324 lines
32 KiB
XML
324 lines
32 KiB
XML
<document id="4F952EAB5A61CCC1BFA9E6BAC16CC54D" ID-DOI="10.4202/app.00784.2020" ID-ISSN="1732-2421" ID-Zenodo-Dep="10626735" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1707250644161" checkinUser="felipe" docAuthor="Fernández, Mercedes, Zimicz, Ana N., Bond, Mariano, Chornogubsky, Laura, Arnal, Michelle, Cárdenas, Magalí & Fernicola, Juan C." docDate="2021" docId="031187C5FFACFF9A4B11F8B9FDC2F83F" docLanguage="en" docName="ActaPalaeontolPol.66.1.85-97.pdf" docOrigin="Acta Palaeontologica Polonica 66 (1)" docSource="http://dx.doi.org/10.4202/app.00784.2020" docStyle="DocumentStyle:E4780F6D34569985EDF9CACFB9BC780E.4:ActaPalaeontolPol.2015-.journal_article" docStyleId="E4780F6D34569985EDF9CACFB9BC780E" docStyleName="ActaPalaeontolPol.2015-.journal_article" docStyleVersion="4" docTitle="Colbertia lumbrerense Bond 1981" docType="treatment" docVersion="4" lastPageNumber="90" masterDocId="FF28FFBDFFA8FF9F484FFFA3FFDAFF98" masterDocTitle="New Eocene South American native ungulates from the Quebrada de los Colorados Formation at Los Cardones National Park, Argentina" masterLastPageNumber="97" masterPageNumber="85" pageNumber="89" updateTime="1714484150498" updateUser="karina" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3EE7C114500B1474AE91BCFB26404A8D">New Eocene South American native ungulates from the Quebrada de los Colorados Formation at Los Cardones National Park, Argentina</mods:title>
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<mods:namePart id="508271B835F233755348836A473353A7">Fernández, Mercedes</mods:namePart>
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<mods:namePart id="DC8BC4F3437A70929650BBA8F33665E1">Zimicz, Ana N.</mods:namePart>
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<mods:namePart id="EF132E40DA2E9E189788422A3060E822">Bond, Mariano</mods:namePart>
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<mods:namePart id="D02C358C8FD70C45C024B2F3DBD8DEBA">Chornogubsky, Laura</mods:namePart>
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<mods:namePart id="F7A6596E69C643D1F803FCE60FBD8C02">Arnal, Michelle</mods:namePart>
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<mods:namePart id="65C7F257851B296C900F0617C61409FE">Cárdenas, Magalí</mods:namePart>
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<mods:namePart id="E10596C2AC0F0A436192E12038AF5D02">Fernicola, Juan C.</mods:namePart>
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<mods:title id="98C1806A58411180A291136F7E167F4F">Acta Palaeontologica Polonica</mods:title>
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<mods:date id="1EF0F5D31C80121F8742699E402CCF3D">2021</mods:date>
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<mods:number id="ACFE7278A200A6CAEF266C9BE375E5B8">2021-02-26</mods:number>
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<mods:url id="63969EC1CEE594078B3AE28B2234C8AC">http://dx.doi.org/10.4202/app.00784.2020</mods:url>
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<mods:identifier id="EEF4D22DB824FA8075C802F8CAD43FC2" type="DOI">10.4202/app.00784.2020</mods:identifier>
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<mods:identifier id="9CD42650FCD38FB5BF275B4516BC8DB8" type="ISSN">1732-2421</mods:identifier>
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<treatment id="031187C5FFACFF9A4B11F8B9FDC2F83F" ID-DOI="http://doi.org/10.5281/zenodo.10626743" ID-Zenodo-Dep="10626743" LSID="urn:lsid:plazi:treatment:031187C5FFACFF9A4B11F8B9FDC2F83F" httpUri="http://treatment.plazi.org/id/031187C5FFACFF9A4B11F8B9FDC2F83F" lastPageId="5" lastPageNumber="90" pageId="4" pageNumber="89">
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<subSubSection id="C3A26558FFACFF9B4B11F8B9FAC1F8AF" box="[862,1307,1818,1847]" pageId="4" pageNumber="89" type="nomenclature">
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<paragraph id="8B0736D3FFACFF9B4B11F8B9FAC1F8AF" blockId="4.[862,1307,1818,1847]" box="[862,1307,1818,1847]" pageId="4" pageNumber="89">
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<heading id="D04F81BFFFACFF9B4B11F8B9FAC1F8AF" box="[862,1307,1818,1847]" fontSize="12" level="2" pageId="4" pageNumber="89" reason="2">
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<taxonomicName id="4CB84D50FFACFF9B4B11F8B9FAC1F8AF" authority="Bond, 1981" authorityName="Bond" authorityYear="1981" box="[862,1307,1818,1847]" class="Magnoliopsida" family="Dilleniaceae" genus="Colbertia" kingdom="Plantae" order="Dilleniales" pageId="4" pageNumber="89" phylum="Tracheophyta" rank="species" species="lumbrerense">
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<emphasis id="B9CCEAC1FFACFF9B4B11F8B9FBA4F8AF" box="[862,1150,1818,1847]" italics="true" pageId="4" pageNumber="89">Colbertia lumbrerense</emphasis>
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Bond, 1981
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3A26558FFACFF9A4B11F8E4FDC2F83F" lastPageId="5" lastPageNumber="90" pageId="4" pageNumber="89" type="description">
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<paragraph id="8B0736D3FFACFF9B4B11F8E4FC13F8C5" blockId="4.[862,969,1863,1885]" box="[862,969,1863,1885]" pageId="4" pageNumber="89">
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<figureCitation id="13832A56FFACFF9B4B11F8E4FC1FF8C5" box="[862,965,1863,1885]" captionStart="Fig" captionStartId="5.[807,840,1851,1873]" captionTargetId="figure-10@6.[153,1517,173,1820]" captionTargetPageId="6" captionText="Fig. 2. New fossil specimens of Typotheria from the Quebrada de Los Colorados Fm. exposed at Quebrada Grande locality, Los Cardones Natio- nal Park (Salta Province), Casamayoran SALMA, Eocene. A, B. Colbertia lumbrerense Bond, 1981. A. IBIGEO-P 45, right maxillary fragment with →" pageId="4" pageNumber="89">Fig. 2A, B</figureCitation>
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.
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</paragraph>
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<paragraph id="8B0736D3FFACFF9B4B11F8D7FA5DF83E" blockId="4.[862,1535,1908,1959]" pageId="4" pageNumber="89">
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<materialsCitation id="3BD03C8EFFACFF9B4B11F8D7FA5DF83E" collectionCode="C" pageId="4" pageNumber="89" specimenCode="PVL 4607, P1, P4, M1, M3" specimenCount="1" typeStatus="holotype">
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<emphasis id="B9CCEAC1FFACFF9B4B11F8D7FC6CF811" box="[862,950,1908,1929]" italics="true" pageId="4" pageNumber="89">
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<typeStatus id="54038871FFACFF9B4B11F8D7FC6CF811" box="[862,950,1908,1929]" pageId="4" pageNumber="89">Holotype</typeStatus>
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</emphasis>
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:
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<specimenCode id="DB1E9EA8FFACFF9B4B8BF8D7FBF1F812" box="[964,1067,1908,1930]" collectionCode="PVL" country="Argentina" httpUri="http://grbio.org/cool/pbkd-hmeb" name="Paleontologia de Vertebrados Lillo" pageId="4" pageNumber="89">PVL 4607</specimenCode>
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, incomplete skull with right
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<collectionCode id="EDA9AE16FFACFF9B4D06F8D7FA81F811" box="[1353,1371,1908,1929]" country="Denmark" name="University of Copenhagen" pageId="4" pageNumber="89" type="Herbarium">C</collectionCode>
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,
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<specimenCode id="DB1E9EA8FFACFF9B4D28F8D7FAA4F811" box="[1383,1406,1908,1929]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="4" pageNumber="89" type="Herbarium">P1</specimenCode>
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–
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<specimenCode id="DB1E9EA8FFACFF9B4DC3F8D7FA72F811" box="[1420,1448,1908,1929]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="4" pageNumber="89" type="Herbarium">P4</specimenCode>
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, and left
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<specimenCode id="DB1E9EA8FFACFF9B4B11F832FCA4F83E" box="[862,894,1937,1958]" collectionCode="M" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="4" pageNumber="89" type="Herbarium">M1</specimenCode>
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–
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<specimenCode id="DB1E9EA8FFACFF9B4BC3F832FC75F83E" box="[908,943,1937,1958]" collectionCode="M" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="4" pageNumber="89" type="Herbarium">M3</specimenCode>
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; and mandible with left i2–m3, and right p2–m3.
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</materialsCitation>
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</paragraph>
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<paragraph id="8B0736D3FFADFF9A482DFF03FD0DFF2D" blockId="5.[98,727,159,181]" box="[98,727,159,181]" pageId="5" pageNumber="90">
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<emphasis id="B9CCEAC1FFADFF9A482DFF03FF06FF2C" box="[98,220,159,181]" italics="true" pageId="5" pageNumber="90">Type locality</emphasis>
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: Estancia Pampa Grande,
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<collectingRegion id="497CF831FFADFF9A4995FF3CFDB2FF2D" box="[474,616,159,181]" country="Argentina" name="Salta" pageId="5" pageNumber="90">Salta Province</collectingRegion>
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,
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<collectingCountry id="F3AF7643FFADFF9A4A3FFF3CFD09FF2D" box="[624,723,159,181]" name="Argentina" pageId="5" pageNumber="90">Argentina</collectingCountry>
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.
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</paragraph>
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<paragraph id="8B0736D3FFADFF9A482DFF61FF4FFF6B" blockId="5.[98,771,193,243]" pageId="5" pageNumber="90">
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<emphasis id="B9CCEAC1FFADFF9A482DFF61FF04FF4E" box="[98,222,193,215]" italics="true" pageId="5" pageNumber="90">Type horizon</emphasis>
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: Lower sections of the Lumbrera Formation, middle Eocene.
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</paragraph>
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<paragraph id="8B0736D3FFADFF9A482DFEA5FE14FE7D" blockId="5.[98,771,262,485]" pageId="5" pageNumber="90">
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<emphasis id="B9CCEAC1FFADFF9A482DFEA5FF1EFEB8" box="[98,196,262,288]" italics="true" pageId="5" pageNumber="90">Material</emphasis>
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.—IBIGEO-P 45 (
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<figureCitation id="13832A56FFADFF9A49D5FEA5FE2CFEB8" box="[410,502,262,288]" captionStart="Fig" captionStartId="5.[807,840,1851,1873]" captionTargetId="figure-10@6.[153,1517,173,1820]" captionTargetPageId="6" captionText="Fig. 2. New fossil specimens of Typotheria from the Quebrada de Los Colorados Fm. exposed at Quebrada Grande locality, Los Cardones Natio- nal Park (Salta Province), Casamayoran SALMA, Eocene. A, B. Colbertia lumbrerense Bond, 1981. A. IBIGEO-P 45, right maxillary fragment with →" pageId="5" pageNumber="90">Fig. 2A</figureCitation>
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), right maxillary fragment with P4 and M3 from the same individual (both teeth were erroneously glued together, but due to their state of preservation they cannot be separated without breaking the material); and IBIGEO-P 56 (
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<figureCitation id="13832A56FFADFF9A4987FE2AFDFEFE3B" box="[456,548,393,419]" captionStart="Fig" captionStartId="5.[807,840,1851,1873]" captionTargetId="figure-10@6.[153,1517,173,1820]" captionTargetPageId="6" captionText="Fig. 2. New fossil specimens of Typotheria from the Quebrada de Los Colorados Fm. exposed at Quebrada Grande locality, Los Cardones Natio- nal Park (Salta Province), Casamayoran SALMA, Eocene. A, B. Colbertia lumbrerense Bond, 1981. A. IBIGEO-P 45, right maxillary fragment with →" pageId="5" pageNumber="90">Fig. 2B</figureCitation>
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), broken right M1 or M2. Quebrada Grande, Los Cardones National Park, Casamayoran SALMA, Eocene.
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</paragraph>
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<paragraph id="8B0736D3FFADFF9A482DFE54FF2DF9F3" blockId="5.[98,771,502,1643]" pageId="5" pageNumber="90">
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<emphasis id="B9CCEAC1FFADFF9A482DFE54FF33FD88" box="[98,233,503,528]" italics="true" pageId="5" pageNumber="90">Description</emphasis>
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.—IBIGEO-P 45 and IBIGEO-P 56 were not found associated; nevertheless, according to their similar structures and size, both are considered here as belonging to the same species. IBIGEO-P 45 presents poorly preserved P4 and M3; both teeth are brachydont and subquadrangular in outline. Unfortunately, not much can be mentioned from the former tooth since only its labial portion, including a broken ectoloph, is preserved. The M3 is less worn than the P4, and preserves its lingual portion; the protoloph is well developed and runs disto-lingually; there is a low mesial cingulum, which is lingually interrupted, whereas the distal cingulum continues more lingually; a reduced hypocone is identified in the metaloph. IBIGEO-P 56 is a low-crowned M1 or M2, subquadrangular in outline; it is also broken and lacks its ectoloph, but it is little worn and the remaining structures are well preserved; the mesio-lingual cingulum is located lower on the crown and well developed, and it is lingually interrupted; there is a reduced, but continuous lingual cingulum that surrounds the lingual region of the protoloph; both protoloph and metaloph are well developed, but the former is wider and more oblique than the latter; the protoloph is curved and runs disto-lingually, whereas the metaloph runs mesio-lingually; the lingual end of the metaloph (hypocone region) unites with the lingual end of the protoloph (protocone region), isolating a central fossette; the central fossette is narrow and oblique, disto-lingually directed; a vestigial lingual sulcus persists to the connection of the protoloph and metaloph; the crochet is short and runs mesio-labially from the metaloph to the ectoloph, isolating the disto-labial fossete, which is shallow in this specimen; the disto-lingual cingulum is higher than the mesio-lingual cingulum, and it is also lingually interrupted. All of these characteristics plus their size (see
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<tableCitation id="C63A0368FFADFF9A4A76F9ACFD56F9B1" box="[569,652,1551,1577]" captionStart="Table 1" captionStartId="4.[862,915,159,181]" captionTargetId="graphics-708@4.[862,1535,257,1083]" captionTargetPageId="4" captionText="Table 1. Measurements (in mm) of the dental specimens. Abbreviations: *, approximate measurements due to the brakeage of the specimen; LLL, labio-lingual length; MDL, mesio-distal length." pageId="5" pageNumber="90">Table 1</tableCitation>
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) allow us to identify IBIGEO-P 45 and IBIGEO-P 56 as
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<emphasis id="B9CCEAC1FFADFF9A4AD9F993FF2DF9F3" italics="true" pageId="5" pageNumber="90">
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<taxonomicName id="4CB84D50FFADFF9A4AD9F993FF29F9F3" authorityName="Bond" authorityYear="1981" class="Magnoliopsida" family="Dilleniaceae" genus="Colbertia" kingdom="Plantae" order="Dilleniales" pageId="5" pageNumber="90" phylum="Tracheophyta" rank="species" species="lumbrerense">Colbertia lumbrerense</taxonomicName>
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.
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</emphasis>
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</paragraph>
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<paragraph id="8B0736D3FFADFF9A482DF9DFFF28F8A3" blockId="5.[98,770,1660,1851]" pageId="5" pageNumber="90">
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<emphasis id="B9CCEAC1FFADFF9A482DF9DFFF1EF90E" box="[98,196,1660,1686]" italics="true" pageId="5" pageNumber="90">Remarks</emphasis>
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.—
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<taxonomicName id="4CB84D50FFADFF9A48A7F9DFFE35F90E" authorityName="Bond" authorityYear="1981" box="[232,495,1660,1686]" class="Magnoliopsida" family="Dilleniaceae" genus="Colbertia" kingdom="Plantae" order="Dilleniales" pageId="5" pageNumber="90" phylum="Tracheophyta" rank="species" species="lumbrerense">
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<emphasis id="B9CCEAC1FFADFF9A48A7F9DFFE35F90E" box="[232,495,1660,1686]" italics="true" pageId="5" pageNumber="90">Colbertia lumbrerense</emphasis>
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</taxonomicName>
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was previously known only for the lower Lumbrera Fm. at Pampa Grande, Salta Bond 1981). In addition,
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<taxonomicName id="4CB84D50FFADFF9A49CCF91DFDCCF940" box="[387,534,1726,1752]" class="Magnoliopsida" family="Dilleniaceae" genus="Colbertia" kingdom="Plantae" order="Dilleniales" pageId="5" pageNumber="90" phylum="Tracheophyta" rank="species" species="undetermined">
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<emphasis id="B9CCEAC1FFADFF9A49CCF91DFE2AF940" box="[387,496,1726,1752]" italics="true" pageId="5" pageNumber="90">Colbertia</emphasis>
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sp.
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</taxonomicName>
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has also been recognized in Geste Fm., Antofagasta de la Sierra (
|
||
<bibRefCitation id="EF294B22FFADFF9A4A24F97CFD2CF961" author="Lopez, G. M." box="[619,758,1759,1785]" pageId="5" pageNumber="90" pagination="61 - 81" refId="ref12349" refString="Lopez, G. M. 1997. Palaeogene faunal assemblage from Antofagasta de la Sierra (Catamarca Province, Argentina). Paleovertebrata 26: 61 - 81." type="journal article" year="1997">López 1997</bibRefCitation>
|
||
), and in the early Eocene of
|
||
<collectingCountry id="F3AF7643FFADFF9A49C9F95CFE16F881" box="[390,460,1791,1817]" name="Brazil" pageId="5" pageNumber="90">Brazil</collectingCountry>
|
||
(Itaboraian SALMA;
|
||
<bibRefCitation id="EF294B22FFADFF9A4A8BF95CFF3CF8A3" author="Paula Couto" pageId="5" pageNumber="90" pagination="1 - 16" refId="ref12690" refString="Paula Couto de, C. 1952. Fossil mammals from the beginning of the Cenozoic in Brazil. Notoungulata. American Museum Novitates 1568: 1 - 16." type="journal article" year="1952">Paula Couto 1952</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B0736D3FFADFF994B68F898FA79F83E" blockId="5.[807,1517,1851,1957]" lastBlockId="6.[151,1535,1853,1961]" lastPageId="6" lastPageNumber="91" pageId="5" pageNumber="90">
|
||
Fig. 2. New fossil specimens of
|
||
<taxonomicName id="4CB84D50FFADFF9A4C2EF898FB19F8C9" authorityName="Zittel" authorityYear="1893" box="[1121,1219,1851,1873]" pageId="5" pageNumber="94" rank="subOrder" subOrder="Typotheria">Typotheria</taxonomicName>
|
||
from the Quebrada de Los Colorados Fm. exposed at Quebrada Grande locality, Los Cardones National Park (Salta Province), Casamayoran SALMA, Eocene.
|
||
<emphasis id="B9CCEAC1FFADFF9A4D72F8D1FA97F81F" bold="true" box="[1341,1357,1906,1927]" pageId="5" pageNumber="90">A</emphasis>
|
||
,
|
||
<emphasis id="B9CCEAC1FFADFF9A4D17F8D0FABDF810" bold="true" box="[1368,1383,1907,1928]" pageId="5" pageNumber="90">B</emphasis>
|
||
.
|
||
<taxonomicName id="4CB84D50FFADFF9A4D3DF8D0FBD1F83C" authority="Bond, 1981" authorityName="Bond" authorityYear="1981" class="Magnoliopsida" family="Dilleniaceae" genus="Colbertia" kingdom="Plantae" order="Dilleniales" pageId="5" pageNumber="90" phylum="Tracheophyta" rank="species" species="lumbrerense">
|
||
<emphasis id="B9CCEAC1FFADFF9A4D3DF8D0FC42F83C" italics="true" pageId="5" pageNumber="90">Colbertia lumbrerense</emphasis>
|
||
Bond, 1981
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B9CCEAC1FFADFF9A4C5AF82DFBFFF83B" bold="true" box="[1045,1061,1934,1955]" pageId="5" pageNumber="90">A</emphasis>
|
||
. IBIGEO-P 45, right maxillary fragment with
|
||
<materialsCitation id="3BD03C8EFFAEFF9948D8F89EFF0CF8CB" box="[151,214,1853,1875]" collectionCode="P" pageId="6" pageNumber="91" specimenCode="P4" specimenCount="1">
|
||
<specimenCode id="DB1E9EA8FFAEFF9948D8F89EFF75F8CA" box="[151,175,1853,1874]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="6" pageNumber="91" type="Herbarium">P4</specimenCode>
|
||
and
|
||
</materialsCitation>
|
||
M
|
||
<quantity id="4C409B36FFAEFF9948BDF89EFECEF8CB" box="[242,276,1853,1875]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="91" unit="in" value="3.0">3 in</quantity>
|
||
occlusal view.
|
||
<materialsCitation id="3BD03C8EFFAEFF9949EEF89EFDE1F8CB" box="[417,571,1853,1875]" collectionCode="B" pageId="6" pageNumber="91" specimenCode="IBIGEO-P 56" specimenCount="1">
|
||
<emphasis id="B9CCEAC1FFAEFF9949EEF89EFE6AF8CA" bold="true" box="[417,432,1853,1874]" pageId="6" pageNumber="91">
|
||
<collectionCode id="EDA9AE16FFAEFF9949EEF89EFE6AF8CA" box="[417,432,1853,1874]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="6" pageNumber="91" type="Herbarium">B</collectionCode>
|
||
</emphasis>
|
||
.
|
||
<specimenCode id="DB1E9EA8FFAEFF9949F3F89EFDE1F8CB" box="[444,571,1853,1875]" collectionCode="IBIGEO-P" pageId="6" pageNumber="91">IBIGEO-P 56</specimenCode>
|
||
</materialsCitation>
|
||
, broken right
|
||
<materialsCitation id="3BD03C8EFFAEFF994AF4F89EFD01F8CA" box="[699,731,1853,1874]" collectionCode="M" pageId="6" pageNumber="91" specimenCode="M1" specimenCount="1">
|
||
<specimenCode id="DB1E9EA8FFAEFF994AF4F89EFD01F8CA" box="[699,731,1853,1874]" collectionCode="M" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="6" pageNumber="91" type="Herbarium">M1</specimenCode>
|
||
</materialsCitation>
|
||
or M
|
||
<quantity id="4C409B36FFAEFF994B40F89EFCE8F8CB" box="[783,818,1853,1875]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="6" pageNumber="91" unit="in" value="2.0">2 in</quantity>
|
||
occlusal view.
|
||
<emphasis id="B9CCEAC1FFAEFF994BF0F89EFC32F8CA" bold="true" box="[959,1000,1853,1875]" pageId="6" pageNumber="91">C–F</emphasis>
|
||
.
|
||
<taxonomicName id="4CB84D50FFAEFF994BBBF89EFBA1F8CA" authority="Fernández & Zimicz & Bond & Chornogubsky & Arnal & Cárdenas & Fernicola, 2021" authorityName="Fernández & Zimicz & Bond & Chornogubsky & Arnal & Cárdenas & Fernicola" authorityYear="2021" box="[1012,1147,1853,1874]" class="Magnoliopsida" family="Dilleniaceae" genus="Colbertia" kingdom="Plantae" order="Dilleniales" pageId="6" pageNumber="91" phylum="Tracheophyta" rank="species" species="falui" status="sp. nov.">
|
||
<emphasis id="B9CCEAC1FFAEFF994BBBF89EFBA1F8CA" box="[1012,1147,1853,1874]" italics="true" pageId="6" pageNumber="91">Colbertia falui</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2FF57BAFFAEFF994CCEF89EFB13F8CA" box="[1153,1225,1853,1874]" pageId="6" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B9CCEAC1FFAEFF994C81F89EFB04F8CB" bold="true" box="[1230,1246,1853,1875]" pageId="6" pageNumber="91">C</emphasis>
|
||
.
|
||
<materialsCitation id="3BD03C8EFFAEFF994CA5F89EFAB3F8CB" box="[1258,1385,1853,1875]" pageId="6" pageNumber="91" specimenCode="IBIGEO-P 67" specimenCount="1">
|
||
<specimenCode id="DB1E9EA8FFAEFF994CA5F89EFAB3F8CB" box="[1258,1385,1853,1875]" collectionCode="IBIGEO-P" pageId="6" pageNumber="91">IBIGEO-P 67</specimenCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3BD03C8EFFAEFF994D3CF89EFC77F8F6" collectionCode="C" pageId="6" pageNumber="91" specimenCode="C1, C3, C4" specimenCount="1" typeStatus="holotype">
|
||
left mandibular fragment with m2–m3 (
|
||
<typeStatus id="54038871FFAEFF994923F8FAFE64F8F7" box="[364,446,1881,1903]" pageId="6" pageNumber="91">holotype</typeStatus>
|
||
) in occlusal (
|
||
<specimenCode id="DB1E9EA8FFAEFF994A79F8FAFD94F8E9" box="[566,590,1881,1905]" collectionCode="C" country="Denmark" name="University of Copenhagen" pageId="6" pageNumber="91" type="Herbarium">
|
||
C
|
||
<subScript id="173C3496FFAEFF994A09F8C0FD94F8E9" attach="left" box="[582,590,1891,1905]" fontSize="6" pageId="6" pageNumber="91">1</subScript>
|
||
</specimenCode>
|
||
), labial (
|
||
<specimenCode id="DB1E9EA8FFAEFF994AD0F8FAFD6CF8E9" box="[671,694,1881,1905]" collectionCode="C" country="Denmark" name="University of Copenhagen" pageId="6" pageNumber="91" type="Herbarium">
|
||
C
|
||
<subScript id="173C3496FFAEFF994AE1F8C0FD6CF8E9" attach="left" box="[686,694,1891,1905]" fontSize="6" pageId="6" pageNumber="91">3</subScript>
|
||
</specimenCode>
|
||
), and lingual (
|
||
<specimenCode id="DB1E9EA8FFAEFF994B76F8FAFC8BF8E9" box="[825,849,1881,1905]" collectionCode="C" country="Denmark" name="University of Copenhagen" pageId="6" pageNumber="91" type="Herbarium">
|
||
C
|
||
<subScript id="173C3496FFAEFF994B06F8C0FC8BF8E9" attach="left" box="[841,849,1891,1905]" fontSize="6" pageId="6" pageNumber="91">4</subScript>
|
||
</specimenCode>
|
||
) views.
|
||
<emphasis id="B9CCEAC1FFAEFF994BD2F8FAFC77F8F6" bold="true" box="[925,941,1881,1902]" pageId="6" pageNumber="91">D</emphasis>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3BD03C8EFFAEFF994BF6F8FAFBEDF8F7" box="[953,1079,1881,1903]" pageId="6" pageNumber="91" specimenCode="IBIGEO-P 55" specimenCount="1" typeStatus="holotype">
|
||
<specimenCode id="DB1E9EA8FFAEFF994BF6F8FAFBEDF8F7" box="[953,1079,1881,1903]" collectionCode="IBIGEO-P" pageId="6" pageNumber="91">IBIGEO-P 55</specimenCode>
|
||
</materialsCitation>
|
||
, right mandibular fragment with p3–p
|
||
<quantity id="4C409B36FFAEFF994DC2F8FAFA6AF8F7" box="[1421,1456,1881,1903]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="6" pageNumber="91" unit="in" value="4.0">4 in</quantity>
|
||
occlusal view.
|
||
<materialsCitation id="3BD03C8EFFAEFF994882F8D6FEBDF813" box="[205,359,1909,1931]" collectionCode="E" pageId="6" pageNumber="91" specimenCode="IBIGEO-P 59" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B9CCEAC1FFAEFF994882F8D6FF06F812" bold="true" box="[205,220,1909,1930]" pageId="6" pageNumber="91">
|
||
<collectionCode id="EDA9AE16FFAEFF994882F8D6FF06F812" box="[205,220,1909,1930]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="6" pageNumber="91" type="Herbarium">E</collectionCode>
|
||
</emphasis>
|
||
.
|
||
<specimenCode id="DB1E9EA8FFAEFF9948A7F8D6FEBDF813" box="[232,359,1909,1931]" collectionCode="IBIGEO-P" pageId="6" pageNumber="91">IBIGEO-P 59</specimenCode>
|
||
</materialsCitation>
|
||
, left mandibular fragment with m2–m
|
||
<quantity id="4C409B36FFAEFF994A8BF8D6FD3FF813" box="[708,741,1909,1931]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="91" unit="in" value="3.0">3 in</quantity>
|
||
occlusal view.
|
||
<emphasis id="B9CCEAC1FFAEFF994B3EF8D6FCA5F812" bold="true" box="[881,895,1909,1930]" pageId="6" pageNumber="91">F</emphasis>
|
||
.
|
||
<materialsCitation id="3BD03C8EFFAEFF994BC4F8D6FBD3F813" box="[907,1033,1909,1931]" pageId="6" pageNumber="91" specimenCode="IBIGEO-P 61" specimenCount="1" typeStatus="holotype">
|
||
<specimenCode id="DB1E9EA8FFAEFF994BC4F8D6FBD3F813" box="[907,1033,1909,1931]" collectionCode="IBIGEO-P" pageId="6" pageNumber="91">IBIGEO-P 61</specimenCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3BD03C8EFFAEFF994C5CF8D6FA44F83E" collectionCode="A, B, C, E, F" collectorName="Photographs" location="Dashed area" pageId="6" pageNumber="91" specimenCode="A1, B1, C1, C3, C4, E1, F1, A2, F2" specimenCount="1" typeStatus="holotype">
|
||
left mandibular fragment with m1 (talonid)–m
|
||
<quantity id="4C409B36FFAEFF994DF4F8D6FA06F813" box="[1467,1500,1909,1931]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="91" unit="in" value="3.0">3 in</quantity>
|
||
occlusal view.
|
||
<collectorName id="264D5305FFAEFF994947F832FEA3F83F" box="[264,377,1937,1959]" pageId="6" pageNumber="91">Photographs</collectorName>
|
||
(
|
||
<specimenCode id="DB1E9EA8FFAEFF9949CAF832FE45F831" box="[389,415,1937,1961]" collectionCode="A" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="6" pageNumber="91" type="Herbarium">
|
||
A
|
||
<subScript id="173C3496FFAEFF9949D8F838FE45F831" attach="right" box="[407,415,1947,1961]" fontSize="6" pageId="6" pageNumber="91">1</subScript>
|
||
</specimenCode>
|
||
,
|
||
<specimenCode id="DB1E9EA8FFAEFF9949E5F832FE1BF831" box="[426,449,1937,1961]" collectionCode="B" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="6" pageNumber="91" type="Herbarium">
|
||
B
|
||
<subScript id="173C3496FFAEFF9949F6F838FE1BF831" attach="left" box="[441,449,1947,1961]" fontSize="6" pageId="6" pageNumber="91">1</subScript>
|
||
</specimenCode>
|
||
,
|
||
<specimenCode id="DB1E9EA8FFAEFF994983F832FE39F831" box="[460,483,1937,1961]" collectionCode="C" country="Denmark" name="University of Copenhagen" pageId="6" pageNumber="91" type="Herbarium">
|
||
C
|
||
<subScript id="173C3496FFAEFF994994F838FE39F831" attach="left" box="[475,483,1947,1961]" fontSize="6" pageId="6" pageNumber="91">1</subScript>
|
||
</specimenCode>
|
||
,
|
||
<specimenCode id="DB1E9EA8FFAEFF9949A0F832FDDCF831" box="[495,518,1937,1961]" collectionCode="C" country="Denmark" name="University of Copenhagen" pageId="6" pageNumber="91" type="Herbarium">
|
||
C
|
||
<subScript id="173C3496FFAEFF9949B1F838FDDCF831" attach="left" box="[510,518,1947,1961]" fontSize="6" pageId="6" pageNumber="91">3</subScript>
|
||
</specimenCode>
|
||
,
|
||
<specimenCode id="DB1E9EA8FFAEFF994A5EF832FDF2F831" box="[529,552,1937,1961]" collectionCode="C" country="Denmark" name="University of Copenhagen" pageId="6" pageNumber="91" type="Herbarium">
|
||
C
|
||
<subScript id="173C3496FFAEFF994A6FF838FDF2F831" attach="left" box="[544,552,1947,1961]" fontSize="6" pageId="6" pageNumber="91">4</subScript>
|
||
</specimenCode>
|
||
, D
|
||
<subScript id="173C3496FFAEFF994A0BF838FD96F831" attach="right" box="[580,588,1947,1961]" fontSize="6" pageId="6" pageNumber="91">1</subScript>
|
||
,
|
||
<specimenCode id="DB1E9EA8FFAEFF994A18F832FDB7F831" box="[599,621,1937,1961]" collectionCode="E" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="6" pageNumber="91" type="Herbarium">
|
||
E
|
||
<subScript id="173C3496FFAEFF994A2AF838FDB7F831" attach="left" box="[613,621,1947,1961]" fontSize="6" pageId="6" pageNumber="91">1</subScript>
|
||
</specimenCode>
|
||
,
|
||
<specimenCode id="DB1E9EA8FFAEFF994A37F832FD57F831" box="[632,653,1937,1961]" collectionCode="F" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="6" pageNumber="91" type="Herbarium">
|
||
F
|
||
<subScript id="173C3496FFAEFF994ACAF838FD57F831" attach="left" box="[645,653,1947,1961]" fontSize="6" pageId="6" pageNumber="91">1</subScript>
|
||
</specimenCode>
|
||
), explanatory drawings (
|
||
<specimenCode id="DB1E9EA8FFAEFF994B20F832FC53F831" box="[879,905,1937,1961]" collectionCode="A" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="6" pageNumber="91" type="Herbarium">
|
||
A
|
||
<subScript id="173C3496FFAEFF994BCEF838FC53F831" attach="right" box="[897,905,1947,1961]" fontSize="6" pageId="6" pageNumber="91">2</subScript>
|
||
</specimenCode>
|
||
–
|
||
<specimenCode id="DB1E9EA8FFAEFF994BDBF832FC73F831" box="[916,937,1937,1961]" collectionCode="F" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="6" pageNumber="91" type="Herbarium">
|
||
F
|
||
<subScript id="173C3496FFAEFF994BEEF838FC73F831" attach="right" box="[929,937,1947,1961]" fontSize="6" pageId="6" pageNumber="91">2</subScript>
|
||
</specimenCode>
|
||
).
|
||
<location id="8E676008FFAEFF994BF3F832FBF1F83E" LSID="urn:lsid:plazi:treatment:031187C5FFACFF9A4B11F8B9FDC2F83F:8E676008FFAEFF994BF3F832FBF1F83E" box="[956,1067,1937,1959]" name="Dashed area" pageId="6" pageNumber="91">Dashed area</location>
|
||
indicates broken or missing dental areas
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B0736D3FFADFF9A482DF8EFFDC2F83F" blockId="5.[98,771,1868,1960]" pageId="5" pageNumber="90">
|
||
<emphasis id="B9CCEAC1FFADFF9A482DF8EFFE20F8FD" box="[98,506,1868,1894]" italics="true" pageId="5" pageNumber="90">Stratigraphic and geographic range</emphasis>
|
||
.—Upper section of the Lumbrera and Quebrada de Los Colorados formations, middle Eocene;
|
||
<collectingRegion id="497CF831FFADFF9A48A3F82EFE49F83F" box="[236,403,1933,1959]" country="Argentina" name="Salta" pageId="5" pageNumber="90">Salta Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3AF7643FFADFF9A49D1F82EFDCEF83F" box="[414,532,1933,1959]" name="Argentina" pageId="5" pageNumber="90">Argentina</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |