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<document id="2FDC6884A91CE5EAE9400B04965F6797" ID-DOI="10.4202/app.00784.2020" ID-ISSN="1732-2421" ID-Zenodo-Dep="10626735" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1707250644161" checkinUser="felipe" docAuthor="Fernández, Mercedes, Zimicz, Ana N., Bond, Mariano, Chornogubsky, Laura, Arnal, Michelle, Cárdenas, Magalí &amp; Fernicola, Juan C." docDate="2021" docId="031187C5FFAFFF964B67FC54FEDAFE24" docLanguage="en" docName="ActaPalaeontolPol.66.1.85-97.pdf" docOrigin="Acta Palaeontologica Polonica 66 (1)" docSource="http://dx.doi.org/10.4202/app.00784.2020" docStyle="DocumentStyle:E4780F6D34569985EDF9CACFB9BC780E.4:ActaPalaeontolPol.2015-.journal_article" docStyleId="E4780F6D34569985EDF9CACFB9BC780E" docStyleName="ActaPalaeontolPol.2015-.journal_article" docStyleVersion="4" docTitle="Pampahippus secundus Deraco and Garcia-Lopez 2016" docType="treatment" docVersion="4" lastPageNumber="94" masterDocId="FF28FFBDFFA8FF9F484FFFA3FFDAFF98" masterDocTitle="New Eocene South American native ungulates from the Quebrada de los Colorados Formation at Los Cardones National Park, Argentina" masterLastPageNumber="97" masterPageNumber="85" pageNumber="92" updateTime="1714484150498" updateUser="karina" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1B0BFA0285D8ACFF43B57C7437EC58DA">New Eocene South American native ungulates from the Quebrada de los Colorados Formation at Los Cardones National Park, Argentina</mods:title>
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<taxonomicName id="4CB84D50FFAFFF984B67FC54FCB2FBA1" authority="Deraco and Garcia-Lopez, 2016" authorityName="Deraco and Garcia-Lopez" authorityYear="2016" class="Mammalia" family="Notohippidae" genus="Pampahippus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="7" pageNumber="92" phylum="Chordata" rank="species" species="secundus">
<emphasis id="B9CCEAC1FFAFFF984B67FC54FB8FFB8C" box="[808,1109,1015,1044]" italics="true" pageId="7" pageNumber="92">Pampahippus secundus</emphasis>
<bibRefCitation id="EF294B22FFAFFF984C12FC5BFCB2FBA1" author="Deraco, V. &amp; Garcia-Lopez, D. A." pageId="7" pageNumber="92" pagination="1037884" refId="ref11377" refString="Deraco, V. and Garcia-Lopez, D. A. 2016. A new Eocene Toxodontia (Mammalia, Notoungulata) from northwestern Argentina. Journal of Vertebrate Paleontology 36: e 1037884." type="journal article" year="2016">Deraco and García-López, 2016</bibRefCitation>
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<paragraph id="8B0736D3FFAFFF984B67FBEBFC48FBC5" blockId="7.[808,914,1096,1118]" box="[808,914,1096,1118]" pageId="7" pageNumber="92">
<figureCitation id="13832A56FFAFFF984B67FBEBFC55FBC5" box="[808,911,1096,1118]" captionStart="Fig" captionStartId="8.[151,184,1111,1133]" captionTargetBox="[156,1526,166,1080]" captionTargetId="figure-538@8.[153,1526,182,1068]" captionTargetPageId="8" captionText="Fig. 3. New fossil specimens of Typotheria and Toxodontia from the Quebrada de Los Colorados Formation exposed at Quebrada Grande locality, Los Cardones National Park (Salta Province), Casamayoran SALMA. A, B. Colbertiasp. A. IBIGEO-P 58a, left maxillary fragment with M2M3 in occlusal view.B. IBIGEO-P 58b, left maxillary fragment with P4M3 in occlusal view.CE. Pampahippus secundus Deraco and García-López, 2016. C. IBIGEO-P 62, right trigonid and incomplete talonid of a lower right molar in occlusal view. D. IBIGEO-P 63, fragmented left p3 or p4 in occlusal view. E. IBIGEO-P 64, left m1? in occlusal (E1), labial (E2), and lingual (E3) views. Photographs (A1E1, E3, E4), explanatory drawings (A2E2). Dashed area indicates broken or missing dental areas." figureDoi="http://doi.org/10.5281/zenodo.10626739" httpUri="https://zenodo.org/record/10626739/files/figure.png" pageId="7" pageNumber="92">Fig. 3CE</figureCitation>
.
</paragraph>
<paragraph id="8B0736D3FFAFFF984B67FBD6FAB1FB12" blockId="7.[808,1387,1141,1163]" box="[808,1387,1141,1163]" pageId="7" pageNumber="92">
<materialsCitation id="3BD03C8EFFAFFF984B67FBD6FAB1FB12" box="[808,1387,1141,1163]" collectionCode="PVL" pageId="7" pageNumber="92" specimenCode="PVL 6426" specimenCount="1" typeStatus="holotype">
<emphasis id="B9CCEAC1FFAFFF984B67FBD6FC5AFB12" box="[808,896,1141,1162]" italics="true" pageId="7" pageNumber="92">
<typeStatus id="54038871FFAFFF984B67FBD6FC5AFB12" box="[808,896,1141,1162]" pageId="7" pageNumber="92">Holotype</typeStatus>
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:
<specimenCode id="DB1E9EA8FFAFFF984BC2FBD6FC2BFB13" box="[909,1009,1141,1163]" collectionCode="PVL" country="Argentina" httpUri="http://grbio.org/cool/pbkd-hmeb" name="Paleontologia de Vertebrados Lillo" pageId="7" pageNumber="92">PVL 6426</specimenCode>
, right maxillary fragment with P2M3.
</materialsCitation>
</paragraph>
<paragraph id="8B0736D3FFAFFF984B67FB34FC55FB51" blockId="7.[808,1481,1175,1225]" pageId="7" pageNumber="92">
<emphasis id="B9CCEAC1FFAFFF984B67FB34FC72FB34" box="[808,936,1175,1196]" italics="true" pageId="7" pageNumber="92">Type locality</emphasis>
: El Simbolar, Guachipas Department,
<collectingRegion id="497CF831FFAFFF984D7CFB34FA1FFB35" box="[1331,1477,1175,1197]" country="Argentina" name="Salta" pageId="7" pageNumber="92">Salta Province</collectingRegion>
,
<collectingCountry id="F3AF7643FFAFFF984B67FB10FC51FB51" box="[808,907,1203,1225]" name="Argentina" pageId="7" pageNumber="92">Argentina</collectingCountry>
.
</paragraph>
<paragraph id="8B0736D3FFAFFF984B67FB75FC81FA9F" blockId="7.[808,1481,1237,1287]" pageId="7" pageNumber="92">
<emphasis id="B9CCEAC1FFAFFF984B67FB75FC7CFB72" box="[808,934,1237,1259]" italics="true" pageId="7" pageNumber="92">Type horizon</emphasis>
: Upper section of the Lumbrera Formation, middle Eocene.
</paragraph>
<paragraph id="8B0736D3FFAFFF984B67FABAFA94FA2D" blockId="7.[808,1481,1305,1461]" pageId="7" pageNumber="92">
<emphasis id="B9CCEAC1FFAFFF984B67FABAFC50FAAB" box="[808,906,1305,1331]" italics="true" pageId="7" pageNumber="92">Material</emphasis>
.—IBIGEO-P 62, right trigonid and incomplete talonid of a lower molar (
<figureCitation id="13832A56FFAFFF984C79FA9AFB54FACC" box="[1078,1166,1337,1364]" captionStart="Fig" captionStartId="8.[151,184,1111,1133]" captionTargetBox="[156,1526,166,1080]" captionTargetId="figure-538@8.[153,1526,182,1068]" captionTargetPageId="8" captionText="Fig. 3. New fossil specimens of Typotheria and Toxodontia from the Quebrada de Los Colorados Formation exposed at Quebrada Grande locality, Los Cardones National Park (Salta Province), Casamayoran SALMA. A, B. Colbertiasp. A. IBIGEO-P 58a, left maxillary fragment with M2M3 in occlusal view.B. IBIGEO-P 58b, left maxillary fragment with P4M3 in occlusal view.CE. Pampahippus secundus Deraco and García-López, 2016. C. IBIGEO-P 62, right trigonid and incomplete talonid of a lower right molar in occlusal view. D. IBIGEO-P 63, fragmented left p3 or p4 in occlusal view. E. IBIGEO-P 64, left m1? in occlusal (E1), labial (E2), and lingual (E3) views. Photographs (A1E1, E3, E4), explanatory drawings (A2E2). Dashed area indicates broken or missing dental areas." figureDoi="http://doi.org/10.5281/zenodo.10626739" httpUri="https://zenodo.org/record/10626739/files/figure.png" pageId="7" pageNumber="92">Fig. 3C</figureCitation>
), and an upper fragmented cheek tooth; IBIGEO-P 63 (
<figureCitation id="13832A56FFAFFF984C27FAF9FB1AFAEC" box="[1128,1216,1370,1396]" captionStart="Fig" captionStartId="8.[151,184,1111,1133]" captionTargetBox="[156,1526,166,1080]" captionTargetId="figure-538@8.[153,1526,182,1068]" captionTargetPageId="8" captionText="Fig. 3. New fossil specimens of Typotheria and Toxodontia from the Quebrada de Los Colorados Formation exposed at Quebrada Grande locality, Los Cardones National Park (Salta Province), Casamayoran SALMA. A, B. Colbertiasp. A. IBIGEO-P 58a, left maxillary fragment with M2M3 in occlusal view.B. IBIGEO-P 58b, left maxillary fragment with P4M3 in occlusal view.CE. Pampahippus secundus Deraco and García-López, 2016. C. IBIGEO-P 62, right trigonid and incomplete talonid of a lower right molar in occlusal view. D. IBIGEO-P 63, fragmented left p3 or p4 in occlusal view. E. IBIGEO-P 64, left m1? in occlusal (E1), labial (E2), and lingual (E3) views. Photographs (A1E1, E3, E4), explanatory drawings (A2E2). Dashed area indicates broken or missing dental areas." figureDoi="http://doi.org/10.5281/zenodo.10626739" httpUri="https://zenodo.org/record/10626739/files/figure.png" pageId="7" pageNumber="92">Fig. 3D</figureCitation>
), fragmented left p3 or p4; IBIGEO-P 64 (
<figureCitation id="13832A56FFAFFF984BB4FAD9FB95FA0D" box="[1019,1103,1402,1429]" captionStart="Fig" captionStartId="8.[151,184,1111,1133]" captionTargetBox="[156,1526,166,1080]" captionTargetId="figure-538@8.[153,1526,182,1068]" captionTargetPageId="8" captionText="Fig. 3. New fossil specimens of Typotheria and Toxodontia from the Quebrada de Los Colorados Formation exposed at Quebrada Grande locality, Los Cardones National Park (Salta Province), Casamayoran SALMA. A, B. Colbertiasp. A. IBIGEO-P 58a, left maxillary fragment with M2M3 in occlusal view.B. IBIGEO-P 58b, left maxillary fragment with P4M3 in occlusal view.CE. Pampahippus secundus Deraco and García-López, 2016. C. IBIGEO-P 62, right trigonid and incomplete talonid of a lower right molar in occlusal view. D. IBIGEO-P 63, fragmented left p3 or p4 in occlusal view. E. IBIGEO-P 64, left m1? in occlusal (E1), labial (E2), and lingual (E3) views. Photographs (A1E1, E3, E4), explanatory drawings (A2E2). Dashed area indicates broken or missing dental areas." figureDoi="http://doi.org/10.5281/zenodo.10626739" httpUri="https://zenodo.org/record/10626739/files/figure.png" pageId="7" pageNumber="92">Fig. 3E</figureCitation>
), left m1?. Quebrada Grande, Los Cardones National Park, Casamayoran SALMA.
</paragraph>
<paragraph id="8B0736D3FFAFFF974B67FA64FB1AF8A2" blockId="7.[808,1481,1478,1960]" lastBlockId="8.[862,1535,1293,1850]" lastPageId="8" lastPageNumber="93" pageId="7" pageNumber="92">
<emphasis id="B9CCEAC1FFAFFF984B67FA64FC6BFA78" box="[808,945,1479,1504]" italics="true" pageId="7" pageNumber="92">Description</emphasis>
.—IBIGEO-P 6264 are brachydont cheek teeth, subrectangular in outline. Although IBIGEO-P 63 is broken, it is probable that this cheek tooth is a p3 or p4 due to the following characteristics; the trigonid and talonid are separated by a deep labial sulcus; the cristid obliqua is connected to the distal wall of the metalophid; the metalophid is straight and disto-lingually directed; the talonid is short and shows a conspicuous bunoid and isolated entoconid; the hypoconulid is located distal to the distal wall of the entoconid. IBIGEO-P 62 is a molariform cheek tooth; due to its breakage it is not possible to determine which tooth it is, but some recognizable features and the absence of others allow us to infer that it could be a lower molar; the trigonid is subrectangular in outline, being wider than long; the protolophid is almost straight; the paralophid is short, narrow, and lingually directed; the mesio-lingual cingulid is reduced and separated from the lingual region of the paralophid, but, based on the height of the cingulid on the crown, its location on the trigonid, and its proximity to the paralophid, with wear, the mesio-lingual cingulid fuses to the paralophid; a small cusp is identified between these structures; the trigonid presents a conspicuous paraconid mesial to the mesio-labial wall of the metalophid; the metalophid is straight, conspicuous, and disto-lingually directed; the cristid obliqua is united to the labial portion of the metalophid; the labial sulcus is transversally and longitudinally deep; the talonid is larger and lower than the trigonid, and subquadrangular in outline; there are no traces of labial and lingual cingulids. In IBIGEO-P 64, the protolophid is almost straight and mesio-labially directed; despite the breakage of the paralophid, it seems to be disto-lingually directed; the metalophid is straight and disto-lingually directed; the paraconid is conspicuous and connected to the mesio-labial face of the metalophid; the cristid obliqua is connected to the labial portion of the metalophid; the labial sulcus is transversally and longitudinally deep; the talonid is labially convex, and larger and slightly lower than the trigonid, but both present almost the same width; the talonid basin is shallow and wide; the lophoid entoconid is well developed and transversally expanded, connected to the hypoconid region, and separated from the hypoconulid by the a small disto-lingual valley, which is limited by a disto-lingual cingulid that connects the hypoconulid and the entoconid; the hypolophid is moderately short and curved as it slightly protrudes from the distal face of the entoconid; there are no traces of labial or lingual cingulids. To sum up, IBIGEO-P 6264 present a combination of characteristics, including their size, the mesio-distal expansion of lower cheek tooth, the identification of a small paraconid mesial to the metaconid, the presence of a well-developed entoconid being bunoid in premolars and lophoid in molars, and the absence of lingual and labial cingulids on molars, all of which allowing their identification as
<taxonomicName id="4CB84D50FFA0FF974BE0F8BCFB63F8A1" authorityName="Deraco and Garcia-Lopez" authorityYear="2016" box="[943,1209,1823,1849]" class="Mammalia" family="Notohippidae" genus="Pampahippus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="93" phylum="Chordata" rank="species" species="secundus">
<emphasis id="B9CCEAC1FFA0FF974BE0F8BCFB63F8A1" box="[943,1209,1823,1849]" italics="true" pageId="8" pageNumber="93">Pampahippus secundus</emphasis>
</taxonomicName>
.
</paragraph>
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<paragraph id="8B0736D3FFA0FF9748D8FBF4FEB6FB61" blockId="8.[151,1535,1111,1273]" pageId="8" pageNumber="93">
Fig. 3. New fossil specimens of
<taxonomicName id="4CB84D50FFA0FF97498EFBF4FDF8FBF5" authorityName="Zittel" authorityYear="1893" box="[449,546,1111,1133]" pageId="8" pageNumber="94" rank="subOrder" subOrder="Typotheria">Typotheria</taxonomicName>
and
<taxonomicName id="4CB84D50FFA0FF974A1EFBF4FD6DFBF5" authorityName="Owen" authorityYear="1853" box="[593,695,1111,1133]" pageId="8" pageNumber="94" rank="subOrder" subOrder="Toxodontia">Toxodontia</taxonomicName>
from the Quebrada de Los Colorados Formation exposed at Quebrada Grande locality, Los Cardones National Park (Salta Province), Casamayoran SALMA.
<emphasis id="B9CCEAC1FFA0FF974AA4FBD0FD21FB10" bold="true" box="[747,763,1139,1160]" pageId="8" pageNumber="93">A</emphasis>
,
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.
<taxonomicName id="4CB84D50FFA0FF974B6DFBD0FC77FB10" box="[802,941,1139,1161]" class="Magnoliopsida" family="Dilleniaceae" genus="Colbertia" kingdom="Plantae" order="Dilleniales" pageId="8" pageNumber="93" phylum="Tracheophyta" rank="species" species="undefined-A">
<emphasis id="B9CCEAC1FFA0FF974B6DFBD0FCA4FB10" box="[802,894,1139,1160]" italics="true" pageId="8" pageNumber="93">Colbertia</emphasis>
sp.
<emphasis id="B9CCEAC1FFA0FF974BD2FBD0FC77FB10" bold="true" box="[925,941,1139,1160]" pageId="8" pageNumber="93">A</emphasis>
</taxonomicName>
. IBIGEO-P 58a, left maxillary fragment with M2M3 in occlusal view.
<emphasis id="B9CCEAC1FFA0FF974884FB2CFF00FB3C" bold="true" box="[203,218,1167,1188]" pageId="8" pageNumber="93">B</emphasis>
. IBIGEO-P 58b, left maxillary fragment with P4M3 in occlusal view.
<emphasis id="B9CCEAC1FFA0FF974B1DFB2CFCA7FB3C" bold="true" box="[850,893,1167,1189]" pageId="8" pageNumber="93">CE</emphasis>
.
<taxonomicName id="4CB84D50FFA0FF974BC8FB2CFAA4FB3D" authority="Deraco and Garcia-Lopez, 2016" authorityName="Deraco and Garcia-Lopez" authorityYear="2016" box="[903,1406,1167,1189]" class="Mammalia" family="Notohippidae" genus="Pampahippus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="93" phylum="Chordata" rank="species" species="secundus">
<emphasis id="B9CCEAC1FFA0FF974BC8FB2CFB83FB3C" box="[903,1113,1167,1188]" italics="true" pageId="8" pageNumber="93">Pampahippus secundus</emphasis>
<bibRefCitation id="EF294B22FFA0FF974C11FB33FAA4FB3D" author="Deraco, V. &amp; Garcia-Lopez, D. A." box="[1118,1406,1167,1189]" pageId="8" pageNumber="93" pagination="1037884" refId="ref11377" refString="Deraco, V. and Garcia-Lopez, D. A. 2016. A new Eocene Toxodontia (Mammalia, Notoungulata) from northwestern Argentina. Journal of Vertebrate Paleontology 36: e 1037884." type="journal article" year="2016">Deraco and García-López, 2016</bibRefCitation>
</taxonomicName>
.
<emphasis id="B9CCEAC1FFA0FF974DCAFB2CFA4FFB3D" bold="true" box="[1413,1429,1167,1189]" pageId="8" pageNumber="93">C</emphasis>
. IBIGEO-P 62, right trigonid and incomplete talonid of a lower right molar in occlusal view.
<emphasis id="B9CCEAC1FFA0FF974B24FB08FCA1FB58" bold="true" box="[875,891,1195,1216]" pageId="8" pageNumber="93">D</emphasis>
. IBIGEO-P 63, fragmented left p3 or p4 in occlusal view.
<emphasis id="B9CCEAC1FFA0FF974DCAFB08FA4EFB58" bold="true" box="[1413,1428,1195,1216]" pageId="8" pageNumber="93">E</emphasis>
. IBIGEO-P 64, left m1? in occlusal (E
<subScript id="173C3496FFA0FF9749CDFB72FE50FB47" attach="left" box="[386,394,1233,1247]" fontSize="6" pageId="8" pageNumber="93">1</subScript>
), labial (E
<subScript id="173C3496FFA0FF9749A6FB72FE2BFB47" attach="left" box="[489,497,1233,1247]" fontSize="6" pageId="8" pageNumber="93">2</subScript>
), and lingual (E
<subScript id="173C3496FFA0FF974ACEFB72FD53FB47" attach="left" box="[641,649,1233,1247]" fontSize="6" pageId="8" pageNumber="93">3</subScript>
) views. Photographs (A
<subScript id="173C3496FFA0FF974B2DFB72FCB0FB47" attach="left" box="[866,874,1233,1247]" fontSize="6" pageId="8" pageNumber="93">1</subScript>
E
<subScript id="173C3496FFA0FF974BCCFB72FC51FB47" attach="left" box="[899,907,1233,1247]" fontSize="6" pageId="8" pageNumber="93">1</subScript>
, E
<subScript id="173C3496FFA0FF974BEBFB72FC76FB47" attach="left" box="[932,940,1233,1247]" fontSize="6" pageId="8" pageNumber="93">3</subScript>
, E
<subScript id="173C3496FFA0FF974B8BFB72FC16FB47" attach="left" box="[964,972,1233,1247]" fontSize="6" pageId="8" pageNumber="93">4</subScript>
), explanatory drawings (A
<subScript id="173C3496FFA0FF974CF2FB72FB1FFB47" attach="left" box="[1213,1221,1233,1247]" fontSize="6" pageId="8" pageNumber="93">2</subScript>
E
<subScript id="173C3496FFA0FF974C91FB72FB3CFB47" attach="right" box="[1246,1254,1233,1247]" fontSize="6" pageId="8" pageNumber="93">2</subScript>
). Dashed area indicates broken or missing dental areas.
</paragraph>
</caption>
<paragraph id="8B0736D3FFA0FF974B11F8E8FA14F83F" blockId="8.[862,1534,1867,1960]" pageId="8" pageNumber="93">
<emphasis id="B9CCEAC1FFA0FF974B11F8E8FAF5F8FD" box="[862,1327,1867,1893]" italics="true" pageId="8" pageNumber="93">Stratigraphic and geographic range.—</emphasis>
Lower and upper sections of the Lumbrera and Quebrada de Los Colorados Formations, middle Eocene;
<collectingRegion id="497CF831FFA0FF974CECF82EFA93F83F" box="[1187,1353,1933,1959]" country="Argentina" name="Salta" pageId="8" pageNumber="93">Salta Province</collectingRegion>
,
<collectingCountry id="F3AF7643FFA0FF974D1AF82EFA11F83F" box="[1365,1483,1933,1959]" name="Argentina" pageId="8" pageNumber="93">Argentina</collectingCountry>
.
</paragraph>
<paragraph id="8B0736D3FFA1FF96482DFF3DFEDAFE24" blockId="9.[98,771,158,444]" pageId="9" pageNumber="94">
<emphasis id="B9CCEAC1FFA1FF96482DFF3DFF1EFF20" box="[98,196,158,184]" italics="true" pageId="9" pageNumber="94">Remarks</emphasis>
.—
<taxonomicName id="4CB84D50FFA1FF9648A8FF3DFE2DFF20" authorityName="Deraco and Garcia-Lopez" authorityYear="2016" box="[231,503,158,184]" class="Mammalia" family="Notohippidae" genus="Pampahippus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="94" phylum="Chordata" rank="species" species="secundus">
<emphasis id="B9CCEAC1FFA1FF9648A8FF3DFE2DFF20" box="[231,503,158,184]" italics="true" pageId="9" pageNumber="94">Pampahippus secundus</emphasis>
</taxonomicName>
has been identified in the lower (
<bibRefCitation id="EF294B22FFA1FF964891FF1CFD90FF41" author="Deraco, V. &amp; Garcia-Lopez, D. A." box="[222,586,190,217]" pageId="9" pageNumber="94" pagination="1037884" refId="ref11377" refString="Deraco, V. and Garcia-Lopez, D. A. 2016. A new Eocene Toxodontia (Mammalia, Notoungulata) from northwestern Argentina. Journal of Vertebrate Paleontology 36: e 1037884." type="journal article" year="2016">Deraco and García-López 2016</bibRefCitation>
) and upper levels (
<bibRefCitation id="EF294B22FFA1FF9648DDFF7CFE6EFF61" author="Garcia-Lopez, D. A. &amp; Deraco, V. &amp; Rougier, G. W. &amp; Del Papa, C. E. &amp; Babot, M. J. &amp; Bertelli, S. &amp; Herrera, C. M. &amp; Giannini, N. P." box="[146,436,223,250]" pageId="9" pageNumber="94" pagination="1582537" refId="ref11735" refString="Garcia-Lopez, D. A., Deraco, V., Rougier, G. W., Del Papa, C. E., Babot, M. J., Bertelli, S., Herrera, C. M., and Giannini, N. P. 2019. New record of Pampahippus secundus (Mammalia, Notoungulata) from the Upper Lumbrera Formation, Eocene of Northwestern Argentina. Journal of Vertebrate Paleontology 39: e 1582537." type="journal article" year="2019">García-López et al. 2019</bibRefCitation>
) of the Lumbrera Fm., at El Simbolar (
<collectingRegion id="497CF831FFA1FF964894FF5CFECCFE82" box="[219,278,255,282]" country="Argentina" name="Salta" pageId="9" pageNumber="94">Salta</collectingRegion>
). In addition,
<taxonomicName id="4CB84D50FFA1FF9649F1FF5CFD83FE81" authorityName="Bond and Lopez" authorityYear="1993" box="[446,601,255,281]" class="Mammalia" family="Notohippidae" genus="Pampahippus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="94" phylum="Chordata" rank="genus">
<emphasis id="B9CCEAC1FFA1FF9649F1FF5CFD83FE81" box="[446,601,255,281]" italics="true" pageId="9" pageNumber="94">Pampahippus</emphasis>
</taxonomicName>
has also been recognized in the lower section of the Lumbrera Fm. at Pampa Grande,
<collectingRegion id="497CF831FFA1FF964957FEE3FE8AFEC2" box="[280,336,320,346]" country="Argentina" name="Salta" pageId="9" pageNumber="94">Salta</collectingRegion>
(Bond and López 1993), Quebrada de Los Colorados Fm. at La Poma,
<collectingRegion id="497CF831FFA1FF964994FEC2FDC9FEE3" box="[475,531,353,379]" country="Argentina" name="Salta" pageId="9" pageNumber="94">Salta</collectingRegion>
(
<bibRefCitation id="EF294B22FFA1FF964A6BFEC2FF40FE03" author="Garcia-Lopez, D. A. &amp; Deraco, V. &amp; Del Papa, C. E." pageId="9" pageNumber="94" pagination="507 - 517" refId="ref11679" refString="Garcia-Lopez, D. A., Deraco, V., and Del Papa, C. E. 2018. Fossil mammals of the Quebrada de Los Colorados Formation (late middle Eocene) at the locality of La Poma, Salta Province, Argentina. Historical Biology 30: 507 - 517." type="journal article" year="2018">García-López et al. 2018</bibRefCitation>
), and Geste Fm. at
<collectingRegion id="497CF831FFA1FF964922FE22FE2CFE03" box="[365,502,385,411]" country="Chile" name="Antofagasta" pageId="9" pageNumber="94">Antofagasta</collectingRegion>
de la Sierra,
<collectingRegion id="497CF831FFA1FF964AC6FE22FCD9FE03" box="[649,771,385,411]" country="Argentina" name="Catamarca" pageId="9" pageNumber="94">Catamarca</collectingRegion>
<bibRefCitation id="EF294B22FFA1FF964826FE02FF2EFE24" author="Lopez, G. M." box="[105,244,417,444]" pageId="9" pageNumber="94" pagination="61 - 81" refId="ref12349" refString="Lopez, G. M. 1997. Palaeogene faunal assemblage from Antofagasta de la Sierra (Catamarca Province, Argentina). Paleovertebrata 26: 61 - 81." type="journal article" year="1997">López 1997</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>