treatments-xml/data/03/80/AC/0380AC3BFF9FFFE7028BFC12FB4929BD.xml

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<mods:title id="FF6032ECBFD6CA533A2F303229850FE0">New and little known species of Halolaelaps (Acari: Mesostigmata: Halolaelapidae) from Ukraine</mods:title>
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<paragraph id="8B961D2DFF9FFFE1028BFC12FD142BCB" blockId="8.[151,717,907,967]" box="[151,717,907,934]" pageId="8" pageNumber="444">
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<taxonomicName id="4C2966AEFF9FFFE1028BFC12FD142BCB" authority="Hirschmann &amp; Gotz" authorityName="Hirschmann &amp; Gotz" box="[151,717,907,934]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="444" phylum="Arthropoda" rank="species" species="saproincisus">
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<emphasis id="B95DC13FFF9FFFE1028BFC12FE1F2BC8" bold="true" box="[151,454,907,933]" italics="true" pageId="8" pageNumber="444">Halolaelaps saproincisus</emphasis>
Hirschmann &amp; Götz
</emphasis>
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</paragraph>
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(
<figureCitation id="131201A8FF9FFFE10283FC37FEC62BAA" box="[159,287,942,967]" captionStart-0="FIGURE 20 – 24" captionStart-1="FIGURES 25 – 28" captionStart-2="FIGURE 29 – 33" captionStart-3="FIGURE 34 – 35" captionStartId-0="9.[151,250,1629,1651]" captionStartId-1="10.[151,264,1774,1797]" captionStartId-2="11.[151,250,1934,1956]" captionStartId-3="12.[151,250,1294,1316]" captionTargetBox-0="[155,1431,203,1599]" captionTargetBox-1="[165,1422,418,1748]" captionTargetBox-2="[157,1426,637,1902]" captionTargetBox-3="[159,1431,199,1268]" captionTargetId-0="figure@9.[151,1436,193,1608]" captionTargetId-1="figure@10.[150,1436,409,1754]" captionTargetId-2="figure@11.[151,1436,625,1913]" captionTargetId-3="figure@12.[150,1436,193,1273]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionText-0="FIGURE 20 – 24. Halolaelaps saproincisus. Adult female. 20. Lateral (antiaxial) view of chelicera; 21. Epistome; 22. Hypostome (without palp tibia and palp tarsus); 23. Dorsal idiosoma; 24. Ventral idiosoma. Scale bar = 50 µm." captionText-1="FIGURES 25 – 28. Halolaelaps saproincisus. Adult female. 25. Leg I (coxa to tibia); 26. Leg II; 27. Leg III; 28. Leg IV. Scale bar = 100 µm." captionText-2="FIGURE 29 – 33. Halolaelaps saproincisus. Adult male. 29. Lateral (antiaxial) view of chelicerae; 30 Epistome; 31. Dorsal idiosoma; 32. Ventral idiosoma; 33. Femur of leg II. Scale bar = 50 µm." captionText-3="FIGURE 34 – 35. Halolaelaps saproincisus. Deutonymph. 34. Dorsal idiosoma; 35. Ventral idiosoma. Scale bar = 50 µm." httpUri-0="https://zenodo.org/record/264418/files/figure.png" httpUri-1="https://zenodo.org/record/264419/files/figure.png" httpUri-2="https://zenodo.org/record/264420/files/figure.png" httpUri-3="https://zenodo.org/record/264421/files/figure.png" pageId="8" pageNumber="444">Figs 20–35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3334EA6FF9FFFE1028BFC6FFB632C05" pageId="8" pageNumber="444" type="reference_group">
<paragraph id="8B961D2DFF9FFFE1028BFC6FFD122C61" blockId="8.[151,1436,1014,1128]" box="[151,715,1014,1037]" pageId="8" pageNumber="444">
<treatmentCitationGroup id="AB393A03FF9FFFE1028BFC6FFD122C61" box="[151,715,1014,1037]" pageId="8" pageNumber="444">
<taxonomicName id="4C2966AEFF9FFFE1028BFC6FFD1E2C61" authority="Hirschmann &amp; Gotz, 1968: 9" authorityName="Hirschmann &amp; Gotz" authorityPageNumber="9" authorityYear="1968" box="[151,711,1014,1037]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="444" phylum="Arthropoda" rank="species" species="saproincisus">
<emphasis id="B95DC13FFF9FFFE1028BFC6FFE412C60" box="[151,408,1014,1037]" italics="true" pageId="8" pageNumber="444">Halolaelaps saproincisus</emphasis>
<treatmentCitation id="0A883B3CFF9FFFE10382FC6FFD1E2C61" author="Hirschmann" box="[414,711,1014,1036]" page="9" pageId="8" pageNumber="444" year="1968">
<bibRefCitation id="EFB860DCFF9FFFE10382FC6FFD762C61" author="Hirschmann" box="[414,687,1014,1036]" pageId="8" pageNumber="444" refString="Hirschmann, W. &amp; Gotz, H. (1968) Neue Halolaelaps - Arten. Acarologie. Schriftenreihe fur Vergleichende Milbenkunde, 11, 7 - 10." type="journal article" year="1968">Hirschmann &amp; Götz, 1968</bibRefCitation>
: 9
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
<paragraph id="8B961D2DFF9FFFE1028BFB8CFCE42C27" blockId="8.[151,1436,1014,1128]" pageId="8" pageNumber="444">
<treatmentCitationGroup id="AB393A03FF9FFFE1028BFB8CFCE42C27" pageId="8" pageNumber="444">
<taxonomicName id="4C2966AEFF9FFFE1028BFB8CFE422C46" box="[151,411,1045,1067]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="444" phylum="Arthropoda" rank="species" species="saproincisus">
<emphasis id="B95DC13FFF9FFFE1028BFB8CFE422C46" box="[151,411,1045,1067]" italics="true" pageId="8" pageNumber="444">Halolaelaps saproincisus</emphasis>
</taxonomicName>
.— 
<treatmentCitation id="0A883B3CFF9FFFE103A6FB8FFDB32C46" author="Karg" box="[442,618,1045,1067]" page="294" pageId="8" pageNumber="444" year="1971">
<bibRefCitation id="EFB860DCFF9FFFE103A6FB8FFDED2C46" author="Karg" box="[442,564,1045,1067]" pageId="8" pageNumber="444" refString="Karg, W. (1971) Acari (Acarina), Milben. Unterordnung Anactinochaeta (Parasitiformes). Die freilebenden Gamasina (Gamasides). Raubmilben. Die Tierwelt Deutschlands, 59, 1 - 475." type="journal article" year="1971">Karg, 1971</bibRefCitation>
: 294
</treatmentCitation>
, 1993: 300; 
<treatmentCitation id="0A883B3CFF9FFFE100EDFB8FFBB12C46" author="Bregetova" box="[753,1128,1045,1067]" page="296" pageId="8" pageNumber="444" year="1977">
<bibRefCitation id="EFB860DCFF9FFFE100EDFB8FFBE82C46" author="Bregetova" box="[753,1073,1045,1067]" pageId="8" pageNumber="444" refString="Bregetova, N. G. &amp; Shcherbak, G. I. (1977) Family Rhodacaridae Oudemans, 1902. In: Ghilyarov, M. S. &amp; Bregetova, N. G. (Eds.), Key to the Soil - Inhabiting Mites. Mesostigmata, Nauka, Leningrad, pp. 256 - 307. [in Russian]" type="book chapter" year="1977">Bregetova &amp; Shcherbak, 1977</bibRefCitation>
: 296
</treatmentCitation>
; 
<treatmentCitation id="0A883B3CFF9FFFE10669FB8CFEFA2C27" author="Koyumdjieva" page="18" pageId="8" pageNumber="444" year="1985">
<bibRefCitation id="EFB860DCFF9FFFE10669FB8CFF242C27" author="Koyumdjieva" pageId="8" pageNumber="444" refString="Koyumdjieva, M. I. &amp; Angelkova, E. B. (1985) Species composition and population dynamics of mites (Gamasina, Mesostigmata and Acaridiae, Astigmata) in a dunghill. Acta Zoologica Bulgarica, 28, 17 - 27." type="journal article" year="1985">Koyumdjieva &amp; Angelkova, 1985</bibRefCitation>
: 18
</treatmentCitation>
; 
<treatmentCitation id="0A883B3CFF9FFFE10331FBADFD582C27" author="Gwiazdowicz" box="[301,641,1076,1098]" page="14" pageId="8" pageNumber="444" year="2004">
<bibRefCitation id="EFB860DCFF9FFFE10331FBADFD822C27" author="Gwiazdowicz" box="[301,603,1076,1098]" pageId="8" pageNumber="444" refString="Gwiazdowicz, D. J. &amp; Klemt, J. (2004) Mesostigmatic mites (Acari, Gamasida) in selected microhabitats of the Biebrza National Park (NE Poland). Biology Letters, 41, 11 - 19." type="journal article" year="2004">Gwiazdowicz &amp; Klemt, 2004</bibRefCitation>
: 14
</treatmentCitation>
; 
<treatmentCitation id="0A883B3CFF9FFFE10097FBACFCE02C27" author="Krantz" box="[651,825,1076,1098]" page="13" pageId="8" pageNumber="444" year="2016">
<bibRefCitation id="EFB860DCFF9FFFE10097FBACFCCA2C27" author="Krantz" box="[651,787,1076,1098]" pageId="8" pageNumber="444" refString="Krantz, G. W. (2016) A new species of Halolaelapidae (Acari: Mesostigmata: Rhodacaroidea) from beach wrack in Yaquina Bay, Oregon, USA, with comments on opisthonotal plasticity and cribral development in the family. Journal of Natural History, 1 - 16. http: // dx. doi. org / 10.1080 / 00222933.2016.1170904" type="book chapter" year="2016">Krantz, 2016</bibRefCitation>
: 13
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B961D2DFF9FFFE1028BFBCBFB632C05" blockId="8.[151,1436,1014,1128]" box="[151,1210,1106,1128]" pageId="8" pageNumber="444">
<treatmentCitationGroup id="AB393A03FF9FFFE1028BFBCBFB632C05" box="[151,1210,1106,1128]" pageId="8" pageNumber="444">
<taxonomicName id="4C2966AEFF9FFFE1028BFBCBFD8A2C05" box="[151,595,1106,1128]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="444" phylum="Arthropoda" rank="species" species="saproincisus" subGenus="Halogamasellus">
<emphasis id="B95DC13FFF9FFFE1028BFBCBFECA2C05" box="[151,275,1106,1128]" italics="true" pageId="8" pageNumber="444">Halolaelaps</emphasis>
(
<emphasis id="B95DC13FFF9FFFE1033DFBCBFE1C2C05" box="[289,453,1106,1128]" italics="true" pageId="8" pageNumber="444">Halogamasellus</emphasis>
) 
<emphasis id="B95DC13FFF9FFFE103C8FBCAFD8A2C05" box="[468,595,1107,1128]" italics="true" pageId="8" pageNumber="444">saproincisus</emphasis>
</taxonomicName>
.— 
<treatmentCitation id="0A883B3CFF9FFFE1006DFBCBFC1F2C05" author="Blaszak" box="[625,966,1106,1128]" page="44" pageId="8" pageNumber="444" year="1995">
<bibRefCitation id="EFB860DCFF9FFFE1006DFBCBFC792C05" author="Blaszak" box="[625,928,1106,1128]" pageId="8" pageNumber="444" refString="Blaszak, C. &amp; Ehrnsberger, R. (1995) Beitrage zur Kenntnis von Halolaelaps (Halogamasellus subgen. nov.), (Acari: Gamasida: Halolaelapidae). Osnabrucker Naturwissenschaftliche Mitteilungen, 20 / 21, 25 - 94." type="journal article" year="1995">Błaszak &amp; Ehrnsberger, 1995</bibRefCitation>
: 44
</treatmentCitation>
; 
<treatmentCitation id="0A883B3CFF9FFFE101CDFBCBFB6F2C05" author="Blaszak" box="[977,1206,1106,1128]" page="6" pageId="8" pageNumber="444" year="2004">
<bibRefCitation id="EFB860DCFF9FFFE101CDFBCBFB442C05" author="Blaszak" box="[977,1181,1106,1128]" pageId="8" pageNumber="444" refString="Blaszak, C., Ehrnsberger, R. &amp; Michalik, J. (2004) Die Milben in der Zoologischen Staatssammlung Munchen. Teil 5. Gattung: Halolaelaps Berlese &amp; Trouessart, 1889 (Acari: Gamasida, Halolaelapidae). Spixiana, 27, 1 - 13." type="journal article" year="2004">
Błaszak 
<emphasis id="B95DC13FFF9FFFE10634FBCAFB8D2C05" box="[1064,1108,1106,1128]" italics="true" pageId="8" pageNumber="444">et al</emphasis>
., 2004
</bibRefCitation>
: 6
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C3334EA6FF9FFFE1028BFB0CFCF72DEB" pageId="8" pageNumber="444" type="diagnosis">
<paragraph id="8B961D2DFF9FFFE1028BFB0CFF172D0F" blockId="8.[151,1437,1173,2026]" pageId="8" pageNumber="444">
<emphasis id="B95DC13FFF9FFFE1028BFB0CFE582CC3" bold="true" box="[151,385,1173,1198]" pageId="8" pageNumber="444">Diagnosis of adults.</emphasis>
Epistome with serrated anterior margin; peritremes long; femur I with 13 setae; coxa II with anterior spur; genu III and IV with nine setae; legs without apophyses. Adult female with 23 pairs of setae on podonotal shield and 14 pairs of setae on opisthonotal shield; anterior margin of opisthonotal shield with narrow and deep notch; most dorsal setae serrated. The adult male with 24 pairs of setae on podonotal shield and 18 pairs of setae on opisthonotal shield; opisthonotal shield without notch and incisions; anal shield with circum-anal setae only.
</paragraph>
<paragraph id="8B961D2DFF9FFFE102DBFAF4FCF72DEB" blockId="8.[151,1437,1173,2026]" box="[199,814,1389,1414]" pageId="8" pageNumber="444">
<emphasis id="B95DC13FFF9FFFE102DBFAF4FEB82DEB" bold="true" box="[199,353,1389,1414]" pageId="8" pageNumber="444">Adult female</emphasis>
(
<figureCitation id="131201A8FF9FFFE1036CFAF4FE2D2DEB" box="[368,500,1389,1414]" captionStart="FIGURE 20 – 24" captionStartId="9.[151,250,1629,1651]" captionTargetBox="[155,1431,203,1599]" captionTargetId="figure@9.[151,1436,193,1608]" captionTargetPageId="9" captionText="FIGURE 20 – 24. Halolaelaps saproincisus. Adult female. 20. Lateral (antiaxial) view of chelicera; 21. Epistome; 22. Hypostome (without palp tibia and palp tarsus); 23. Dorsal idiosoma; 24. Ventral idiosoma. Scale bar = 50 µm." httpUri="https://zenodo.org/record/264418/files/figure.png" pageId="8" pageNumber="444">Figs 20–24</figureCitation>
). Five specimens measured.
</paragraph>
</subSubSection>
<subSubSection id="C3334EA6FF9FFFE402DBFA08FEE62C18" lastPageId="13" lastPageNumber="449" pageId="8" pageNumber="444" type="description">
<paragraph id="8B961D2DFF9FFFE102DBFA08FE832ECB" blockId="8.[151,1437,1173,2026]" pageId="8" pageNumber="444">
<emphasis id="B95DC13FFF9FFFE102DBFA08FE862DC7" bold="true" box="[199,351,1425,1450]" pageId="8" pageNumber="444">Gnathosoma</emphasis>
. Second cheliceral segment 88–95 long, fixed cheliceral digit with three proximal teeth and one or two small preapical teeth in addition to apical tooth and setiform pilus dentilis, movable cheliceral digit with three teeth in addition to apical tooth (
<figureCitation id="131201A8FF9FFFE10048FA40FD722D9F" box="[596,683,1497,1522]" captionStart="FIGURE 20 – 24" captionStartId="9.[151,250,1629,1651]" captionTargetBox="[155,1431,203,1599]" captionTargetId="figure@9.[151,1436,193,1608]" captionTargetPageId="9" captionText="FIGURE 20 – 24. Halolaelaps saproincisus. Adult female. 20. Lateral (antiaxial) view of chelicera; 21. Epistome; 22. Hypostome (without palp tibia and palp tarsus); 23. Dorsal idiosoma; 24. Ventral idiosoma. Scale bar = 50 µm." httpUri="https://zenodo.org/record/264418/files/figure.png" pageId="8" pageNumber="444">Fig. 20</figureCitation>
). Epistome with serrated anterior margin (
<figureCitation id="131201A8FF9FFFE1068EFA40FB312D9F" box="[1170,1256,1497,1522]" captionStart="FIGURE 20 – 24" captionStartId="9.[151,250,1629,1651]" captionTargetBox="[155,1431,203,1599]" captionTargetId="figure@9.[151,1436,193,1608]" captionTargetPageId="9" captionText="FIGURE 20 – 24. Halolaelaps saproincisus. Adult female. 20. Lateral (antiaxial) view of chelicera; 21. Epistome; 22. Hypostome (without palp tibia and palp tarsus); 23. Dorsal idiosoma; 24. Ventral idiosoma. Scale bar = 50 µm." httpUri="https://zenodo.org/record/264418/files/figure.png" pageId="8" pageNumber="444">Fig. 21</figureCitation>
). Palp 132–145 long; number of setae (trocanter to tarsus): 
<date id="FF973BEDFF9FFFE10099FA64FD332E7B" box="[645,746,1533,1558]" pageId="8" pageNumber="444" value="2014-05-02" valueMax="2014-06-02" valueMin="2014-05-02">2-5-6-14</date>
-15; apotele 3-tined; seta 
<emphasis id="B95DC13FFF9FFFE10615FA67FBFD2E7B" box="[1033,1060,1534,1558]" italics="true" pageId="8" pageNumber="444">
<date id="FF973BEDFF9FFFE10615FA67FBFD2E7B" box="[1033,1060,1534,1558]" pageId="8" pageNumber="444">v2</date>
</emphasis>
on trochanter, 
<emphasis id="B95DC13FFF9FFFE106CCFA64FB3C2E7B" box="[1232,1253,1533,1558]" italics="true" pageId="8" pageNumber="444">al</emphasis>
on femur, setae 
<emphasis id="B95DC13FFF9FFFE1028BF9B8FF622E57" box="[151,187,1569,1594]" italics="true" pageId="8" pageNumber="444">al1</emphasis>
and 
<emphasis id="B95DC13FFF9FFFE102E9F9B8FEC12E57" box="[245,280,1569,1594]" italics="true" pageId="8" pageNumber="444">al2</emphasis>
on genu slightly thickened. Subcapitulum 86–101 wide at widest level (
<figureCitation id="131201A8FF9FFFE1064BF9B8FB772E57" box="[1111,1198,1569,1594]" captionStart="FIGURE 20 – 24" captionStartId="9.[151,250,1629,1651]" captionTargetBox="[155,1431,203,1599]" captionTargetId="figure@9.[151,1436,193,1608]" captionTargetPageId="9" captionText="FIGURE 20 – 24. Halolaelaps saproincisus. Adult female. 20. Lateral (antiaxial) view of chelicera; 21. Epistome; 22. Hypostome (without palp tibia and palp tarsus); 23. Dorsal idiosoma; 24. Ventral idiosoma. Scale bar = 50 µm." httpUri="https://zenodo.org/record/264418/files/figure.png" pageId="8" pageNumber="444">Fig. 22</figureCitation>
). Deutosternum with seven (rarely six) rows of deutosternal denticles (4–13 denticles per row). Internal malae short, outer branch of each internal mala and inner branch smooth. Corniculus horn-like. Measurements of setae: 
<emphasis id="B95DC13FFF9FFFE1068FF9F0FB772EEF" box="[1171,1198,1641,1666]" italics="true" pageId="8" pageNumber="444">
h 
<date id="FF973BEDFF9FFFE106BEF9F0FB772EEF" box="[1186,1198,1641,1666]" pageId="8" pageNumber="444" value="1923-01-27">1</date>
</emphasis>
23–27, 
<emphasis id="B95DC13FFF9FFFE10715F9F0FAFF2EEF" box="[1289,1318,1641,1666]" italics="true" pageId="8" pageNumber="444">
h 
<date id="FF973BEDFF9FFFE10704F9F0FAFF2EEF" box="[1304,1318,1641,1666]" pageId="8" pageNumber="444" value="2013-02-15">2</date>
</emphasis>
13–15, 
<emphasis id="B95DC13FFF9FFFE1079CF9F0FA452EEF" box="[1408,1436,1641,1666]" italics="true" pageId="8" pageNumber="444">
h 
<date id="FF973BEDFF9FFFE10792F9F0FA452EEF" box="[1422,1436,1641,1666]" pageId="8" pageNumber="444" value="1932-03-25">3</date>
</emphasis>
25–32, 
<emphasis id="B95DC13FFF9FFFE102F7F917FEDC2EC8" box="[235,261,1678,1701]" italics="true" pageId="8" pageNumber="444">pc</emphasis>
19–23.
</paragraph>
<paragraph id="8B961D2DFF9FFFE102DBF928FDB62F87" blockId="8.[151,1437,1173,2026]" pageId="8" pageNumber="444">
<emphasis id="B95DC13FFF9FFFE102DBF928FE532EA7" bold="true" box="[199,394,1713,1738]" pageId="8" pageNumber="444">Dorsal idiosoma</emphasis>
(
<figureCitation id="131201A8FF9FFFE10386F928FE362EA7" box="[410,495,1713,1738]" captionStart="FIGURE 20 – 24" captionStartId="9.[151,250,1629,1651]" captionTargetBox="[155,1431,203,1599]" captionTargetId="figure@9.[151,1436,193,1608]" captionTargetPageId="9" captionText="FIGURE 20 – 24. Halolaelaps saproincisus. Adult female. 20. Lateral (antiaxial) view of chelicera; 21. Epistome; 22. Hypostome (without palp tibia and palp tarsus); 23. Dorsal idiosoma; 24. Ventral idiosoma. Scale bar = 50 µm." httpUri="https://zenodo.org/record/264418/files/figure.png" pageId="8" pageNumber="444">Fig. 23</figureCitation>
). Idiosoma ovate. Podonotal shield 218–244 long and 244–281 wide at level of 
<emphasis id="B95DC13FFF9FFFE10760F92BFA4D2EA7" box="[1404,1428,1714,1738]" italics="true" pageId="8" pageNumber="444">s4</emphasis>
, strongly tapering anteriorly from 
<emphasis id="B95DC13FFF9FFFE10010F94FFDFC2E83" box="[524,549,1750,1774]" italics="true" pageId="8" pageNumber="444">r3</emphasis>
; reticulated; with 23 pairs of setae (
<emphasis id="B95DC13FFF9FFFE101A9F94FFC362E83" box="[949,1007,1749,1774]" italics="true" pageId="8" pageNumber="444">j1–j6</emphasis>
, 
<emphasis id="B95DC13FFF9FFFE101E0F94FFBE52E83" box="[1020,1084,1749,1774]" italics="true" pageId="8" pageNumber="444">z1–z6</emphasis>
, 
<emphasis id="B95DC13FFF9FFFE10656F94FFB532E83" box="[1098,1162,1749,1774]" italics="true" pageId="8" pageNumber="444">s1–s6</emphasis>
, 
<emphasis id="B95DC13FFF9FFFE1068BF94FFB0E2E83" box="[1175,1239,1750,1774]" italics="true" pageId="8" pageNumber="444">r1–r5</emphasis>
) and five pairs of distinguishable pores. Unsclerotised cuticle laterad of podonotal shield with setae 
<emphasis id="B95DC13FFF9FFFE1062DF960FB902F7F" box="[1073,1097,1785,1810]" italics="true" pageId="8" pageNumber="444">r6</emphasis>
. Opisthonotal shield 244–286 long and 239–277 wide at level 
<emphasis id="B95DC13FFF9FFFE103E2F887FDC32F5B" box="[510,538,1822,1846]" italics="true" pageId="8" pageNumber="444">S3</emphasis>
; reticulated; anterior margin with narrow and deep notch; posterior margin with cribrum; rear corners of shield curved ventrally; with 14 pairs of setae (
<emphasis id="B95DC13FFF9FFFE101A0F8DBFC0F2F37" box="[956,982,1858,1882]" italics="true" pageId="8" pageNumber="444">J1</emphasis>
, 
<emphasis id="B95DC13FFF9FFFE101F9F8DBFC262F37" box="[997,1023,1858,1882]" italics="true" pageId="8" pageNumber="444">J2</emphasis>
, 
<emphasis id="B95DC13FFF9FFFE10612F8DBFBF12F37" box="[1038,1064,1858,1882]" italics="true" pageId="8" pageNumber="444">J4</emphasis>
, 
<emphasis id="B95DC13FFF9FFFE1062AF8DBFB882F34" box="[1078,1105,1858,1881]" italics="true" pageId="8" pageNumber="444">J5</emphasis>
, 
<emphasis id="B95DC13FFF9FFFE10643F8DBFB712F34" box="[1119,1192,1858,1882]" italics="true" pageId="8" pageNumber="444">Z1–Z5</emphasis>
, 
<emphasis id="B95DC13FFF9FFFE106ABF8DBFB252F37" box="[1207,1276,1858,1882]" italics="true" pageId="8" pageNumber="444">S1–S5</emphasis>
) and six pairs of distinguishable pores. Unsclerotised cuticle laterad of opisthonotal shield with five pairs of setae (
<emphasis id="B95DC13FFF9FFFE10703F8FFFAB52F10" box="[1311,1388,1894,1918]" italics="true" pageId="8" pageNumber="444">R1–R5</emphasis>
; 
<emphasis id="B95DC13FFF9FFFE10761F8FFFA452F10" box="[1405,1436,1894,1917]" italics="true" pageId="8" pageNumber="444">R5</emphasis>
visible in the ventrolateral regions). Measurements of setae: 
<emphasis id="B95DC13FFF9FFFE10150F813FCBA2FCF" box="[844,867,1930,1954]" italics="true" pageId="8" pageNumber="444">
j 
<date id="FF973BEDFF9FFFE10144F813FCBA2FCF" box="[856,867,1930,1954]" pageId="8" pageNumber="444" value="2015-01-19">1</date>
</emphasis>
15–19, 
<emphasis id="B95DC13FFF9FFFE101DDF813FC002FCF" box="[961,985,1930,1954]" italics="true" pageId="8" pageNumber="444">
z 
<date id="FF973BEDFF9FFFE101D1F813FC002FCF" box="[973,985,1930,1954]" pageId="8" pageNumber="444" value="2013-11-01">1</date>
</emphasis>
11–13, other setae on podonotal shield 23–32, setae on opisthonotal shiels 25–34, setae on unsclerotised cuticle 21–27. Setae 
<emphasis id="B95DC13FFF9FFFE10694F837FB462FAB" box="[1160,1183,1966,1990]" italics="true" pageId="8" pageNumber="444">j1</emphasis>
and 
<emphasis id="B95DC13FFF9FFFE106C7F837FB2D2FAB" box="[1243,1268,1966,1990]" italics="true" pageId="8" pageNumber="444">z1</emphasis>
smooth, other dorsal setae more or less serrated apically.
</paragraph>
<caption id="DF564DA5FF9EFFE0028BF9C4FB232EFF" httpUri="https://zenodo.org/record/264418/files/figure.png" pageId="9" pageNumber="445" targetBox="[155,1431,203,1599]" targetPageId="9">
<paragraph id="8B961D2DFF9EFFE0028BF9C4FB232EFF" blockId="9.[151,1436,1629,1682]" pageId="9" pageNumber="445">
<emphasis id="B95DC13FFF9EFFE0028BF9C4FE942E1E" bold="true" box="[151,333,1629,1651]" pageId="9" pageNumber="445">FIGURE 20–24.</emphasis>
<taxonomicName id="4C2966AEFF9EFFE00345F9C4FDB82E19" box="[345,609,1629,1652]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="445" phylum="Arthropoda" rank="species" species="saproincisus">
<emphasis id="B95DC13FFF9EFFE00345F9C4FDB82E19" box="[345,609,1629,1652]" italics="true" pageId="9" pageNumber="445">Halolaelaps saproincisus</emphasis>
</taxonomicName>
. Adult female. 20. Lateral (antiaxial) view of chelicera; 21. Epistome; 22. Hypostome (without palp tibia and palp tarsus); 23. Dorsal idiosoma; 24. Ventral idiosoma. Scale bar = 50 µm.
</paragraph>
</caption>
<paragraph id="8B961D2DFF9EFFE302DBF920FE0E2908" blockId="9.[151,1437,1721,2034]" lastBlockId="10.[151,1436,151,357]" lastPageId="10" lastPageNumber="446" pageId="9" pageNumber="445">
<emphasis id="B95DC13FFF9EFFE002DBF920FE4D2EBF" bold="true" box="[199,404,1721,1746]" pageId="9" pageNumber="445">Ventral idiosoma</emphasis>
(
<figureCitation id="131201A8FF9EFFE003BFF920FE212EBF" box="[419,504,1721,1746]" captionStart="FIGURE 20 – 24" captionStartId="9.[151,250,1629,1651]" captionTargetBox="[155,1431,203,1599]" captionTargetId="figure@9.[151,1436,193,1608]" captionTargetPageId="9" captionText="FIGURE 20 – 24. Halolaelaps saproincisus. Adult female. 20. Lateral (antiaxial) view of chelicera; 21. Epistome; 22. Hypostome (without palp tibia and palp tarsus); 23. Dorsal idiosoma; 24. Ventral idiosoma. Scale bar = 50 µm." httpUri="https://zenodo.org/record/264418/files/figure.png" pageId="9" pageNumber="445">Fig. 24</figureCitation>
). Tritosternum with wide trapezoidal base and pilose laciniae, base of tritosternum length 21–27, laciniae length 42–55. Presternal area lightly sclerotised. Sternal shield rectangular; almost smooth; approximately 82–97 long (without presternal area) and 65–78 wide at widest level; with three pairs of setae and two pairs of lyrifissures. Setae 
<emphasis id="B95DC13FFF9EFFE003EBF8BFFDC12F53" box="[503,536,1830,1854]" italics="true" pageId="9" pageNumber="445">st4</emphasis>
and lyrifissures 
<emphasis id="B95DC13FFF9EFFE000C9F8BFFD212F53" box="[725,760,1830,1854]" italics="true" pageId="9" pageNumber="445">
<date id="FF973BEDFF9EFFE000C9F8BFFD212F53" box="[725,760,1830,1854]" pageId="9" pageNumber="445">iv3</date>
</emphasis>
on metasternal plates. Genital shield smooth; with seta 
<emphasis id="B95DC13FFF9EFFE00768F8BFFA4D2F50" box="[1396,1428,1830,1853]" italics="true" pageId="9" pageNumber="445">st5</emphasis>
; longer than wide, slightly narrowed anterior to 
<emphasis id="B95DC13FFF9EFFE000B0F8D3FD152F0C" box="[684,716,1866,1889]" italics="true" pageId="9" pageNumber="445">st5</emphasis>
; approximately 88–107 long and 71–80 wide at at widest level. Two pairs of pores on unsclerotised cuticle, posterolaterad of 
<emphasis id="B95DC13FFF9EFFE0015FF8F7FCBA2FE8" box="[835,867,1902,1925]" italics="true" pageId="9" pageNumber="445">st5</emphasis>
. Endopodal plates between coxae I and II, coxae II and III, coxae III and IV, and thin parapodal platelets free. Anal shield oval; 92–111 long and 82–99 wide at widest level; anus located slightly anterior to mid-level of shield; with circum-anal setae and a pair of pores; cribrum well developed. Unsclerotised cuticle around of anal shield with two pair of elongate metapodal plates (the larger 19–25 long, 8–11 wide, the smaller 6–11 long, 2–4 wide), 10 pairs of setae (
<emphasis id="B95DC13FFF9DFFE301A8FF01FBC828C2" box="[948,1041,152,176]" italics="true" pageId="10" pageNumber="446">Jv1–Jv5</emphasis>
, 
<emphasis id="B95DC13FFF9DFFE3063CFF01FB5B28C2" box="[1056,1154,152,176]" italics="true" pageId="10" pageNumber="446">Zv1–Zv5</emphasis>
) and five pairs of pores. Peritreme 206–216 long, extending to dorsal setae 
<emphasis id="B95DC13FFF9DFFE300D0FF25FD3F28B9" box="[716,742,188,212]" italics="true" pageId="10" pageNumber="446">s1</emphasis>
; peritrematal shields extending almost to 
<emphasis id="B95DC13FFF9DFFE306A4FF25FB0828B9" box="[1208,1233,188,212]" italics="true" pageId="10" pageNumber="446">z1</emphasis>
; one plate located between peritrematal shields and podonotal shield fused with peritrematal shields near coxae II–III. Spermathecal structures indiscernible. Measurements of setae: 
<emphasis id="B95DC13FFF9DFFE300C2FE9DFCF42971" box="[734,813,260,284]" italics="true" pageId="10" pageNumber="446">
st1–st 
<date id="FF973BEDFF9DFFE30103FE9CFCF42971" box="[799,813,261,284]" pageId="10" pageNumber="446" value="1917-05-23">5</date>
</emphasis>
17–23, para-anal 19–23, post-anal 29–36, setae on unsclerotised cuticle 23–34. Setae 
<emphasis id="B95DC13FFF9DFFE30034FEB1FD93292D" box="[552,586,296,320]" italics="true" pageId="10" pageNumber="446">st2</emphasis>
thick, blunt and serrated, post-anal seta and setae 
<emphasis id="B95DC13FFF9DFFE30686FEB1FB18292D" box="[1178,1217,296,320]" italics="true" pageId="10" pageNumber="446">Jv4</emphasis>
, 
<emphasis id="B95DC13FFF9DFFE306CEFEB1FB232952" box="[1234,1274,296,319]" italics="true" pageId="10" pageNumber="446">Jv5</emphasis>
, 
<emphasis id="B95DC13FFF9DFFE30717FEB1FAEF292D" box="[1291,1334,296,320]" italics="true" pageId="10" pageNumber="446">Zv4</emphasis>
and 
<emphasis id="B95DC13FFF9DFFE3076EFEB1FA452952" box="[1394,1436,296,319]" italics="true" pageId="10" pageNumber="446">Zv5</emphasis>
serrated, other setae smooth.
</paragraph>
<caption id="DF564DA5FF9DFFE3028BF977FEFE2F49" httpUri="https://zenodo.org/record/264419/files/figure.png" pageId="10" pageNumber="446" targetBox="[165,1422,418,1748]" targetPageId="10">
<paragraph id="8B961D2DFF9DFFE3028BF977FEFE2F49" blockId="10.[151,1436,1774,1828]" pageId="10" pageNumber="446">
<emphasis id="B95DC13FFF9DFFE3028BF977FE942F69" bold="true" box="[151,333,1774,1797]" pageId="10" pageNumber="446">FIGURES 25–28</emphasis>
. 
<taxonomicName id="4C2966AEFF9DFFE30347F976FD852F68" box="[347,604,1775,1797]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="446" phylum="Arthropoda" rank="species" species="saproincisus">
<emphasis id="B95DC13FFF9DFFE30347F976FD852F68" box="[347,604,1775,1797]" italics="true" pageId="10" pageNumber="446">Halolaelaps saproincisus</emphasis>
</taxonomicName>
. Adult female. 25. Leg I (coxa to tibia); 26. Leg II; 27. Leg III; 28. Leg IV. Scale bar = 100 µm.
</paragraph>
</caption>
<paragraph id="8B961D2DFF9DFFE202DBF8D2FB32292D" blockId="10.[151,1436,1867,2036]" lastBlockId="11.[151,1437,151,572]" lastPageId="11" lastPageNumber="447" pageId="10" pageNumber="446">
<emphasis id="B95DC13FFF9DFFE302DBF8D2FF262F09" bold="true" box="[199,255,1867,1892]" pageId="10" pageNumber="446">Legs</emphasis>
(
<figureCitation id="131201A8FF9DFFE3030DF8D2FE4E2F09" box="[273,407,1867,1892]" captionStart="FIGURES 25 – 28" captionStartId="10.[151,264,1774,1797]" captionTargetBox="[165,1422,418,1748]" captionTargetId="figure@10.[150,1436,409,1754]" captionTargetPageId="10" captionText="FIGURES 25 – 28. Halolaelaps saproincisus. Adult female. 25. Leg I (coxa to tibia); 26. Leg II; 27. Leg III; 28. Leg IV. Scale bar = 100 µm." httpUri="https://zenodo.org/record/264419/files/figure.png" pageId="10" pageNumber="446">Figs 25–28</figureCitation>
). Lengths: I 344–363, II 279–307, III 251–288, IV 344–372. Chaetotaxy: Leg I: coxa 2, trochanter 6 (1 0/1 1/2 1), femur 13 (2 3/1 2/3 2), genu 12 (2 3/2 2/1 2), tibia 12 (2 3/2 2/1 2); Leg II: coxa 2, trochanter 5 (1 0/1 0/2 1), femur 11 (2 3/1 2/2 1), genu 11 (2 3/1 2/1 2), tibia 10 (2 2/1 2/1 2), tarsus 18 (3 3/3 1/1 2/ 2 3); Leg III: coxa 2, trochanter 5 (1 0/1 0/2 1), femur 6 (1 2/1 1/0 1), genu 9 (2 2/1 2/1 1), tibia 8 (2 1/1 2/1 1), tarsus 18 (3 3/3 1/1 2/2 3); Leg IV: coxa 1, trochanter 5 (1 0/1 0/2 1), femur 6 (1 2/1 1/0 1), genu 9 (2 2/1 3/0 1), tibia 8 (2 1/1 2/1 1), tarsus 18 (3 3/3 1/1 2/2 3). Most setae smooth and pointed, some slightly thickened, some (
<emphasis id="B95DC13FFF9CFFE202BCFF25FF1328B8" box="[160,202,188,213]" italics="true" pageId="11" pageNumber="447">pd1</emphasis>
, 
<emphasis id="B95DC13FFF9CFFE202C4FF25FEDB28B8" box="[216,258,188,213]" italics="true" pageId="11" pageNumber="447">pd2</emphasis>
and 
<emphasis id="B95DC13FFF9CFFE20324FF25FEB828B9" box="[312,353,188,212]" italics="true" pageId="11" pageNumber="447">pv1</emphasis>
on femur I, 
<emphasis id="B95DC13FFF9CFFE203F7FF25FDCC28B8" box="[491,533,188,213]" italics="true" pageId="11" pageNumber="447">ad1</emphasis>
, 
<emphasis id="B95DC13FFF9CFFE2003FFF25FD9428B8" box="[547,589,188,213]" italics="true" pageId="11" pageNumber="447">pd1</emphasis>
, 
<emphasis id="B95DC13FFF9CFFE20047FF25FDA728B8" box="[603,638,188,213]" italics="true" pageId="11" pageNumber="447">pl1</emphasis>
and 
<emphasis id="B95DC13FFF9CFFE200A8FF25FD0028B8" box="[692,729,188,213]" italics="true" pageId="11" pageNumber="447">pl2</emphasis>
on genu I, 
<emphasis id="B95DC13FFF9CFFE2014AFF25FCA328B8" box="[854,890,188,213]" italics="true" pageId="11" pageNumber="447">pl1</emphasis>
on tibia I; 
<emphasis id="B95DC13FFF9CFFE201E8FF25FBC428B8" box="[1012,1053,188,213]" italics="true" pageId="11" pageNumber="447">pd1</emphasis>
on femur II, 
<emphasis id="B95DC13FFF9CFFE206AEFF25FB0228B8" box="[1202,1243,188,213]" italics="true" pageId="11" pageNumber="447">ad1</emphasis>
and 
<emphasis id="B95DC13FFF9CFFE2070EFF25FAE528B8" box="[1298,1340,188,213]" italics="true" pageId="11" pageNumber="447">pd1</emphasis>
on genu 
<date id="FF973BEDFF9CFFE2028BFF79FF612895" box="[151,184,224,248]" pageId="11" pageNumber="447">II4</date>
<emphasis id="B95DC13FFF9CFFE202A2FF46FF302895" box="[190,233,223,248]" italics="true" pageId="11" pageNumber="447">ad1</emphasis>
on genu III, 
<emphasis id="B95DC13FFF9CFFE20366FF46FE7D2895" box="[378,420,223,248]" italics="true" pageId="11" pageNumber="447">pd2</emphasis>
on tibia II; 
<emphasis id="B95DC13FFF9CFFE20034FF46FD8B2895" box="[552,594,223,248]" italics="true" pageId="11" pageNumber="447">ad2</emphasis>
, 
<emphasis id="B95DC13FFF9CFFE2007CFF46FD532895" box="[608,650,223,248]" italics="true" pageId="11" pageNumber="447">pd1</emphasis>
, 
<emphasis id="B95DC13FFF9CFFE20084FF46FD182895" box="[664,705,223,248]" italics="true" pageId="11" pageNumber="447">pd3</emphasis>
on genu IV, 
<emphasis id="B95DC13FFF9CFFE2014DFF46FCA52895" box="[849,892,223,248]" italics="true" pageId="11" pageNumber="447">pd1</emphasis>
and 
<emphasis id="B95DC13FFF9CFFE201ADFF46FC022895" box="[945,987,223,248]" italics="true" pageId="11" pageNumber="447">pd2</emphasis>
on tibia IV) often more or less serrated. Coxa II with anterior spine. Pretarsi I-IV similar in shape; consisting of an ambulacral stalk, a pair of sclerotised claws, four rounded pulvillar lobes and two acuminate ventro-lateral processes (obscure on tarsi I).
</paragraph>
<paragraph id="8B961D2DFF9CFFE202DBFED5FCCE2908" blockId="11.[151,1437,151,572]" box="[199,791,332,357]" pageId="11" pageNumber="447">
<emphasis id="B95DC13FFF9CFFE202DBFED5FE952908" bold="true" box="[199,332,332,357]" pageId="11" pageNumber="447">Adult male</emphasis>
(
<figureCitation id="131201A8FF9CFFE20346FED5FE072909" box="[346,478,332,357]" captionStart="FIGURE 29 – 33" captionStartId="11.[151,250,1934,1956]" captionTargetBox="[157,1426,637,1902]" captionTargetId="figure@11.[151,1436,625,1913]" captionTargetPageId="11" captionText="FIGURE 29 – 33. Halolaelaps saproincisus. Adult male. 29. Lateral (antiaxial) view of chelicerae; 30 Epistome; 31. Dorsal idiosoma; 32. Ventral idiosoma; 33. Femur of leg II. Scale bar = 50 µm." httpUri="https://zenodo.org/record/264420/files/figure.png" pageId="11" pageNumber="447">Figs 29–33</figureCitation>
). Five specimens measured.
</paragraph>
<paragraph id="8B961D2DFF9CFFE202DBFEF6FF632A51" blockId="11.[151,1437,151,572]" pageId="11" pageNumber="447">
<emphasis id="B95DC13FFF9CFFE202DBFEF6FE8629E5" bold="true" box="[199,351,367,392]" pageId="11" pageNumber="447">Gnathosoma</emphasis>
. Second cheliceral segment 86–95 long, fixed cheliceral digit with one tooth in addition to apical tooth and setiform pilus dentilis; movable cheliceral digit with one tooth in addition to apical tooth; dorsal cheliceral seta obscure (
<figureCitation id="131201A8FF9CFFE203A8FE2EFDD729BD" box="[436,526,439,464]" captionStart="FIGURE 29 – 33" captionStartId="11.[151,250,1934,1956]" captionTargetBox="[157,1426,637,1902]" captionTargetId="figure@11.[151,1436,625,1913]" captionTargetPageId="11" captionText="FIGURE 29 – 33. Halolaelaps saproincisus. Adult male. 29. Lateral (antiaxial) view of chelicerae; 30 Epistome; 31. Dorsal idiosoma; 32. Ventral idiosoma; 33. Femur of leg II. Scale bar = 50 µm." httpUri="https://zenodo.org/record/264420/files/figure.png" pageId="11" pageNumber="447">Fig. 29</figureCitation>
). Spermatodactyl 32–36 long, sinuous, longer than movable digit. Arthrodial process of chelicera, palp chaetotaxy, apotele, epistome, deutosternum and shape of hypostomal setae as in adult female. Subcapitulum 76–80 wide at widest level. Measurements of setae: 
<emphasis id="B95DC13FFF9CFFE201F9FE66FBD82A75" box="[997,1025,511,536]" italics="true" pageId="11" pageNumber="447">
h 
<date id="FF973BEDFF9CFFE201E9FE66FBD82A75" box="[1013,1025,511,536]" pageId="11" pageNumber="447" value="1917-01-21">1</date>
</emphasis>
17–21, 
<emphasis id="B95DC13FFF9CFFE20642FE66FBA32A75" box="[1118,1146,511,536]" italics="true" pageId="11" pageNumber="447">
h 
<date id="FF973BEDFF9CFFE20670FE66FBA32A75" box="[1132,1146,511,536]" pageId="11" pageNumber="447" value="2013-02-15">2</date>
</emphasis>
13–15, 
<emphasis id="B95DC13FFF9CFFE206CBFE66FB2A2A75" box="[1239,1267,511,536]" italics="true" pageId="11" pageNumber="447">
h 
<date id="FF973BEDFF9CFFE206F9FE66FB2A2A75" box="[1253,1267,511,536]" pageId="11" pageNumber="447" value="1919-03-23">3</date>
</emphasis>
19–23, 
<emphasis id="B95DC13FFF9CFFE2074CFD98FAB32A75" box="[1360,1386,513,536]" italics="true" pageId="11" pageNumber="447">pc</emphasis>
17– 19.
</paragraph>
<caption id="DF564DA5FF9CFFE2028BF817FCB42FAE" httpUri="https://zenodo.org/record/264420/files/figure.png" pageId="11" pageNumber="447" targetBox="[157,1426,637,1902]" targetPageId="11">
<paragraph id="8B961D2DFF9CFFE2028BF817FCB42FAE" blockId="11.[151,1436,1934,1987]" pageId="11" pageNumber="447">
<emphasis id="B95DC13FFF9CFFE2028BF817FE912FC9" bold="true" box="[151,328,1934,1956]" pageId="11" pageNumber="447">FIGURE 29–33.</emphasis>
<taxonomicName id="4C2966AEFF9CFFE2034CF817FD8D2FC8" box="[336,596,1934,1957]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="447" phylum="Arthropoda" rank="species" species="saproincisus">
<emphasis id="B95DC13FFF9CFFE2034CF817FD8D2FC8" box="[336,596,1934,1957]" italics="true" pageId="11" pageNumber="447">Halolaelaps saproincisus</emphasis>
</taxonomicName>
. Adult male. 29. Lateral (antiaxial) view of chelicerae; 30 Epistome; 31. Dorsal idiosoma; 32. Ventral idiosoma; 33. Femur of leg II. Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="DF564DA5FF9BFFE5028BFA97FABA2D49" box="[151,1379,1294,1316]" httpUri="https://zenodo.org/record/264421/files/figure.png" pageId="12" pageNumber="448" targetBox="[159,1431,199,1268]" targetPageId="12">
<paragraph id="8B961D2DFF9BFFE5028BFA97FABA2D49" blockId="12.[151,1379,1294,1316]" box="[151,1379,1294,1316]" pageId="12" pageNumber="448">
<emphasis id="B95DC13FFF9BFFE5028BFA97FE9C2D4E" bold="true" box="[151,325,1294,1316]" pageId="12" pageNumber="448">FIGURE 34–35.</emphasis>
<taxonomicName id="4C2966AEFF9BFFE50350FA97FD952D49" box="[332,588,1294,1316]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="448" phylum="Arthropoda" rank="species" species="saproincisus">
<emphasis id="B95DC13FFF9BFFE50350FA97FD952D49" box="[332,588,1294,1316]" italics="true" pageId="12" pageNumber="448">Halolaelaps saproincisus</emphasis>
</taxonomicName>
. Deutonymph. 34. Dorsal idiosoma; 35. Ventral idiosoma. Scale bar = 50 µm.
</paragraph>
</caption>
<paragraph id="8B961D2DFF9BFFE502DBFAC8FCB72E73" blockId="12.[151,1436,1361,2034]" pageId="12" pageNumber="448">
<emphasis id="B95DC13FFF9BFFE502DBFAC8FE502D07" bold="true" box="[199,393,1361,1386]" pageId="12" pageNumber="448">Dorsal idiosoma</emphasis>
(
<figureCitation id="131201A8FF9BFFE50386FAC8FE372D04" box="[410,494,1361,1386]" captionStart="FIGURE 29 – 33" captionStartId="11.[151,250,1934,1956]" captionTargetBox="[157,1426,637,1902]" captionTargetId="figure@11.[151,1436,625,1913]" captionTargetPageId="11" captionText="FIGURE 29 – 33. Halolaelaps saproincisus. Adult male. 29. Lateral (antiaxial) view of chelicerae; 30 Epistome; 31. Dorsal idiosoma; 32. Ventral idiosoma; 33. Femur of leg II. Scale bar = 50 µm." httpUri="https://zenodo.org/record/264420/files/figure.png" pageId="12" pageNumber="448">Fig. 31</figureCitation>
). Idiosoma ovoid. Podonotal shield 189–210 long and 218–260 wide at level of 
<emphasis id="B95DC13FFF9BFFE50767FACBFA4D2D04" box="[1403,1428,1362,1385]" italics="true" pageId="12" pageNumber="448">r5</emphasis>
; reticulated; with 24 pairs of setae (
<emphasis id="B95DC13FFF9BFFE50004FAEFFD8B2DE3" box="[536,594,1397,1422]" italics="true" pageId="12" pageNumber="448">j1–j6</emphasis>
, 
<emphasis id="B95DC13FFF9BFFE50043FAEFFD462DE3" box="[607,671,1397,1422]" italics="true" pageId="12" pageNumber="448">z1–z6</emphasis>
, 
<emphasis id="B95DC13FFF9BFFE500B0FAEFFD342DE3" box="[684,749,1397,1422]" italics="true" pageId="12" pageNumber="448">s1–s6</emphasis>
, 
<emphasis id="B95DC13FFF9BFFE500E6FAEFFCE02DE3" box="[762,825,1397,1422]" italics="true" pageId="12" pageNumber="448">r1–r6</emphasis>
) and three pairs of distinguishable pores. Opisthonotal shield 160–181 long and 218–260 wide at level of 
<emphasis id="B95DC13FFF9BFFE500CDFA00FD292DDC" box="[721,752,1433,1457]" italics="true" pageId="12" pageNumber="448">R1</emphasis>
; reticulated; anterior margin without notch; posterior margin with cribrum; with 19 pairs of setae (
<emphasis id="B95DC13FFF9BFFE50027FA27FD8C2DBB" box="[571,597,1470,1494]" italics="true" pageId="12" pageNumber="448">J1</emphasis>
, 
<emphasis id="B95DC13FFF9BFFE50078FA27FDA72DBB" box="[612,638,1470,1494]" italics="true" pageId="12" pageNumber="448">J2</emphasis>
, 
<emphasis id="B95DC13FFF9BFFE50091FA27FD7E2DBB" box="[653,679,1470,1494]" italics="true" pageId="12" pageNumber="448">J4</emphasis>
, 
<emphasis id="B95DC13FFF9BFFE500AAFA27FD092DB8" box="[694,720,1470,1493]" italics="true" pageId="12" pageNumber="448">J5</emphasis>
, 
<emphasis id="B95DC13FFF9BFFE500C3FA27FCF12DB8" box="[735,808,1470,1494]" italics="true" pageId="12" pageNumber="448">Z1–Z5</emphasis>
, 
<emphasis id="B95DC13FFF9BFFE5012BFA27FCA42DB8" box="[823,893,1470,1494]" italics="true" pageId="12" pageNumber="448">S1–S5</emphasis>
, 
<emphasis id="B95DC13FFF9BFFE50197FA27FC012DB8" box="[907,984,1470,1494]" italics="true" pageId="12" pageNumber="448">R1–R5</emphasis>
) and two pairs of distinguishable pores. Measurements of setae: 
<emphasis id="B95DC13FFF9BFFE503BBFA78FE642D94" box="[423,445,1505,1529]" italics="true" pageId="12" pageNumber="448">
j 
<date id="FF973BEDFF9BFFE503AEFA78FE642D94" box="[434,445,1505,1529]" pageId="12" pageNumber="448" value="1917-01-19">1</date>
</emphasis>
17–19, 
<emphasis id="B95DC13FFF9BFFE50004FA78FDE82D94" box="[536,561,1505,1529]" italics="true" pageId="12" pageNumber="448">
z 
<date id="FF973BEDFF9BFFE50039FA78FDE82D94" box="[549,561,1505,1529]" pageId="12" pageNumber="448" value="2013-11-01">1</date>
</emphasis>
11–13, other setae on podonotal shield 21–34, 
<emphasis id="B95DC13FFF9BFFE50658FA7BFB862D94" box="[1092,1119,1506,1529]" italics="true" pageId="12" pageNumber="448">
J 
<date id="FF973BEDFF9BFFE5064EFA7BFB862D94" box="[1106,1119,1506,1529]" pageId="12" pageNumber="448" value="2011-08-05">5</date>
</emphasis>
8–11, 
<emphasis id="B95DC13FFF9BFFE506B7FA7BFB102D94" box="[1195,1225,1506,1529]" italics="true" pageId="12" pageNumber="448">
Z 
<date id="FF973BEDFF9BFFE506A0FA7BFB102D94" box="[1212,1225,1506,1529]" pageId="12" pageNumber="448" value="2013-11-05">5</date>
</emphasis>
11–13, other setae on opisthonotal shield 17–32. All dorsal idiosomal setae smooth.
</paragraph>
<paragraph id="8B961D2DFF9BFFE502DBF9B0FEE92FEB" blockId="12.[151,1436,1361,2034]" pageId="12" pageNumber="448">
<emphasis id="B95DC13FFF9BFFE502DBF9B0FE4D2E2F" bold="true" box="[199,404,1577,1602]" pageId="12" pageNumber="448">Ventral idiosoma</emphasis>
(
<figureCitation id="131201A8FF9BFFE503B9F9B0FE222E2C" box="[421,507,1577,1602]" captionStart="FIGURE 29 – 33" captionStartId="11.[151,250,1934,1956]" captionTargetBox="[157,1426,637,1902]" captionTargetId="figure@11.[151,1436,625,1913]" captionTargetPageId="11" captionText="FIGURE 29 – 33. Halolaelaps saproincisus. Adult male. 29. Lateral (antiaxial) view of chelicerae; 30 Epistome; 31. Dorsal idiosoma; 32. Ventral idiosoma; 33. Femur of leg II. Scale bar = 50 µm." httpUri="https://zenodo.org/record/264420/files/figure.png" pageId="12" pageNumber="448">Fig. 32</figureCitation>
). Tritosternum with wide trapezoidal base and fine lightly pilose laciniae, base of tritosternum length 11–13, laciniae length 25–38. Presternal area lightly sclerotised. Sternogenital shield smooth; approximately 179–189 long (without presternal area); with elongate antero-lateral corners; with five pairs of setae and three pairs of lyrifissures. Endopodal plates fused with sternal shield. Anal shield oval; 59–67 long and 67–76 wide at widest level; anus located slightly anterior to mid-level of shield; with circum-anal setae and one pair of distinguishable pores; cribrum well developed. Unsclerotised cuticle around of anal shield with lineate ornamentation; with a pair of small metapodal plates (11–15 long, 4–8 wide) and ten pairs of setae (
<emphasis id="B95DC13FFF9BFFE506E6F898FA8F2F74" box="[1274,1366,1793,1817]" italics="true" pageId="12" pageNumber="448">Jv1–Jv5</emphasis>
, 
<emphasis id="B95DC13FFF9BFFE50778F898FF192F50" italics="true" pageId="12" pageNumber="448">Zv1– Zv5</emphasis>
). Peritrematal shields, peritremes and adjacent plates as in female, peritreme 193–206 long. Measurements of setae: 
<emphasis id="B95DC13FFF9BFFE502C2F8D0FEF42F0C" box="[222,301,1865,1889]" italics="true" pageId="12" pageNumber="448">
st1–st 
<date id="FF973BEDFF9BFFE50303F8D3FEF42F0C" box="[287,301,1866,1889]" pageId="12" pageNumber="448" value="1917-05-23">5</date>
</emphasis>
17–23, para-anal 15–19, post-anal 17–19, setae on unsclerotised cuticle 15–25. All ventral idiosoma setae smooth.
</paragraph>
<paragraph id="8B961D2DFF9BFFE502DBF808FEF32F9F" blockId="12.[151,1436,1361,2034]" pageId="12" pageNumber="448">
<emphasis id="B95DC13FFF9BFFE502DBF808FF262FC7" bold="true" box="[199,255,1937,1962]" pageId="12" pageNumber="448">Legs</emphasis>
. Legths: I 344–363, II 270–288, 
<date id="FF973BEDFF9BFFE50067F80BFD102FC7" box="[635,713,1937,1962]" pageId="12" pageNumber="448" value="1960-03-02">III 2 60</date>
–270, IV 344–391. Leg chaetotaxy as in adult female. Shape of setae similar to those of adult females, seta 
<emphasis id="B95DC13FFF9BFFE5009AF82FFD772FA3" box="[646,686,1974,1998]" italics="true" pageId="12" pageNumber="448">pv1</emphasis>
on femur II strongly spinose (
<figureCitation id="131201A8FF9BFFE5061BF82CFB822FA3" box="[1031,1115,1973,1998]" captionStart="FIGURE 29 – 33" captionStartId="11.[151,250,1934,1956]" captionTargetBox="[157,1426,637,1902]" captionTargetId="figure@11.[151,1436,625,1913]" captionTargetPageId="11" captionText="FIGURE 29 – 33. Halolaelaps saproincisus. Adult male. 29. Lateral (antiaxial) view of chelicerae; 30 Epistome; 31. Dorsal idiosoma; 32. Ventral idiosoma; 33. Femur of leg II. Scale bar = 50 µm." httpUri="https://zenodo.org/record/264420/files/figure.png" pageId="12" pageNumber="448">Fig. 33</figureCitation>
). Pretarsi similar to those of adult female.
</paragraph>
<paragraph id="8B961D2DFF9AFFE402DBFF0EFCF628DD" blockId="13.[151,1437,151,2035]" box="[199,815,151,176]" pageId="13" pageNumber="449">
<emphasis id="B95DC13FFF9AFFE402DBFF0EFEBA28DD" bold="true" box="[199,355,151,176]" pageId="13" pageNumber="449">Deutonymph</emphasis>
(
<figureCitation id="131201A8FF9AFFE4036DFF0EFE2C28DD" box="[369,501,151,176]" captionStart="FIGURE 34 – 35" captionStartId="12.[151,250,1294,1316]" captionTargetBox="[159,1431,199,1268]" captionTargetId="figure@12.[150,1436,193,1273]" captionTargetPageId="12" captionText="FIGURE 34 – 35. Halolaelaps saproincisus. Deutonymph. 34. Dorsal idiosoma; 35. Ventral idiosoma. Scale bar = 50 µm." httpUri="https://zenodo.org/record/264421/files/figure.png" pageId="13" pageNumber="449">Figs 34–35</figureCitation>
). Five specimens measured.
</paragraph>
<paragraph id="8B961D2DFF9AFFE402DBFF22FD342952" blockId="13.[151,1437,151,2035]" pageId="13" pageNumber="449">
<emphasis id="B95DC13FFF9AFFE402DBFF22FE8628B9" bold="true" box="[199,351,187,212]" pageId="13" pageNumber="449">Gnathosoma</emphasis>
. Mostly as in adult female. Second cheliceral segment 78–84 long. Palp 105–116 long. Subcapitulum 80–90 wide at widest level. Deutosternum with seven (rarely six) rows of deutosternal denticles (0–5 denticles per row). Measurements of setae: 
<emphasis id="B95DC13FFF9AFFE40063FE9AFD432971" box="[639,666,259,284]" italics="true" pageId="13" pageNumber="449">
h 
<date id="FF973BEDFF9AFFE40092FE9AFD432971" box="[654,666,259,284]" pageId="13" pageNumber="449" value="2013-11-01">1</date>
</emphasis>
11–13, 
<emphasis id="B95DC13FFF9AFFE400E9FE9AFCC82971" box="[757,785,259,284]" italics="true" pageId="13" pageNumber="449">
h 
<date id="FF973BEDFF9AFFE4011FFE9AFCC82971" box="[771,785,259,284]" pageId="13" pageNumber="449" value="1917-02-21">2</date>
</emphasis>
17–21, 
<emphasis id="B95DC13FFF9AFFE40170FE9AFC512971" box="[876,904,259,284]" italics="true" pageId="13" pageNumber="449">
h 
<date id="FF973BEDFF9AFFE40166FE9AFC512971" box="[890,904,259,284]" pageId="13" pageNumber="449" value="2015-03-17">3</date>
</emphasis>
15–17, 
<emphasis id="B95DC13FFF9AFFE401FFFE9DFC272976" box="[995,1022,260,283]" italics="true" pageId="13" pageNumber="449">pc</emphasis>
17–21. Setae 
<emphasis id="B95DC13FFF9AFFE40680FE9AFB3B2971" box="[1180,1250,259,284]" italics="true" pageId="13" pageNumber="449">h1–h3</emphasis>
simple, setae 
<emphasis id="B95DC13FFF9AFFE4079DFE9DFA422976" box="[1409,1435,260,283]" italics="true" pageId="13" pageNumber="449">pc</emphasis>
variable, normal (as 
<emphasis id="B95DC13FFF9AFFE40361FEBFFE1A2952" box="[381,451,294,319]" italics="true" pageId="13" pageNumber="449">h1–h3</emphasis>
) or thickened (as 
<emphasis id="B95DC13FFF9AFFE40091FEBEFD052953" box="[653,732,295,319]" italics="true" pageId="13" pageNumber="449">st1–st5</emphasis>
).
</paragraph>
<paragraph id="8B961D2DFF9AFFE402DBFED3FF282AEC" blockId="13.[151,1437,151,2035]" pageId="13" pageNumber="449">
<emphasis id="B95DC13FFF9AFFE402DBFED3FE55290E" bold="true" box="[199,396,330,355]" pageId="13" pageNumber="449">Dorsal idiosoma</emphasis>
(
<figureCitation id="131201A8FF9AFFE40383FED3FE2E290E" box="[415,503,330,355]" captionStart="FIGURE 34 – 35" captionStartId="12.[151,250,1294,1316]" captionTargetBox="[159,1431,199,1268]" captionTargetId="figure@12.[150,1436,193,1273]" captionTargetPageId="12" captionText="FIGURE 34 – 35. Halolaelaps saproincisus. Deutonymph. 34. Dorsal idiosoma; 35. Ventral idiosoma. Scale bar = 50 µm." httpUri="https://zenodo.org/record/264421/files/figure.png" pageId="13" pageNumber="449">Fig. 34</figureCitation>
). Idiosoma ovate. Podonotal shield 185–210 long and 185–218 wide at level of 
<emphasis id="B95DC13FFF9AFFE4028BFEF7FF0F29EA" box="[151,214,366,391]" italics="true" pageId="13" pageNumber="449">r3–r6</emphasis>
, tapering anteriorly to 
<emphasis id="B95DC13FFF9AFFE403FAFEF6FE2629EA" box="[486,511,367,391]" italics="true" pageId="13" pageNumber="449">r3</emphasis>
; reticulated stronger than in adults; with 24 pairs of setae (
<emphasis id="B95DC13FFF9AFFE406A5FEF6FB2B29EA" box="[1209,1266,366,391]" italics="true" pageId="13" pageNumber="449">j1–j6</emphasis>
, 
<emphasis id="B95DC13FFF9AFFE40718FEF6FA9A29EA" box="[1284,1347,366,391]" italics="true" pageId="13" pageNumber="449">z1–z6</emphasis>
, 
<emphasis id="B95DC13FFF9AFFE40749FEF6FA4C29EA" box="[1365,1429,366,391]" italics="true" pageId="13" pageNumber="449">s1–s6</emphasis>
, 
<emphasis id="B95DC13FFF9AFFE4028BFE0BFF0F29C6" box="[151,214,402,427]" italics="true" pageId="13" pageNumber="449">r1–r6</emphasis>
) (in some specimens 
<emphasis id="B95DC13FFF9AFFE403D7FE0BFE3D29C6" box="[459,484,402,427]" italics="true" pageId="13" pageNumber="449">r6</emphasis>
on unsclerotised cuticle) and four pairs of distinguishable pores. Opisthonotal shield 147–173 long and 176–202 wide at level of setae 
<emphasis id="B95DC13FFF9AFFE400C8FE2FFD2929A3" box="[724,752,438,462]" italics="true" pageId="13" pageNumber="449">S1</emphasis>
; reticulated stronger than in adults; anterior margin without notch; with 14 pairs of setae (
<emphasis id="B95DC13FFF9AFFE403F5FE43FDDA299F" box="[489,515,474,498]" italics="true" pageId="13" pageNumber="449">J1</emphasis>
, 
<emphasis id="B95DC13FFF9AFFE4000EFE43FDF5299F" box="[530,556,474,498]" italics="true" pageId="13" pageNumber="449">J2</emphasis>
, 
<emphasis id="B95DC13FFF9AFFE40020FE43FD8F299F" box="[572,598,474,498]" italics="true" pageId="13" pageNumber="449">J4</emphasis>
, 
<emphasis id="B95DC13FFF9AFFE40079FE43FDA6299C" box="[613,639,474,497]" italics="true" pageId="13" pageNumber="449">J5</emphasis>
, 
<emphasis id="B95DC13FFF9AFFE40092FE43FD0E299C" box="[654,727,474,498]" italics="true" pageId="13" pageNumber="449">Z1–Z5</emphasis>
, 
<emphasis id="B95DC13FFF9AFFE400FAFE43FCF5299C" box="[742,812,474,498]" italics="true" pageId="13" pageNumber="449">S1–S5</emphasis>
; 
<emphasis id="B95DC13FFF9AFFE40120FE43FC8E299F" box="[828,855,474,498]" italics="true" pageId="13" pageNumber="449">J3</emphasis>
absent) and with six pairs of distinguishable pores. Unsclerotised cuticle laterad of opisthonotal shield with five pairs of setae (
<emphasis id="B95DC13FFF9AFFE40605FE67FBBC2A78" box="[1049,1125,510,534]" italics="true" pageId="13" pageNumber="449">R1–R5</emphasis>
; 
<emphasis id="B95DC13FFF9AFFE40664FE67FB412A7B" box="[1144,1176,510,534]" italics="true" pageId="13" pageNumber="449">R4</emphasis>
and 
<emphasis id="B95DC13FFF9AFFE406CBFE67FB2E2A78" box="[1239,1271,510,533]" italics="true" pageId="13" pageNumber="449">R5</emphasis>
visible in the ventrolateral regions). Measurements of setae: 
<emphasis id="B95DC13FFF9AFFE400BBFDB8FD642A54" box="[679,701,545,569]" italics="true" pageId="13" pageNumber="449">
j 
<date id="FF973BEDFF9AFFE400AFFDB8FD642A54" box="[691,701,545,569]" pageId="13" pageNumber="449" value="2013-11-01">1</date>
</emphasis>
11–13, 
<emphasis id="B95DC13FFF9AFFE40104FDB8FCE82A54" box="[792,817,545,569]" italics="true" pageId="13" pageNumber="449">
z 
<date id="FF973BEDFF9AFFE40139FDB8FCE82A54" box="[805,817,545,569]" pageId="13" pageNumber="449" value="2013-08-01">1</date>
</emphasis>
8–13, other setae on podonotal shield 17–23, 
<emphasis id="B95DC13FFF9AFFE40727FDBBFA8C2A54" box="[1339,1365,546,569]" italics="true" pageId="13" pageNumber="449">
J 
<date id="FF973BEDFF9AFFE40754FDBBFA8C2A54" box="[1352,1365,546,569]" pageId="13" pageNumber="449" value="2013-08-05">5</date>
</emphasis>
8–13, 
<emphasis id="B95DC13FFF9AFFE4028BFDDCFF6C2A31" box="[151,181,581,604]" italics="true" pageId="13" pageNumber="449">
Z 
<date id="FF973BEDFF9AFFE402B4FDDCFF6C2A31" box="[168,181,581,604]" pageId="13" pageNumber="449" value="2013-08-05">5</date>
</emphasis>
8–13, other setae on opisthonotal shield 13–19, setae on unsclerotised cuticle 13–15. All dorsal idiosomal setae smooth.
</paragraph>
<paragraph id="8B961D2DFF9AFFE402DBFD15FF302B8B" blockId="13.[151,1437,151,2035]" pageId="13" pageNumber="449">
<emphasis id="B95DC13FFF9AFFE402DBFD15FE452AC8" bold="true" box="[199,412,652,677]" pageId="13" pageNumber="449">Ventral idiosoma</emphasis>
(
<figureCitation id="131201A8FF9AFFE403AFFD15FDC92AC8" box="[435,528,652,677]" captionStart="FIGURE 34 – 35" captionStartId="12.[151,250,1294,1316]" captionTargetBox="[159,1431,199,1268]" captionTargetId="figure@12.[150,1436,193,1273]" captionTargetPageId="12" captionText="FIGURE 34 – 35. Halolaelaps saproincisus. Deutonymph. 34. Dorsal idiosoma; 35. Ventral idiosoma. Scale bar = 50 µm." httpUri="https://zenodo.org/record/264421/files/figure.png" pageId="13" pageNumber="449">Fig. 35</figureCitation>
). Tritosternum with elongate trapezoidal base and pilose laciniae, base of tritosternum length 21–23, laciniae length 29–34. Presternal area fused with sternal shield. Sternal shield reticulated; approximately 168–183 long (with presternal area); with five pairs of setae and three pairs of lyrifissures. Endopodal plates between coxae I and II, coxae II and III, and coxae III and IV free. Anal shield reticulated; obovate; 55–65 long and 63–78 wide at widest level; with circum-anal setae and a pair of pores; cribrum well developed. Unsclerotised cuticle around of anal shield with ten pairs of setae (
<emphasis id="B95DC13FFF9AFFE406B6FCA6FADF2B3B" box="[1194,1286,831,855]" italics="true" pageId="13" pageNumber="449">Jv1–Jv5</emphasis>
, 
<emphasis id="B95DC13FFF9AFFE40709FCA6FAAF2B3B" box="[1301,1398,831,855]" italics="true" pageId="13" pageNumber="449">Zv1–Zv5</emphasis>
), a pair of small metapodal plates (17–25 long, 4–6 wide) and four pairs of pores. Peritreme 200–221 long, extending anteriorly to level of coxa I (region of 
<emphasis id="B95DC13FFF9AFFE40043FC1EFDA12BF2" box="[607,632,903,927]" italics="true" pageId="13" pageNumber="449">s1</emphasis>
). Peritrematal shield reduced. Measurements of setae: 
<emphasis id="B95DC13FFF9AFFE406E9FC1EFA9D2BF3" box="[1269,1348,903,927]" italics="true" pageId="13" pageNumber="449">
st1–st 
<date id="FF973BEDFF9AFFE4072AFC1EFA9D2BF3" box="[1334,1348,903,926]" pageId="13" pageNumber="449" value="1919-05-23">5</date>
</emphasis>
19–23, para-anal 15–17, post-anal 15–17, other setae on ventrianal shield 15–23. Setae 
<emphasis id="B95DC13FFF9AFFE40659FC33FB4D2BAF" box="[1093,1172,938,962]" italics="true" pageId="13" pageNumber="449">st1–st5</emphasis>
thickened, other setae simple.
</paragraph>
<paragraph id="8B961D2DFF9AFFE402DBFC68FEE62C18" blockId="13.[151,1437,151,2035]" pageId="13" pageNumber="449">
<emphasis id="B95DC13FFF9AFFE402DBFC68FF262C67" bold="true" box="[199,255,1009,1034]" pageId="13" pageNumber="449">Legs</emphasis>
. Lengths: I 279–307, II 242–260, 
<date id="FF973BEDFF9AFFE4008CFC6BFD062C67" box="[656,735,1009,1034]" pageId="13" pageNumber="449" value="1922-03-03">III 22 3</date>
–251, IV 298–316. Mostly as in adult female, but acuminate ventro-lateral processes on pretarsi obscure. Setae 
<emphasis id="B95DC13FFF9AFFE400FFFB8FFD272C40" box="[739,766,1046,1069]" italics="true" pageId="13" pageNumber="449">pv</emphasis>
of coxae II and III modified, club-like and asymmetrical (some with acuminate tip) (
<figureCitation id="131201A8FF9AFFE403D5FBA0FDC52C3F" box="[457,540,1081,1106]" captionStart="FIGURE 34 – 35" captionStartId="12.[151,250,1294,1316]" captionTargetBox="[159,1431,199,1268]" captionTargetId="figure@12.[150,1436,193,1273]" captionTargetPageId="12" captionText="FIGURE 34 – 35. Halolaelaps saproincisus. Deutonymph. 34. Dorsal idiosoma; 35. Ventral idiosoma. Scale bar = 50 µm." httpUri="https://zenodo.org/record/264421/files/figure.png" pageId="13" pageNumber="449">Fig. 35</figureCitation>
), other setae simple and pointed, setae of 
<emphasis id="B95DC13FFF9AFFE401F7FBA0FBDE2C3F" box="[1003,1031,1081,1106]" italics="true" pageId="13" pageNumber="449">ad</emphasis>
-, 
<emphasis id="B95DC13FFF9AFFE40602FBA0FBE32C3F" box="[1054,1082,1081,1106]" italics="true" pageId="13" pageNumber="449">pd</emphasis>
- and 
<emphasis id="B95DC13FFF9AFFE40665FBA0FB562C3F" box="[1145,1167,1081,1106]" italics="true" pageId="13" pageNumber="449">pl</emphasis>
-series relatively thicker than in female.
</paragraph>
</subSubSection>
<subSubSection id="C3334EA6FF9AFFE402DBFB19FED72DB6" pageId="13" pageNumber="449" type="materials_examined">
<paragraph id="8B961D2DFF9AFFE402DBFB19FED72DB6" blockId="13.[151,1437,151,2035]" pageId="13" pageNumber="449">
<materialsCitation id="3B411770FF9AFFE402DBFB19FAA42CD0" ID-GBIF-Occurrence="1291862184" collectingDate="2004-05-12" country="Ukraine" county="Material" latitude="49.9" location="Ivano-Frankove" longLatPrecision="1100" longitude="23.75" municipality="Yavorivsky district" pageId="13" pageNumber="449" specimenCount="1" stateProvince="Lviv">
<emphasis id="B95DC13FFF9AFFE402DBFB19FE6C2CF4" bold="true" box="[199,437,1152,1177]" pageId="13" pageNumber="449">
<collectingCounty id="62F765A1FF9AFFE402DBFB19FEF62CF4" box="[199,303,1152,1177]" pageId="13" pageNumber="449">Material</collectingCounty>
examined.
</emphasis>
1 deutonymph, 
<collectingCountry id="F33E5DBDFF9AFFE4006BFB19FD0D2CF4" box="[631,724,1152,1177]" name="Ukraine" pageId="13" pageNumber="449">Ukraine</collectingCountry>
, 
<collectingRegion id="49EDD3CFFF9AFFE400F8FB19FCC02CF4" box="[740,793,1152,1177]" country="Ukraine" name="L'viv" pageId="13" pageNumber="449">Lviv</collectingRegion>
oblast, 
<collectingMunicipality id="6BF28757FF9AFFE40166FB19FB8F2CF4" box="[890,1110,1152,1177]" pageId="13" pageNumber="449">Yavorivsky district</collectingMunicipality>
, vicinity of 
<location id="8EF64BF6FF9AFFE406F5FB19FA452CF4" LSID="urn:lsid:plazi:treatment:0380AC3BFF9FFFE7028BFC12FB4929BD:8EF64BF6FF9AFFE406F5FB19FA452CF4" box="[1257,1436,1152,1177]" country="Ukraine" county="Material" latitude="49.9" longLatPrecision="1100" longitude="23.75" municipality="Yavorivsky district" name="Ivano-Frankove" pageId="13" pageNumber="449" stateProvince="Lviv">Ivano-Frankove</location>
(
<geoCoordinate id="EE1D7BEAFF9AFFE40283FB3DFED22CD1" box="[159,267,1188,1213]" direction="north" orientation="latitude" pageId="13" pageNumber="449" precision="925" value="49.9">49°54' N</geoCoordinate>
, 
<geoCoordinate id="EE1D7BEAFF9AFFE40300FB3DFE5D2CD0" box="[284,388,1188,1213]" direction="east" orientation="longitude" pageId="13" pageNumber="449" precision="925" value="23.75">23°45' E</geoCoordinate>
), 
<collectedFrom id="38355ED0FF9AFFE40382FB3CFCA72CD0" box="[414,894,1188,1213]" pageId="13" pageNumber="449">
on 
<taxonomicName id="4C2966AEFF9AFFE403DAFB3DFCA72CD0" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[454,894,1188,1213]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="449" phylum="Arthropoda" rank="species" species="fimetarius">
<emphasis id="B95DC13FFF9AFFE403DAFB3DFD772CD0" box="[454,686,1188,1213]" italics="true" pageId="13" pageNumber="449">Aphodius fimetarius</emphasis>
(Linnaeus, 1758)
</taxonomicName>
</collectedFrom>
(
<taxonomicName id="4C2966AEFF9AFFE4018EFB3DFBF52CD0" box="[914,1068,1188,1213]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="449" phylum="Arthropoda" rank="family">Scarabaeidae</taxonomicName>
: 
<taxonomicName id="4C2966AEFF9AFFE40627FB3DFB1B2CD0" box="[1083,1218,1188,1213]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="449" phylum="Arthropoda" rank="subFamily" subFamily="Aphodiinae">Aphodiinae</taxonomicName>
), 
<date id="FF973BEDFF9AFFE406C6FB3CFAA42CD0" box="[1242,1405,1189,1213]" pageId="13" pageNumber="449" value="2004-05-12">
<collectingDate id="EFD3C205FF9AFFE406C6FB3CFAA42CD0" box="[1242,1405,1189,1213]" pageId="13" pageNumber="449" value="2004-05-12">12 May 2004</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B411770FF9AFFE40792FB3CFB022D69" ID-GBIF-Occurrence="1291862177" collectingDate="2008-04-22" country="Ukraine" latitude="46.716667" location="Kirovo village" longLatPrecision="1121" longitude="30.266666" municipality="Razdelnyansky district" pageId="13" pageNumber="449" specimenCount="10" specimenCount-female="3" specimenCount-male="7" stateProvince="Odessa">
<specimenCount id="9D2FD6A4FF9AFFE40792FB3CFF282C8D" pageId="13" pageNumber="449" type="female">3 females</specimenCount>
, 
<specimenCount id="9D2FD6A4FF9AFFE4031CFB51FE872C8D" box="[256,350,1223,1248]" pageId="13" pageNumber="449" type="male">7 males</specimenCount>
, 43 deutonymphs, 
<collectingCountry id="F33E5DBDFF9AFFE40022FB5EFD422C8D" box="[574,667,1223,1248]" name="Ukraine" pageId="13" pageNumber="449">Ukraine</collectingCountry>
, 
<collectingRegion id="49EDD3CFFF9AFFE400B6FB5EFD222C8D" box="[682,763,1223,1248]" country="Ukraine" name="Odesa" pageId="13" pageNumber="449">Odessa</collectingRegion>
oblast, 
<collectingMunicipality id="6BF28757FF9AFFE40147FB5EFBBA2C8D" box="[859,1123,1223,1248]" pageId="13" pageNumber="449">Razdelnyansky district</collectingMunicipality>
, 
<location id="8EF64BF6FF9AFFE4066EFB5EFAC02C8D" LSID="urn:lsid:plazi:treatment:0380AC3BFF9FFFE7028BFC12FB4929BD:8EF64BF6FF9AFFE4066EFB5EFAC02C8D" box="[1138,1305,1223,1248]" country="Ukraine" latitude="46.716667" longLatPrecision="1121" longitude="30.266666" municipality="Razdelnyansky district" name="Kirovo village" pageId="13" pageNumber="449" stateProvince="Odessa">Kirovo village</location>
(
<geoCoordinate id="EE1D7BEAFF9AFFE40730FB5EFA412CB2" box="[1324,1432,1223,1248]" direction="north" orientation="latitude" pageId="13" pageNumber="449" precision="925" value="46.716667">46°43' N</geoCoordinate>
, 
<geoCoordinate id="EE1D7BEAFF9AFFE4028BFB72FED92D69" box="[151,256,1259,1284]" direction="east" orientation="longitude" pageId="13" pageNumber="449" precision="925" value="30.266666">30°16' E</geoCoordinate>
), in soil with bird faeces from the chicken coops (with chickens), 
<date id="FF973BEDFF9AFFE40631FB75FB022D69" box="[1069,1243,1259,1284]" pageId="13" pageNumber="449" value="2008-04-22">
<collectingDate id="EFD3C205FF9AFFE40631FB75FB022D69" box="[1069,1243,1259,1284]" pageId="13" pageNumber="449" value="2008-04-22">22 April 2008</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B411770FF9AFFE406F0FB75FA412D45" ID-GBIF-Occurrence="1291862179" collectingDate="2008-04-22" country="Ukraine" latitude="46.716667" location="Kirovo village" longLatPrecision="1121" longitude="30.266666" municipality="Razdelnyansky district" pageId="13" pageNumber="449" specimenCount="2" specimenCount-female="2" stateProvince="Odessa">
<specimenCount id="9D2FD6A4FF9AFFE406F0FB75FAB82D69" box="[1260,1377,1259,1284]" pageId="13" pageNumber="449" type="female">2 females</specimenCount>
, 120 deutonymphs, same locality and date, in soil with bird faeces from the chicken coops (with geese); 2 deutonymphs
</materialsCitation>
, 
<materialsCitation id="3B411770FF9AFFE4028BFAADFD7C2D26" ID-GBIF-Occurrence="1291862174" box="[151,677,1331,1356]" collectingDate="2008-05-18" country="Ukraine" latitude="46.716667" location="Kirovo village" longLatPrecision="1121" longitude="30.266666" municipality="Razdelnyansky district" pageId="13" pageNumber="449" specimenCount="1" stateProvince="Odessa">
same locality, in old cow dung, 
<date id="FF973BEDFF9AFFE40014FAAAFD7C2D26" box="[520,677,1331,1356]" pageId="13" pageNumber="449" value="2008-05-18">
<collectingDate id="EFD3C205FF9AFFE40014FAAAFD7C2D26" box="[520,677,1331,1356]" pageId="13" pageNumber="449" value="2008-05-18">18 May 2008</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B411770FF9AFFE400AFFAAAFDFA2DFE" ID-GBIF-Occurrence="1291862176" collectingDate="2014-01-12" collectorName="P. Yatsenko" country="Ukraine" latitude="46.383335" location="Udobnoe village" longLatPrecision="1124" longitude="30.05" municipality="Belgorod-Dnestrovsky district" pageId="13" pageNumber="449" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Odessa">
<specimenCount id="9D2FD6A4FF9AFFE400AFFAAAFCC32D21" box="[691,794,1331,1356]" pageId="13" pageNumber="449" type="female">1 female</specimenCount>
, 
<specimenCount id="9D2FD6A4FF9AFFE4013BFAAAFCA12D21" box="[807,888,1331,1356]" pageId="13" pageNumber="449" type="male">1 male</specimenCount>
, 
<collectingCountry id="F33E5DBDFF9AFFE40199FAAAFC3B2D21" box="[901,994,1331,1356]" name="Ukraine" pageId="13" pageNumber="449">Ukraine</collectingCountry>
, 
<collectingRegion id="49EDD3CFFF9AFFE401ECFAAAFB982D21" box="[1008,1089,1331,1356]" country="Ukraine" name="Odesa" pageId="13" pageNumber="449">Odessa</collectingRegion>
oblast, 
<collectingMunicipality id="6BF28757FF9AFFE40681FAAAFF312D02" pageId="13" pageNumber="449">Belgorod-Dnestrovsky district</collectingMunicipality>
, 
<location id="8EF64BF6FF9AFFE402EBFACFFE6F2D02" LSID="urn:lsid:plazi:treatment:0380AC3BFF9FFFE7028BFC12FB4929BD:8EF64BF6FF9AFFE402EBFACFFE6F2D02" box="[247,438,1366,1391]" country="Ukraine" latitude="46.383335" longLatPrecision="1124" longitude="30.05" municipality="Belgorod-Dnestrovsky district" name="Udobnoe village" pageId="13" pageNumber="449" stateProvince="Odessa">Udobnoe village</location>
(
<geoCoordinate id="EE1D7BEAFF9AFFE403D5FACFFDED2D02" box="[457,564,1366,1391]" direction="north" orientation="latitude" pageId="13" pageNumber="449" precision="925" value="46.383335">46°23' N</geoCoordinate>
, 
<geoCoordinate id="EE1D7BEAFF9AFFE40058FACEFD722D02" box="[580,683,1367,1391]" direction="east" orientation="longitude" pageId="13" pageNumber="449" precision="925" value="30.05">30°03' E</geoCoordinate>
), soil with bird faeces from the chicken coops (with ducks), 
<date id="FF973BEDFF9AFFE4079CFACEFEE82DFE" pageId="13" pageNumber="449" value="2014-01-12">
<collectingDate id="EFD3C205FF9AFFE4079CFACEFEE82DFE" pageId="13" pageNumber="449" value="2014-01-12">12 January 2014</collectingDate>
</date>
, Yu. 
<collectorName id="26DC78FBFF9AFFE40378FAE2FE3E2DFE" box="[356,487,1402,1427]" pageId="13" pageNumber="449">P. Yatsenko</collectorName>
coll.
</materialsCitation>
; 
<materialsCitation id="3B411770FF9AFFE4002CFAE2FED72DB6" ID-GBIF-Occurrence="1291862175" collectingDate="2014-04-23" collectorName="P. Yatsenko" country="Ukraine" latitude="46.383335" location="Udobnoe village" longLatPrecision="1124" longitude="30.05" municipality="Belgorod-Dnestrovsky district" pageId="13" pageNumber="449" specimenCount="1" specimenCount-male="1" stateProvince="Odessa">
<specimenCount id="9D2FD6A4FF9AFFE4002CFAE2FDA62DFE" box="[560,639,1402,1427]" pageId="13" pageNumber="449" type="male">1 male</specimenCount>
, 4 deutonymphs, same locality, soil with bird faeces from the chicken coops (with chickens), 
<date id="FF973BEDFF9AFFE40381FA06FDE62DDA" box="[413,575,1438,1463]" pageId="13" pageNumber="449" value="2014-04-23">
<collectingDate id="EFD3C205FF9AFFE40381FA06FDE62DDA" box="[413,575,1438,1463]" pageId="13" pageNumber="449" value="2014-04-23">23 April 2014</collectingDate>
</date>
, Yu. 
<collectorName id="26DC78FBFF9AFFE4006DFA06FD2D2DDA" box="[625,756,1438,1463]" pageId="13" pageNumber="449">P. Yatsenko</collectorName>
coll. Unless otherwise stated, all material were collected by the author.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3334EA6FF9AFFE402DBFA7CFB1C2F9F" pageId="13" pageNumber="449" type="discussion">
<paragraph id="8B961D2DFF9AFFE402DBFA7CFE222E04" blockId="13.[151,1437,151,2035]" pageId="13" pageNumber="449">
<emphasis id="B95DC13FFF9AFFE402DBFA7CFEE22D93" bold="true" box="[199,315,1509,1534]" pageId="13" pageNumber="449">Remarks.</emphasis>
<taxonomicName id="4C2966AEFF9AFFE4035AFA7CFDB12D93" box="[326,616,1509,1534]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="449" phylum="Arthropoda" rank="species" species="saproincisus">
<emphasis id="B95DC13FFF9AFFE4035AFA7CFDB12D93" box="[326,616,1509,1534]" italics="true" pageId="13" pageNumber="449">Halolaelaps saproincisus</emphasis>
</taxonomicName>
was described from cattle dung in 
<collectingCountry id="F33E5DBDFF9AFFE4060FFA7FFBA22D93" box="[1043,1147,1510,1534]" name="Germany" pageId="13" pageNumber="449">Germany</collectingCountry>
by 
<bibRefCitation id="EFB860DCFF9AFFE406B3FA7CFF3C2E4F" author="Hirschmann" pageId="13" pageNumber="449" refString="Hirschmann, W. &amp; Gotz, H. (1968) Neue Halolaelaps - Arten. Acarologie. Schriftenreihe fur Vergleichende Milbenkunde, 11, 7 - 10." type="journal article" year="1968">Hirschmann &amp; Götz (1968)</bibRefCitation>
. It has also been recorded from compost and dung mixed with soil in 
<collectingCountry id="F33E5DBDFF9AFFE4063EF990FBB42E4F" box="[1058,1133,1545,1570]" name="Russia" pageId="13" pageNumber="449">Russia</collectingCountry>
(
<bibRefCitation id="EFB860DCFF9AFFE40663F993FF0A2E2B" author="Bregetova" pageId="13" pageNumber="449" refString="Bregetova, N. G. &amp; Shcherbak, G. I. (1977) Family Rhodacaridae Oudemans, 1902. In: Ghilyarov, M. S. &amp; Bregetova, N. G. (Eds.), Key to the Soil - Inhabiting Mites. Mesostigmata, Nauka, Leningrad, pp. 256 - 307. [in Russian]" type="book chapter" year="1977">Bregetova &amp; Shcherbak, 1977</bibRefCitation>
), from dung in 
<collectingCountry id="F33E5DBDFF9AFFE403B9F9B4FDDC2E2B" box="[421,517,1581,1606]" name="Bulgaria" pageId="13" pageNumber="449">Bulgaria</collectingCountry>
(
<bibRefCitation id="EFB860DCFF9AFFE40001F9B4FC622E2B" author="Koyumdjieva" box="[541,955,1581,1606]" pageId="13" pageNumber="449" refString="Koyumdjieva, M. I. &amp; Angelkova, E. B. (1985) Species composition and population dynamics of mites (Gamasina, Mesostigmata and Acaridiae, Astigmata) in a dunghill. Acta Zoologica Bulgarica, 28, 17 - 27." type="journal article" year="1985">Koyumdjieva &amp; Angelkova, 1985</bibRefCitation>
), from unknown excrements in 
<collectingCountry id="F33E5DBDFF9AFFE40752F9B4FA452E2B" box="[1358,1436,1581,1606]" name="Poland" pageId="13" pageNumber="449">Poland</collectingCountry>
(
<bibRefCitation id="EFB860DCFF9AFFE40282F9C8FE362E04" author="Gwiazdowicz" box="[158,495,1617,1642]" pageId="13" pageNumber="449" refString="Gwiazdowicz, D. J. &amp; Klemt, J. (2004) Mesostigmatic mites (Acari, Gamasida) in selected microhabitats of the Biebrza National Park (NE Poland). Biology Letters, 41, 11 - 19." type="journal article" year="2004">Gwiazdowicz &amp; Klemt, 2004</bibRefCitation>
).
</paragraph>
<paragraph id="8B961D2DFF9AFFE402DBF9ECFBE22F2D" blockId="13.[151,1437,151,2035]" pageId="13" pageNumber="449">
By the presence of club-like setae on the coxae, deutonymphs of 
<taxonomicName id="4C2966AEFF9AFFE401E5F9EFFB762EE0" box="[1017,1199,1653,1677]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="449" phylum="Arthropoda" rank="species" species="saproincisus">
<emphasis id="B95DC13FFF9AFFE401E5F9EFFBD42EE0" box="[1017,1037,1654,1677]" italics="true" pageId="13" pageNumber="449">H</emphasis>
. 
<emphasis id="B95DC13FFF9AFFE4063FF9EFFB762EE0" box="[1059,1199,1654,1677]" italics="true" pageId="13" pageNumber="449">saproincisus</emphasis>
</taxonomicName>
is similar to other 
<taxonomicName id="4C2966AEFF9AFFE4028BF901FEF92EDC" box="[151,288,1688,1713]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="449" phylum="Arthropoda" rank="genus">
<emphasis id="B95DC13FFF9AFFE4028BF901FEF92EDC" box="[151,288,1688,1713]" italics="true" pageId="13" pageNumber="449">Halolaelaps</emphasis>
</taxonomicName>
species: 
<taxonomicName id="4C2966AEFF9AFFE40387F900FCE12EDC" authority="Oudemans, 1902" authorityName="Oudemans" authorityYear="1902" box="[411,824,1688,1713]" class="Arachnida" family="Parasitidae" genus="Parasitus" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="449" phylum="Arthropoda" rank="species" species="sexclavatus">
<emphasis id="B95DC13FFF9AFFE40387F900FE762EDD" box="[411,431,1689,1712]" italics="true" pageId="13" pageNumber="449">H</emphasis>
. 
<emphasis id="B95DC13FFF9AFFE403D4F901FD902EDC" box="[456,585,1688,1713]" italics="true" pageId="13" pageNumber="449">sexclavatus</emphasis>
(Oudemans, 1902)
</taxonomicName>
, 
<taxonomicName id="4C2966AEFF9AFFE40152F900FB3E2EDC" authority="Vitzthum, 1918" authorityName="Vitzthum" authorityYear="1918" box="[846,1255,1688,1713]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="449" phylum="Arthropoda" rank="species" species="octoclavatus">
<emphasis id="B95DC13FFF9AFFE40152F900FCBB2EDD" box="[846,866,1689,1712]" italics="true" pageId="13" pageNumber="449">H</emphasis>
. 
<emphasis id="B95DC13FFF9AFFE40167F901FBD12EDC" box="[891,1032,1688,1713]" italics="true" pageId="13" pageNumber="449">octoclavatus</emphasis>
(
<bibRefCitation id="EFB860DCFF9AFFE4063DF901FB062EDC" author="Vitzthum" box="[1057,1247,1688,1713]" pageId="13" pageNumber="449" refString="Vitzthum, H. G. (1918) Acarologische Beobachtungen. Zweite Reihe. Archiv fur Naturgeschichte, Series A, 84 (6), 1 - 40." type="journal article" year="1918">Vitzthum, 1918</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C2966AEFF9AFFE406E1F900FEBC2EB8" authority="Schweizer, 1949" authorityName="Schweizer" authorityYear="1949" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="449" phylum="Arthropoda" rank="species" species="orbinellus">
<emphasis id="B95DC13FFF9AFFE406E1F900FAC82EDD" box="[1277,1297,1689,1712]" italics="true" pageId="13" pageNumber="449">H</emphasis>
. 
<emphasis id="B95DC13FFF9AFFE40736F901FA452EDC" box="[1322,1436,1688,1713]" italics="true" pageId="13" pageNumber="449">orbinellus</emphasis>
(
<bibRefCitation id="EFB860DCFF9AFFE40282F925FE872EB8" author="Schweizer" box="[158,350,1724,1749]" pageId="13" pageNumber="449" refString="Schweizer, J. (1949) Die Landmilben des Schweizerischen Nationalparkes. 1. Teil: Parasitiformes Reuter 1909. Ergebnisse der Wissenschaftlichen Untersuchung des Schweizerischen Nationalparks, 2, 1 - 99." type="journal article" year="1949">Schweizer, 1949</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C2966AEFF9AFFE4036DF924FD122EB8" authority="Ryke, 1961" authorityName="Ryke" authorityYear="1961" box="[369,715,1724,1749]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="449" phylum="Arthropoda" rank="species" species="neoorbinellus">
<emphasis id="B95DC13FFF9AFFE4036DF924FE5C2EB9" box="[369,389,1725,1748]" italics="true" pageId="13" pageNumber="449">H</emphasis>
. 
<emphasis id="B95DC13FFF9AFFE40388F925FDF72EB8" box="[404,558,1724,1749]" italics="true" pageId="13" pageNumber="449">neoorbinellus</emphasis>
(
<bibRefCitation id="EFB860DCFF9AFFE40021F925FD182EB8" author="Ryke" box="[573,705,1724,1749]" pageId="13" pageNumber="449" refString="Ryke, P. A. J. (1961) A review of the genus Saintdidieria Oudemans (Acarina: Rhodacaridae) with remarks on the genus Lobocephalus Kramer. Acarologia, 3, 250 - 255." type="journal article" year="1961">Ryke, 1961</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C2966AEFF9AFFE400CAF924FBC62EB8" authority="Elsen, 1974" authorityName="Elsen" authorityYear="1974" box="[726,1055,1724,1749]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="449" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="B95DC13FFF9AFFE400CAF924FD332EB9" box="[726,746,1725,1748]" italics="true" pageId="13" pageNumber="449">H</emphasis>
. 
<emphasis id="B95DC13FFF9AFFE400E5F925FCA42EB8" box="[761,893,1724,1749]" italics="true" pageId="13" pageNumber="449">intermedius</emphasis>
(
<bibRefCitation id="EFB860DCFF9AFFE40190F925FBCF2EB8" author="Elsen" box="[908,1046,1724,1749]" pageId="13" pageNumber="449" refString="Elsen, P. (1974) Saintdidieria intermedia spec. nov. phoretique d'un Coleoptere d'Afrique Centrale (Acarina: Rhodacaridae). Revue de Zoologie Africaine, 88, 453 - 456." type="journal article" year="1974">Elsen, 1974</bibRefCitation>
)
</taxonomicName>
. Deutonymphs of 
<taxonomicName id="4C2966AEFF9AFFE406F1F924FA452EB9" box="[1261,1436,1725,1748]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="449" phylum="Arthropoda" rank="species" species="saproincisus">
<emphasis id="B95DC13FFF9AFFE406F1F924FAD82EB9" box="[1261,1281,1725,1748]" italics="true" pageId="13" pageNumber="449">H</emphasis>
. 
<emphasis id="B95DC13FFF9AFFE40713F924FA452EB9" box="[1295,1436,1725,1748]" italics="true" pageId="13" pageNumber="449">saproincisus</emphasis>
</taxonomicName>
differs from other species of 
<taxonomicName id="4C2966AEFF9AFFE403C1F946FDBF2E95" box="[477,614,1759,1784]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="449" phylum="Arthropoda" rank="genus">
<emphasis id="B95DC13FFF9AFFE403C1F946FDBF2E95" box="[477,614,1759,1784]" italics="true" pageId="13" pageNumber="449">Halolaelaps</emphasis>
</taxonomicName>
by normal 
<emphasis id="B95DC13FFF9AFFE400F4F978FCDA2E95" box="[744,771,1761,1784]" italics="true" pageId="13" pageNumber="449">pv</emphasis>
seta on coxa I and unmodified setae 
<emphasis id="B95DC13FFF9AFFE406B9F978FB192E95" box="[1189,1216,1761,1784]" italics="true" pageId="13" pageNumber="449">pc</emphasis>
on the gnathosoma (
<taxonomicName id="4C2966AEFF9AFFE402BCF89CFE9C2F71" box="[160,325,1795,1820]" class="Arachnida" family="Parasitidae" genus="Parasitus" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="449" phylum="Arthropoda" rank="species" species="sexclavatus">
<emphasis id="B95DC13FFF9AFFE402BCF89CFF6D2F71" box="[160,180,1797,1820]" italics="true" pageId="13" pageNumber="449">H</emphasis>
. 
<emphasis id="B95DC13FFF9AFFE402D8F89AFE9C2F71" box="[196,325,1795,1820]" italics="true" pageId="13" pageNumber="449">sexclavatus</emphasis>
</taxonomicName>
also has 
<emphasis id="B95DC13FFF9AFFE403ADF89CFE152F71" box="[433,460,1797,1820]" italics="true" pageId="13" pageNumber="449">pv</emphasis>
seta on coxa I not-modified, but seta 
<emphasis id="B95DC13FFF9AFFE40161F89CFC412F71" box="[893,920,1797,1820]" italics="true" pageId="13" pageNumber="449">pc</emphasis>
is club-like). Moreover, modified setae in 
<taxonomicName id="4C2966AEFF9AFFE4079DF89CFEFA2F52" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="449" phylum="Arthropoda" rank="species" species="saproincisus">
<emphasis id="B95DC13FFF9AFFE4079DF89CFA4C2F71" box="[1409,1429,1797,1820]" italics="true" pageId="13" pageNumber="449">H</emphasis>
. 
<emphasis id="B95DC13FFF9AFFE4028BF8B1FEFA2F52" box="[151,291,1832,1855]" italics="true" pageId="13" pageNumber="449">saproincisus</emphasis>
</taxonomicName>
are asymmetrical and in other species club-like setae are symmetrical.
</paragraph>
<paragraph id="8B961D2DFF9AFFE402DBF8D5FB1C2F9F" blockId="13.[151,1437,151,2035]" pageId="13" pageNumber="449">
It should be noted that thick or club-like setae (
<emphasis id="B95DC13FFF9AFFE400C5F8D5FD2D2F0E" box="[729,756,1868,1891]" italics="true" pageId="13" pageNumber="449">pc</emphasis>
, 
<emphasis id="B95DC13FFF9AFFE4011DF8D2FCE72F0E" box="[769,830,1867,1891]" italics="true" pageId="13" pageNumber="449">st1–5</emphasis>
, 
<emphasis id="B95DC13FFF9AFFE40150F8D5FCBE2F0E" box="[844,871,1868,1891]" italics="true" pageId="13" pageNumber="449">pv</emphasis>
on coxae I–III) are present in a number of beetleassociated 
<taxonomicName id="4C2966AEFF9AFFE4030BF8F7FE532FEA" box="[279,394,1902,1927]" class="Arachnida" higherTaxonomySource="CoL" infraOrder="Gamasina" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="449" phylum="Arthropoda" rank="infraOrder">Gamasina</taxonomicName>
, e.g. some species of the genera 
<taxonomicName id="4C2966AEFF9AFFE4013FF8F7FC662FEA" box="[803,959,1902,1927]" class="Arachnida" family="Parasitidae" genus="Poecilochirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="449" phylum="Arthropoda" rank="genus">
<emphasis id="B95DC13FFF9AFFE4013FF8F7FC662FEA" box="[803,959,1902,1927]" italics="true" pageId="13" pageNumber="449">Poecilochirus</emphasis>
</taxonomicName>
G. &amp; R. Canestrini, 1882 (
<taxonomicName id="4C2966AEFF9AFFE40713F8F7FA492FEA" box="[1295,1424,1902,1927]" class="Arachnida" family="Parasitidae" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="449" phylum="Arthropoda" rank="family">Parasitidae</taxonomicName>
), 
<taxonomicName id="4C2966AEFF9AFFE4028BF80BFD722FC6" authority="Willmann, 1949" authorityName="Willmann" authorityYear="1949" box="[151,683,1938,1963]" class="Arachnida" family="Ascidae" genus="Panteniphis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="449" phylum="Arthropoda" rank="genus">
<emphasis id="B95DC13FFF9AFFE4028BF80BFEC42FC6" box="[151,285,1938,1963]" italics="true" pageId="13" pageNumber="449">Panteniphis</emphasis>
Willmann, 1949 (Digamasellidae)
</taxonomicName>
, 
<taxonomicName id="4C2966AEFF9AFFE400A5F80BFCEC2FC6" box="[697,821,1938,1963]" class="Arachnida" family="Ologamasidae" genus="Stylochirus" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="449" phylum="Arthropoda" rank="genus">
<emphasis id="B95DC13FFF9AFFE400A5F80BFCEC2FC6" box="[697,821,1938,1963]" italics="true" pageId="13" pageNumber="449">Stylochirus</emphasis>
</taxonomicName>
G. &amp; R. Canestrini, 1882 (
<taxonomicName id="4C2966AEFF9AFFE4066CF80BFACF2FC6" box="[1136,1302,1938,1963]" class="Arachnida" family="Ologamasidae" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="449" phylum="Arthropoda" rank="family">Ologamasidae</taxonomicName>
), 
<taxonomicName id="4C2966AEFF9AFFE40730F80BFE392FA2" authority="Berlese, 1910" authorityName="Berlese" authorityYear="1910" class="Arachnida" family="Eviphididae" genus="Copriphis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="449" phylum="Arthropoda" rank="genus">
<emphasis id="B95DC13FFF9AFFE40730F80BFA452FC6" box="[1324,1436,1938,1963]" italics="true" pageId="13" pageNumber="449">Copriphis</emphasis>
Berlese, 1910 (Eviphididae)
</taxonomicName>
, 
<taxonomicName id="4C2966AEFF9AFFE403EDF82EFCEB2FA2" authority="Berlese, 1916" authorityName="Berlese" authorityYear="1916" box="[497,818,1974,1999]" class="Arachnida" family="Ascidae" genus="Antennoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="449" phylum="Arthropoda" rank="genus">
<emphasis id="B95DC13FFF9AFFE403EDF82EFD5F2FA3" box="[497,646,1975,1998]" italics="true" pageId="13" pageNumber="449">Antennoseius</emphasis>
Berlese, 1916
</taxonomicName>
and 
<taxonomicName id="4C2966AEFF9AFFE40168F82FFAF32FA2" authority="Berlese, 1911" authorityName="Berlese" authorityYear="1911" box="[884,1322,1974,1999]" class="Arachnida" family="Ascidae" genus="Anystipalpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="449" phylum="Arthropoda" rank="genus">
<emphasis id="B95DC13FFF9AFFE40168F82FFBDC2FA2" box="[884,1029,1974,1999]" italics="true" pageId="13" pageNumber="449">Anystipalpus</emphasis>
Berlese, 1911 (Ascidae)
</taxonomicName>
(
<bibRefCitation id="EFB860DCFF9AFFE40722F82FFEF12F9E" author="Ryke" pageId="13" pageNumber="449" refString="Ryke, P. A. J. &amp; Meyer, M. K. P. (1957) Eviphidinae Berlese 1913 (Mesostigmata: Acarina) associated with South African beetles. Annals and Magazine of Natural History, Series 12, 10, 593 - 618. http: // dx. doi. org / 10.1080 / 00222935708656003" type="journal article" year="1957">Ryke &amp; Meyer, 1957</bibRefCitation>
; Bregetova, 1977; 
<bibRefCitation id="EFB860DCFF9AFFE403E7F843FD492F9F" author="Masan" box="[507,656,2010,2034]" pageId="13" pageNumber="449" refString="Masan, P. (1999) Mites (Acarina) associated with burying and carrion beetles (Coleoptera, Silphidae) and description of Poecilochirus mrciaki sp. n. (Mesostigmata, Gamasina). Biologia, Bratislava, 54, 515 - 524." type="journal article" year="1999">Mašán, 1999</bibRefCitation>
; 
<bibRefCitation id="EFB860DCFF9AFFE40087F843FCA62F9F" author="Gwiazdowicz" box="[667,895,2010,2035]" pageId="13" pageNumber="449" refString="Gwiazdowicz, D. J. (2000) Description of deutonymph of Panteniphis mirandus (Acari: Rhodacaridae) from Poland. Biologia, Bratislava, 55, 465 - 470." type="journal article" year="2000">Gwiazdowicz, 2000</bibRefCitation>
; 
<bibRefCitation id="EFB860DCFF9AFFE40196F843FB602F9F" author="Lindquist" box="[906,1209,2010,2035]" pageId="13" pageNumber="449" refString="Lindquist, E. E. &amp; Moraza, M. L. (2009) Anystipalpus, Antennoseius and Vitzthumia: a taxonomic and nomenclatural conundrum of genera (Acari: Mesostigmata: Dermanyssina), with description of four species of Anystipalpus. Zootaxa, 2243, 1 - 39." type="journal article" year="2009">Lindquist &amp; Moraza, 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3334EA6FF99FFE702DBFF0EFB4929BD" pageId="14" pageNumber="450" type="reference_group">
<paragraph id="8B961D2DFF99FFE702DBFF0EFD3E2908" blockId="14.[151,1436,151,464]" pageId="14" pageNumber="450">
<taxonomicName id="4C2966AEFF99FFE702DBFF0EFE3228DD" box="[199,491,151,176]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="450" phylum="Arthropoda" rank="species" species="octoclavatus">
<emphasis id="B95DC13FFF99FFE702DBFF0EFE3228DD" box="[199,491,151,176]" italics="true" pageId="14" pageNumber="450">Halolaelaps octoclavatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2966AEFF99FFE7001DFF00FD4228DD" box="[513,667,151,176]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="450" phylum="Arthropoda" rank="species" species="orbinellus">
<emphasis id="B95DC13FFF99FFE7001DFF00FDCC28DD" box="[513,533,153,176]" italics="true" pageId="14" pageNumber="450">H</emphasis>
. 
<emphasis id="B95DC13FFF99FFE70036FF0EFD4228DD" box="[554,667,151,176]" italics="true" pageId="14" pageNumber="450">orbinellus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2966AEFF99FFE700ADFF00FCAD28DD" box="[689,884,151,176]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="450" phylum="Arthropoda" rank="species" species="neoorbinellus">
<emphasis id="B95DC13FFF99FFE700ADFF00FD1C28DD" box="[689,709,153,176]" italics="true" pageId="14" pageNumber="450">H</emphasis>
. 
<emphasis id="B95DC13FFF99FFE700C6FF0EFCAD28DD" box="[730,884,151,176]" italics="true" pageId="14" pageNumber="450">neoorbinellus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2966AEFF99FFE701A5FF00FB8428DD" box="[953,1117,151,176]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="450" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B95DC13FFF99FFE701A5FF00FC1428DD" box="[953,973,153,176]" italics="true" pageId="14" pageNumber="450">H</emphasis>
. 
<emphasis id="B95DC13FFF99FFE701FFFF0EFB8428DD" box="[995,1117,151,176]" italics="true" pageId="14" pageNumber="450">intermedia</emphasis>
</taxonomicName>
are known only from the deutonymph. Adults are described only in 
<taxonomicName id="4C2966AEFF99FFE7009EFF24FCF128B8" box="[642,808,188,213]" class="Arachnida" family="Parasitidae" genus="Parasitus" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="450" phylum="Arthropoda" rank="species" species="sexclavatus">
<emphasis id="B95DC13FFF99FFE7009EFF24FD4F28B9" box="[642,662,189,212]" italics="true" pageId="14" pageNumber="450">H</emphasis>
. 
<emphasis id="B95DC13FFF99FFE700BBFF25FCF128B8" box="[679,808,188,213]" italics="true" pageId="14" pageNumber="450">sexclavatus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB860DCFF99FFE70125FF25FC3628B9" author="Vitzthum" box="[825,1007,188,213]" pageId="14" pageNumber="450" refString="Vitzthum, H. G. (1918) Acarologische Beobachtungen. Zweite Reihe. Archiv fur Naturgeschichte, Series A, 84 (6), 1 - 40." type="journal article" year="1918">Vitzthum, 1918</bibRefCitation>
; 
<bibRefCitation id="EFB860DCFF99FFE701E1FF25FB6528B9" author="Schweizer" box="[1021,1212,188,213]" pageId="14" pageNumber="450" refString="Schweizer, J. (1949) Die Landmilben des Schweizerischen Nationalparkes. 1. Teil: Parasitiformes Reuter 1909. Ergebnisse der Wissenschaftlichen Untersuchung des Schweizerischen Nationalparks, 2, 1 - 99." type="journal article" year="1949">Schweizer, 1949</bibRefCitation>
; 
<bibRefCitation id="EFB860DCFF99FFE706D6FF25FA9628B8" author="Ryke" box="[1226,1359,188,213]" pageId="14" pageNumber="450" refString="Ryke, P. A. J. (1961) A review of the genus Saintdidieria Oudemans (Acarina: Rhodacaridae) with remarks on the genus Lobocephalus Kramer. Acarologia, 3, 250 - 255." type="journal article" year="1961">Ryke, 1961</bibRefCitation>
; 
<bibRefCitation id="EFB860DCFF99FFE70742FF24FF0B2895" author="Karg" pageId="14" pageNumber="450" refString="Karg, W. (1965) Larvalsystematische und phylogenetische untersuchung sowie revision des systems der Gamasina Leach, 1915 (Acarina, Parasitiformes). Mitteilungen aus dem Museum fur Naturkunde in Berlin, 41, 193 - 340. http: // dx. doi. org / 10.1002 / mmnz. 19650410207" type="journal article" year="1965">Karg, 1965</bibRefCitation>
; 
<bibRefCitation id="EFB860DCFF99FFE702C2FF46FEBC2895" author="Elsen" box="[222,357,223,248]" pageId="14" pageNumber="450" refString="Elsen, P. (1974) Saintdidieria intermedia spec. nov. phoretique d'un Coleoptere d'Afrique Centrale (Acarina: Rhodacaridae). Revue de Zoologie Africaine, 88, 453 - 456." type="journal article" year="1974">Elsen, 1974</bibRefCitation>
; 
<bibRefCitation id="EFB860DCFF99FFE7036DFF46FD102895" author="Blaszak" box="[369,713,223,248]" pageId="14" pageNumber="450" refString="Blaszak, C. &amp; Ehrnsberger, R. (1998 c) Eine neue Untergattung Haloseius subgen. nov. in der gattung Halolaelaps Berlese &amp; Trouessart, 1889 (Acari: Gamasida: Halolaelapidae). Genus, 9, 421 - 429." type="journal article" year="1998" yearSuffix="c">Błaszak &amp; Ehrnsberger, 1998c</bibRefCitation>
, 
<bibRefCitation id="EFB860DCFF99FFE700C9FF79FCC82895" author="Ehrnsberger" box="[725,785,224,248]" pageId="14" pageNumber="450" refString="Blaszak, C. &amp; Ehrnsberger, R. (2001) Beschreibung des Mannchens von Halolaelaps (Haloseius) sexclavatus (Oudemans, 1902) (Acari, Gamasida: Halolaelapidae). Osnabrucker Naturwissenschaftliche Mitteilungen, 27, 99 - 101." type="journal article" year="2001">2001</bibRefCitation>
). Adult females and males of 
<taxonomicName id="4C2966AEFF99FFE7067CFF78FAD62895" box="[1120,1295,224,248]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="450" phylum="Arthropoda" rank="species" species="saproincisus">
<emphasis id="B95DC13FFF99FFE7067CFF78FBAD2895" box="[1120,1140,225,248]" italics="true" pageId="14" pageNumber="450">H</emphasis>
. 
<emphasis id="B95DC13FFF99FFE7069EFF78FAD62895" box="[1154,1295,225,248]" italics="true" pageId="14" pageNumber="450">saproincisus</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C2966AEFF99FFE7028BFE9CFEE22970" box="[151,315,260,285]" class="Arachnida" family="Parasitidae" genus="Parasitus" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="450" phylum="Arthropoda" rank="species" species="sexclavatus">
<emphasis id="B95DC13FFF99FFE7028BFE9CFF722971" box="[151,171,261,284]" italics="true" pageId="14" pageNumber="450">H</emphasis>
. 
<emphasis id="B95DC13FFF99FFE702A6FE9DFEE22970" box="[186,315,260,285]" italics="true" pageId="14" pageNumber="450">sexclavatus</emphasis>
</taxonomicName>
in a number of important characters (e. g. by shape of epistome, number of setae on the podonotal shield, measurements and shape of setae on dorsal shields, shape of the anterior margin of opisthonotal shield, leg chaetotaxy) and these species are not closely related.
</paragraph>
<paragraph id="8B961D2DFF99FFE702DBFEF6FB4929BD" blockId="14.[151,1436,151,464]" pageId="14" pageNumber="450">
The phoretic associations of 
<taxonomicName id="4C2966AEFF99FFE70015FEE8FD6129E5" box="[521,696,368,392]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="450" phylum="Arthropoda" rank="species" species="saproincisus">
<emphasis id="B95DC13FFF99FFE70015FEE8FDC429E5" box="[521,541,369,392]" italics="true" pageId="14" pageNumber="450">H</emphasis>
. 
<emphasis id="B95DC13FFF99FFE70037FEE8FD6129E5" box="[555,696,369,392]" italics="true" pageId="14" pageNumber="450">saproincisus</emphasis>
</taxonomicName>
are not clear. Dung-beetles do not occur in the chicken coops that were sampled and dung-beetles collected in the vicinity of the chicken coop did not carry 
<taxonomicName id="4C2966AEFF99FFE706DFFE0CFAAC29C1" box="[1219,1397,405,428]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="450" phylum="Arthropoda" rank="species" species="saproincisus">
<emphasis id="B95DC13FFF99FFE706DFFE0CFB0E29C1" box="[1219,1239,405,428]" italics="true" pageId="14" pageNumber="450">H</emphasis>
. 
<emphasis id="B95DC13FFF99FFE706F5FE0CFAAC29C1" box="[1257,1397,405,428]" italics="true" pageId="14" pageNumber="450">saproincisus</emphasis>
</taxonomicName>
. Is possible that mites are associated with some other insects that have not yet been identified.
</paragraph>
</subSubSection>
</treatment>
</document>