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<document id="02F90D97330A83A9C50058420A37F5E9" ID-CLB-Dataset="24558" ID-DOI="10.11646/zootaxa.4712.1.2" ID-GBIF-Dataset="19829187-393b-46c5-8e83-c132a5742fb3" ID-ISSN="1175-5326" ID-Zenodo-Dep="3586816" ID-ZooBank="6C18DE3D-8B51-473D-B528-98337BF5730E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576824202715" checkinUser="plazi" docAuthor="Perreault, Ray T. &amp; Buckeridge, John S." docDate="2019" docId="03F87725FFAE890C5CA9FC4595D8F979" docLanguage="en" docName="zootaxa.4712.1.2.pdf" docOrigin="Zootaxa 4712 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Verruca alabamensis Perreault &amp; Buckeridge 2019, sp. nov." docType="treatment" docUuid="FD071DD7-1907-416A-81ED-6C5F4AAC8273" docUuidSource="ZooBank" docVersion="8" lastPageNumber="50" masterDocId="FFC10F5DFFA889045C3EFF909770FFE2" masterDocTitle="Paleogene Verrucidae (Cirripedia: Verrucomorpha) of North America, with descriptions of three new species" masterLastPageNumber="50" masterPageNumber="34" pageNumber="34" updateTime="1698775395024" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0563EAC982DA13777A9419A368078760">Paleogene Verrucidae (Cirripedia: Verrucomorpha) of North America, with descriptions of three new species</mods:title>
</mods:titleInfo>
<mods:name id="C405DA87D2970DE156EF5EF826C9EED0" type="personal">
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<mods:namePart id="C3534004E3629AFD5ACF42FE040AE545">Perreault, Ray T.</mods:namePart>
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<mods:namePart id="B43F164718C8819FA6E0B2002BBCDA76">Buckeridge, John S.</mods:namePart>
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<mods:typeOfResource id="558DA8D4ECD92507A4DCAD4BA71648FB">text</mods:typeOfResource>
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<mods:title id="048EE490EA80371183066202108142D3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="06302ACA0DD2F28C18ECD16C40AE3D81">
<mods:date id="CACC4208E1E39C5D3E10D1282460FBCE">2019</mods:date>
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<mods:number id="5EBDF0C00F664F7BC7A53BC7D65720B8">2019-12-18</mods:number>
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<mods:number id="6BA752B8BAC1B49B23360D38FD93F95F">4712</mods:number>
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<mods:classification id="CB210AA508A075D7F08ECB7F5B87DD71">journal article</mods:classification>
<mods:identifier id="5828C0BFE37384C55AD5A67D38847A65" type="CLB-Dataset">24558</mods:identifier>
<mods:identifier id="52B6599C860BBB3927E4A95FEB57E367" type="DOI">10.11646/zootaxa.4712.1.2</mods:identifier>
<mods:identifier id="ED1AAE27050E4D9CEAD95143D46BA60E" type="GBIF-Dataset">19829187-393b-46c5-8e83-c132a5742fb3</mods:identifier>
<mods:identifier id="F953C2E1B16692867BB4AE9E7B5042A8" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03F87725FFAE890C5CA9FC4595D8F979" ID-GBIF-Taxon="160797953" LSID="urn:lsid:zoobank.org:act:FD071DD7-1907-416A-81ED-6C5F4AAC8273" httpUri="http://treatment.plazi.org/id/03F87725FFAE890C5CA9FC4595D8F979" lastPageId="8" pageId="6">
<subSubSection id="C34B95B8FFAE89025CA9FC459642FBF1" pageId="6" type="nomenclature">
<paragraph id="8BEEC633FFAE89025CA9FC459445FC0D" blockId="6.[151,975,981,1079]" box="[151,821,981,1008]" pageId="6">
<heading id="D0A6715FFFAE89025CA9FC459445FC0D" bold="true" box="[151,821,981,1008]" fontSize="11" level="1" pageId="6" reason="1">
<emphasis id="B9251A21FFAE89025CA9FC459445FC0D" bold="true" box="[151,821,981,1008]" pageId="6">
<taxonomicName id="4C51BDB0FFAE89025CA9FC4595BAFC0D" authority="Perreault &amp; Buckeridge" authorityName="Perreault &amp; Buckeridge" authorityYear="2019" box="[151,714,981,1008]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="6" phylum="Arthropoda" rank="species" species="alabamensis" status="sp. nov.">
<emphasis id="B9251A21FFAE89025CA9FC4596E5FC12" bold="true" box="[151,405,981,1008]" italics="true" pageId="6">Verruca alabamensis</emphasis>
Perreault &amp; Buckeridge
</taxonomicName>
,
<taxonomicNameLabel id="A216A75AFFAE89025EE9FC459445FC0D" box="[727,821,981,1007]" pageId="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEEC633FFAE89025CA9FC699642FBF1" blockId="6.[151,975,981,1079]" box="[151,306,1017,1043]" pageId="6">Pl. 2, figs aq</paragraph>
</subSubSection>
<subSubSection id="C34B95B8FFAE89025CA9FB8C94BFFBD5" box="[151,975,1052,1079]" pageId="6" type="description">
<paragraph id="8BEEC633FFAE89025CA9FB8C94BFFBD5" blockId="6.[151,975,981,1079]" box="[151,975,1052,1079]" pageId="6">
<uri id="FFC0CA31FFAE89025CA9FB8C94BFFBD5" box="[151,975,1052,1079]" pageId="6">
urn:lsid:zoobank.org:act:
<uuid id="FFF7FCE6FFAE89025D97FB8C94BFFBD5" box="[425,975,1052,1079]" pageId="6">FD071DD7-1907-416A-81ED-6C5F4AAC8273</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C34B95B8FFAE89025CA9FBF496A6FB09" pageId="6" type="diagnosis">
<paragraph id="8BEEC633FFAE89025CA9FBF496A6FB09" blockId="6.[151,1437,1124,2015]" pageId="6">
<emphasis id="B9251A21FFAE89025CA9FBF49662FB9D" bold="true" box="[151,274,1124,1151]" pageId="6">Diagnosis.</emphasis>
All plates non-punctate; wall plates with ribbing prominent near plate margins, crossed with fine growth lines; rostral-carinal articulation with five ridges; movable scutum with one distinct articular rib; first and second articular ridges of movable tergum indistinct, about equal width; fixed scutal radio-alar wing extending one-third basal width of fixed scutum.
</paragraph>
</subSubSection>
<subSubSection id="C34B95B8FFAE89025CF9FB659597F9EE" pageId="6" type="materials_examined">
<paragraph id="8BEEC633FFAE89025CF9FB659605FAED" blockId="6.[151,1437,1124,2015]" box="[199,373,1268,1295]" pageId="6">
<emphasis id="B9251A21FFAE89025CF9FB659605FAED" bold="true" box="[199,373,1268,1295]" pageId="6">Type material.</emphasis>
</paragraph>
<paragraph id="8BEEC633FFAE89025CF9FA8994E9FAD1" blockId="6.[151,1437,1124,2015]" box="[199,921,1305,1331]" pageId="6">
<materialsCitation id="3B39CC6EFFAE89025CF9FA8994E9FAD1" ID-GBIF-Occurrence="2521263189" box="[199,921,1305,1331]" collectionCode="LSUMG" pageId="6" specimenCode="I-10727" specimenCount="1" typeStatus="holotype">
<emphasis id="B9251A21FFAE89025CF9FA89964BFAD1" bold="true" box="[199,315,1305,1331]" pageId="6">
<typeStatus id="54EA7891FFAE89025CF9FA899646FAD1" box="[199,310,1305,1331]" pageId="6" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED405EF6FFAE89025D7CFA8996D4FAD1" box="[322,420,1305,1331]" pageId="6">LSUMG</collectionCode>
<specimenCode id="DBF76E48FFAE89025D95FA899577FAD1" box="[427,519,1305,1331]" collectionCode="I-" pageId="6">I-10727</specimenCode>
, a movable scutum (pl. 2, figs o, p).
</materialsCitation>
</paragraph>
<paragraph id="8BEEC633FFAE89025CF9FAAD94E3FA26" blockId="6.[151,1437,1124,2015]" pageId="6">
<emphasis id="B9251A21FFAE89025CF9FAAD9636FAB5" bold="true" box="[199,326,1341,1367]" pageId="6">
<typeStatus id="54EA7891FFAE89025CF9FAAD9631FAB5" box="[199,321,1341,1367]" pageId="6" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Figured
<typeStatus id="54EA7891FFAE89025D8FFAAD9551FAB5" box="[433,545,1341,1367]" pageId="6" type="paratype">paratypes</typeStatus>
:
<materialsCitation id="3B39CC6EFFAE89025E10FAAD9318FABA" ID-GBIF-Occurrence="2521263201" box="[558,1128,1341,1368]" collectionCode="LSUMG" pageId="6" specimenCode="I-10723" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED405EF6FFAE89025E10FAAD95E0FAB5" box="[558,656,1341,1367]" pageId="6">LSUMG</collectionCode>
<specimenCode id="DBF76E48FFAE89025EA4FAAD9585FAB5" box="[666,757,1341,1367]" collectionCode="I-" pageId="6">I-10723</specimenCode>
, a fixed scutum (pl. 2, figs a,b)
</materialsCitation>
;
<materialsCitation id="3B39CC6EFFAE89025849FAAD96D4FA9E" ID-GBIF-Occurrence="2521263200" collectionCode="LSUMG" pageId="6" specimenCode="I-10724" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED405EF6FFAE89025849FAAD93A9FAB5" box="[1143,1241,1341,1367]" pageId="6">LSUMG</collectionCode>
<specimenCode id="DBF76E48FFAE890258DDFAAD924EFAB5" box="[1251,1342,1341,1367]" collectionCode="I-" pageId="6">I-10724</specimenCode>
, a fixed tergum (pl. 2, figs c, d)
</materialsCitation>
;
<materialsCitation id="3B39CC6EFFAE89025D8CFAF194D4FA9E" ID-GBIF-Occurrence="2521263185" box="[434,932,1376,1404]" collectionCode="LSUMG" pageId="6" specimenCode="I-10725" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED405EF6FFAE89025D8CFAF19564FA99" box="[434,532,1377,1403]" pageId="6">LSUMG</collectionCode>
<specimenCode id="DBF76E48FFAE89025E22FAF09508FA99" box="[540,632,1376,1403]" collectionCode="I-" pageId="6">I-10725</specimenCode>
, a rostrum (pl. 2, figs i, j)
</materialsCitation>
;
<materialsCitation id="3B39CC6EFFAE89025F8FFAF192E7FA9E" ID-GBIF-Occurrence="2521263187" box="[945,1431,1377,1404]" collectionCode="LSUMG" pageId="6" specimenCode="I-10726" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED405EF6FFAE89025F8FFAF19363FA99" box="[945,1043,1377,1403]" pageId="6">LSUMG</collectionCode>
<specimenCode id="DBF76E48FFAE89025822FAF19308FA99" box="[1052,1144,1377,1403]" collectionCode="I-" pageId="6">I-10726</specimenCode>
, a carina (pl. 2, figs k, l)
</materialsCitation>
;
<materialsCitation id="3B39CC6EFFAE89025CA9FA1595A7FA42" ID-GBIF-Occurrence="2521263196" box="[151,727,1413,1440]" collectionCode="LSUMG" pageId="6" specimenCode="I-10728" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED405EF6FFAE89025CA9FA159789FA7D" box="[151,249,1413,1439]" pageId="6">LSUMG</collectionCode>
<specimenCode id="DBF76E48FFAE89025D3EFA15962BFA7D" box="[256,347,1413,1439]" collectionCode="I-" pageId="6">I-10728</specimenCode>
, a movable tergum (pl. 2, fig g, h)
</materialsCitation>
;
<materialsCitation id="3B39CC6EFFAE89025EDCFA1597A5FA21" ID-GBIF-Occurrence="2521263183" collectionCode="LSUMG" pageId="6" specimenCode="I-10729" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED405EF6FFAE89025EDCFA159434FA7D" box="[738,836,1413,1439]" pageId="6">LSUMG</collectionCode>
<specimenCode id="DBF76E48FFAE89025F74FA1594D6FA7D" box="[842,934,1413,1439]" collectionCode="I-" pageId="6">I-10729</specimenCode>
, an articulated carina-rostrum pair (pl. 2, figs m, n)
</materialsCitation>
;
<materialsCitation id="3B39CC6EFFAE89025CDFFA3994FFFA26" ID-GBIF-Occurrence="2521263188" box="[225,911,1449,1476]" collectionCode="LSUMG" pageId="6" specimenCode="I-10730" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED405EF6FFAE89025CDFFA399633FA21" box="[225,323,1449,1475]" pageId="6">LSUMG</collectionCode>
<specimenCode id="DBF76E48FFAE89025D74FA3996D5FA21" box="[330,421,1449,1475]" collectionCode="I-" pageId="6">I-10730</specimenCode>
, a juvenile movable scutum (pl. 2, figs e, f)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEEC633FFAE89025CF9FA5D94C5FA0A" blockId="6.[151,1437,1124,2015]" box="[199,949,1484,1512]" pageId="6">
<emphasis id="B9251A21FFAE89025CF9FA5D9615FA05" bold="true" box="[199,357,1484,1511]" pageId="6">Type locality.</emphasis>
<collectionCode id="ED405EF6FFAE89025D52FA5D96BEFA05" box="[364,462,1485,1511]" pageId="6">LSUMG</collectionCode>
locality
<collectionCode id="ED405EF6FFAE89025E0EFA5D9539FA05" box="[560,585,1485,1511]" pageId="6">IL</collectionCode>
1974K (= HVH locality 319-6).
</paragraph>
<paragraph id="8BEEC633FFAE89025CF9FA609597F9EE" blockId="6.[151,1437,1124,2015]" box="[199,743,1520,1548]" pageId="6">
<emphasis id="B9251A21FFAE89025CF9FA6096CAF9E9" bold="true" box="[199,442,1520,1547]" pageId="6">Stratigraphic range.</emphasis>
Late Eocene (Priabonian).
</paragraph>
</subSubSection>
<subSubSection id="C34B95B8FFAE89025CF9F984979EF979" pageId="6" type="distribution">
<paragraph id="8BEEC633FFAE89025CF9F9849569F9CD" blockId="6.[151,1437,1124,2015]" box="[199,537,1556,1583]" pageId="6">
<emphasis id="B9251A21FFAE89025CF9F9849611F9CD" bold="true" box="[199,353,1556,1583]" pageId="6">Distribution.</emphasis>
<collectingRegion id="499508D1FFAE89025D58F98596BEF9CD" box="[358,462,1557,1583]" country="United States of America" name="Alabama" pageId="6">Alabama</collectingRegion>
,
<collectingCountry id="F34686A3FFAE89025DE7F9859565F9CD" box="[473,533,1557,1583]" name="United States of America" pageId="6">USA</collectingCountry>
.
</paragraph>
<paragraph id="8BEEC633FFAE89025CF9F9A8979EF979" blockId="6.[151,1437,1124,2015]" pageId="6">
<emphasis id="B9251A21FFAE89025CF9F9A896CEF9B1" bold="true" box="[199,446,1592,1619]" pageId="6">Additional material.</emphasis>
8 carinae, 10 rostra, 4 fixed scuta, 4 fixed terga. 2 movable scuta, 2 movable terga (LSUMG IL 1974K); 1 carina, 1 rostrum, 1 fixed scutum (Shubuta Member, UF locality ZA039) (Cat. No. UF- 70252).
</paragraph>
</subSubSection>
<subSubSection id="C34B95B8FFAE890C5CF9F934958FFC0D" lastPageId="8" pageId="6" type="description">
<paragraph id="8BEEC633FFAE89025CF9F93496D8F8E5" blockId="6.[151,1437,1124,2015]" pageId="6">
<emphasis id="B9251A21FFAE89025CF9F9349629F95D" bold="true" box="[199,345,1700,1727]" pageId="6">Description.</emphasis>
Shell ovate, elongated carino-rostrally with D-shaped orifice; all plates without ribbing or punctae, with fine growth lines; some wall plates provided with thickened base and calcareous internal rim, clearly shown in plate 2, fig. m.
</paragraph>
<paragraph id="8BEEC633FFAE89025CF9F88195DBF891" blockId="6.[151,1437,1124,2015]" pageId="6">Carina strongly asymmetrical, with tilt toward fixed tergal side; fixed tergal margin with three articular ribs, distinct, with lowermost very small; rostral margin with five articular ribs, evenly lengthening apically; interior provided with dependent sheath widest at apex.</paragraph>
<paragraph id="8BEEC633FFAE89025CA9F8ED9621F83D" blockId="6.[151,1437,1124,2015]" pageId="6">Rostrum strongly asymmetrical, with three articular ribs on fixed scutal margin, five ribs on carinal margin; ornamented by fine growth lines only; interior provided with narrow sheath, dependent, with furrow to receive free opercular plates.</paragraph>
<caption id="DF2E96BBFFAF89035CA9F903934EF829" ID-DOI="http://doi.org/10.5281/zenodo.3586820" ID-Zenodo-Dep="3586820" httpUri="https://zenodo.org/record/3586820/files/figure.png" pageId="7" startId="7.[151,234,1683,1708]" targetBox="[240,1347,181,1657]" targetPageId="7">
<paragraph id="8BEEC633FFAF89035CA9F903934EF829" blockId="7.[151,1436,1683,1996]" pageId="7">
<emphasis id="B9251A21FFAF89035CA9F903978DF94E" bold="true" box="[151,253,1683,1708]" pageId="7">PLATE 2</emphasis>
.
<taxonomicName id="4C51BDB0FFAF89035D34F9049690F94E" authority="Perreault &amp; Buckeridge, 2019" authorityName="Perreault &amp; Buckeridge" authorityYear="2019" box="[266,480,1684,1708]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="7" phylum="Arthropoda" rank="species" species="alabamensis" status="sp. nov.">
<emphasis id="B9251A21FFAF89035D34F9049690F94E" box="[266,480,1684,1708]" italics="true" pageId="7">Verruca alabamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A216A75AFFAF89035DD8F9049545F949" box="[486,565,1684,1707]" pageId="7" rank="species">sp. nov.</taxonomicNameLabel>
a. Paratype I-10723, fixed scutum (interior); b. paratype I-10723, fixed scutum (exterior); c. paratype I-10724, fixed tergum (interior); d. paratype I-10724, fixed tergum (exterior); e. paratype I-10730, movable scutum, juvenile (interior); f. paratype I-10730, movable scutum, juvenile (exterior); g. paratype I-10728, movable tergum (interior); h. paratype I-10728, movable tergum (exterior); i. paratype I-10725, rostrum (exterior); j. paratype I-10725, rostrum (interior); k. paratype I-10726, carina (interior); l. paratype I-10726, carina (exterior); m. paratype I-10729, articulated carina and rostrum, (basal view) showing inward extension of shell; n. paratype I-10729, articulated carina and rostrum, apical view; o. holotype I-10727, movable scutum (interior); p. holotype I-10727, movable scutum (exterior); q. inset of o, boring of unknown origin, located just below adductor scar. All specimens are from the Yazoo Formation, Pachuta Marl Member, Henry V. Howe Location HVH 319-6, Clarke County, Alabama. Scale bar for q = 0.25 mm, all others 1 mm.
</paragraph>
</caption>
<paragraph id="8BEEC633FFA0890C5CF9FF0996CDFEA1" blockId="8.[151,1437,153,1691]" pageId="8">Fixed scutum convex, smooth, wider than high, with narrow, crescentic radio-alar wing extending about onethird its own distance from rostral margin, ornamented with fine, transverse growth lines only; rostral articulation with two articulating ribs; fixed tergal margin convex with node one third to base, with two ribs below; interior with bordering ridge on fixed tergal margin; straight broad articular ridge extending almost to apex with one rib, tilting basally; myophore absent.</paragraph>
<paragraph id="8BEEC633FFA0890C5CF9FEDD96D9FE15" blockId="8.[151,1437,153,1691]" pageId="8">Fixed tergum convex, smooth, higher than wide, flared basally, ornamented by fine growth lines only; fixed scutal margin straight or slightly convex, with wide internal border ridge; protruding ala short, curved with small indentation at base; carinal margin with two coarse, vague articular ribs and prominent upward curving ala slightly protruding from carinal margin and prominent indentation below; interior strongly concave, apical shelf dependent, strongly sloping basally.</paragraph>
<paragraph id="8BEEC633FFA0890C5CF9FD919253FCF5" blockId="8.[151,1437,153,1691]" pageId="8">Movable scutum slightly bowed, plate moderately curved tergally; exterior ornamented by regularly spaced incised growth lines, each forming very low tooth on highly convex occludent margin; tergal margin strongly concave, less curved than occludent margin; margin provided with low, rounded border ridge, fading basally; one prominent, wide articular ridge extending three-quarters to basal margin; ridge protruding from tergal margin and separated from plate by shallow, distinct furrow with incised borders; basal margin very slightly convex, basi-tergal angle nearly 90˚, basi-occludent angle broadly rounded. Interior smooth, all margins provided with low, rounded ridge; adductor scar large, sub-rounded, vaguely defined, placed close to occludent margin well apically of center; apex with prominent overfold for articulation with movable tergum, marked by distinct bordering ridge.</paragraph>
<paragraph id="8BEEC633FFA0890C5CF9FCB1958FFC0D" blockId="8.[151,1437,153,1691]" pageId="8">Movable tergum flat, rhomboidal, with wide, prominent apico-basal ridge; ridge raised, demarked by incised edges, slightly overhanging, protruding slightly more than its own width from basiscutal angle, merging into plate near apex; basal margin nearly straight, carinal margin gently convex, meeting at nearly right angle; basi-scutal angle acute, scutal margin undulating; carinal portion triangular, with prominent growth ridges at nearly right angles to carinal margin basally, sloping basally in apical half; growth ridges converging slightly towards apico-basal ridge; two articular ridges, vague, coarse; interior featureless.</paragraph>
</subSubSection>
<subSubSection id="C34B95B8FFA0890C5CF9FC68920BF9E9" pageId="8" type="discussion">
<paragraph id="8BEEC633FFA0890C5CF9FC689604FBD5" blockId="8.[151,1437,153,1691]" pageId="8">
<emphasis id="B9251A21FFA0890C5CF9FC68963CFBF1" bold="true" box="[199,332,1016,1043]" pageId="8">Discussion.</emphasis>
The lack of punctae and ribbing differentiates this species from
<taxonomicName id="4C51BDB0FFA0890C5829FC6A9262FBF1" authority="Perreault &amp; Buckeridge, 2019" authorityName="Perreault &amp; Buckeridge" authorityYear="2019" box="[1047,1298,1017,1043]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="8" phylum="Arthropoda" rank="species" species="gailgoedertae" status="sp. nov.">
<emphasis id="B9251A21FFA0890C5829FC6A9262FBF1" box="[1047,1298,1017,1043]" italics="true" pageId="8">Verruca gailgoedertae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A216A75AFFA0890C5926FC69921EFBF1" box="[1304,1390,1017,1043]" pageId="8" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C51BDB0FFA0890C5CA9FB8E9667FBD5" authority="Perreault &amp; Buckeridge, 2019" authorityName="Perreault &amp; Buckeridge" authorityYear="2019" box="[151,279,1053,1079]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="8" phylum="Arthropoda" rank="species" species="sorrellae" status="sp. nov.">
<emphasis id="B9251A21FFA0890C5CA9FB8E9667FBD5" box="[151,279,1053,1079]" italics="true" pageId="8">V. sorrellae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A216A75AFFA0890C5D20FB8D9604FBD5" box="[286,372,1053,1079]" pageId="8" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BEEC633FFA0890C5CF9FBD295ACFB25" blockId="8.[151,1437,153,1691]" pageId="8">
<taxonomicName id="4C51BDB0FFA0890C5CF9FBD296C1FBB9" authority="Perreault &amp; Buckeridge, 2019" authorityName="Perreault &amp; Buckeridge" authorityYear="2019" box="[199,433,1089,1115]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="8" phylum="Arthropoda" rank="species" species="alabamensis" status="sp. nov.">
<emphasis id="B9251A21FFA0890C5CF9FBD296C1FBB9" box="[199,433,1089,1115]" italics="true" pageId="8">Verruca alabamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A216A75AFFA0890C5D89FBD1957CFBB9" box="[439,524,1089,1115]" pageId="8" rank="species">sp. nov.</taxonomicNameLabel>
may be distinguished from other known North American Paleogene
<taxonomicName id="4C51BDB0FFA0890C593FFBD1920AFBB9" authorityName="Darwin" authorityYear="1854" box="[1281,1402,1089,1115]" class="Maxillopoda" family="Verrucidae" kingdom="Animalia" order="Sessilia" pageId="8" phylum="Arthropoda" rank="family">Verrucidae</taxonomicName>
by its inturned calcareous rim on some plates. This feature does not seem to be related to size or ontogeny. All of these in-turned rims, and most basal margins, show a preferred attachment to ribbed substrates. Most likely, plicatulid bivalves are candidates, by the shape of the ribbing.
</paragraph>
<paragraph id="8BEEC633FFA0890C5CF9FB429569FA21" blockId="8.[151,1437,153,1691]" pageId="8">
<taxonomicName id="4C51BDB0FFA0890C5CF9FB4296C2FB09" authority="Perreault &amp; Buckeridge, 2019" authorityName="Perreault &amp; Buckeridge" authorityYear="2019" box="[199,434,1233,1259]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="8" phylum="Arthropoda" rank="species" species="alabamensis" status="sp. nov.">
<emphasis id="B9251A21FFA0890C5CF9FB4296C2FB09" box="[199,434,1233,1259]" italics="true" pageId="8">Verruca alabamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A216A75AFFA0890C5D86FB41957EFB09" box="[440,526,1233,1259]" pageId="8" rank="species">sp. nov.</taxonomicNameLabel>
is uncommon, and must have occurred under restricted environmental conditions. Both known localities are characterized by abundant branching bryozoans, crinoids, gorgonian axes and brachiopods (dominantly
<taxonomicName id="4C51BDB0FFA0890C5D5CFA89951BFAD1" baseAuthorityName="Morton" baseAuthorityYear="1834" box="[354,619,1305,1331]" class="Rhynchonellata" family="Cancellothyrididae" genus="Terebratulina" kingdom="Animalia" order="Terebratulida" pageId="8" phylum="Brachiopoda" rank="species" species="lachryma">
<emphasis id="B9251A21FFA0890C5D5CFA89951BFAD1" box="[354,619,1305,1331]" italics="true" pageId="8">Terebratulina lachryma</emphasis>
</taxonomicName>
). In both localities, aragonitic remains are completely leached, and phosphate is present. At the
<typeStatus id="54EA7891FFA0890C5DA5FAAD96BBFAB5" box="[411,459,1341,1367]" pageId="8">type</typeStatus>
locality, phosphate occurs as internal molds of molluscs and solitary corals. Corals may be identified to genus on the basis of these internal molds. At the Shubuta locality, phosphate occurs additionally as small nodules. These nodules are sometimes encrusted with bryozoan colonies, indicating nodule formation was contemporaneous with deposition.
</paragraph>
<paragraph id="8BEEC633FFA0890C5CF9FA5D920BF9E9" blockId="8.[151,1437,153,1691]" pageId="8">
Boring traces are common at the type locality. Several wall plates show naticoid drill holes, some fully penetrating the shell. The
<typeStatus id="54EA7891FFA0890C5D5EFA6196B2F9E9" box="[352,450,1521,1547]" pageId="8" type="holotype">holotype</typeStatus>
scutum features a minute, spearhead-shaped boring of unknown origin (Pl. 2, fig. q).
</paragraph>
</subSubSection>
<subSubSection id="C34B95B8FFA0890C5CF9F98595A4F9CD" box="[199,724,1557,1583]" pageId="8" type="etymology">
<paragraph id="8BEEC633FFA0890C5CF9F98595A4F9CD" blockId="8.[151,1437,153,1691]" box="[199,724,1557,1583]" pageId="8">
<emphasis id="B9251A21FFA0890C5CF9F985963EF9CD" bold="true" box="[199,334,1557,1583]" pageId="8">Etymology.</emphasis>
Named after the state of
<collectingRegion id="499508D1FFA0890C5E59F98595A0F9CD" box="[615,720,1557,1583]" country="United States of America" name="Alabama" pageId="8">Alabama</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C34B95B8FFA0890C5CF9F9A995D8F979" pageId="8" type="materials_examined">
<paragraph id="8BEEC633FFA0890C5CF9F9A995D8F979" blockId="8.[151,1437,153,1691]" pageId="8">
<emphasis id="B9251A21FFA0890C5CF9F9A996D6F9B1" bold="true" box="[199,422,1592,1619]" pageId="8">Type Repositories.</emphasis>
<materialsCitation id="3B39CC6EFFA0890C5D90F9A995D8F979" ID-GBIF-Occurrence="2521263198" collectionCode="LSUMG" collectorName="Baton Rouge, LA" county="All" location="Invertebrates" municipality="Collection of Fossil Protists" pageId="8" specimenCode="I-10727, I-10723-10726, 10728-10730, I-10731-10737" specimenCount="2" stateProvince="Louisiana" typeStatus="holotype">
<typeStatus id="54EA7891FFA0890C5D90F9A99566F9B1" box="[430,534,1593,1619]" pageId="8" type="holotype">Holotype</typeStatus>
<collectionCode id="ED405EF6FFA0890C5E21F9A995F1F9B1" box="[543,641,1593,1619]" pageId="8">LSUMG</collectionCode>
<specimenCode id="DBF76E48FFA0890C5EB4F9A99596F9B1" box="[650,742,1593,1619]" collectionCode="I-" pageId="8">I-10727</specimenCode>
,
<typeStatus id="54EA7891FFA0890C5ECDF9A9942EF9B1" box="[755,862,1593,1619]" pageId="8" type="paratype">paratypes</typeStatus>
<collectionCode id="ED405EF6FFA0890C5F59F9A994B9F9B1" box="[871,969,1593,1619]" pageId="8">LSUMG</collectionCode>
<specimenCode id="DBF76E48FFA0890C5FECF9A9930CF9B1" box="[978,1148,1593,1619]" collectionCode="I-" pageId="8">I-10723-10726</specimenCode>
,
<specimenCode id="DBF76E48FFA0890C58B7F9A99252F9B1" box="[1161,1314,1593,1619]" collectionCode="I-" pageId="8">10728-10730</specimenCode>
; topotypes
<collectionCode id="ED405EF6FFA0890C5CA9F9CD9789F995" box="[151,249,1629,1655]" pageId="8">LSUMG</collectionCode>
<specimenCode id="DBF76E48FFA0890C5D3DF9CD96DEF995" box="[259,430,1629,1655]" collectionCode="I-" pageId="8">I-10731-10737</specimenCode>
.
<collectingCounty id="628FBEBFFFA0890C5D84F9CD96AEF995" box="[442,478,1629,1655]" pageId="8">All</collectingCounty>
held by
<collectingMunicipality id="6B8A5C49FFA0890C5E76F9CD94F8F995" box="[584,904,1629,1655]" pageId="8">Collection of Fossil Protists</collectingMunicipality>
and
<location id="8E8E90E8FFA0890C5FFAF9CD9329F995" LSID="urn:lsid:plazi:treatment:03F87725FFAE890C5CA9FC4595D8F979:8E8E90E8FFA0890C5FFAF9CD9329F995" box="[964,1113,1629,1655]" county="All" municipality="Collection of Fossil Protists" name="Invertebrates" pageId="8" stateProvince="Louisiana">Invertebrates</location>
,
<collectingRegion id="499508D1FFA0890C5859F9CD9267F995" box="[1127,1303,1629,1655]" country="United States of America" name="Louisiana" pageId="8">Louisiana State</collectingRegion>
<location id="8E8E90E8FFA0890C591FF9CD92E8F995" LSID="urn:lsid:plazi:treatment:03F87725FFAE890C5CA9FC4595D8F979:8E8E90E8FFA0890C591FF9CD92E8F995" box="[1313,1432,1629,1655]" county="All" municipality="Collection of Fossil Protists" name="University" pageId="8" stateProvince="Louisiana">University</location>
,
<location id="8E8E90E8FFA0890C5CA9F91196BEF979" LSID="urn:lsid:plazi:treatment:03F87725FFAE890C5CA9FC4595D8F979:8E8E90E8FFA0890C5CA9F91196BEF979" box="[151,462,1665,1691]" county="All" municipality="Collection of Fossil Protists" name="Museum of Natural History" pageId="8" stateProvince="Louisiana">Museum of Natural History</location>
,
<collectorName id="26A4A3E5FFA0890C5DE4F91195D4F979" box="[474,676,1665,1691]" pageId="8">Baton Rouge, LA</collectorName>
.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>