treatments-xml/data/03/F6/A2/03F6A238AB10FFA6FF7BFE8AFB4E5D99.xml
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<document id="3E1DBDA9E84DD00F346493EC38278996" ID-DOI="10.5281/zenodo.5352378" ID-ISSN="1942-1354" ID-Zenodo-Dep="5352378" ID-ZooBank="5E18383A-4237-44CD-8BC2-86829DDBBA9D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630208073887" checkinUser="marcus" docAuthor="Springer, Charles A. &amp; Waller, Deborah A." docDate="2021" docId="03F6A238AB10FFA6FF7BFE8AFB4E5D99" docLanguage="en" docName="InsectaMundi.2021.0882.1-9.pdf" docOrigin="Insecta Mundi 2021 (882)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Scirtes goodrichi Springer and Waller 2021, new species" docType="treatment" docVersion="4" lastPageNumber="7" masterDocId="FFCFDA40AB14FFAEFFBBFF9EFFC25F35" masterDocTitle="A new species of Scirtes Illiger, 1807 (Coleoptera: Scirtidae) from Virginia and South Carolina, U. S. A." masterLastPageNumber="9" masterPageNumber="1" pageNumber="3" updateTime="1699185587170" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="702EE45FA8B064BA536ADCA4C6973DD6">A new species of Scirtes Illiger, 1807 (Coleoptera: Scirtidae) from Virginia and South Carolina, U. S. A.</mods:title>
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<mods:namePart id="76557A82DACF23EAF1BB1220EC206195">Springer, Charles A.</mods:namePart>
<mods:affiliation id="F6B94D1162C60C94AC862402BAF35AA0">1300 Cloncurry Road Norfolk, Virginia 23505</mods:affiliation>
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<mods:namePart id="1FBEA613AB128B43F3607AAE584984F8">Waller, Deborah A.</mods:namePart>
<mods:affiliation id="5248C6FB671143D78B9BF0DDBA57EE61">Department of Biology Old Dominion University Norfolk, VA 23529</mods:affiliation>
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<mods:title id="492CCB3028950C676D8F8927B677CCBD">Insecta Mundi</mods:title>
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<mods:part id="25C15BB0746151A2AE0D85FAC659B344">
<mods:date id="8F8402C6A363AAFDC638ED6FF9BC9FDC">2021</mods:date>
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<mods:identifier id="27384CB5C10EE295CD2031E8E777E840" type="DOI">10.5281/zenodo.5352378</mods:identifier>
<mods:identifier id="A3831685586DDC1CE8F0CB5D3097F6F6" type="ISSN">1942-1354</mods:identifier>
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<treatment id="03F6A238AB10FFA6FF7BFE8AFB4E5D99" ID-DOI="http://doi.org/10.5281/zenodo.5450469" ID-Zenodo-Dep="5450469" LSID="urn:lsid:plazi:treatment:03F6A238AB10FFA6FF7BFE8AFB4E5D99" httpUri="http://treatment.plazi.org/id/03F6A238AB10FFA6FF7BFE8AFB4E5D99" lastPageId="8" lastPageNumber="7" pageId="4" pageNumber="3">
<subSubSection id="C34540A5AB10FFAAFF7BFE8AFCFB5E04" box="[192,825,275,305]" pageId="4" pageNumber="3" type="nomenclature">
<paragraph id="8BE0132EAB10FFAAFF7BFE8AFCFB5E04" blockId="4.[192,825,275,305]" box="[192,825,275,305]" pageId="4" pageNumber="3">
<heading id="D0A8A442AB10FFAAFF7BFE8AFCFB5E04" box="[192,825,275,305]" fontSize="12" level="2" pageId="4" pageNumber="3" reason="2">
<emphasis id="B92BCF3CAB10FFAAFF7BFE8AFCFB5E04" bold="true" box="[192,825,275,305]" pageId="4" pageNumber="3">
<taxonomicName id="4C5F68ADAB10FFAAFF7BFE8AFD5A5E05" authority="Springer and Waller" authorityName="Springer and Waller" authorityYear="2021" box="[192,664,275,305]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="goodrichi" status="sp. nov.">
<emphasis id="B92BCF3CAB10FFAAFF7BFE8AFE525E05" bold="true" box="[192,400,275,304]" italics="true" pageId="4" pageNumber="3">Scirtes goodrichi</emphasis>
Springer and Waller
</taxonomicName>
,
<taxonomicNameLabel id="A2187247AB10FFAAFD1FFE8AFCFB5E04" box="[676,825,276,305]" pageId="4" pageNumber="3" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34540A5AB10FFAAFF7BFEDEFC355C6E" pageId="4" pageNumber="3" type="diagnosis">
<paragraph id="8BE0132EAB10FFAAFF7BFEDEFDDB5D12" blockId="4.[191,1440,320,859]" pageId="4" pageNumber="3">
<emphasis id="B92BCF3CAB10FFAAFF7BFEDEFEFE5E6F" bold="true" box="[192,316,320,346]" pageId="4" pageNumber="3">Diagnosis.</emphasis>
<taxonomicName id="4C5F68ADAB10FFAAFEF8FEDEFE355E6F" authorityName="Springer and Waller" authorityYear="2021" box="[323,503,320,346]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="goodrichi">
<emphasis id="B92BCF3CAB10FFAAFEF8FEDEFE355E6F" box="[323,503,320,346]" italics="true" pageId="4" pageNumber="3">Scirtes goodrichi</emphasis>
</taxonomicName>
(
<figureCitation id="13640FABAB10FFAAFDBDFEDEFD875E6F" box="[518,581,320,346]" captionStart="Figures 14" captionStartId="6.[193,280,1866,1892]" captionTargetBox="[274,1341,567,1824]" captionTargetId="figure-174@6.[274,1359,567,1837]" captionTargetPageId="6" captionText="Figures 14. Scirtes goodrichi. 1) Adult dorsal habitus. 2) Female prehensor. 3) Mandible. 4) Prosternal process lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5352380" httpUri="https://zenodo.org/record/5352380/files/figure.png" pageId="4" pageNumber="3">Fig. 1</figureCitation>
) is distinguished from the five other species of
<taxonomicName id="4C5F68ADAB10FFAAFBE6FEDEFB665E6F" authorityName="Illiger" authorityYear="1807" box="[1117,1188,320,346]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B92BCF3CAB10FFAAFBE6FEDEFB665E6F" box="[1117,1188,320,346]" italics="true" pageId="4" pageNumber="3">Scirtes</emphasis>
</taxonomicName>
in our area by the following combination of characters that define this species: head width over 2.31 times head length; antennomere II arises from apex of I; clypeus with anterolateral angles not recurved; head, pronotum and proepisternum dark, without white, tan or orange markings; length of elytra 4.105.58 times mid-elytral height; elytra never vittate but usually with orange-brown sutural stripe; lateral margin of elytra slightly explanate, width of turned-out rim less than width of protibia; shape of prosternal process, male genitalia, tergite VIII (except
<taxonomicName id="4C5F68ADAB10FFAAFB3AFE75FAD75D30" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[1153,1301,491,517]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="orbiculatus">
<emphasis id="B92BCF3CAB10FFAAFB3AFE75FAD75D30" box="[1153,1301,491,517]" italics="true" pageId="4" pageNumber="3">S. orbiculatus</emphasis>
</taxonomicName>
) and female prehensor also species specific.
</paragraph>
<paragraph id="8BE0132EAB10FFAAFEBBFDAEFC355C6E" blockId="4.[191,1440,320,859]" pageId="4" pageNumber="3">
<taxonomicName id="4C5F68ADAB10FFAAFEBBFDAEFE775D7C" authorityName="Springer and Waller" authorityYear="2021" box="[256,437,559,586]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="goodrichi">
<emphasis id="B92BCF3CAB10FFAAFEBBFDAEFE775D7C" box="[256,437,559,586]" italics="true" pageId="4" pageNumber="3">Scirtes goodrichi</emphasis>
</taxonomicName>
without elytral stripe resemble the general habitus of
<taxonomicName id="4C5F68ADAB10FFAAFBA5FDAEFB465D7C" authorityName="Guerin-Meneville" authorityYear="1843" box="[1054,1156,559,586]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B92BCF3CAB10FFAAFBA5FDAEFB465D7C" box="[1054,1156,559,586]" italics="true" pageId="4" pageNumber="3">S. tibialis</emphasis>
</taxonomicName>
but can be distinguished externally by the clypeal anterolateral margins not recurved, head width over 2.31 times head length; long overlapping mandibular teeth, and shape of prosternal process. Internally,
<taxonomicName id="4C5F68ADAB10FFAAFC73FDEAFB8A5DB8" authorityName="Springer and Waller" authorityYear="2021" box="[968,1096,627,654]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="goodrichi">
<emphasis id="B92BCF3CAB10FFAAFC73FDEAFB8A5DB8" box="[968,1096,627,654]" italics="true" pageId="4" pageNumber="3">S. goodrichi</emphasis>
</taxonomicName>
differ markedly from
<taxonomicName id="4C5F68ADAB10FFAAFA87FDEAFA625DB8" authorityName="Guerin-Meneville" authorityYear="1843" box="[1340,1440,627,654]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B92BCF3CAB10FFAAFA87FDEAFA625DB8" box="[1340,1440,627,654]" italics="true" pageId="4" pageNumber="3">S. tibialis</emphasis>
</taxonomicName>
by the male genitalia, male tergites VIII and IX and female prehensor (
<figureCitation id="13640FABAB10FFAAFC6FFD08FBD45D85" box="[980,1046,662,688]" captionStart="Figures 14" captionStartId="6.[193,280,1866,1892]" captionTargetBox="[274,1341,567,1824]" captionTargetId="figure-174@6.[274,1359,567,1837]" captionTargetPageId="6" captionText="Figures 14. Scirtes goodrichi. 1) Adult dorsal habitus. 2) Female prehensor. 3) Mandible. 4) Prosternal process lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5352380" httpUri="https://zenodo.org/record/5352380/files/figure.png" pageId="4" pageNumber="3">Fig. 2</figureCitation>
). If a completely dark or melanistic phase of
<taxonomicName id="4C5F68ADAB10FFAAFE9BFD26FE765DE7" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[288,436,696,722]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="orbiculatus">
<emphasis id="B92BCF3CAB10FFAAFE9BFD26FE765DE7" box="[288,436,696,722]" italics="true" pageId="4" pageNumber="3">S. orbiculatus</emphasis>
</taxonomicName>
is found, yet unknown to the authors, differences in elytral convexity, body length to width ratio and shape of prosternal lateral aspect will separate them from
<taxonomicName id="4C5F68ADAB10FFAAFC7DFD44FB8A5DC1" authorityName="Springer and Waller" authorityYear="2021" box="[966,1096,730,756]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="goodrichi">
<emphasis id="B92BCF3CAB10FFAAFC7DFD44FB8A5DC1" box="[966,1096,730,756]" italics="true" pageId="4" pageNumber="3">S. goodrichi</emphasis>
</taxonomicName>
externally, differences in male genitalia, male tergite IX, and female prehensor will separate them internally.
<taxonomicName id="4C5F68ADAB10FFAAFB87FD63FB335C23" authorityName="Springer and Waller" authorityYear="2021" box="[1084,1265,764,791]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="goodrichi">
<emphasis id="B92BCF3CAB10FFAAFB87FD63FB335C23" box="[1084,1265,764,791]" italics="true" pageId="4" pageNumber="3">Scirtes goodrichi</emphasis>
</taxonomicName>
is most closely related to
<taxonomicName id="4C5F68ADAB10FFAAFE88FC81FE0A5C0D" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[307,456,798,825]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="orbiculatus">
<emphasis id="B92BCF3CAB10FFAAFE88FC81FE0A5C0D" box="[307,456,798,825]" italics="true" pageId="4" pageNumber="3">S. orbiculatus</emphasis>
</taxonomicName>
based on similarities of the clypeus, head width to head length ratio, mandibular tooth size, elytral stripe, male genitalia, male tergite VIII, and female prehensor.
</paragraph>
</subSubSection>
<subSubSection id="C34540A5AB10FFABFF7BFCF4FEBC5D56" lastPageId="5" lastPageNumber="4" pageId="4" pageNumber="3" type="description">
<paragraph id="8BE0132EAB10FFABFF7BFCF4FEBC5D56" blockId="4.[191,1442,874,1927]" lastBlockId="5.[191,1441,214,611]" lastPageId="5" lastPageNumber="4" pageId="4" pageNumber="3">
<emphasis id="B92BCF3CAB10FFAAFF7BFCF4FE205CB0" bold="true" box="[192,482,874,901]" pageId="4" pageNumber="3">
Description of
<typeStatus id="54E4AD8CAB10FFAAFECFFCF5FE1F5CB0" box="[372,477,875,901]" pageId="4" pageNumber="3" type="holotype">holotype</typeStatus>
.
</emphasis>
<emphasis id="B92BCF3CAB10FFAAFE57FCF5FDEB5CB0" box="[492,553,875,901]" italics="true" pageId="4" pageNumber="3">Body.</emphasis>
Elongate oval, elytra slightly convex, head and pronotum deflexed in normal relaxed position, length from anterior margin of pronotum to apex of elytra
<quantity id="4CA7BECBAB10FFAAFBA4FC13FB475C92" box="[1055,1157,909,936]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" pageId="4" pageNumber="3" unit="mm" value="2.75">2.75 mm</quantity>
, width across both elytra
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, length 1.62 times width. Head width across both eyes
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, pronotum width across posterior margin
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, pronotum 1.54 times wider than head, elytra 1.32 times wider than pronotum. Maximum lengths: head from anterior margin of clypeus to base of eyes
<quantity id="4CA7BECBAB10FFAAFCAFFC6DFCB85B3B" box="[788,890,1011,1038]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="3" unit="mm" value="0.35">0.35 mm</quantity>
; pronotum
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; elytra
<quantity id="4CA7BECBAB10FFAAFB0FFC6AFAD85B3B" box="[1204,1306,1012,1038]" metricMagnitude="-3" metricUnit="m" metricValue="2.38" pageId="4" pageNumber="3" unit="mm" value="2.38">2.38 mm</quantity>
; total maximum length
<quantity id="4CA7BECBAB10FFAAFEEAFB88FE755B05" box="[337,439,1046,1072]" metricMagnitude="-3" metricUnit="m" metricValue="3.24" pageId="4" pageNumber="3" unit="mm" value="3.24">3.24 mm</quantity>
; total length 1.92 times width.
<emphasis id="B92BCF3CAB10FFAAFCA9FB88FC4C5B05" box="[786,910,1046,1072]" italics="true" pageId="4" pageNumber="3">Coloration.</emphasis>
Entire dorsum shiny, dark brown except for an orange-brown longitudinal stripe on the middle 1/3 of the elytral suture, not extending outward over disc, width of stripe subequal to one dorsal eye width on each elytron, color and width of stripe decrease anteriorly and posteriorly. Prosternum, mesosternum, metasternum, all coxae and femora brown; abdominal sternum with irregular areas of dark and light brown. Proepisterna and elytral epipleura light brown. All palps, tibiae and tarsi light brown to stramineous. Antennomere I brown, IIVIII light brown, IXXI fuscous.
<emphasis id="B92BCF3CAB10FFAAFBE5FB5FFB1C5BEF" box="[1118,1246,1217,1242]" italics="true" pageId="4" pageNumber="3">Punctation.</emphasis>
Setigerous punctures on head, pronotum and scutellar shield very fine, subequal in size to one eye facet diameter; head punctures separated by one to three puncture diameters; punctures on pronotum and scutellum separated by 35 puncture diameters, areas between punctures smooth, shiny. Elytral punctures larger, equal in size to 3 eye facet diameters, separated by one-half to one puncture diameter, producing a coarser surface than on head and pronotum.
<emphasis id="B92BCF3CAB10FFAAFAD9FAD4FF315AB0" italics="true" pageId="4" pageNumber="3">Vestiture.</emphasis>
Most setae on dorsum pale yellow and semi-erect; setae on head directed forward, setae on pronotum and scutellar shield directed backward, all above setae subequal in length to width of prothoracic tarsomere I; setae on elytra directed backwards, longer, subequal in length to width of metathoracic tarsomere I. Setae on venter and appendages shorter than on head, pale yellow and recumbent.
<emphasis id="B92BCF3CAB10FFAAFCCEFA4CFC755AD9" box="[885,951,1490,1516]" italics="true" pageId="4" pageNumber="3">Head.</emphasis>
Deflexed, inserted into prothorax to base of eyes, wider than long, width across eyes
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, length from anterior margin of clypeus to posterior margin of eyes
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, width 2.38 times length. Dorsal interocular distance
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, dorsal eye width
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, dorsal interocular distance 3.2 times one dorsal eye width. Eyes dark grey in middle with argenteous rim two to three eye facets wide. Subocular carina well developed under entire eye, carina height equals two to three eye facet diameters. Clypeus with lateral margins straight and directed anteriorly, anterolateral angles rounded, anterior margin slightly emarginate, width 14 times longer than depth. Antennae filiform,
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long, 1.49 times width of head across eyes, and 0.45 times body length. Antennomere II attached to apex of I, antennomeres I to III decrease in size, III is the smallest overall, IV to X subequal, XI longest.
<emphasis id="B92BCF3CAB10FFAAFC66F97AFB9159C8" box="[989,1107,1764,1789]" italics="true" pageId="4" pageNumber="3">Pronotum.</emphasis>
Slightly deflexed, trapezoidal, very transverse,
<quantity id="4CA7BECBAB10FFAAFECCF89BFE195815" box="[375,475,1797,1824]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" pageId="4" pageNumber="3" unit="mm" value="0.51">0.51 mm</quantity>
long at midline, anterior margin
<quantity id="4CA7BECBAB10FFAAFCEFF89BFC7A5815" box="[852,952,1797,1824]" metricMagnitude="-4" metricUnit="m" metricValue="8.6" pageId="4" pageNumber="3" unit="mm" value="0.86">0.86 mm</quantity>
wide, posterior margin
<quantity id="4CA7BECBAB10FFAAFB73F898FAEC5815" box="[1224,1326,1798,1824]" metricMagnitude="-3" metricUnit="m" metricValue="1.28" pageId="4" pageNumber="3" unit="mm" value="1.28">1.28 mm</quantity>
, posterior margin width 2.5 times length; anterior margin broadly emarginate, lateral margin slightly convex, angles backward at about 114 degrees, posterior margin sinuate, pronotal width 1.54 times head width.
<emphasis id="B92BCF3CAB10FFAAFB6CF8D4FAEF5851" box="[1239,1325,1866,1892]" italics="true" pageId="4" pageNumber="3">Thorax.</emphasis>
Length of prosternal process equal to but does not exceed apices of procoxae; metathoracic femur enlarged, saltatorial, length 1.84 times width, compared to mesothoracic femur length 2.58 times width; metatibial long spur 2.6 times longer than short spur; metatibial long spur 1.0 times metatarsomeres IIV combined length. Metatarsomere 1 length 1.3 times metatarsomeres IIV combined length.
<emphasis id="B92BCF3CAB11FFABFCFAFE87FC405E06" box="[833,898,281,307]" italics="true" pageId="5" pageNumber="4">Elytra</emphasis>
. Length
<quantity id="4CA7BECBAB11FFABFC5CFE87FB8C5E01" box="[999,1102,281,308]" metricMagnitude="-3" metricUnit="m" metricValue="2.38" pageId="5" pageNumber="4" unit="mm" value="2.38">2.38 mm</quantity>
, width across both elytra
<quantity id="4CA7BECBAB11FFABFACAFE87FF335E63" metricMagnitude="-3" metricUnit="m" metricValue="1.69" pageId="5" pageNumber="4" unit="mm" value="1.69">1.69 mm</quantity>
, length 1.41 times width; elytral width 2.03 times head width; elytral width 1.32 times pronotal width; elytral height at middle
<quantity id="4CA7BECBAB11FFABFE3FFEC3FE295E42" box="[388,491,349,376]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="5" pageNumber="4" unit="mm" value="0.56">0.56 mm</quantity>
; length 4.25 times longer than height (convexity); lateral margin slightly explanate, greatest width of turned-out rim on anterior 1/3 equals width of antennomere IV, reduced posteriorly; elytral epipleuron width subequal to one dorsal eye width on anterior 1/3, reduced posteriorly.
<emphasis id="B92BCF3CAB11FFABFBCFFE3EFAD55E8F" box="[1140,1303,416,442]" italics="true" pageId="5" pageNumber="4">Scutellar shield</emphasis>
. Triangular, width
<quantity id="4CA7BECBAB11FFABFEB3FE5CFEAF5EE8" box="[264,365,450,477]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="5" pageNumber="4" unit="mm" value="0.32">0.32 mm</quantity>
, length
<quantity id="4CA7BECBAB11FFABFE7FFE5CFDEB5EE8" box="[452,553,450,477]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="5" pageNumber="4" unit="mm" value="0.22">0.22 mm</quantity>
, width 1.43 times length.
<emphasis id="B92BCF3CAB11FFABFCFEFE5CFC6D5EE9" box="[837,943,450,476]" italics="true" pageId="5" pageNumber="4">Abdomen</emphasis>
. Fifth visible abdominal sternite emarginate,
<quantity id="4CA7BECBAB11FFABFF7BFE7AFEE65ECB" box="[192,292,484,510]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="4" unit="mm" value="0.24">0.24 mm</quantity>
wide,
<quantity id="4CA7BECBAB11FFABFECBFE7AFE165ECB" box="[368,468,484,510]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="4" unit="mm" value="0.04">0.04 mm</quantity>
deep, width 6.0 times depth.
<emphasis id="B92BCF3CAB11FFABFC9DFE7AFC245ECB" box="[806,998,484,510]" italics="true" pageId="5" pageNumber="4">Female prehensor</emphasis>
. The female prehensor (bursal sclerite) (
<figureCitation id="13640FABAB11FFABFF71FD98FECB5D15" box="[202,265,518,544]" captionStart="Figures 14" captionStartId="6.[193,280,1866,1892]" captionTargetBox="[274,1341,567,1824]" captionTargetId="figure-174@6.[274,1359,567,1837]" captionTargetPageId="6" captionText="Figures 14. Scirtes goodrichi. 1) Adult dorsal habitus. 2) Female prehensor. 3) Mandible. 4) Prosternal process lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5352380" httpUri="https://zenodo.org/record/5352380/files/figure.png" pageId="5" pageNumber="4">Fig. 2</figureCitation>
) has two pairs of processes directed anteriorly, one pair directed posteriorly and a single tooth at the dorsal midline directed anteriorly. Prehensor width
<quantity id="4CA7BECBAB11FFABFD0AFDB9FCD45D77" box="[689,790,551,578]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="4" unit="mm" value="0.4">0.40 mm</quantity>
, width across both elytra
<quantity id="4CA7BECBAB11FFABFB95FDB9FB505D77" box="[1070,1170,551,578]" metricMagnitude="-3" metricUnit="m" metricValue="1.69" pageId="5" pageNumber="4" unit="mm" value="1.69">1.69 mm</quantity>
, elytral width 4.23 times prehensor width.
</paragraph>
</subSubSection>
<subSubSection id="C34540A5AB11FFABFF7BFDEDFCA45A49" pageId="5" pageNumber="4" type="materials_examined">
<paragraph id="8BE0132EAB11FFABFF7BFDEDFEF25BD9" blockId="5.[191,1442,627,1261]" pageId="5" pageNumber="4">
<emphasis id="B92BCF3CAB11FFABFF7BFDEDFEA85DB8" bold="true" box="[192,362,627,653]" pageId="5" pageNumber="4">Type material.</emphasis>
<materialsCitation id="3B371973AB11FFABFECAFDEDFB725CAE" collectingDate="2011-05-28" collectingDateMax="2020-08-03" collectingDateMin="2011-05-28" collectionCode="UV, TYPE, LOCALITY, NMNH" collectorName="C. A. Springer" county="National Museum of Natural History." latitude="36.913612" location="Female" longLatPrecision="1" longitude="-76.31416" municipality="Norfolk" pageId="5" pageNumber="4" specimenCount="56" specimenCount-female="13" specimenCount-male="12" stateProvince="South Carolina" typeStatus="holotype">
A total of
<specimenCount id="9D59D8A7AB11FFABFE58FDEDFDB85DB8" box="[483,634,627,653]" pageId="5" pageNumber="4" type="generic">24 specimens</specimenCount>
was examined, 17 from the type locality,
<specimenCount id="9D59D8A7AB11FFABFBF5FDEDFB665DB8" box="[1102,1188,627,653]" pageId="5" pageNumber="4" type="male">9 males</specimenCount>
and
<specimenCount id="9D59D8A7AB11FFABFB67FDEDFA8A5DB8" box="[1244,1352,627,653]" pageId="5" pageNumber="4" type="female">8 females</specimenCount>
, all dissected. The
<specimenCount id="9D59D8A7AB11FFABFEFAFD0BFE0B5D9A" box="[321,457,661,687]" pageId="5" pageNumber="4" type="generic">7 specimens</specimenCount>
from
<collectingRegion id="499BDDCCAB11FFABFDB1FD0BFD715D9A" box="[522,691,661,687]" country="United States of America" name="South Carolina" pageId="5" pageNumber="4">South Carolina</collectingRegion>
included
<specimenCount id="9D59D8A7AB11FFABFCA4FD0BFCB15D9A" box="[799,883,661,687]" pageId="5" pageNumber="4" type="male">3 males</specimenCount>
and
<specimenCount id="9D59D8A7AB11FFABFC13FD0BFBD35D9A" box="[936,1041,661,687]" pageId="5" pageNumber="4" type="female">4 females</specimenCount>
, all dissected.
<typeStatus id="54E4AD8CAB11FFABFB16FD0BFA835D9A" box="[1197,1345,661,687]" pageId="5" pageNumber="4" type="holotype">HOLOTYPE</typeStatus>
.
<location id="8E8045F5AB11FFABFAF2FD0BFA5F5D9A" LSID="urn:lsid:plazi:treatment:03F6A238AB10FFA6FF7BFE8AFB4E5D99:8E8045F5AB11FFABFAF2FD0BFA5F5D9A" box="[1353,1437,661,687]" county="National Museum of Natural History." latitude="36.913612" longLatPrecision="1" longitude="-76.31416" municipality="Norfolk" name="Female" pageId="5" pageNumber="4" stateProvince="South Carolina">Female</location>
: labeled, “VA:Norfolk
<date id="FFE135EEAB11FFABFE0FFD29FDE95DE4" box="[436,555,695,721]" pageId="5" pageNumber="4" value="2017-05-19">
<collectingDate id="EFA5CC06AB11FFABFE0FFD29FDE95DE4" box="[436,555,695,721]" pageId="5" pageNumber="4" value="2017-05-19">19-V-2017</collectingDate>
</date>
<collectorName id="26AA76F8AB11FFABFD8FFD29FD115DE5" box="[564,723,694,721]" pageId="5" pageNumber="4">C.A. Springer</collectorName>
”/”
<collectionCode id="ED4E8BEBAB11FFABFD55FD28FCD55DE5" box="[750,791,694,720]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="5" pageNumber="4">UV</collectionCode>
Light”/red label “
<typeStatus id="54E4AD8CAB11FFABFC5EFD29FBB55DE4" box="[997,1143,695,721]" pageId="5" pageNumber="4" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C5F68ADAB11FFABFB3AFD29FEE75DC7" authority="Springer &amp; Waller" authorityName="Springer &amp; Waller" authorityYear="2021" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="goodrichi">
<emphasis id="B92BCF3CAB11FFABFB3AFD29FAFA5DE5" box="[1153,1336,694,721]" italics="true" pageId="5" pageNumber="4">Scirtes goodrichi</emphasis>
Springer &amp; Waller
</taxonomicName>
”.
<location id="8E8045F5AB11FFABFE82FD46FD595DC6" LSID="urn:lsid:plazi:treatment:03F6A238AB10FFA6FF7BFE8AFB4E5D99:8E8045F5AB11FFABFE82FD46FD595DC6" box="[313,667,728,755]" county="National Museum of Natural History." latitude="36.913612" longLatPrecision="1" longitude="-76.31416" municipality="Norfolk" name="Deposited in National Museum" pageId="5" pageNumber="4" stateProvince="South Carolina">
Deposited in
<location id="8E8045F5AB11FFABFE6BFD46FD595DC6" LSID="urn:lsid:plazi:treatment:03F6A238AB10FFA6FF7BFE8AFB4E5D99:8E8045F5AB11FFABFE6BFD46FD595DC6" box="[464,667,728,755]" county="National Museum of Natural History." latitude="36.913612" longLatPrecision="1" longitude="-76.31416" municipality="Norfolk" name="National Museum of Natural History" pageId="5" pageNumber="4" stateProvince="South Carolina">National Museum</location>
</location>
of Natural History,
<collectingRegion id="499BDDCCAB11FFABFCC7FD46FBC75DC7" box="[892,1029,728,754]" country="United States of America" name="Washington" pageId="5" pageNumber="4">Washington</collectingRegion>
, DC.
<collectionCode id="ED4E8BEBAB11FFABFBFDFD47FADA5DC7" box="[1094,1304,728,755]" pageId="5" pageNumber="4">TYPE LOCALITY</collectionCode>
. 1300
<location id="8E8045F5AB11FFABFAE4FD46FE815C21" LSID="urn:lsid:plazi:treatment:03F6A238AB10FFA6FF7BFE8AFB4E5D99:8E8045F5AB11FFABFAE4FD46FE815C21" county="National Museum of Natural History." latitude="36.913612" longLatPrecision="1" longitude="-76.31416" municipality="Norfolk" name="Cloncurry Road" pageId="5" pageNumber="4" stateProvince="South Carolina">Cloncurry Road</location>
,
<collectingMunicipality id="6B848954AB11FFABFEF4FD64FE6B5C21" box="[335,425,762,788]" pageId="5" pageNumber="4">Norfolk</collectingMunicipality>
,
<collectingRegion id="499BDDCCAB11FFABFE0EFD64FDD35C21" box="[437,529,762,788]" country="United States of America" name="Virginia" pageId="5" pageNumber="4">Virginia</collectingRegion>
23505 (
<geoCoordinate id="EE6B75E9AB11FFABFDD7FD64FCE65C21" box="[620,804,762,788]" degrees="36.91361111" direction="north" orientation="latitude" pageId="5" pageNumber="4" precision="1" value="36.913612">36.91361111 °N</geoCoordinate>
,
<geoCoordinate id="EE6B75E9AB11FFABFC8BFD64FC2C5C21" box="[816,1006,762,788]" degrees="76.31416667" direction="west" orientation="longitude" pageId="5" pageNumber="4" precision="1" value="-76.31416">76.31416667 °W</geoCoordinate>
). Elevation: 3.0 m.
<typeStatus id="54E4AD8CAB11FFABFB72FD64FA905C21" box="[1225,1362,762,788]" pageId="5" pageNumber="4" type="allotype">ALLOTYPE</typeStatus>
. Male: labeled “VA: Norfolk
<date id="FFE135EEAB11FFABFE15FC82FDF75C03" box="[430,565,796,822]" pageId="5" pageNumber="4" value="2020-08-03">
<collectingDate id="EFA5CC06AB11FFABFE15FC82FDF75C03" box="[430,565,796,822]" pageId="5" pageNumber="4" value="2020-08-03">3-VIII-2020</collectingDate>
</date>
<collectorName id="26AA76F8AB11FFABFD80FC82FD145C03" box="[571,726,796,822]" pageId="5" pageNumber="4">C.A. Springer</collectorName>
”/”
<collectionCode id="ED4E8BEBAB11FFABFD4AFC82FCD85C03" box="[753,794,796,822]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="5" pageNumber="4">UV</collectionCode>
Light”/blue label “
<typeStatus id="54E4AD8CAB11FFABFC5EFC82FBAF5C03" box="[997,1133,796,822]" pageId="5" pageNumber="4" type="allotype">ALLOTYPE</typeStatus>
<taxonomicName id="4C5F68ADAB11FFABFBD4FC82FECB5C62" authority="Springer &amp; Waller" authorityName="Springer &amp; Waller" authorityYear="2021" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="goodrichi">
<emphasis id="B92BCF3CAB11FFABFBD4FC82FAE05C03" box="[1135,1314,796,822]" italics="true" pageId="5" pageNumber="4">Scirtes goodrichi</emphasis>
Springer &amp; Waller
</taxonomicName>
”. Deposited in the
<collectingCounty id="62816BA2AB11FFABFE62FCA3FCBB5C62" box="[473,889,829,856]" pageId="5" pageNumber="4">National Museum of Natural History.</collectingCounty>
<typeStatus id="54E4AD8CAB11FFABFCC5FCA3FC1C5C62" box="[894,990,829,855]" pageId="5" pageNumber="4" type="allotype">Allotype</typeStatus>
locality same as holotype.
<typeStatus id="54E4AD8CAB11FFABFABFFCA0FA5E5C6D" box="[1284,1436,830,856]" pageId="5" pageNumber="4" type="paratype">PARATYPES</typeStatus>
. The following
<typeStatus id="54E4AD8CAB11FFABFEE6FCFEFE0B5C4F" box="[349,457,864,890]" pageId="5" pageNumber="4" type="paratype">paratypes</typeStatus>
from
<emphasis id="B92BCF3CAB11FFABFDB0FCC1FD175C4C" bold="true" box="[523,725,863,889]" pageId="5" pageNumber="4">
Norfolk,
<collectingRegion id="499BDDCCAB11FFABFDC8FCC1FD175C4C" box="[627,725,863,889]" country="United States of America" name="Virginia" pageId="5" pageNumber="4">Virginia</collectingRegion>
</emphasis>
all share the type locality, collector, collection technique, and are deposited in
<collectionCode id="ED4E8BEBAB11FFABFEEAFC1CFE6E5CA9" box="[337,428,898,924]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="4" type="Museum">NMNH</collectionCode>
. The genders and collection dates are as follows:
<specimenCount id="9D59D8A7AB11FFABFC69FC1FFBF25CAE" box="[978,1072,897,923]" pageId="5" pageNumber="4" type="female">1 female</specimenCount>
<date id="FFE135EEAB11FFABFB83FC1FFB725CAE" box="[1080,1200,897,923]" pageId="5" pageNumber="4" value="2011-05-28">
<collectingDate id="EFA5CC06AB11FFABFB83FC1FFB725CAE" box="[1080,1200,897,923]" pageId="5" pageNumber="4" value="2011-05-28">28-V-2011</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B371973AB11FFABFB07FC1FFA595CAE" box="[1212,1435,897,923]" collectingDate="2013-05-31" collectingDateMax="2020-08-03" collectingDateMin="2011-05-28" collectionCode="UV, TYPE, LOCALITY, NMNH" collectorName="C. A. Springer" county="National Museum of Natural History." isTailEnumeration="true" latitude="36.913612" location="Female" longLatPrecision="1" longitude="-76.31416" municipality="Norfolk" pageId="5" pageNumber="4" specimenCount="1" specimenCount-female="1" stateProvince="South Carolina" typeStatus="holotype">
<specimenCount id="9D59D8A7AB11FFABFB07FC1FFAD95CAE" box="[1212,1307,897,923]" pageId="5" pageNumber="4" type="female">1 female</specimenCount>
<date id="FFE135EEAB11FFABFA99FC1FFA595CAE" box="[1314,1435,897,923]" pageId="5" pageNumber="4" value="2013-05-31">
<collectingDate id="EFA5CC06AB11FFABFA99FC1FFA595CAE" box="[1314,1435,897,923]" pageId="5" pageNumber="4" value="2013-05-31">31-V-2013</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="9D59D8A7AB11FFABFF7AFC3DFEE25C88" box="[193,288,931,957]" pageId="5" pageNumber="4" type="female">1 female</specimenCount>
<date id="FFE135EEAB11FFABFE92FC3DFE7D5C88" box="[297,447,931,957]" pageId="5" pageNumber="4" value="2013-08-28">28-VIII-2013</date>
;
<specimenCount id="9D59D8A7AB11FFABFE77FC3DFDE95C88" box="[460,555,931,957]" pageId="5" pageNumber="4" type="female">1 female</specimenCount>
<date id="FFE135EEAB11FFABFD8FFC3DFD7A5C88" box="[564,696,931,957]" pageId="5" pageNumber="4" value="2014-06-11">11-VI-2014</date>
,
<specimenCount id="9D59D8A7AB11FFABFD7FFC3DFCCD5C88" box="[708,783,931,957]" pageId="5" pageNumber="4" type="male">1 male</specimenCount>
<date id="FFE135EEAB11FFABFCACFC3DFC595C88" box="[791,923,931,957]" pageId="5" pageNumber="4" value="2014-06-17">17-VI-2014</date>
;
<materialsCitation id="3B371973AB11FFABFC13FC3DFB515C88" box="[936,1171,931,957]" collectingDate="2014-06-18" collectingDateMax="2020-08-03" collectingDateMin="2011-05-28" collectionCode="UV, TYPE, LOCALITY, NMNH" collectorName="C. A. Springer" county="National Museum of Natural History." isTailEnumeration="true" latitude="36.913612" location="Female" longLatPrecision="1" longitude="-76.31416" municipality="Norfolk" pageId="5" pageNumber="4" specimenCount="1" specimenCount-female="1" stateProvince="South Carolina" typeStatus="holotype">
<specimenCount id="9D59D8A7AB11FFABFC13FC3DFBC55C88" box="[936,1031,931,957]" pageId="5" pageNumber="4" type="female">1 female</specimenCount>
<date id="FFE135EEAB11FFABFBB4FC3DFB515C88" box="[1039,1171,931,957]" pageId="5" pageNumber="4" value="2014-06-18">
<collectingDate id="EFA5CC06AB11FFABFBB4FC3DFB515C88" box="[1039,1171,931,957]" pageId="5" pageNumber="4" value="2014-06-18">18-VI-2014</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="9D59D8A7AB11FFABFB1BFC3DFB3D5C88" box="[1184,1279,931,957]" pageId="5" pageNumber="4" type="female">1 female</specimenCount>
<date id="FFE135EEAB11FFABFABCFC3DFA455C88" box="[1287,1415,931,957]" pageId="5" pageNumber="4" value="2014-07-03">3-VII-2014</date>
;
<specimenCount id="9D59D8A7AB11FFABFA28FC3DFF3A5CEB" pageId="5" pageNumber="4" type="male">1 male</specimenCount>
<date id="FFE135EEAB11FFABFF44FC5BFE4F5CEA" box="[255,397,965,991]" pageId="5" pageNumber="4" value="2014-07-29">29-VII-2014</date>
;
<specimenCount id="9D59D8A7AB11FFABFE23FC5BFE355CEB" box="[408,503,964,991]" pageId="5" pageNumber="4" type="female">1 female</specimenCount>
<date id="FFE135EEAB11FFABFE44FC5BFD415CEA" box="[511,643,965,991]" pageId="5" pageNumber="4" value="2020-06-24">24-VI-2020</date>
;
<specimenCount id="9D59D8A7AB11FFABFD35FC5BFD1B5CEB" box="[654,729,964,991]" pageId="5" pageNumber="4" type="male">1 male</specimenCount>
<date id="FFE135EEAB11FFABFD5AFC5BFCAD5CEA" box="[737,879,965,991]" pageId="5" pageNumber="4" value="2020-07-12">12-VII-2020</date>
;
<specimenCount id="9D59D8A7AB11FFABFCC1FC5BFC075CEB" box="[890,965,964,991]" pageId="5" pageNumber="4" type="male">1 male</specimenCount>
<date id="FFE135EEAB11FFABFC77FC5BFB945CEA" box="[972,1110,965,991]" pageId="5" pageNumber="4" value="2020-08-03">3-VIII-2020</date>
;
<specimenCount id="9D59D8A7AB11FFABFBDAFC5BFB6E5CEB" box="[1121,1196,964,991]" pageId="5" pageNumber="4" type="male">1 male</specimenCount>
<date id="FFE135EEAB11FFABFB0FFC5BFA895CEA" box="[1204,1355,965,991]" pageId="5" pageNumber="4" value="2020-08-11">11-VIII-2020</date>
;
<specimenCount id="9D59D8A7AB11FFABFAECFC5BFA635CEB" box="[1367,1441,964,991]" pageId="5" pageNumber="4" type="male">1 male</specimenCount>
<date id="FFE135EEAB11FFABFF7AFC78FE9A5B35" box="[193,344,998,1024]" pageId="5" pageNumber="4" value="2020-08-12">12-VIII-2020</date>
;
<specimenCount id="9D59D8A7AB11FFABFED9FC78FE695B35" box="[354,427,998,1024]" pageId="5" pageNumber="4" type="male">1 male</specimenCount>
<date id="FFE135EEAB11FFABFE0AFC78FD8A5B35" box="[433,584,998,1024]" pageId="5" pageNumber="4" value="2020-08-18">18-VIII-2020</date>
;
<materialsCitation id="3B371973AB11FFABFDE9FC78FDCC5B71" collectingDate="2020-08-22" collectorName="M. Caterino &amp; A. Deczynski" country="United States of America" county="Georgetown Co." latitude="33.3302" location="Georgetown Co." longLatPrecision="6" longitude="-79.2469" pageId="5" pageNumber="4" specimenCount="2" specimenCount-male="1" stateProvince="South Carolina" typeStatus="holotype">
<specimenCount id="9D59D8A7AB11FFABFDE9FC78FD5E5B35" box="[594,668,998,1024]" pageId="5" pageNumber="4" type="male">1 male</specimenCount>
<date id="FFE135EEAB11FFABFD19FC78FCF85B35" box="[674,826,998,1024]" pageId="5" pageNumber="4" value="2020-08-22">
<collectingDate id="EFA5CC06AB11FFABFD19FC78FCF85B35" box="[674,826,998,1024]" pageId="5" pageNumber="4" value="2020-08-22">22-VIII-2020</collectingDate>
</date>
.
<emphasis id="B92BCF3CAB11FFABFCFAFC78FC315B35" bold="true" box="[833,1011,998,1024]" pageId="5" pageNumber="4">
<collectingRegion id="499BDDCCAB11FFABFCFAFC78FC315B35" box="[833,1011,998,1024]" country="United States of America" name="South Carolina" pageId="5" pageNumber="4">South Carolina</collectingRegion>
</emphasis>
: the next
<specimenCount id="9D59D8A7AB11FFABFBE4FC78FACB5B35" box="[1119,1289,998,1024]" pageId="5" pageNumber="4" type="generic">four specimens</specimenCount>
share the following collection data: “
<collectingCountry id="F34853BEAB11FFABFE74FB96FDCB5B17" box="[463,521,1032,1058]" name="United States of America" pageId="5" pageNumber="4">USA</collectingCountry>
: SC:
<collectingCounty id="62816BA2AB11FFABFDFBFB96FD3B5B17" box="[576,761,1032,1058]" pageId="5" pageNumber="4">Georgetown Co.</collectingCounty>
<geoCoordinate id="EE6B75E9AB11FFABFCBBFB96FCBD5B16" box="[768,895,1032,1059]" degrees="33.3302" direction="north" orientation="latitude" pageId="5" pageNumber="4" precision="5" value="33.3302">33.3302 °N</geoCoordinate>
,
<geoCoordinate id="EE6B75E9AB11FFABFC31FB96FBCF5B16" box="[906,1037,1032,1059]" degrees="79.2469" direction="west" orientation="longitude" pageId="5" pageNumber="4" precision="5" value="-79.2469">79.2469 °W</geoCoordinate>
Baruch/Hobcaw Barony VII.9.2015
<collectorName id="26AA76F8AB11FFABFF7AFBB4FE885B71" box="[193,330,1066,1092]" pageId="5" pageNumber="4">M. Caterino</collectorName>
&amp;
<collectorName id="26AA76F8AB11FFABFED6FBB4FDC45B71" box="[365,518,1066,1092]" pageId="5" pageNumber="4">A. Deczynski</collectorName>
</materialsCitation>
;
<materialsCitation id="3B371973AB11FFABFDA1FBB4FBC05B71" box="[538,1026,1066,1093]" collectionCode="CUAC" country="United States of America" pageId="5" pageNumber="4" specimenCount="1" specimenCount-male="1" stateProvince="South Carolina" typeStatus="holotype">
<specimenCount id="9D59D8A7AB11FFABFDA1FBB4FDA75B71" box="[538,613,1066,1092]" pageId="5" pageNumber="4" type="male">1 male</specimenCount>
“Clemson University 13450”,
<collectionCode id="ED4E8BEBAB11FFABFC0EFBB4FBC05B71" box="[949,1026,1066,1092]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" pageId="5" pageNumber="4" type="University or college">CUAC</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B371973AB11FFABFBB6FBB4FECC5B53" collectionCode="CUAC" country="United States of America" pageId="5" pageNumber="4" specimenCount="1" specimenCount-male="1" stateProvince="South Carolina" typeStatus="holotype">
<specimenCount id="9D59D8A7AB11FFABFBB6FBB4FB9A5B71" box="[1037,1112,1066,1092]" pageId="5" pageNumber="4" type="male">1 male</specimenCount>
“Clemson University 13021”,
<collectionCode id="ED4E8BEBAB11FFABFF7AFBD2FECC5B53" box="[193,270,1100,1126]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" pageId="5" pageNumber="4" type="University or college">CUAC</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B371973AB11FFABFEA3FBD2FD3E5B53" box="[280,764,1099,1126]" collectionCode="CUAC" country="United States of America" pageId="5" pageNumber="4" specimenCount="1" specimenCount-male="1" stateProvince="South Carolina" typeStatus="holotype">
<specimenCount id="9D59D8A7AB11FFABFEA3FBD2FEA05B50" box="[280,354,1099,1126]" pageId="5" pageNumber="4" type="male">1 male</specimenCount>
“Clemson University 13275”,
<collectionCode id="ED4E8BEBAB11FFABFD14FBD2FD3E5B53" box="[687,764,1100,1126]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" pageId="5" pageNumber="4" type="University or college">CUAC</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B371973AB11FFABFCBDFBD2FAC05B53" box="[774,1282,1099,1126]" collectionCode="CUCA" country="United States of America" pageId="5" pageNumber="4" specimenCount="1" specimenCount-female="1" stateProvince="South Carolina" typeStatus="holotype">
<specimenCount id="9D59D8A7AB11FFABFCBDFBD2FCA75B50" box="[774,869,1099,1126]" pageId="5" pageNumber="4" type="female">1 female</specimenCount>
“Clemson University 13023”,
<collectionCode id="ED4E8BEBAB11FFABFB09FBD2FAC05B53" box="[1202,1282,1100,1126]" pageId="5" pageNumber="4">CUCA</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B371973AB11FFABFAB7FBD2FB575BB2" collectionCode="V, CUAC" collectorName="Berkeley Co. Cypress Gardens &amp; Coll. J. C. Ciegler" country="United States of America" pageId="5" pageNumber="4" specimenCount="1" specimenCount-female="1" stateProvince="South Carolina" typeStatus="holotype">
<specimenCount id="9D59D8A7AB11FFABFAB7FBD2FAA85B50" box="[1292,1386,1099,1126]" pageId="5" pageNumber="4" type="female">1 female</specimenCount>
“SC:
<collectorName id="26AA76F8AB11FFABFF7BFBF3FDCD5BB2" box="[192,527,1133,1159]" pageId="5" pageNumber="4">Berkeley Co. Cypress Gardens</collectorName>
PIN-V-8/9-98”/”At U.
<collectionCode id="ED4E8BEBAB11FFABFCBDFBF0FCD85BBD" box="[774,794,1134,1160]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="4" type="Museum">V</collectionCode>
. Light”/”
<collectorName id="26AA76F8AB11FFABFCC5FBF3FBF45BB2" box="[894,1078,1133,1159]" pageId="5" pageNumber="4">Coll. J.C. Ciegler</collectorName>
”,
<collectionCode id="ED4E8BEBAB11FFABFBF3FBF3FB575BB2" box="[1096,1173,1133,1159]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" pageId="5" pageNumber="4" type="University or college">CUAC</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B371973AB11FFABFB25FBF3FCB35B9C" collectingDate="2004-09-17" collectionCode="V" collectorName="Coll. J. C. Ciegler" country="United States of America" location="Light" pageId="5" pageNumber="4" specimenCount="1" specimenCount-female="1" stateProvince="South Carolina" typeStatus="holotype">
<specimenCount id="9D59D8A7AB11FFABFB25FBF3FB3E5BB2" box="[1182,1276,1133,1159]" pageId="5" pageNumber="4" type="female">1 female</specimenCount>
“Moncks Corner, S.C.
<date id="FFE135EEAB11FFABFE92FB11FE5A5B9C" box="[297,408,1167,1193]" pageId="5" pageNumber="4" value="2004-09-17">
<collectingDate id="EFA5CC06AB11FFABFE92FB11FE5A5B9C" box="[297,408,1167,1193]" pageId="5" pageNumber="4" value="2004-09-17">9/17/2004</collectingDate>
</date>
”/”At U.
<collectionCode id="ED4E8BEBAB11FFABFE4AFB0EFDC75B9F" box="[497,517,1168,1194]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="4" type="Museum">V</collectionCode>
.
<location id="8E8045F5AB11FFABFDAAFB11FD925B9C" LSID="urn:lsid:plazi:treatment:03F6A238AB10FFA6FF7BFE8AFB4E5D99:8E8045F5AB11FFABFDAAFB11FD925B9C" box="[529,592,1167,1193]" country="United States of America" name="Light" pageId="5" pageNumber="4" stateProvince="South Carolina">Light</location>
”/”
<collectorName id="26AA76F8AB11FFABFDD7FB11FCE85B9C" box="[620,810,1167,1193]" pageId="5" pageNumber="4">Coll. J.C. Ciegler</collectorName>
”, JCC
</materialsCitation>
;
<materialsCitation id="3B371973AB11FFABFCC5FB11FEEE5BD9" collectionCode="V" collectorName="Coll. J. C. Ciegler &amp; All" country="United States of America" location="Light" pageId="5" pageNumber="4" specimenCount="1" specimenCount-female="1" stateProvince="South Carolina" typeStatus="paratype">
<specimenCount id="9D59D8A7AB11FFABFCC5FB11FC1C5B9C" box="[894,990,1167,1193]" pageId="5" pageNumber="4" type="female">1 female</specimenCount>
“Moncks Corner, S.C. 5/8/98”/”At U.
<collectionCode id="ED4E8BEBAB11FFABFA32FB0EFA5F5B9F" box="[1417,1437,1168,1194]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="4" type="Museum">V</collectionCode>
.
<location id="8E8045F5AB11FFABFF7BFB2FFF3D5BFE" LSID="urn:lsid:plazi:treatment:03F6A238AB10FFA6FF7BFE8AFB4E5D99:8E8045F5AB11FFABFF7BFB2FFF3D5BFE" box="[192,255,1201,1227]" country="United States of America" name="Light" pageId="5" pageNumber="4" stateProvince="South Carolina">Light</location>
”/”
<collectorName id="26AA76F8AB11FFABFEA1FB2FFE135BFE" box="[282,465,1201,1227]" pageId="5" pageNumber="4">Coll. J.C. Ciegler</collectorName>
”, JCC.
<collectorName id="26AA76F8AB11FFABFDA6FB2FFDFD5BFE" box="[541,575,1201,1227]" pageId="5" pageNumber="4">All</collectorName>
specimens with yellow
<typeStatus id="54E4AD8CAB11FFABFCF9FB2FFC615BFE" box="[834,931,1201,1227]" pageId="5" pageNumber="4" type="paratype">paratype</typeStatus>
labels, “
<typeStatus id="54E4AD8CAB11FFABFC40FB2FFB465BFE" box="[1019,1156,1201,1227]" pageId="5" pageNumber="4" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C5F68ADAB11FFABFB32FB2FFEE15BD9" authority="Springer &amp; Waller" authorityName="Springer &amp; Waller" authorityYear="2021" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="goodrichi">
<emphasis id="B92BCF3CAB11FFABFB32FB2FFAF95BFE" box="[1161,1339,1201,1227]" italics="true" pageId="5" pageNumber="4">Scirtes goodrichi</emphasis>
Springer &amp; Waller
</taxonomicName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BE0132EAB11FFABFF7BFB62FCA45A49" blockId="5.[192,1441,1276,1404]" pageId="5" pageNumber="4">
<emphasis id="B92BCF3CAB11FFABFF7BFB62FE6C5A23" bold="true" box="[192,430,1276,1302]" pageId="5" pageNumber="4">Geographical range.</emphasis>
Known from only two states,
<taxonomicName id="4C5F68ADAB11FFABFCBAFB63FC405A23" authorityName="Springer &amp; Waller" authorityYear="2021" box="[769,898,1276,1303]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="goodrichi">
<emphasis id="B92BCF3CAB11FFABFCBAFB63FC405A23" box="[769,898,1276,1303]" italics="true" pageId="5" pageNumber="4">S. goodrichi</emphasis>
</taxonomicName>
first came to our attention with specimens from the
<typeStatus id="54E4AD8CAB11FFABFF54FA81FEDD5A0C" box="[239,287,1311,1337]" pageId="5" pageNumber="4">type</typeStatus>
locality in
<collectingRegion id="499BDDCCAB11FFABFE13FA80FDC05A0D" box="[424,514,1310,1336]" country="United Kingdom" name="Norfolk" pageId="5" pageNumber="4">Norfolk</collectingRegion>
,
<collectingRegion id="499BDDCCAB11FFABFDAAFA80FDAD5A0D" box="[529,623,1310,1336]" country="United States of America" name="Virginia" pageId="5" pageNumber="4">Virginia</collectingRegion>
, that eventually totaled 17. Loans from Clemson University and Janet Cieglers personal collection yielded seven more specimens from eastern
<collectingRegion id="499BDDCCAB11FFABFC55FADEFB585A6F" box="[1006,1178,1344,1370]" country="United States of America" name="South Carolina" pageId="5" pageNumber="4">South Carolina</collectingRegion>
, including one location in Berkeley County and one location in Georgetown County.
</paragraph>
</subSubSection>
<subSubSection id="C34540A5AB11FFA6FF7BFA12FB4E5D99" lastPageId="8" lastPageNumber="7" pageId="5" pageNumber="4" type="etymology">
<paragraph id="8BE0132EAB11FFABFF7BFA12FA495AFD" blockId="5.[191,1440,1419,1480]" pageId="5" pageNumber="4">
<emphasis id="B92BCF3CAB11FFABFF7BFA12FE805A93" bold="true" box="[192,322,1420,1446]" pageId="5" pageNumber="4">Etymology.</emphasis>
The species is named for the late Michael Goodrich, Eastern
<collectingRegion id="499BDDCCAB11FFABFC6DFA15FBE05A90" box="[982,1058,1419,1445]" country="United States of America" name="Illinois" pageId="5" pageNumber="4">Illinois</collectingRegion>
University, Charleston,
<collectingRegion id="499BDDCCAB11FFABFA9EFA15FAB15A90" box="[1317,1395,1419,1445]" country="United States of America" name="Illinois" pageId="5" pageNumber="4">Illinois</collectingRegion>
, the senior authors friend and research associate from 19612015. The name is a masculine noun in the genitive case.
</paragraph>
<paragraph id="8BE0132EAB11FFA6FF7BFA49FE945E8C" blockId="5.[191,1442,1495,1927]" lastBlockId="8.[191,1441,214,441]" lastPageId="8" lastPageNumber="7" pageId="5" pageNumber="4">
<emphasis id="B92BCF3CAB11FFABFF7BFA49FDFB5AC4" bold="true" box="[192,569,1495,1521]" pageId="5" pageNumber="4">Range of intraspecific variation.</emphasis>
<emphasis id="B92BCF3CAB11FFABFDFBFA49FDBA5AC4" box="[576,632,1495,1521]" italics="true" pageId="5" pageNumber="4">Body</emphasis>
. Smallest specimen: male, Georgetown Co.,
<collectingRegion id="499BDDCCAB11FFABFBD0FA49FAD55AC4" box="[1131,1303,1495,1521]" country="United States of America" name="South Carolina" pageId="5" pageNumber="4">South Carolina</collectingRegion>
, VII.9.2015,
<geoCoordinate id="EE6B75E9AB11FFABFF7AFA67FE885926" box="[193,330,1529,1555]" degrees="33. 3302" direction="north" orientation="latitude" pageId="5" pageNumber="4" precision="5" value="33.3302">33. 3302 °N</geoCoordinate>
<geoCoordinate id="EE6B75E9AB11FFABFEE8FA67FE205926" box="[339,482,1529,1555]" degrees="79. 2469" direction="west" orientation="longitude" pageId="5" pageNumber="4" precision="5" value="-79.2469">79. 2469 °W</geoCoordinate>
, Baruch/Hobcaw Barony, M. Caterino &amp; A. Deczynski, Clemson University 13021. Largest specimen: female, VA:
<collectingRegion id="499BDDCCAB11FFABFDA9F984FDAE5901" box="[530,620,1562,1588]" country="United Kingdom" name="Norfolk" pageId="5" pageNumber="4">Norfolk</collectingRegion>
,
<date id="FFE135EEAB11FFABFDCCF985FD395900" box="[631,763,1563,1589]" pageId="5" pageNumber="4" value="2014-06-11">11-VI-2014</date>
, C. A. Springer. Size range in normal deflexed position from anterior margin of pronotum to apex of elytra; body length
<quantity id="4CA7BECBAB11FFABFCD1F9A2FBCD5962" box="[874,1039,1596,1623]" metricMagnitude="-3" metricUnit="m" metricValue="2.745" metricValueMax="3.1" metricValueMin="2.39" pageId="5" pageNumber="4" unit="mm" value="2.745" valueMax="3.1" valueMin="2.39">2.393.10 mm</quantity>
, width across both elytra
<quantity id="4CA7BECBAB11FFABFA8EF9A2FF30594D" metricMagnitude="-3" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.97" metricValueMin="1.63" pageId="5" pageNumber="4" unit="mm" value="1.7999999999999998" valueMax="1.97" valueMin="1.63">1.631.97 mm</quantity>
, length 1.471.65 times width. Head width across both eyes
<quantity id="4CA7BECBAB11FFABFC2EF9C0FBFA594D" box="[917,1080,1630,1656]" metricMagnitude="-4" metricUnit="m" metricValue="8.25" metricValueMax="8.8" metricValueMin="7.7" pageId="5" pageNumber="4" unit="mm" value="0.825" valueMax="0.88" valueMin="0.77">0.770.88 mm</quantity>
, pronotal width across posterior margin
<quantity id="4CA7BECBAB11FFABFEACF91EFE7B59AF" box="[279,441,1664,1690]" metricMagnitude="-3" metricUnit="m" metricValue="1.29" metricValueMax="1.38" metricValueMin="1.2" pageId="5" pageNumber="4" unit="mm" value="1.29" valueMax="1.38" valueMin="1.2">1.201.38 mm</quantity>
, pronotal width 1.521.62 times head width, elytral width 1.301.47 times pronotal width.
<emphasis id="B92BCF3CAB11FFABFF04F93FFEF6598E" box="[191,308,1697,1723]" italics="true" pageId="5" pageNumber="4">Coloration</emphasis>
. Base color dorsally ranges from brown to black. Width of orange-brown sutural stripe varies from two dorsal eye widths on each elytron to completely absent.
<emphasis id="B92BCF3CAB11FFABFCE1F95DFC5759E8" box="[858,917,1731,1757]" italics="true" pageId="5" pageNumber="4">Head</emphasis>
. Clypeal emargination 1325 times wider than deep; dorsal interocular distance 2.753.75 times one dorsal eye width; subocular carina height equals one to three eye facet diameters, more strongly developed laterally than medially; antennal length
<quantity id="4CA7BECBAB11FFABFB16F899FA8C5814" box="[1197,1358,1799,1825]" metricMagnitude="-3" metricUnit="m" metricValue="1.175" metricValueMax="1.28" metricValueMin="1.07" pageId="5" pageNumber="4" unit="mm" value="1.175" valueMax="1.28" valueMin="1.07">1.071.28 mm</quantity>
, antennal length 1.311.66 times head width, body length 1.952.70 times antennal length, elytral width 2.032.36 times head width; eye color highly variable, can be all argenteous, black, grey or magenta; bicolored eyes include a background of the previous colors with contrasting spots or variegations, or black with argenteous or magenta rim.
<emphasis id="B92BCF3CAB12FFA8FF4CFF48FEBA5FC5" box="[247,376,214,240]" italics="true" pageId="6" pageNumber="5">Mouthparts</emphasis>
. Labrum 1.902.20 times wider than long, anterolateral angles broadly rounded, anterior margin slightly convex. Mandibles unidentate (
<figureCitation id="13640FABAB12FFA8FD12FF69FD2F5E24" box="[681,749,247,273]" captionStart="Figures 14" captionStartId="6.[193,280,1866,1892]" captionTargetBox="[274,1341,567,1824]" captionTargetId="figure-174@6.[274,1359,567,1837]" captionTargetPageId="6" captionText="Figures 14. Scirtes goodrichi. 1) Adult dorsal habitus. 2) Female prehensor. 3) Mandible. 4) Prosternal process lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5352380" httpUri="https://zenodo.org/record/5352380/files/figure.png" pageId="6" pageNumber="5">Fig. 3</figureCitation>
), tooth length equals 1/41/3 greatest width of mandible, teeth overlap each other (left over right) in closed position, denticles absent. Maxillary palps with 4 palpomeres, I shortest, II widest of all and usually slightly longer than, or subequal to III; IV longest, 1.25 times longer than II, attenuate at apex. Labial palps with 3 palpomeres, I shortest, II longest, more than 2 times longer than I and 1.25 times longer than III, III arises subapically from middle of II on medial side, II and III rounded at apex, all palpomeres subequal in greatest width.
<emphasis id="B92BCF3CAB12FFA8FD3CFE01FD175E8C" box="[647,725,415,441]" italics="true" pageId="6" pageNumber="5">Thorax</emphasis>
. Prosternal process (
<figureCitation id="13640FABAB12FFA8FC7EFE01FBCA5E8C" box="[965,1032,415,441]" captionStart="Figures 14" captionStartId="6.[193,280,1866,1892]" captionTargetBox="[274,1341,567,1824]" captionTargetId="figure-174@6.[274,1359,567,1837]" captionTargetPageId="6" captionText="Figures 14. Scirtes goodrichi. 1) Adult dorsal habitus. 2) Female prehensor. 3) Mandible. 4) Prosternal process lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5352380" httpUri="https://zenodo.org/record/5352380/files/figure.png" pageId="6" pageNumber="5">Fig. 4</figureCitation>
) with anterior margin convex, posterior margin concave, both converging posteriorly to a rounded apex, length of prosternal process equal to, but does not exceed, apices of procoxae; metafemoral length 1.761.97 times width; metatibial long spur length 1.972.60 times short spur length; metatibial long spur length 0.691.16 times metatarsomeres IIV combined length; metatarsomere 1 length 1.231.48 times metatarsomeres IIV combined length; all femora and tibiae can be lighter in color at both ends, as well as all trochanters.
<emphasis id="B92BCF3CAB13FFA9FCC0FE87FC7F5E06" box="[891,957,281,307]" italics="true" pageId="7" pageNumber="6">Elytra</emphasis>
. Elytral length 4.105.58 times mid-elytral height (convexity).
<emphasis id="B92BCF3CAB13FFA9FE25FEA4FD875E61" box="[414,581,314,340]" italics="true" pageId="7" pageNumber="6">Scutellar shield</emphasis>
. Triangular, range of width
<quantity id="4CA7BECBAB13FFA9FC30FEA5FBF35E60" box="[907,1073,315,341]" metricMagnitude="-4" metricUnit="m" metricValue="2.95" metricValueMax="3.2" metricValueMin="2.7" pageId="7" pageNumber="6" unit="mm" value="0.295" valueMax="0.32" valueMin="0.27">0.270.32 mm</quantity>
, range of length
<quantity id="4CA7BECBAB13FFA9FB4DFEA4FA5E5E60" box="[1270,1436,314,341]" metricMagnitude="-4" metricUnit="m" metricValue="2.3499999999999996" metricValueMax="2.5" metricValueMin="2.2" pageId="7" pageNumber="6" unit="mm" value="0.235" valueMax="0.25" valueMin="0.22">0.220.25 mm</quantity>
, width 1.161.50 times length.
<emphasis id="B92BCF3CAB13FFA9FDA8FEC2FDBE5E43" box="[531,636,348,374]" italics="true" pageId="7" pageNumber="6">Abdomen</emphasis>
. Emargination of 5th abdominal sternite, width for males 4.68.6 times depth and females 5.815.0 times depth.
<emphasis id="B92BCF3CAB13FFA9FD37FEE0FC595EAD" box="[652,923,382,408]" italics="true" pageId="7" pageNumber="6">Secondary sex characters</emphasis>
. No secondary sex characteristics were recognized.
<emphasis id="B92BCF3CAB13FFA9FEB1FE01FE675E8C" box="[266,421,415,441]" italics="true" pageId="7" pageNumber="6">Male genitalia</emphasis>
. The aedeagus is composed of an outer tegmen (
<figureCitation id="13640FABAB13FFA9FC7BFE01FBDF5E8F" box="[960,1053,415,442]" captionStart="Figures 510" captionStartId="7.[193,280,1765,1791]" captionTargetBox="[204,1425,836,1742]" captionTargetId="figure-305@7.[189,1440,810,1749]" captionTargetPageId="7" captionText="Figures 510. Scirtes goodrichi. 5) Male tegmen lateral view illustrating lateral lobe (paramere) and strut. 6) Male tegmen dorsal view illustrating lateral lobe (paramere) and strut. 7) Male penis, lateral view illustrating basal body (pala), median lobe (parameroid) and serrate process (trigonium). 8) Male penis, dorsal view illustrating basal body (pala), median lobe (parameroid) and serrate process (trigonium). 9) Male tergite VIII. 10) Male tergite IX. Scale lines = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5352382" httpUri="https://zenodo.org/record/5352382/files/figure.png" pageId="7" pageNumber="6">Fig. 56</figureCitation>
) surrounding the inner penis (
<figureCitation id="13640FABAB13FFA9FACFFE01FF2F5EEE" captionStart="Figures 510" captionStartId="7.[193,280,1765,1791]" captionTargetBox="[204,1425,836,1742]" captionTargetId="figure-305@7.[189,1440,810,1749]" captionTargetPageId="7" captionText="Figures 510. Scirtes goodrichi. 5) Male tegmen lateral view illustrating lateral lobe (paramere) and strut. 6) Male tegmen dorsal view illustrating lateral lobe (paramere) and strut. 7) Male penis, lateral view illustrating basal body (pala), median lobe (parameroid) and serrate process (trigonium). 8) Male penis, dorsal view illustrating basal body (pala), median lobe (parameroid) and serrate process (trigonium). 9) Male tergite VIII. 10) Male tergite IX. Scale lines = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5352382" httpUri="https://zenodo.org/record/5352382/files/figure.png" pageId="7" pageNumber="6">Fig. 78</figureCitation>
). Tegmen laterally flattened, with two lateral lobes (parameres) directed posteriorly and a dorsal strut directed anteriorly, tegmen closed dorsally, open ventrally. The penis is composed of a large supporting basal body (pala) with two movable processes directed posteriorly. The median lobe (parameroid) arises subapically from the right side of basal body, with a large apical tooth directed dorsally, tooth is 1.01.7 times width of median lobe. The serrate process (trigonium) arises apically from the left side of basal body, with a row of 1518 small ventral teeth, apex rounded, without large apical tooth. Males dissected, 12.
<emphasis id="B92BCF3CAB13FFA9FC3AFDF7FB5A5DB6" box="[897,1176,617,643]" italics="true" pageId="7" pageNumber="6">Male tergites VIII and IX</emphasis>
. Male tergite VIII (
<figureCitation id="13640FABAB13FFA9FACFFDF7FF125D90" captionStart="Figures 510" captionStartId="7.[193,280,1765,1791]" captionTargetBox="[204,1425,836,1742]" captionTargetId="figure-305@7.[189,1440,810,1749]" captionTargetPageId="7" captionText="Figures 510. Scirtes goodrichi. 5) Male tegmen lateral view illustrating lateral lobe (paramere) and strut. 6) Male tegmen dorsal view illustrating lateral lobe (paramere) and strut. 7) Male penis, lateral view illustrating basal body (pala), median lobe (parameroid) and serrate process (trigonium). 8) Male penis, dorsal view illustrating basal body (pala), median lobe (parameroid) and serrate process (trigonium). 9) Male tergite VIII. 10) Male tergite IX. Scale lines = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5352382" httpUri="https://zenodo.org/record/5352382/files/figure.png" pageId="7" pageNumber="6">Fig. 9</figureCitation>
), has two longitudinal apodemes that curve ventrally at apex, connected by a cross-bar at their midpoint, these three rods support an apical membrane between them with posterior margin deeply concave; tergite IX (
<figureCitation id="13640FABAB13FFA9FAF2FD32FA555DF3" box="[1353,1431,684,710]" captionStart="Figures 510" captionStartId="7.[193,280,1765,1791]" captionTargetBox="[204,1425,836,1742]" captionTargetId="figure-305@7.[189,1440,810,1749]" captionTargetPageId="7" captionText="Figures 510. Scirtes goodrichi. 5) Male tegmen lateral view illustrating lateral lobe (paramere) and strut. 6) Male tegmen dorsal view illustrating lateral lobe (paramere) and strut. 7) Male penis, lateral view illustrating basal body (pala), median lobe (parameroid) and serrate process (trigonium). 8) Male penis, dorsal view illustrating basal body (pala), median lobe (parameroid) and serrate process (trigonium). 9) Male tergite VIII. 10) Male tergite IX. Scale lines = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5352382" httpUri="https://zenodo.org/record/5352382/files/figure.png" pageId="7" pageNumber="6">Fig. 10</figureCitation>
) has two longitudinal apodemes supporting an apical membrane, with posterior margin convex, anterior margin concave. Tergite VIII is different from all other
<taxonomicName id="4C5F68ADAB13FFA9FD6DFD6EFCDF5C3F" authorityName="Illiger" authorityYear="1807" box="[726,797,752,778]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B92BCF3CAB13FFA9FD6DFD6EFCDF5C3F" box="[726,797,752,778]" italics="true" pageId="7" pageNumber="6">Scirtes</emphasis>
</taxonomicName>
in our area except
<taxonomicName id="4C5F68ADAB13FFA9FC4EFD6EFB4A5C3C" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[1013,1160,751,778]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="orbiculatus">
<emphasis id="B92BCF3CAB13FFA9FC4EFD6EFB4A5C3C" box="[1013,1160,751,778]" italics="true" pageId="7" pageNumber="6">S. orbiculatus</emphasis>
</taxonomicName>
, and tergite IX is unique among
<taxonomicName id="4C5F68ADAB1CFFA6FEA9FF48FE9B5FC5" authorityName="Illiger" authorityYear="1807" box="[274,345,214,240]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B92BCF3CAB1CFFA6FEA9FF48FE9B5FC5" box="[274,345,214,240]" italics="true" pageId="8" pageNumber="7">Scirtes</emphasis>
</taxonomicName>
.
<emphasis id="B92BCF3CAB1CFFA6FEDEFF48FDE65FC5" box="[357,548,214,240]" italics="true" pageId="8" pageNumber="7">Female prehensor</emphasis>
. The female prehensor (bursal sclerite) (
<figureCitation id="13640FABAB1CFFA6FC65FF48FBDC5FC5" box="[990,1054,214,240]" captionStart="Figures 14" captionStartId="6.[193,280,1866,1892]" captionTargetBox="[274,1341,567,1824]" captionTargetId="figure-174@6.[274,1359,567,1837]" captionTargetPageId="6" captionText="Figures 14. Scirtes goodrichi. 1) Adult dorsal habitus. 2) Female prehensor. 3) Mandible. 4) Prosternal process lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5352380" httpUri="https://zenodo.org/record/5352380/files/figure.png" pageId="8" pageNumber="7">Fig. 2</figureCitation>
) is a well sclerotized structure usually found at the midline within abdominal segments two to four, but in gravid individuals the presence of eggs can displace it to other areas. The central structure is a transverse bar that supports three pairs of processes and one unpaired tooth; the inner anterior pair of processes are the longest, the lateral anterior pair are the smallest, and the lateral posterior pair are intermediate in size; a single dorsoventrally flattened tooth at the dorsal midline points anteriorly. Prehensor width range
<quantity id="4CA7BECBAB1CFFA6FD28FEE0FCF55EAD" box="[659,823,382,408]" metricMagnitude="-4" metricUnit="m" metricValue="3.65" metricValueMax="4.1" metricValueMin="3.2" pageId="8" pageNumber="7" unit="mm" value="0.365" valueMax="0.41" valueMin="0.32">0.320.41 mm</quantity>
, elytral width 4.05.5 times prehensor width. Females dissected, 12.
</paragraph>
<caption id="DF2043A6AB12FFA8FF7AF8D4FE9B58B2" ID-DOI="http://doi.org/10.5281/zenodo.5352380" ID-Zenodo-Dep="5352380" httpUri="https://zenodo.org/record/5352380/files/figure.png" pageId="6" pageNumber="5" startId="6.[193,280,1866,1892]" targetBox="[274,1341,567,1824]" targetPageId="6">
<paragraph id="8BE0132EAB12FFA8FF7AF8D4FE9B58B2" blockId="6.[193,1440,1866,1927]" pageId="6" pageNumber="5">
<emphasis id="B92BCF3CAB12FFA8FF7AF8D4FE935851" bold="true" box="[193,337,1866,1892]" pageId="6" pageNumber="5">Figures 14.</emphasis>
<taxonomicName id="4C5F68ADAB12FFA8FEECF8D5FDC95851" authorityName="Springer &amp; Waller" authorityYear="2021" box="[343,523,1866,1893]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="goodrichi">
<emphasis id="B92BCF3CAB12FFA8FEECF8D5FDC95851" box="[343,523,1866,1893]" italics="true" pageId="6" pageNumber="5">Scirtes goodrichi</emphasis>
</taxonomicName>
.
<emphasis id="B92BCF3CAB12FFA8FDA3F8D4FDF25851" bold="true" box="[536,560,1866,1892]" pageId="6" pageNumber="5">1)</emphasis>
Adult dorsal habitus.
<emphasis id="B92BCF3CAB12FFA8FC93F8D4FC825851" bold="true" box="[808,832,1866,1892]" pageId="6" pageNumber="5">2)</emphasis>
Female prehensor.
<emphasis id="B92BCF3CAB12FFA8FBA7F8D4FBF65851" bold="true" box="[1052,1076,1866,1892]" pageId="6" pageNumber="5">3)</emphasis>
Mandible.
<emphasis id="B92BCF3CAB12FFA8FB08F8D4FB0E5851" bold="true" box="[1203,1228,1866,1892]" pageId="6" pageNumber="5">4)</emphasis>
Prosternal process lateral aspect.
</paragraph>
</caption>
<caption id="DF2043A6AB13FFA9FF7AF97BFDC658B2" ID-DOI="http://doi.org/10.5281/zenodo.5352382" ID-Zenodo-Dep="5352382" httpUri="https://zenodo.org/record/5352382/files/figure.png" pageId="7" pageNumber="6" startId="7.[193,280,1765,1791]" targetBox="[204,1425,836,1742]" targetPageId="7">
<paragraph id="8BE0132EAB13FFA9FF7AF97BFDC658B2" blockId="7.[192,1440,1765,1927]" pageId="7" pageNumber="6">
<emphasis id="B92BCF3CAB13FFA9FF7AF97BFE9E59CA" bold="true" box="[193,348,1765,1791]" pageId="7" pageNumber="6">Figures 510.</emphasis>
<taxonomicName id="4C5F68ADAB13FFA9FEDAF978FDD65835" authorityName="Springer &amp; Waller" authorityYear="2021" box="[353,532,1766,1792]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="goodrichi">
<emphasis id="B92BCF3CAB13FFA9FEDAF978FDD65835" box="[353,532,1766,1792]" italics="true" pageId="7" pageNumber="6">Scirtes goodrichi</emphasis>
</taxonomicName>
.
<emphasis id="B92BCF3CAB13FFA9FD9BF97BFDFA59CA" bold="true" box="[544,568,1765,1791]" pageId="7" pageNumber="6">5)</emphasis>
Male tegmen lateral view illustrating lateral lobe (paramere) and strut.
<emphasis id="B92BCF3CAB13FFA9FAF0F97BFAA159CA" bold="true" box="[1355,1379,1765,1791]" pageId="7" pageNumber="6">6)</emphasis>
Male tegmen dorsal view illustrating lateral lobe (paramere) and strut.
<emphasis id="B92BCF3CAB13FFA9FC16F899FC045814" bold="true" box="[941,966,1799,1825]" pageId="7" pageNumber="6">7)</emphasis>
Male penis, lateral view illustrating basal body (pala), median lobe (parameroid) and serrate process (trigonium).
<emphasis id="B92BCF3CAB13FFA9FC4EF8B6FBCF5877" bold="true" box="[1013,1037,1832,1858]" pageId="7" pageNumber="6">8)</emphasis>
Male penis, dorsal view illustrating basal body (pala), median lobe (parameroid) and serrate process (trigonium).
<emphasis id="B92BCF3CAB13FFA9FB9CF8D4FBFD5851" bold="true" box="[1063,1087,1866,1892]" pageId="7" pageNumber="6">9)</emphasis>
Male tergite VIII.
<emphasis id="B92BCF3CAB13FFA9FAB5F8D4FAF15851" bold="true" box="[1294,1331,1866,1892]" pageId="7" pageNumber="6">10)</emphasis>
Male tergite IX. Scale lines = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BE0132EAB1CFFA6FF7BFE57FB4E5D99" blockId="8.[192,1442,457,685]" pageId="8" pageNumber="7">
<emphasis id="B92BCF3CAB1CFFA6FF7BFE57FE8E5ED6" bold="true" box="[192,332,457,483]" pageId="8" pageNumber="7">Life history.</emphasis>
Little is known about the ecology and life history of
<taxonomicName id="4C5F68ADAB1CFFA6FC20FE57FBD95ED6" authorityName="Springer &amp; Waller" authorityYear="2021" box="[923,1051,457,483]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="goodrichi">
<emphasis id="B92BCF3CAB1CFFA6FC20FE57FBD95ED6" box="[923,1051,457,483]" italics="true" pageId="8" pageNumber="7">S. goodrichi</emphasis>
</taxonomicName>
. Specimens have been taken at the
<typeStatus id="54E4AD8CAB1CFFA6FF7BFE75FF325D30" box="[192,240,491,517]" pageId="8" pageNumber="7">type</typeStatus>
locality every month from May to August, all by UV light trap. In
<collectingRegion id="499BDDCCAB1CFFA6FC54FE74FB5E5D31" box="[1007,1180,490,516]" country="United States of America" name="South Carolina" pageId="8" pageNumber="7">South Carolina</collectingRegion>
, they were collected in May, July and September, at UV light and by sweeping near water, but no other collection or ecological data are provided. However, two well-known species in the genus,
<taxonomicName id="4C5F68ADAB1CFFA6FCF8FDB0FC155D7D" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[835,983,558,584]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="orbiculatus">
<emphasis id="B92BCF3CAB1CFFA6FCF8FDB0FC155D7D" box="[835,983,558,584]" italics="true" pageId="8" pageNumber="7">S. orbiculatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5F68ADAB1CFFA6FBB5FDB0FBB35D7D" authorityName="Guerin-Meneville" authorityYear="1843" box="[1038,1137,558,584]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B92BCF3CAB1CFFA6FBB5FDB0FBB35D7D" box="[1038,1137,558,584]" italics="true" pageId="8" pageNumber="7">S. tibialis</emphasis>
</taxonomicName>
, sympatric with each other and
<taxonomicName id="4C5F68ADAB1CFFA6FF48FDCEFEB45D5C" authorityName="Springer &amp; Waller" authorityYear="2021" box="[243,374,591,618]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="goodrichi">
<emphasis id="B92BCF3CAB1CFFA6FF48FDCEFEB45D5C" box="[243,374,591,618]" italics="true" pageId="8" pageNumber="7">S. goodrichi</emphasis>
</taxonomicName>
, have aquatic larvae associated with decomposing plant material in marshes, ponds, springs, swamps, tree holes and phytotelmata (
<bibRefCitation id="EFCE6EDFAB1CFFA6FDCDFDEFFCC15DBE" author="Kraatz WC" box="[630,771,625,651]" pageId="8" pageNumber="7" pagination="393 - 400" refId="ref7092" refString="Kraatz WC. 1918. Scirtes tibialis, Guer., (Coleoptera-Dascyllidae), with observations on its life history. Annals of the Entomological Society of America 11: 393 - 400." type="journal article" year="1918">Kraatz 1918</bibRefCitation>
;
<bibRefCitation id="EFCE6EDFAB1CFFA6FCABFDEFFC0F5DBE" author="Beerbower FV" box="[784,973,625,651]" pageId="8" pageNumber="7" pagination="672 - 680" refId="ref6780" refString="Beerbower FV. 1943. Life history of Scirtes orbiculatus Fabricius (Coleoptera: Helodidae). Annals of the Entomological Society of America 36: 672 - 680." type="journal article" year="1943">Beerbower 1943</bibRefCitation>
). The suggestion is strong therefore, that when studied,
<taxonomicName id="4C5F68ADAB1CFFA6FED8FD0DFE215D99" authorityName="Springer &amp; Waller" authorityYear="2021" box="[355,483,658,685]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="goodrichi">
<emphasis id="B92BCF3CAB1CFFA6FED8FD0DFE215D99" box="[355,483,658,685]" italics="true" pageId="8" pageNumber="7">S. goodrichi</emphasis>
</taxonomicName>
will exhibit similar ecological and life history characteristics.
</paragraph>
</subSubSection>
</treatment>
</document>