
<emphasis id="B96B52619F571C44FFDCFAD7FE13FA88" box="[113,435,1381,1404]" italics="true" pageId="13" pageNumber="638">Thoracic pleuron and sternum</emphasis>
. Mesosternal carina distinctly surpassing base of head, highly elevated, rather broadly rounded anteriorly; metathoracic scent gland ostiole with a long, slightly arched peritreme.
</paragraph>
<paragraph id="8BA08E739F571C47FF3DFA77FD61F9CA" blockId="13.[113,1090,485,1596]" lastBlockId="14.[151,1127,1415,1598]" lastPageId="14" lastPageNumber="639" pageId="13" pageNumber="638">
<emphasis id="B96B52619F571C44FF3DFA77FDA6FA28" box="[144,518,1477,1500]" italics="true" pageId="13" pageNumber="638">Male and female terminalia. Male</emphasis>
. Genital capsule (
<figureCitation id="132492F69F571C44FD63FA77FCBDFA28" box="[718,797,1477,1500]" captionStart="Figs 1–10" captionStartId="10.[151,188,1450,1469]" captionTargetBox="[154,1125,221,1407]" captionTargetId="figure-10@10.[151,1126,221,1420]" captionTargetPageId="10" captionText="Figs 1–10.Acanthosoma spp., males. 1 – A.atayal sp.nov., posterior part of abdomen, ventral view; 2 – same, genital capsule and exposed part of segment VIII, ventral view; 3 – same, genital capsule, lateral view; 4 – A. haemorrhoidale formosanum subsp. nov., holotype, genital capsule, posterior view; 5 – A.haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM), genital capsule, posterior view; 6 – A. asahinai Ishihara, 1943, posterior part of abdomen, ventral view; 7 – same, genital capsule, ventral view; 8 – A. fallax sp. nov., posterior part of abdomen, ventral view; 9 – same, genital capsule, ventral view; 10 – A. pugnax sp. nov., genital capsule and exposed part of segment VIII, ventral view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303573" httpUri="https://zenodo.org/record/5303573/files/figure.png" pageId="13" pageNumber="638">Fig. 10</figureCitation>
) with short, broad, widely diverging posterolateral projections, provided with a tuft of dense setae apically; dorsal rim broadly V-shaped, with a heavily sclerotized and pigmented ridge along its margin; ventral rim with a concave incision medially, provided with setal tufts, its infolding densely pilose. Paramere (
<figureCitation id="132492F69F541C47FEA7FA35FE23FA6A" box="[266,387,1415,1438]" captionStart="Figs 20–44" captionStartId="14.[151,188,1196,1215]" captionTargetBox="[151,1126,231,1174]" captionTargetId="figure-84@14.[151,1126,231,1178]" captionTargetPageId="14" captionText="Figs 20–44. Acanthosoma spp., right parameres in different views. 20–21 – A. atayal sp. nov.; 22 – A. haemorrhoidale formosanum subsp. nov., holotype; 23 – A. haemorrhoidale ouchii Ishihara, 1950 (China: Sichuan, Mt. Emei, USNM); 24 – A. haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM); 25–28 – A. asahinai Ishihara, 1943; 29–32 – A. fallax sp. nov.; 33–36 – A. pugnax sp. nov.; 37–39 – A. axicia sp. nov.; 40–41 – A. laevicorne Dallas, 1851; 42–44 – A. expansum Horváth, 1905. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303577" httpUri="https://zenodo.org/record/5303577/files/figure.png" pageId="14" pageNumber="639">Figs 33–36</figureCitation>
) bifurcated and heavily pigmented apically, internal surface concave, upper branch elongate with a digitiform process, lower branch short with a fin-shaped process, stem with a protrusion subapically.
<emphasis id="B96B52619F541C47FDBBFA75FDC6FA2A" box="[534,614,1479,1502]" italics="true" pageId="14" pageNumber="639">Female</emphasis>
(
<figureCitation id="132492F69F541C47FDDBFA75FD66FA2A" box="[630,710,1479,1502]" captionStart="Figs 52–55" captionStartId="18.[151,188,1522,1541]" captionTargetBox="[151,1124,840,1491]" captionTargetId="figure-10@18.[151,1126,839,1496]" captionTargetPageId="18" captionText="Figs 52–55. Acanthosoma spp., female terminalia, posteroventral view. 52 – A. axicia sp. nov.; 53 – A. pugnax sp. nov.; 54 – A. laevicorne Dallas, 1851; 55 – A. expansum Horváth, 1905. PO6, PO7 = Pendergrast’s organs of ventrites VI and VII, respectively. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303581" httpUri="https://zenodo.org/record/5303581/files/figure.png" pageId="14" pageNumber="639">Fig. 52</figureCitation>
). Posterior margin of ventrite VII with a deep, U-shaped median incision margined with black at bases of valvifers VIII; posterior margin of laterotergites VIII moderately rounded and prominent; Pendergrast’s organs on ventrite VII much smaller than those on ventrite VI.
</paragraph>
<caption id="DF60DEFB9F541C47FF3AFB1EFCF2FADE" ID-DOI="http://doi.org/10.5281/zenodo.5303577" ID-Zenodo-Dep="5303577" httpUri="https://zenodo.org/record/5303577/files/figure.png" pageId="14" pageNumber="639" startId="14.[151,188,1196,1215]" targetBox="[151,1126,231,1174]" targetPageId="14">
<paragraph id="8BA08E739F541C47FF3AFB1EFCF2FADE" blockId="14.[151,1128,1196,1322]" pageId="14" pageNumber="639">
Figs 20–44.
<taxonomicName id="4C1FF5F09F541C47FEA9FB1EFE04FB4B" box="[260,420,1196,1215]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="639" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B96B52619F541C47FEA9FB1EFED8FB4B" box="[260,376,1196,1215]" italics="true" pageId="14" pageNumber="639">Acanthosoma</emphasis>
spp.
</taxonomicName>
, right parameres in different views. 20–21 –
<taxonomicName id="4C1FF5F09F541C47FC81FB1EFCDAFB4B" authority="Tsai &amp; Rédei, 2015" authorityName="Tsai &amp; Rédei" authorityYear="2015" box="[812,890,1196,1215]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="639" phylum="Arthropoda" rank="species" species="atayal" status="sp. nov.">
<emphasis id="B96B52619F541C47FC81FB1EFCDAFB4B" box="[812,890,1196,1215]" italics="true" pageId="14" pageNumber="639">A. atayal</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A258EF1A9F541C47FC2CFB1EFC67FB4B" box="[897,967,1196,1215]" pageId="14" pageNumber="639" rank="species">sp. nov.</taxonomicNameLabel>
; 22 –
<taxonomicName id="4C1FF5F09F541C47FC50FB1EFEEAFB2E" authority="Tsai &amp; Rédei, 2015" authorityName="Tsai &amp; Rédei" authorityYear="2015" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="639" phylum="Arthropoda" rank="subSpecies" species="haemorrhoidale" status="subsp. nov." subSpecies="formosanum">
<emphasis id="B96B52619F541C47FC50FB1EFEEAFB2E" italics="true" pageId="14" pageNumber="639">A. haemorrhoidale formosanum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A258EF1A9F541C47FEFCFB75FE15FB2E" box="[337,437,1223,1242]" pageId="14" pageNumber="639" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
, holotype; 23 –
<taxonomicName id="4C1FF5F09F541C47FDE8FB75FC08FB2E" authority="Ishihara, 1950" authorityName="Ishihara" authorityYear="1950" box="[581,936,1223,1242]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="639" phylum="Arthropoda" rank="subSpecies" species="haemorrhoidale" subSpecies="ouchii">
<emphasis id="B96B52619F541C47FDE8FB75FC84FB2E" box="[581,804,1223,1242]" italics="true" pageId="14" pageNumber="639">A. haemorrhoidale ouchii</emphasis>
Ishihara, 1950
</taxonomicName>
(China: Sichuan, Mt. Emei, USNM); 24 –
<taxonomicName id="4C1FF5F09F541C47FEE5FB50FCADFB01" authority="haemorrhoidale (Linnaeus, 1758)" authorityName="haemorrhoidale (Linnaeus" authorityYear="1758" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[328,781,1250,1269]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="639" phylum="Arthropoda" rank="subSpecies" species="haemorrhoidale" subSpecies="haemorrhoidale">
<emphasis id="B96B52619F541C47FEE5FB50FDD7FB01" box="[328,631,1250,1269]" italics="true" pageId="14" pageNumber="639">A. haemorrhoidale haemorrhoidale</emphasis>
(Linnaeus, 1758)
</taxonomicName>
(Spain: Fuente Dé, HNHM); 25–28 –
<taxonomicName id="4C1FF5F09F541C47FBF9FB50FEC3FAFB" authority="Ishihara, 1943" authorityName="Ishihara" authorityYear="1943" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="639" phylum="Arthropoda" rank="species" species="asahinai">
<emphasis id="B96B52619F541C47FBF9FB50FF40FAFB" italics="true" pageId="14" pageNumber="639">A. asahinai</emphasis>
<bibRefCitation id="EF8EF3829F541C47FF4BFB4EFEC3FAFB" author="ISHIHARA T." box="[230,355,1276,1295]" pageId="14" pageNumber="639" pagination="495 - 497" refId="ref19649" refString="ISHIHARA T. 1943: Zwei Acanthosoma-arten der Insel Formosa (Hemiptera: Pentatomidae). Transactions of the Natural History Society of Taiwan 33: 495 - 497." type="journal article" year="1943">Ishihara, 1943</bibRefCitation>
</taxonomicName>
; 29–32 –
<taxonomicName id="4C1FF5F09F541C47FE1AFB4EFDA0FAFB" authority="Tsai &amp; Rédei, 2015" authorityName="Tsai &amp; Rédei" authorityYear="2015" box="[439,512,1276,1295]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="639" phylum="Arthropoda" rank="species" species="fallax" status="sp. nov.">
<emphasis id="B96B52619F541C47FE1AFB4EFDA0FAFB" box="[439,512,1276,1295]" italics="true" pageId="14" pageNumber="639">A. fallax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A258EF1A9F541C47FDA8FB4EFDEBFAFB" box="[517,587,1276,1295]" pageId="14" pageNumber="639" rank="species">sp. nov.</taxonomicNameLabel>
; 33–36 –
<taxonomicName id="4C1FF5F09F541C47FD33FB4EFD54FAFB" authority="Tsai &amp; Rédei, 2015" authorityName="Tsai &amp; Rédei" authorityYear="2015" box="[670,756,1276,1295]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="639" phylum="Arthropoda" rank="species" species="pugnax" status="sp. nov.">
<emphasis id="B96B52619F541C47FD33FB4EFD54FAFB" box="[670,756,1276,1295]" italics="true" pageId="14" pageNumber="639">A. pugnax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A258EF1A9F541C47FD57FB4EFC9FFAFB" box="[762,831,1276,1295]" pageId="14" pageNumber="639" rank="species">sp. nov.</taxonomicNameLabel>
; 37–39 –
<taxonomicName id="4C1FF5F09F541C47FC3FFB4EFC7EFAFB" authority="Tsai &amp; Rédei, 2015" authorityName="Tsai &amp; Rédei" authorityYear="2015" box="[914,990,1276,1295]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="639" phylum="Arthropoda" rank="species" species="axicia" status="sp. nov.">
<emphasis id="B96B52619F541C47FC3FFB4EFC7EFAFB" box="[914,990,1276,1295]" italics="true" pageId="14" pageNumber="639">A. axicia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A258EF1A9F541C47FC49FB4EFB89FAFB" box="[996,1065,1276,1295]" pageId="14" pageNumber="639" rank="species">sp. nov.</taxonomicNameLabel>
; 40–41 –
<taxonomicName id="4C1FF5F09F541C47FF0AFAA5FE2CFADE" authority="Dallas, 1851" authorityName="Dallas" authorityYear="1851" box="[167,396,1303,1322]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="639" phylum="Arthropoda" rank="species" species="laevicorne">
<emphasis id="B96B52619F541C47FF0AFAA5FEB9FADE" box="[167,281,1303,1322]" italics="true" pageId="14" pageNumber="639">A. laevicorne</emphasis>
<bibRefCitation id="EF8EF3829F541C47FEB3FAA5FE2CFADE" author="DALLAS W. S." box="[286,396,1303,1322]" pageId="14" pageNumber="639" refId="ref18627" refString="DALLAS W. S. 1851: List of the specimens of hemipterous insects in the collection of the British Museum. Part 1. Trustees of the British Museum, London, 368 pp, XI plates." type="book" year="1851">Dallas, 1851</bibRefCitation>
</taxonomicName>
; 42–44 –
<taxonomicName id="4C1FF5F09F541C47FE4DFAA5FD70FADE" authority="Horvath, 1905" authorityName="Horvath" authorityYear="1905" box="[480,720,1303,1322]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="639" phylum="Arthropoda" rank="species" species="expansum">
<emphasis id="B96B52619F541C47FE4DFAA5FDEDFADE" box="[480,589,1303,1322]" italics="true" pageId="14" pageNumber="639">A. expansum</emphasis>
<bibRefCitation id="EF8EF3829F541C47FDFFFAA5FD70FADE" author="HORVATH G." box="[594,720,1303,1322]" pageId="14" pageNumber="639" pagination="413 - 423" refId="ref19504" refString="HORVATH G. 1905: Hemipteres nouveaux de Japon. Annales Musei Nationalis Hungarici 3: 413 - 423." type="journal article" year="1905">Horváth, 1905</bibRefCitation>
</taxonomicName>
. Scales in mm.
</paragraph>
</caption>
<paragraph id="8BA08E739F551C46FF3DFF50FBE1FE6D" blockId="15.[113,1089,219,921]" pageId="15" pageNumber="640">
<emphasis id="B96B52619F551C46FF3DFF50FE91FF0D" bold="true" box="[144,305,226,249]" italics="true" pageId="15" pageNumber="640">Measurements</emphasis>
(
<typeStatus id="54A430D19F551C46FE93FF50FE3DFF0D" box="[318,413,226,249]" pageId="15" pageNumber="640" type="holotype">holotype</typeStatus>
J /
<typeStatus id="54A430D19F551C46FE6BFF50FD82FF0D" box="[454,546,226,249]" pageId="15" pageNumber="640" type="paratype">paratype</typeStatus>
<specimenCount id="9D1945FA9F551C46FD8BFF69FD99FF0E" box="[550,569,219,250]" pageId="15" pageNumber="640" type="female">♀</specimenCount>
) (in mm). Body length from apex of head to apex of membrane 13.3 / 13.8 (
<specimenCount id="9D1945FA9F551C46FE2EFF49FE36FEEE" box="[387,406,251,282]" pageId="15" pageNumber="640" type="female">♀</specimenCount>
); greatest width of body posteriad of humeral processes – / 6.41; median length of head 1.95 / 1.85, width across eyes 2.30 / 2.30, interocular distance 1.45 / 1.50; length of antennal segments (I) 1.65 / 1.65: (IIa) 1.65 / 1.60: (IIb) 1.75 / 1.40: (III) 1.35 / 1.85: (IV) 2.15 / 1.55; median length of pronotum 2.30 / 2.65, greatest width (across tips of humeri) 10.6 / 10.6; median length of scutellum 3.70 / 3.95, width at base 3.35 / 3.45.
</paragraph>
</subSubSection>
<subSubSection id="C305DDF89F551C46FFDCFE10FC83FE0D" pageId="15" pageNumber="640" type="etymology">
<paragraph id="8BA08E739F551C46FFDCFE10FCBDFE2D" blockId="15.[113,1089,219,921]" pageId="15" pageNumber="640">
<emphasis id="B96B52619F551C46FFDCFE10FF52FE4D" bold="true" box="[113,242,418,441]" pageId="15" pageNumber="640">Etymology.</emphasis>
The specific epithet is the Latin adjective
<taxonomicName id="4C1FF5F09F551C46FD15FE10FCA7FE4D" authorityName="Tsai &amp; Rédei" authorityYear="2015" box="[696,775,418,441]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="640" phylum="Arthropoda" rank="species" species="pugnax">
<emphasis id="B96B52619F551C46FD15FE10FCA7FE4D" box="[696,775,418,441]" italics="true" pageId="15" pageNumber="640">pugnax</emphasis>
</taxonomicName>
(m, f, n) ‘combative, fond of fighting’, referring to the produced, sharp humeri of the species.
</paragraph>
<paragraph id="8BA08E739F551C46FFDCFE50FC83FE0D" blockId="15.[113,1089,219,921]" box="[113,803,482,505]" pageId="15" pageNumber="640">
<emphasis id="B96B52619F551C46FFDCFE50FF4FFE0D" bold="true" box="[113,239,482,505]" pageId="15" pageNumber="640">Bionomics.</emphasis>
Unknown. The male
<typeStatus id="54A430D19F551C46FE7BFE50FD93FE0D" box="[470,563,482,505]" pageId="15" pageNumber="640" type="holotype">holotype</typeStatus>
was collected at light.
</paragraph>
</subSubSection>
<subSubSection id="C305DDF89F551C46FFDCFDB0FD95FDAD" pageId="15" pageNumber="640" type="distribution">
<paragraph id="8BA08E739F551C46FFDCFDB0FD95FDAD" blockId="15.[113,1089,219,921]" pageId="15" pageNumber="640">
<emphasis id="B96B52619F551C46FFDCFDB0FEA1FDED" bold="true" box="[113,257,514,537]" pageId="15" pageNumber="640">Distribution.</emphasis>
This species is apparently endemic to
<collectingCountry id="F308CEE39F551C46FD21FDB0FD7DFDED" box="[652,733,514,537]" name="Taiwan" pageId="15" pageNumber="640">Taiwan</collectingCountry>
. Specimens are rare in collections, so far only the
<typeStatus id="54A430D19F551C46FEBFFD90FECFFDCD" box="[274,367,546,569]" pageId="15" pageNumber="640" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D1945FA9F551C46FE09FD90FD8DFDCD" box="[420,557,546,569]" pageId="15" pageNumber="640" type="generic" typeStatus="paratype">one paratype</specimenCount>
are known, both collected in the central, mountainous region of the main island of
<collectingCountry id="F308CEE39F551C46FE72FDF0FD91FDAD" box="[479,561,578,601]" name="Taiwan" pageId="15" pageNumber="640">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C305DDF89F551C46FFDCFDD0FD09FC6D" pageId="15" pageNumber="640" type="discussion">
<paragraph id="8BA08E739F551C46FFDCFDD0FD7AFD4D" blockId="15.[113,1089,219,921]" pageId="15" pageNumber="640">
<emphasis id="B96B52619F551C46FFDCFDD0FF40FD8D" bold="true" box="[113,224,610,633]" pageId="15" pageNumber="640">Remarks.</emphasis>
Although the single male available is badly damaged (
<figureCitation id="132492F69F551C46FCB6FDD0FC36FD8D" box="[795,918,610,633]" captionStart="Figs 72–75" captionStartId="26.[151,188,1547,1566]" captionTargetBox="[194,1071,246,1496]" captionTargetId="figure-10@26.[151,1126,217,1516]" captionTargetPageId="26" captionText="Figs 72–75. Acanthosoma pugnax sp. nov. 72–73 – male (holotype); 74–75 – female (paratype). Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303591" httpUri="https://zenodo.org/record/5303591/files/figure.png" pageId="15" pageNumber="640">Figs 72–73</figureCitation>
), we judged that it, together with an obviously conspecific female, will make it possible to prepare a description which will allow unambiguous recognition of the species.
</paragraph>
<paragraph id="8BA08E739F551C46FF3DFD70FD09FC6D" blockId="15.[113,1089,219,921]" pageId="15" pageNumber="640">
The new species is taxonomically closely related to
<taxonomicName id="4C1FF5F09F551C46FD0AFD70FC5AFD2D" authority="Hsiao &amp; Liu, 1977" authorityName="Hsiao &amp; Liu" authorityYear="1977" box="[679,1018,706,729]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="640" phylum="Arthropoda" rank="species" species="shensiense">
<emphasis id="B96B52619F551C46FD0AFD70FC94FD2D" box="[679,820,706,729]" italics="true" pageId="15" pageNumber="640">A. shensiense</emphasis>
<bibRefCitation id="EF8EF3829F551C46FC94FD70FC5AFD2D" author="HSIAO T. Y. &amp; LIU S. L." box="[825,1018,706,729]" pageId="15" pageNumber="640" pagination="300 - 302" refId="ref19524" refString="HSIAO T. Y. &amp; LIU S. L. 1977: Family Acanthosomatidae. Pp. 159 - 180, 300 - 302, plates 25 - 29. In: HSIAO T. Y., REN S. Z., ZHENG L. Y., JING H. L. &amp; LIU S. L.: A handbook for the determination of the Chinese Hemiptera-Heteroptera. Volume I. Science Press, Beijing, iii + 330 pp, 52 plates (in Chinese, English summary)." type="book chapter" year="1977">Hsiao &amp; Liu, 1977</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C1FF5F09F551C46FB87FD70FEFBFD0D" authority="(Liu, 1987)" baseAuthorityName="Liu" baseAuthorityYear="1987" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="640" phylum="Arthropoda" rank="species" species="montanum">
<emphasis id="B96B52619F551C46FB87FD70FF40FD0D" italics="true" pageId="15" pageNumber="640">A. montanum</emphasis>
(
<bibRefCitation id="EF8EF3829F551C46FF40FD50FEF3FD0D" author="LIU S. L." box="[237,339,738,761]" pageId="15" pageNumber="640" pagination="149 - 155" refId="ref20589" refString="LIU S. L. 1987: Acanthosomatidae. Pp. 149 - 155. In: HUANG F. S. (ed.): Forest insects of Yunnan. Yunnan Science and Technology Press, Kunming, 9 + 1622 pp, 16 plates (in Chinese)." type="book chapter" year="1987">Liu, 1987</bibRefCitation>
)
</taxonomicName>
as indicated by the presence of a sclerotized and pigmented longitudinal ridge along the dorsal rim of the genital capsule of the male (a potential synapomorphy) and the similar shape of the paramere. Regarding
<taxonomicName id="4C1FF5F09F551C46FDCFFC90FD52FCCD" baseAuthorityName="Liu" baseAuthorityYear="1987" box="[610,754,802,825]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="640" phylum="Arthropoda" rank="species" species="montanum">
<emphasis id="B96B52619F551C46FDCFFC90FD52FCCD" box="[610,754,802,825]" italics="true" pageId="15" pageNumber="640">A. montanum</emphasis>
</taxonomicName>
, the new species can be easily distinguished by the apically sharp humeri (apically obtuse, recurved in
<taxonomicName id="4C1FF5F09F551C46FCD7FCF0FBA9FCAD" baseAuthorityName="Liu" baseAuthorityYear="1987" box="[890,1033,834,857]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="640" phylum="Arthropoda" rank="species" species="montanum">
<emphasis id="B96B52619F551C46FCD7FCF0FBA9FCAD" box="[890,1033,834,857]" italics="true" pageId="15" pageNumber="640">A. montanum</emphasis>
</taxonomicName>
), the different parameres, the more prominent posterior margin of laterotergites VIII of the female, and the reduced Pendergrast’s organs on ventrite VII.
</paragraph>
</subSubSection>
</treatment>
</document>
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