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<document id="C7534E42950A4D1EDEE83963A1FC48BD" ID-DOI="10.5281/zenodo.179949" ID-GBIF-Dataset="109e1b5c-dd7c-49da-8828-4e0ba2d07603" ID-ISSN="1175-5326" ID-Zenodo-Dep="179949" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460020948282" checkinUser="plazi" docAuthor="Godunko, Roman J." docDate="2007" docId="0380C10F1776E266FF03F8EBFE5DBC62" docLanguage="en" docName="zt01661p068.pdf" docOrigin="Zootaxa 1661" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Ecdyonurus (Nestormeus) groehnorum Godunko, 2007, sp. nov." docType="treatment" docVersion="8" lastPageNumber="68" masterDocId="FFB9B9771776E263FF94FF81FFB7BE20" masterDocTitle="Contribution to the knowledge of the fossil subgenus Nestormeus Godunko, 2004 (Ephemeroptera: Heptageniidae: Ecdyonurus) from the Baltic amber (Eocene)" masterLastPageNumber="68" masterPageNumber="63" pageNumber="63" updateTime="1698233743044" updateUser="plazi">
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<mods:title id="110BB42D3B530671351649862E6FDC4F">Contribution to the knowledge of the fossil subgenus Nestormeus Godunko, 2004 (Ephemeroptera: Heptageniidae: Ecdyonurus) from the Baltic amber (Eocene)</mods:title>
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<mods:namePart id="760650103C254A47A10AA100F8422A36">Godunko, Roman J.</mods:namePart>
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<mods:date id="89F1EC06486A7FB78BFC4A5E2F38FC1C">2007</mods:date>
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<subSubSection id="C33323921776E263FF03F8EBFEA7B987" pageId="0" pageNumber="63" type="nomenclature">
<paragraph id="8B9670191776E263FF03F8EBFD61B9A4" blockId="0.[151,726,1898,1959]" box="[151,726,1898,1924]" pageId="0" pageNumber="63">
<heading id="D0DEC7751776E263FF03F8EBFD61B9A4" bold="true" box="[151,726,1898,1924]" fontSize="11" level="1" pageId="0" pageNumber="63" reason="1">
<emphasis id="B95DAC0B1776E263FF03F8EBFD61B9A4" bold="true" box="[151,726,1898,1924]" pageId="0" pageNumber="63">
<taxonomicName id="4C290B9A1776E263FF03F8EBFDC4B9A4" box="[151,627,1898,1924]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="groehnorum" status="sp. nov." subGenus="Nestormeus">
<emphasis id="B95DAC0B1776E263FF03F8EBFE9DB9A4" bold="true" box="[151,298,1898,1924]" italics="true" pageId="0" pageNumber="63">Ecdyonurus</emphasis>
(
<emphasis id="B95DAC0B1776E263FEAEF8EBFE7EB9A4" bold="true" box="[314,457,1898,1924]" italics="true" pageId="0" pageNumber="63">Nestormeus</emphasis>
)
<emphasis id="B95DAC0B1776E263FE4FF8EBFDC4B9A4" bold="true" box="[475,627,1898,1924]" italics="true" pageId="0" pageNumber="63">groehnorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E11701776E263FDEDF8EBFD61B9A4" box="[633,726,1898,1924]" pageId="0" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9670191776E263FF03F80CFEA7B987" blockId="0.[151,726,1898,1959]" box="[151,272,1933,1959]" pageId="0" pageNumber="63">
(
<figureCitation id="13126C9C1776E263FF0BF80CFEB0B987" box="[159,263,1933,1959]" captionStart-0="FIGURES 1 3" captionStart-1="FIGURES 4 6" captionStartId-0="3.[151,269,1726,1750]" captionStartId-1="4.[151,269,1846,1870]" captionTargetBox-0="[165,1431,217,1671]" captionTargetBox-1="[432,1149,200,1811]" captionTargetId-0="figure@3.[150,1456,191,1702]" captionTargetId-1="figure@4.[419,1170,194,1820]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 1 3. Ecdyonurus (Nestormeus) groehnorum sp. nov., holotype, female imago: 1, head and thorax laterally and dorsally; 2, body ventrally; 3, tip of abdomen ventrally; 1 2: scale bars = 1 mm; 3: scale bar = 0.5 mm." captionText-1="FIGURES 4 6. Ecdyonurus (Nestormeus) groehnorum sp. nov., holotype, female imago: 4, tip of abdomen ventrally; 5, mesothorax ventrally; 6, body ventrally." httpUri-0="https://zenodo.org/record/179950/files/figure.png" httpUri-1="https://zenodo.org/record/179951/files/figure.png" pageId="0" pageNumber="63">Figs 16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33323921776E266FF03F856FD1CBF72" lastPageId="5" lastPageNumber="68" pageId="0" pageNumber="63" type="description">
<paragraph id="8B9670191776E263FF03F856FC87B9D2" blockId="0.[151,816,2007,2034]" box="[151,816,2007,2034]" pageId="0" pageNumber="63">
<emphasis id="B95DAC0B1776E263FF03F856FE9FB9D1" bold="true" box="[151,296,2007,2033]" pageId="0" pageNumber="63">Description</emphasis>
. Female imago. Measurements: see
<tableCitation id="C6AB45A21776E263FD47F859FC9CB9D2" box="[723,811,2008,2034]" captionStart="TABLE 1" captionStartId="1.[151,239,151,175]" captionText="TABLE 1. Morphometrics of holotype (female imago) of Ecdyonurus (Nestormeus) groehnorum sp. nov. from the Baltic amber (Eocene)." pageId="0" pageNumber="63">Table 1</tableCitation>
.
</paragraph>
<caption id="DF5620911777E262FF03FF16FEDEBEF1" pageId="1" pageNumber="64">
<paragraph id="8B9670191777E262FF03FF16FEDEBEF1" blockId="1.[151,1436,151,209]" pageId="1" pageNumber="64">
<emphasis id="B95DAC0B1777E262FF03FF16FEBEBE8F" bold="true" box="[151,265,151,175]" pageId="1" pageNumber="64">TABLE 1.</emphasis>
Morphometrics of holotype (female imago) of
<taxonomicName id="4C290B9A1777E262FC93FF19FB10BE8F" box="[775,1191,152,175]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="species" species="groehnorum" status="sp. nov." subGenus="Nestormeus">
<emphasis id="B95DAC0B1777E262FC93FF19FC31BE8F" box="[775,902,152,175]" italics="true" pageId="1" pageNumber="64">Ecdyonurus</emphasis>
(
<emphasis id="B95DAC0B1777E262FC03FF19FBA2BE8F" box="[919,1045,152,175]" italics="true" pageId="1" pageNumber="64">Nestormeus</emphasis>
)
<emphasis id="B95DAC0B1777E262FBB0FF19FB10BE8F" box="[1060,1191,152,175]" italics="true" pageId="1" pageNumber="64">groehnorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E11701777E262FB3AFF16FAB2BE8F" box="[1198,1285,151,175]" pageId="1" pageNumber="64" rank="species">
<emphasis id="B95DAC0B1777E262FB3AFF16FAB2BE8F" bold="true" box="[1198,1285,151,175]" pageId="1" pageNumber="64">sp. nov.</emphasis>
</taxonomicNameLabel>
from the Baltic amber (Eocene).
</paragraph>
</caption>
<paragraph id="8B9670191777E262FF0BFF73FDC9BF14" blockId="1.[159,1430,242,2015]" pageId="1" pageNumber="64">Characters (mm) Length of body 5.33</paragraph>
<paragraph id="8B9670191777E262FF0BFEC9FDC9BF7F" blockId="1.[159,1430,242,2015]" box="[159,638,328,351]" pageId="1" pageNumber="64">Length of right fore leg 3.95</paragraph>
<paragraph id="8B9670191777E262FF0BFEF3FDC9BFA9" blockId="1.[159,1430,242,2015]" box="[159,638,370,393]" pageId="1" pageNumber="64">Length of femur 1.53</paragraph>
<paragraph id="8B9670191777E262FF0BFE1CFDC9BF94" blockId="1.[159,1430,242,2015]" box="[159,638,413,436]" pageId="1" pageNumber="64">Length of tibia 1.38</paragraph>
<paragraph id="8B9670191777E262FF0BFE49FDC9BFFF" blockId="1.[159,1430,242,2015]" box="[159,638,456,479]" pageId="1" pageNumber="64">Length of tarsus 1.04</paragraph>
<paragraph id="8B9670191777E262FF0BFE73FDC9BC29" blockId="1.[159,1430,242,2015]" box="[159,638,498,521]" pageId="1" pageNumber="64">Segment I 0.15</paragraph>
<paragraph id="8B9670191777E262FF0BFD9CFDCABC14" blockId="1.[159,1430,242,2015]" box="[159,637,541,564]" pageId="1" pageNumber="64">Segment II 0.28</paragraph>
<paragraph id="8B9670191777E262FF0BFDC9FDC9BC7F" blockId="1.[159,1430,242,2015]" box="[159,638,584,607]" pageId="1" pageNumber="64">Segment III 0.25</paragraph>
<paragraph id="8B9670191777E262FF0BFDF3FDCABCA9" blockId="1.[159,1430,242,2015]" box="[159,637,626,649]" pageId="1" pageNumber="64">Segment IV 0.18</paragraph>
<paragraph id="8B9670191777E262FF0BFD1CFDC9BC94" blockId="1.[159,1430,242,2015]" box="[159,638,669,692]" pageId="1" pageNumber="64">Segment V 0.18</paragraph>
<paragraph id="8B9670191777E262FF0BFD49FDC9BCFF" blockId="1.[159,1430,242,2015]" box="[159,638,712,735]" pageId="1" pageNumber="64">Length of left fore leg 3.86</paragraph>
<paragraph id="8B9670191777E262FF0BFD73FDC9BD29" blockId="1.[159,1430,242,2015]" box="[159,638,754,777]" pageId="1" pageNumber="64">Length of femur 1.50</paragraph>
<paragraph id="8B9670191777E262FF0BFC9CFDC9BD14" blockId="1.[159,1430,242,2015]" box="[159,638,797,820]" pageId="1" pageNumber="64">Length of tibia 1.40</paragraph>
<paragraph id="8B9670191777E262FF0BFCC9FDC9BD7F" blockId="1.[159,1430,242,2015]" box="[159,638,840,863]" pageId="1" pageNumber="64">Length of tarsus 0.96</paragraph>
<paragraph id="8B9670191777E262FF0BFCF3FDC9BDA9" blockId="1.[159,1430,242,2015]" box="[159,638,882,905]" pageId="1" pageNumber="64">Segment I 0.15</paragraph>
<paragraph id="8B9670191777E262FF0BFC1CFDCABD94" blockId="1.[159,1430,242,2015]" box="[159,637,925,948]" pageId="1" pageNumber="64">Segment II 0.28</paragraph>
<paragraph id="8B9670191777E262FF0BFC49FDC9BDFF" blockId="1.[159,1430,242,2015]" box="[159,638,968,991]" pageId="1" pageNumber="64">Segment III 0.23</paragraph>
<paragraph id="8B9670191777E262FF0BFC73FDCABA29" blockId="1.[159,1430,242,2015]" box="[159,637,1010,1033]" pageId="1" pageNumber="64">Segment IV 0.15</paragraph>
<paragraph id="8B9670191777E262FF0BFB9CFDC9BA14" blockId="1.[159,1430,242,2015]" box="[159,638,1053,1076]" pageId="1" pageNumber="64">Segment V 0.15</paragraph>
<paragraph id="8B9670191777E262FF0BFBC9FDC9BA7F" blockId="1.[159,1430,242,2015]" box="[159,638,1096,1119]" pageId="1" pageNumber="64">Length of right middle leg 3.52</paragraph>
<paragraph id="8B9670191777E262FF0BFBF3FDC9BAA9" blockId="1.[159,1430,242,2015]" box="[159,638,1138,1161]" pageId="1" pageNumber="64">Length of femur 1.63</paragraph>
<paragraph id="8B9670191777E262FF0BFB1CFDC9BA94" blockId="1.[159,1430,242,2015]" box="[159,638,1181,1204]" pageId="1" pageNumber="64">Length of tibia 1.23</paragraph>
<paragraph id="8B9670191777E262FF0BFB49FDC9BAFF" blockId="1.[159,1430,242,2015]" box="[159,638,1224,1247]" pageId="1" pageNumber="64">Length of tarsus 0.66</paragraph>
<paragraph id="8B9670191777E262FF0BFB73FDC9BB29" blockId="1.[159,1430,242,2015]" box="[159,638,1266,1289]" pageId="1" pageNumber="64">Segment I 0.13</paragraph>
<paragraph id="8B9670191777E262FF0BFA9CFDCABB14" blockId="1.[159,1430,242,2015]" box="[159,637,1309,1332]" pageId="1" pageNumber="64">Segment II 0.15</paragraph>
<paragraph id="8B9670191777E262FF0BFAC9FDC9BB7F" blockId="1.[159,1430,242,2015]" box="[159,638,1352,1375]" pageId="1" pageNumber="64">Segment III 0.13</paragraph>
<paragraph id="8B9670191777E262FF0BFAF3FDCABBA9" blockId="1.[159,1430,242,2015]" box="[159,637,1394,1417]" pageId="1" pageNumber="64">Segment IV 0.10</paragraph>
<paragraph id="8B9670191777E262FF0BFA1CFDC9BB94" blockId="1.[159,1430,242,2015]" box="[159,638,1437,1460]" pageId="1" pageNumber="64">Segment V 0.15</paragraph>
<paragraph id="8B9670191777E262FF0BFA49FDC9BBFF" blockId="1.[159,1430,242,2015]" box="[159,638,1480,1503]" pageId="1" pageNumber="64">Length of left middle leg 3.50</paragraph>
<paragraph id="8B9670191777E262FF0BFA73FDC9B829" blockId="1.[159,1430,242,2015]" box="[159,638,1522,1545]" pageId="1" pageNumber="64">Length of femur 1.58</paragraph>
<paragraph id="8B9670191777E262FF0BF99CFDC9B814" blockId="1.[159,1430,242,2015]" box="[159,638,1565,1588]" pageId="1" pageNumber="64">Length of tibia 1.28</paragraph>
<paragraph id="8B9670191777E262FF0BF9C9FDC9B87F" blockId="1.[159,1430,242,2015]" box="[159,638,1608,1631]" pageId="1" pageNumber="64">Length of tarsus 0.64</paragraph>
<paragraph id="8B9670191777E262FF0BF9F3FDC9B8A9" blockId="1.[159,1430,242,2015]" box="[159,638,1650,1673]" pageId="1" pageNumber="64">Segment I 0.13</paragraph>
<paragraph id="8B9670191777E262FF0BF91CFDCAB894" blockId="1.[159,1430,242,2015]" box="[159,637,1693,1716]" pageId="1" pageNumber="64">Segment II 0.13</paragraph>
<paragraph id="8B9670191777E262FF0BF949FDC9B8FF" blockId="1.[159,1430,242,2015]" box="[159,638,1736,1759]" pageId="1" pageNumber="64">Segment III 0.13</paragraph>
<paragraph id="8B9670191777E262FF0BF973FDCAB929" blockId="1.[159,1430,242,2015]" box="[159,637,1778,1801]" pageId="1" pageNumber="64">Segment IV 0.10</paragraph>
<paragraph id="8B9670191777E262FF0BF89CFDC9B914" blockId="1.[159,1430,242,2015]" box="[159,638,1821,1844]" pageId="1" pageNumber="64">Segment V 0.15</paragraph>
<paragraph id="8B9670191777E262FF0BF8C9FDC9B97F" blockId="1.[159,1430,242,2015]" box="[159,638,1864,1887]" pageId="1" pageNumber="64">Length of right hind leg 3.20</paragraph>
<paragraph id="8B9670191777E262FF0BF8F3FDC9B9A9" blockId="1.[159,1430,242,2015]" box="[159,638,1906,1929]" pageId="1" pageNumber="64">Length of femur 1.58</paragraph>
<paragraph id="8B9670191777E262FF0BF81CFDC9B994" blockId="1.[159,1430,242,2015]" box="[159,638,1949,1972]" pageId="1" pageNumber="64">Length of tibia 1.05</paragraph>
<paragraph id="8B9670191777E262FB7EF849FA21B9FF" blockId="1.[159,1430,242,2015]" box="[1258,1430,1992,2015]" pageId="1" pageNumber="64">to be continued.</paragraph>
<caption id="DF5620911774E261FF0BFF1EFE24BE97" box="[159,403,159,183]" pageId="2" pageNumber="65">
<paragraph id="8B9670191774E261FF0BFF1EFE24BE97" blockId="2.[159,651,159,737]" box="[159,403,159,183]" pageId="2" pageNumber="65">
<emphasis id="B95DAC0B1774E261FF0BFF1EFEA7BE97" bold="true" box="[159,272,159,183]" pageId="2" pageNumber="65">TABLE 1.</emphasis>
(continued)
</paragraph>
</caption>
<paragraph id="8B9670191774E261FF0BFF4BFDEFBC41" blockId="2.[159,651,159,737]" pageId="2" pageNumber="65">Characters (mm) Length of tarsus 0.57 Segment I 0.13 Segment II 0.13 Segment III 0.10 Segment IV 0.08 Segment V 0.13 Length of right fore wing 5.93 Length of left fore wing 5.75 Length of right hind wing -</paragraph>
<paragraph id="8B9670191774E261FF0BFDF4FD3CBCC1" blockId="2.[159,651,159,737]" pageId="2" pageNumber="65">
Length of left hind wing 2.13
<collectingCountry id="F33E30891774E261FF0BFD21FF64BC97" box="[159,211,672,695]" name="India" pageId="2" pageNumber="65">Hind</collectingCountry>
/fore wings length ratio 0.36 Length of cerci 8.00*
</paragraph>
<paragraph id="8B9670191774E261FF03FC83FB6ABD39" blockId="2.[151,1245,770,793]" box="[151,1245,770,793]" pageId="2" pageNumber="65">
*Completely preserved right cercus only. Left cercus damaged, the length of preserved part is
<quantity id="4CD1DDFC1774E261FB10FC83FB6EBD39" box="[1156,1241,770,793]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="65" unit="mm" value="2.4">2.4 mm</quantity>
.
</paragraph>
<paragraph id="8B9670191774E261FF53FCEBFB61BDA4" blockId="2.[151,1437,874,1980]" box="[199,1238,874,900]" pageId="2" pageNumber="65">Body pale. Head uniformly brownish pigmented. Eyes noncontiguous, widely separated.</paragraph>
<paragraph id="8B9670191774E261FF53FC13FD2EBABC" blockId="2.[151,1437,874,1980]" pageId="2" pageNumber="65">
Thorax light brown to brown with paler pronotum. Some brownish smudges around mesonotal sutures. Ventral side of thorax with the same color as dorsal one. Mesonotal suture deep, well visible, stretched backward medially. Medioparapsidal suture is well visible from the left side of the body only. Lateroparapsidal sutures deep and slightly curved, noncontiguous with medioparapsidal sutures (
<figureCitation id="13126C9C1774E261FBFDFB8BFB04BA04" box="[1129,1203,1034,1060]" captionStart="FIGURES 1 3" captionStartId="3.[151,269,1726,1750]" captionTargetBox="[165,1431,217,1671]" captionTargetId="figure@3.[150,1456,191,1702]" captionTargetPageId="3" captionText="FIGURES 1 3. Ecdyonurus (Nestormeus) groehnorum sp. nov., holotype, female imago: 1, head and thorax laterally and dorsally; 2, body ventrally; 3, tip of abdomen ventrally; 1 2: scale bars = 1 mm; 3: scale bar = 0.5 mm." httpUri="https://zenodo.org/record/179950/files/figure.png" pageId="2" pageNumber="65">Fig. 1</figureCitation>
). Proximal part of mesonotum invisible from dorsal view. Median impression of furcasternum of mesothorax with sub-parallel margins, narrowed posteriorly and distinctly divergent in anterior part. Prosternum without any transversal ridge, typical of
<taxonomicName id="4C290B9A1774E261FECCFB02FE53BABC" box="[344,484,1155,1180]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B95DAC0B1774E261FECCFB02FE53BABC" box="[344,484,1155,1180]" italics="true" pageId="2" pageNumber="65">Ecdyonurus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E11701774E261FE78FB03FDA2BABC" box="[492,533,1154,1180]" pageId="2" pageNumber="65" sensu="lato">s. l.</taxonomicNameLabel>
(
<figureCitation id="13126C9C1774E261FDB1FB03FDDABABC" box="[549,621,1154,1180]" captionStart="FIGURES 1 3" captionStartId="3.[151,269,1726,1750]" captionTargetBox="[165,1431,217,1671]" captionTargetId="figure@3.[150,1456,191,1702]" captionTargetPageId="3" captionText="FIGURES 1 3. Ecdyonurus (Nestormeus) groehnorum sp. nov., holotype, female imago: 1, head and thorax laterally and dorsally; 2, body ventrally; 3, tip of abdomen ventrally; 1 2: scale bars = 1 mm; 3: scale bar = 0.5 mm." httpUri="https://zenodo.org/record/179950/files/figure.png" pageId="2" pageNumber="65">Fig. 2</figureCitation>
,
<figureCitation id="13126C9C1774E261FDEDFB03FD3DBABC" box="[633,650,1154,1180]" captionStart="FIGURES 4 6" captionStartId="4.[151,269,1846,1870]" captionTargetBox="[432,1149,200,1811]" captionTargetId="figure@4.[419,1170,194,1820]" captionTargetPageId="4" captionText="FIGURES 4 6. Ecdyonurus (Nestormeus) groehnorum sp. nov., holotype, female imago: 4, tip of abdomen ventrally; 5, mesothorax ventrally; 6, body ventrally." httpUri="https://zenodo.org/record/179951/files/figure.png" pageId="2" pageNumber="65">5</figureCitation>
).
</paragraph>
<paragraph id="8B9670191774E261FF53FB2BFE30BB44" blockId="2.[151,1437,874,1980]" pageId="2" pageNumber="65">
Both pairs of wings translucent and opaque (
<figureCitation id="13126C9C1774E261FD41FB2BFCABBAE4" box="[725,796,1194,1220]" captionStart="FIGURES 4 6" captionStartId="4.[151,269,1846,1870]" captionTargetBox="[432,1149,200,1811]" captionTargetId="figure@4.[419,1170,194,1820]" captionTargetPageId="4" captionText="FIGURES 4 6. Ecdyonurus (Nestormeus) groehnorum sp. nov., holotype, female imago: 4, tip of abdomen ventrally; 5, mesothorax ventrally; 6, body ventrally." httpUri="https://zenodo.org/record/179951/files/figure.png" pageId="2" pageNumber="65">Fig. 6</figureCitation>
). Cubital field of fore wings with two pairs of intercalary veins.
<collectingCountry id="F33E30891774E261FE8CFB53FEE4BACC" box="[280,339,1234,1260]" name="India" pageId="2" pageNumber="65">Hind</collectingCountry>
wings with RS, MA and MP triads.
<collectingCountry id="F33E30891774E261FD69FB53FC8FBACC" box="[765,824,1234,1260]" name="India" pageId="2" pageNumber="65">Hind</collectingCountry>
right wing damaged. Wing venation well preserved, with typical structure of the family
<taxonomicName id="4C290B9A1774E261FDAEFB7BFD54BB34" box="[570,739,1274,1300]" class="Insecta" family="Heptageniidae" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="family">Heptageniidae</taxonomicName>
(see
<bibRefCitation id="EFB80DE81774E261FCB6FB7BFC06BB34" author="Kluge" box="[802,945,1274,1300]" pageId="2" pageNumber="68" refString="Kluge, N. J. (2004) The Phylogenetic System of Ephemeroptera (the first experience in consistently non-ranking taxonomy) Volume 1. Ephemeroptera except for Turbanoculata and Leptophlebia / fg 1. Kluwer Academic Publishers, Dordrecht-Hardbound, 456 pp." type="book" year="2004">Kluge 2004</bibRefCitation>
: 380384). Patella-tibial suture is distinct enough in middle and hind legs. Tarsi of all legs are 5-segmented, with dissimilar tarsal claws (one pointed and one blunt claw).
</paragraph>
<paragraph id="8B9670191774E261FF53FAF3FCCFBBFC" blockId="2.[151,1437,874,1980]" pageId="2" pageNumber="65">
Abdominal segments slightly deformed. Abdominal terga and sterna paler than thorax. Subgenital plate large, well developed, with visible apically impression. Subanal plate elongate, rounded, apically with marked tip (
<figureCitation id="13126C9C1774E261FF51FA43FEA7BBFC" box="[197,272,1474,1500]" captionStart="FIGURES 1 3" captionStartId="3.[151,269,1726,1750]" captionTargetBox="[165,1431,217,1671]" captionTargetId="figure@3.[150,1456,191,1702]" captionTargetPageId="3" captionText="FIGURES 1 3. Ecdyonurus (Nestormeus) groehnorum sp. nov., holotype, female imago: 1, head and thorax laterally and dorsally; 2, body ventrally; 3, tip of abdomen ventrally; 1 2: scale bars = 1 mm; 3: scale bar = 0.5 mm." httpUri="https://zenodo.org/record/179950/files/figure.png" pageId="2" pageNumber="65">Figs 3</figureCitation>
,
<figureCitation id="13126C9C1774E261FE8AFA43FE87BBFC" box="[286,304,1474,1500]" captionStart="FIGURES 4 6" captionStartId="4.[151,269,1846,1870]" captionTargetBox="[432,1149,200,1811]" captionTargetId="figure@4.[419,1170,194,1820]" captionTargetPageId="4" captionText="FIGURES 4 6. Ecdyonurus (Nestormeus) groehnorum sp. nov., holotype, female imago: 4, tip of abdomen ventrally; 5, mesothorax ventrally; 6, body ventrally." httpUri="https://zenodo.org/record/179951/files/figure.png" pageId="2" pageNumber="65">4</figureCitation>
). Cerci preserved, longer than body (see
<tableCitation id="C6AB45A21774E261FC84FA43FCD1BBFC" box="[784,870,1474,1500]" captionStart="TABLE 1" captionStartId="1.[151,239,151,175]" captionText="TABLE 1. Morphometrics of holotype (female imago) of Ecdyonurus (Nestormeus) groehnorum sp. nov. from the Baltic amber (Eocene)." pageId="2" pageNumber="65">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8B9670191774E261FF53FA6BFB27B884" blockId="2.[151,1437,874,1980]" pageId="2" pageNumber="65">
<emphasis id="B95DAC0B1774E261FF53FA6BFEF5B824" bold="true" box="[199,322,1514,1540]" pageId="2" pageNumber="65">Affinities.</emphasis>
<taxonomicName id="4C290B9A1774E261FEDDFA6AFDDAB824" box="[329,621,1515,1540]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="groehnorum" status="sp. nov.">
<emphasis id="B95DAC0B1774E261FEDDFA6AFDDAB824" box="[329,621,1515,1540]" italics="true" pageId="2" pageNumber="65">Ecdyonurus groehnorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E11701774E261FDE0FA6BFD65B824" box="[628,722,1514,1540]" pageId="2" pageNumber="65" rank="species">
<emphasis id="B95DAC0B1774E261FDE0FA6BFD65B824" bold="true" box="[628,722,1514,1540]" pageId="2" pageNumber="65">sp. nov.</emphasis>
</taxonomicNameLabel>
belongs to the subgenus
<taxonomicName id="4C290B9A1774E261FC68FA6AFB30B824" box="[1020,1159,1515,1540]" class="Insecta" family="Heptageniidae" genus="Nestormeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B95DAC0B1774E261FC68FA6AFB30B824" box="[1020,1159,1515,1540]" italics="true" pageId="2" pageNumber="65">Nestormeus</emphasis>
</taxonomicName>
within the genus
<taxonomicName id="4C290B9A1774E261FAC3FA6AFF5FB80C" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B95DAC0B1774E261FAC3FA6AFF5FB80C" italics="true" pageId="2" pageNumber="65">Ecdyonurus</emphasis>
</taxonomicName>
, which can be distinguished from the other representatives of this genus by the combination of the following characters: (1) median impression of furcasternum of mesothorax with sub-parallel margins, narrowed posteriorly and distinctly divergent in its anterior part (see
<bibRefCitation id="EFB80DE81774E261FCDDF9E3FC60B85C" author="Kluge" box="[841,983,1634,1660]" pageId="2" pageNumber="68" refString="Kluge, N. J. (2004) The Phylogenetic System of Ephemeroptera (the first experience in consistently non-ranking taxonomy) Volume 1. Ephemeroptera except for Turbanoculata and Leptophlebia / fg 1. Kluwer Academic Publishers, Dordrecht-Hardbound, 456 pp." type="book" year="2004">Kluge 2004</bibRefCitation>
), and (2) lateroparapsidal sutures deep and slightly curved, noncontiguous with medioparapsidal sutures (see
<bibRefCitation id="EFB80DE81774E261FC5BF90BFB34B884" author="Godunko" box="[975,1155,1674,1700]" pageId="2" pageNumber="68" refString="Godunko, R. J. (2004) A new fossil subgenus and species of the genus Ecdyonurus Eaton, 1868 from Eocene Baltic amber (Ephemeroptera: Heptageniidae). Genus, 15 (3), 323 - 328." type="journal article" year="2004">Godunko 2004</bibRefCitation>
).
</paragraph>
<paragraph id="8B9670191774E261FF53F933FEFCB99C" blockId="2.[151,1437,874,1980]" pageId="2" pageNumber="65">
The new species can be easily distinguished from the earlier described
<taxonomicName id="4C290B9A1774E261FB9DF932FB01B8EC" box="[1033,1206,1715,1740]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="leopoliensis">
<emphasis id="B95DAC0B1774E261FB9DF932FB01B8EC" box="[1033,1206,1715,1740]" italics="true" pageId="2" pageNumber="65">E. leopoliensis</emphasis>
</taxonomicName>
by the shape of the subanal and (especially) the subgenital plates. In contrast to
<taxonomicName id="4C290B9A1774E261FCC1F95AFBB5B8D4" box="[853,1026,1755,1780]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="leopoliensis">
<emphasis id="B95DAC0B1774E261FCC1F95AFBB5B8D4" box="[853,1026,1755,1780]" italics="true" pageId="2" pageNumber="65">E. leopoliensis</emphasis>
</taxonomicName>
, which can be characterized by the presence of a rounded apically subgenital plate without any impression,
<taxonomicName id="4C290B9A1774E261FC79F882FB29B93C" box="[1005,1182,1795,1820]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="groehnorum" status="sp. nov.">
<emphasis id="B95DAC0B1774E261FC79F882FB29B93C" box="[1005,1182,1795,1820]" italics="true" pageId="2" pageNumber="65">E. groehnorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E11701774E261FB32F883FAB2B93C" box="[1190,1285,1794,1820]" pageId="2" pageNumber="65" rank="species">
<emphasis id="B95DAC0B1774E261FB32F883FAB2B93C" bold="true" box="[1190,1285,1794,1820]" pageId="2" pageNumber="65">sp. nov.</emphasis>
</taxonomicNameLabel>
bears a visible impression on the tip of the subgenital plate. The subanal plate of the new species is elongate with the tip marked apically, in contrast to the short and uniformly rounded subanal plate of
<taxonomicName id="4C290B9A1774E261FBD5F8D2FB5AB94C" box="[1089,1261,1875,1900]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="leopoliensis">
<emphasis id="B95DAC0B1774E261FBD5F8D2FB5AB94C" box="[1089,1261,1875,1900]" italics="true" pageId="2" pageNumber="65">E. leopoliensis</emphasis>
</taxonomicName>
. Other characters of the
<typeStatus id="5492CEBB1774E261FE81F8FBFECCB9B4" box="[277,379,1914,1940]" pageId="2" pageNumber="65" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C290B9A1774E261FE30F8FAFDE2B9B4" box="[420,597,1915,1940]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="groehnorum">
<emphasis id="B95DAC0B1774E261FE30F8FAFDE2B9B4" box="[420,597,1915,1940]" italics="true" pageId="2" pageNumber="65">E. groehnorum</emphasis>
</taxonomicName>
, i.e. body measurements and general aspects of coloration, are close to
<emphasis id="B95DAC0B1774E261FF03F822FEFCB99C" box="[151,331,1955,1980]" italics="true" pageId="2" pageNumber="65">
<taxonomicName id="4C290B9A1774E261FF03F822FEF1B99C" box="[151,326,1955,1980]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="leopoliensis">E. leopoliensis</taxonomicName>
.
</emphasis>
</paragraph>
<caption id="DF5620911775E260FF03F93FFAA7B8DC" httpUri="https://zenodo.org/record/179950/files/figure.png" pageId="3" pageNumber="66" targetBox="[165,1431,217,1671]" targetPageId="3">
<paragraph id="8B9670191775E260FF03F93FFAA7B8DC" blockId="3.[151,1435,1726,1788]" pageId="3" pageNumber="66">
<emphasis id="B95DAC0B1775E260FF03F93FFEF2B8F6" bold="true" box="[151,325,1726,1750]" pageId="3" pageNumber="66">FIGURES 13.</emphasis>
<taxonomicName id="4C290B9A1775E260FED9F93EFD46B8F6" box="[333,753,1726,1750]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="groehnorum" status="sp. nov." subGenus="Nestormeus">
<emphasis id="B95DAC0B1775E260FED9F93EFE7BB8F6" box="[333,460,1727,1750]" italics="true" pageId="3" pageNumber="66">Ecdyonurus</emphasis>
(
<emphasis id="B95DAC0B1775E260FE4AF93EFDEBB8F6" box="[478,604,1727,1750]" italics="true" pageId="3" pageNumber="66">Nestormeus</emphasis>
)
<emphasis id="B95DAC0B1775E260FDFAF93EFD46B8F6" box="[622,753,1727,1750]" italics="true" pageId="3" pageNumber="66">groehnorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E11701775E260FD6DF93FFCE6B8F6" box="[761,849,1726,1750]" pageId="3" pageNumber="66" rank="species">
<emphasis id="B95DAC0B1775E260FD6DF93FFCE6B8F6" bold="true" box="[761,849,1726,1750]" pageId="3" pageNumber="66">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, female imago: 1, head and thorax laterally and dorsally; 2, body ventrally; 3, tip of abdomen ventrally; 12: scale bars = 1 mm; 3: scale bar = 0.5 mm.
</paragraph>
</caption>
<caption id="DF5620911772E267FF03F8B7FDF5B950" httpUri="https://zenodo.org/record/179951/files/figure.png" pageId="4" pageNumber="67" targetBox="[432,1149,200,1811]" targetPageId="4">
<paragraph id="8B9670191772E267FF03F8B7FDF5B950" blockId="4.[151,1436,1846,1904]" pageId="4" pageNumber="67">
<emphasis id="B95DAC0B1772E267FF03F8B7FEF5B96E" bold="true" box="[151,322,1846,1870]" pageId="4" pageNumber="67">FIGURES 46.</emphasis>
<taxonomicName id="4C290B9A1772E267FEDDF8B6FD50B96E" box="[329,743,1846,1870]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="groehnorum" status="sp. nov." subGenus="Nestormeus">
<emphasis id="B95DAC0B1772E267FEDDF8B6FE7FB96E" box="[329,456,1847,1870]" italics="true" pageId="4" pageNumber="67">Ecdyonurus</emphasis>
(
<emphasis id="B95DAC0B1772E267FE43F8B6FDE2B96E" box="[471,597,1847,1870]" italics="true" pageId="4" pageNumber="67">Nestormeus</emphasis>
)
<emphasis id="B95DAC0B1772E267FDF0F8B6FD50B96E" box="[612,743,1847,1870]" italics="true" pageId="4" pageNumber="67">groehnorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E11701772E267FD7AF8B7FCF4B96E" box="[750,835,1846,1870]" pageId="4" pageNumber="67" rank="species">
<emphasis id="B95DAC0B1772E267FD7AF8B7FCF4B96E" bold="true" box="[750,835,1846,1870]" pageId="4" pageNumber="67">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, female imago: 4, tip of abdomen ventrally; 5, mesothorax ventrally; 6, body ventrally.
</paragraph>
</caption>
<paragraph id="8B9670191773E266FF53FF19FD1CBF72" blockId="5.[151,1436,152,578]" pageId="5" pageNumber="68">
In the previous publication on the subgenus
<taxonomicName id="4C290B9A1773E266FD42FF19FCD6BE91" box="[726,865,152,177]" class="Insecta" family="Heptageniidae" genus="Nestormeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis id="B95DAC0B1773E266FD42FF19FCD6BE91" box="[726,865,152,177]" italics="true" pageId="5" pageNumber="68">Nestormeus</emphasis>
</taxonomicName>
, the prothorax structure has not been described, because the anterior part of the
<typeStatus id="5492CEBB1773E266FD9AFF41FDF7BEFA" box="[526,576,192,218]" pageId="5" pageNumber="68">type</typeStatus>
specimen of
<taxonomicName id="4C290B9A1773E266FD75FF41FC39BEF9" box="[737,910,192,217]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="leopoliensis">
<emphasis id="B95DAC0B1773E266FD75FF41FC39BEF9" box="[737,910,192,217]" italics="true" pageId="5" pageNumber="68">E. leopoliensis</emphasis>
</taxonomicName>
is invisible in ventral view (
<bibRefCitation id="EFB80DE81773E266FB76FF41FEAFBF22" author="Godunko" pageId="5" pageNumber="68" refString="Godunko, R. J. (2004) A new fossil subgenus and species of the genus Ecdyonurus Eaton, 1868 from Eocene Baltic amber (Ephemeroptera: Heptageniidae). Genus, 15 (3), 323 - 328." type="journal article" year="2004">
Godunko 2004: 326,
<figureCitation id="13126C9C1773E266FF45FF69FEAFBF22" box="[209,280,232,258]" captionStart="FIGURES 1 3" captionStartId="3.[151,269,1726,1750]" captionTargetBox="[165,1431,217,1671]" captionTargetId="figure@3.[150,1456,191,1702]" captionTargetPageId="3" captionText="FIGURES 1 3. Ecdyonurus (Nestormeus) groehnorum sp. nov., holotype, female imago: 1, head and thorax laterally and dorsally; 2, body ventrally; 3, tip of abdomen ventrally; 1 2: scale bars = 1 mm; 3: scale bar = 0.5 mm." httpUri="https://zenodo.org/record/179950/files/figure.png" pageId="5" pageNumber="68">Fig. 2</figureCitation>
</bibRefCitation>
). Thus, the examination of the prothorax peculiarities in
<taxonomicName id="4C290B9A1773E266FC26FF69FBD5BF21" box="[946,1122,232,257]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="groehnorum" status="sp. nov.">
<emphasis id="B95DAC0B1773E266FC26FF69FBD5BF21" box="[946,1122,232,257]" italics="true" pageId="5" pageNumber="68">E. groehnorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E11701773E266FBFDFF66FB71BF21" box="[1129,1222,231,257]" pageId="5" pageNumber="68" rank="species">
<emphasis id="B95DAC0B1773E266FBFDFF66FB71BF21" bold="true" box="[1129,1222,231,257]" pageId="5" pageNumber="68">sp. nov.</emphasis>
</taxonomicNameLabel>
may contribute to the diagnosis of this fossil subgenus, showing the presences of the prosternum without a transversal ridge (typical of
<taxonomicName id="4C290B9A1773E266FE83FEB9FE14BF71" box="[279,419,312,337]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B95DAC0B1773E266FE83FEB9FE14BF71" box="[279,419,312,337]" italics="true" pageId="5" pageNumber="68">Ecdyonurus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E11701773E266FE3FFEB9FE62BF72" box="[427,469,312,338]" pageId="5" pageNumber="68" sensu="lato">s. l.</taxonomicNameLabel>
, see
<bibRefCitation id="EFB80DE81773E266FD84FEB9FD29BF72" author="Kluge" box="[528,670,312,338]" pageId="5" pageNumber="68" refString="Kluge, N. J. (2004) The Phylogenetic System of Ephemeroptera (the first experience in consistently non-ranking taxonomy) Volume 1. Ephemeroptera except for Turbanoculata and Leptophlebia / fg 1. Kluwer Academic Publishers, Dordrecht-Hardbound, 456 pp." type="book" year="2004">Kluge 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33323921773E266FF53FEDEFE13BFD2" pageId="5" pageNumber="68" type="materials_examined">
<paragraph id="8B9670191773E266FF53FEDEFE13BFD2" blockId="5.[151,1436,152,578]" pageId="5" pageNumber="68">
<emphasis id="B95DAC0B1773E266FF53FEDEFEBEBF59" bold="true" box="[199,265,351,377]" pageId="5" pageNumber="68">
<typeStatus id="5492CEBB1773E266FF53FEDEFEB5BF59" box="[199,258,351,377]" pageId="5" pageNumber="68">Type</typeStatus>
.
</emphasis>
<typeStatus id="5492CEBB1773E266FE87FEE1FE32BF5A" box="[275,389,352,378]" pageId="5" pageNumber="68" type="holotype">Holotype</typeStatus>
: female imago in the Baltic amber (Eocene), with well preserved body, being well visible from ventral view, and partly from dorsal and lateral views. The
<typeStatus id="5492CEBB1773E266FC37FE09FBBEBF82" box="[931,1033,392,418]" pageId="5" pageNumber="68" type="holotype">holotype</typeStatus>
is housed in the collection of the Geological-Palaeontological Museum, University of Hamburg (
<collectingCountry id="F33E30891773E266FC33FE31FBADBFEA" box="[935,1050,432,458]" name="Germany" pageId="5" pageNumber="68">Germany</collectingCountry>
), S.G.P.I.H. no. 4451 (ex coll. Carsten Gröhn no. 76).
</paragraph>
</subSubSection>
<subSubSection id="C33323921773E266FF53FE7EFE5DBC62" pageId="5" pageNumber="68" type="etymology">
<paragraph id="8B9670191773E266FF53FE7EFE5DBC62" blockId="5.[151,1436,152,578]" pageId="5" pageNumber="68">
<emphasis id="B95DAC0B1773E266FF53FE7EFEE4BC39" bold="true" box="[199,339,511,537]" pageId="5" pageNumber="68">Etymology.</emphasis>
The new species is named in honor of Jutta and Carsten Gröhn with hearty gratitude for supporting my stay in Hamburg.
</paragraph>
</subSubSection>
</treatment>
</document>