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<document id="D03580E577C6E8F0127C67ED923460B1" ID-DOI="10.5281/zenodo.198060" ID-GBIF-Dataset="ed312cce-318a-473e-9e60-a9b097373666" ID-ISSN="1175-5326" ID-Zenodo-Dep="198060" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460283733665" checkinUser="plazi" docAuthor="Zhao, Zony-Yi &amp; Zhou, Hong-Zhang" docDate="2010" docId="03EF87FABD10FFEE9DD3DCC8FE9876CE" docLanguage="en" docName="zt02619p038.pdf" docOrigin="Zootaxa 2619" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Indoquedius radius Zhao &amp; Zhou, 2010, sp. nov." docType="treatment" docVersion="9" lastPageNumber="37" masterDocId="FFD6FF82BD19FFE49D44DF6FFFF3742C" masterDocTitle="Taxonomy of the genus Indoquedius Blackwelder (Coleoptera: Staphylinidae: Staphylininae) of China with description of four new species" masterLastPageNumber="38" masterPageNumber="27" pageNumber="36" updateTime="1698249979089" updateUser="plazi">
<mods:mods id="FD8A239C2F68FFCDFF1098963C6AB09E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DE134639E2414ADA0A746A9B3D9B5304">
<mods:title id="8F4771AFFD158BDD8E3EDB6B3482D06B">Taxonomy of the genus Indoquedius Blackwelder (Coleoptera: Staphylinidae: Staphylininae) of China with description of four new species</mods:title>
</mods:titleInfo>
<mods:name id="AA0A2E420BE4489733CF1D10866ED1C1" type="personal">
<mods:role id="E65681E4CB6FC1EE6536B645F037BE22">
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<mods:namePart id="DE8B5C06E40E45A714E898C9BD70BED3">Zhao, Zony-Yi</mods:namePart>
</mods:name>
<mods:name id="E0D7D072E1BAA89BFCCEFADAA0D3DF29" type="personal">
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<mods:namePart id="3253E07AAF4BF48C9A557B73B0ED15E6">Zhou, Hong-Zhang</mods:namePart>
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<mods:typeOfResource id="B9A0A2C64A7DC4B52B00CBEF51B8E323">text</mods:typeOfResource>
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<mods:title id="9FAF5DDA3C20E01911AF27DF5D41E2EB">Zootaxa</mods:title>
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<mods:part id="EE30D8A4FD155690987092B24CFF0E01">
<mods:date id="7F94288BDF7D5D73F757480C25C428F6">2010</mods:date>
<mods:detail id="147E672E3F57455E6DE7D27CEAEA4F30" type="volume">
<mods:number id="F6E7BBCFE2A765D31C4161E073492FDE">2619</mods:number>
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<mods:identifier id="F1FCC0E3534F73CC659020353B0EF4C4" type="DOI">10.5281/zenodo.198060</mods:identifier>
<mods:identifier id="18A39ED2C97D3B4E72E98BD7B5DA1099" type="GBIF-Dataset">ed312cce-318a-473e-9e60-a9b097373666</mods:identifier>
<mods:identifier id="127CA87427C68197593FF9EB7C8D9848" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="2ABF54BAA91FF7F7835935EBD69EAE13" type="Zenodo-Dep">198060</mods:identifier>
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<treatment id="03EF87FABD10FFEE9DD3DCC8FE9876CE" ID-DOI="http://doi.org/10.5281/zenodo.6210142" ID-GBIF-Taxon="119397514" ID-Zenodo-Dep="6210142" LSID="urn:lsid:plazi:treatment:03EF87FABD10FFEE9DD3DCC8FE9876CE" httpUri="http://treatment.plazi.org/id/03EF87FABD10FFEE9DD3DCC8FE9876CE" lastPageId="10" lastPageNumber="37" pageId="9" pageNumber="36">
<subSubSection id="C35C6567BD10FFED9DD3DCC8FDE177ED" box="[151,530,935,961]" pageId="9" pageNumber="36" type="nomenclature">
<paragraph id="8BF936ECBD10FFED9DD3DCC8FDE177ED" blockId="9.[151,530,935,996]" box="[151,530,935,961]" pageId="9" pageNumber="36">
<heading id="D0B18180BD10FFED9DD3DCC8FDE177ED" bold="true" box="[151,530,935,961]" fontSize="11" level="1" pageId="9" pageNumber="36" reason="1">
<emphasis id="B932EAFEBD10FFED9DD3DCC8FDE177ED" bold="true" box="[151,530,935,961]" pageId="9" pageNumber="36">
10.
<taxonomicName id="4C464D6FBD10FFED9D87DCC8FE5E77ED" ID-CoL="3PNKN" box="[195,429,935,961]" class="Insecta" family="Staphylinidae" genus="Indoquedius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="36" phylum="Arthropoda" rank="species" species="radius" status="sp. nov.">
<emphasis id="B932EAFEBD10FFED9D87DCC8FE5E77ED" bold="true" box="[195,429,935,961]" italics="true" pageId="9" pageNumber="36">Indoquedius radius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2015785BD10FFED9CF1DCC8FDE177ED" box="[437,530,935,961]" pageId="9" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35C6567BD10FFED9DD3DCA5FF1C77C8" box="[151,239,970,996]" pageId="9" pageNumber="36" type="description">
<paragraph id="8BF936ECBD10FFED9DD3DCA5FF1C77C8" blockId="9.[151,530,935,996]" box="[151,239,970,996]" pageId="9" pageNumber="36">
(
<figureCitation id="137D2A69BD10FFED9DDBDCA5FF1677C8" box="[159,229,970,996]" captionStart="FIGURE 7" captionStartId="10.[151,258,1712,1736]" captionTargetBox="[152,1421,785,1711]" captionTargetId="figure@10.[151,1436,772,1715]" captionTargetPageId="10" captionText="FIGURE 7. Indoquedius radius sp. nov.: A, male sternite VII; B, male tergite VIII; C, male sternite VIII; D, male sternite IX; E, male tergite X; F, aedeagus in lateral view; G, aedeagus in parameral view; H, underside of paramere. Scales = 0.5 mm." httpUri="https://zenodo.org/record/198067/files/figure.png" pageId="9" pageNumber="36">Fig. 7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35C6567BD10FFED9DD3DB7AFB3F7003" box="[151,1228,1044,1071]" pageId="9" pageNumber="36" type="materials_examined">
<paragraph id="8BF936ECBD10FFED9DD3DB7AFB3F7003" blockId="9.[151,1437,1044,2004]" box="[151,1228,1044,1071]" pageId="9" pageNumber="36">
<emphasis id="B932EAFEBD10FFED9DD3DB7AFD5A7003" bold="true" box="[151,681,1045,1071]" pageId="9" pageNumber="36">
<typeStatus id="54FD884EBD10FFED9DD3DB7AFF277003" box="[151,212,1045,1071]" pageId="9" pageNumber="36">Type</typeStatus>
material:
<typeStatus id="54FD884EBD10FFED9C12DB7AFE3F7003" box="[342,460,1045,1071]" pageId="9" pageNumber="36" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F351767CBD10FFED9C9DDB7AFDCC7003" box="[473,575,1045,1071]" name="China" pageId="9" pageNumber="36">CHINA</collectingCountry>
: Fujian:
</emphasis>
ɗ, Wuyishan Mountain,
<date id="FFF8102CBD10FFED9E94DB7AFBB57003" box="[976,1094,1045,1071]" pageId="9" pageNumber="36" value="1973-06-06">6.VI.1973</date>
(
<emphasis id="B932EAFEBD10FFED991CDB7AFB487003" bold="true" box="[1112,1211,1045,1071]" pageId="9" pageNumber="36">IZ-CAS</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C35C6567BD10FFEE9D81DB55FC9F760B" lastPageId="10" lastPageNumber="37" pageId="9" pageNumber="36" type="description">
<paragraph id="8BF936ECBD10FFED9D81DB55FB3170B3" blockId="9.[151,1437,1044,2004]" pageId="9" pageNumber="36">
<emphasis id="B932EAFEBD10FFED9D81DB55FEAE7078" bold="true" box="[197,349,1082,1108]" pageId="9" pageNumber="36">Description.</emphasis>
Head, pronotum and elytra nearly black; abdomen dark brown, iridescent; antennomeres III brown, IIIVI darker apically, VIIIIX gradually becoming paler, XXI milky yellow; labrum entirely brown, mandible reddish-brown, maxillary and labial palpi entirely brown; all legs entirely brown.
</paragraph>
<paragraph id="8BF936ECBD10FFED9D81DBC5FA9970E8" blockId="9.[151,1437,1044,2004]" box="[197,1386,1194,1220]" pageId="9" pageNumber="36">BL = 10.0 mm, BW = 2.0 mm, HL/PL/EL = 1.00:1.29:1.43, HW/PW/EW/AW = 1.00:1.06:1.24:1.00.</paragraph>
<paragraph id="8BF936ECBD10FFED9D81DBBFFE277188" blockId="9.[151,1437,1044,2004]" pageId="9" pageNumber="36">Head round, wider than long, HW/HL = 1.33; eye very large and convex, nearly occupying the whole lateral side of head, in dorsal view HEL/HTL = 10.00; posterior frontal setiferous puncture situated before level of posterior side of eye and with socket touching the eye; two setiferous punctures between anterior and posterior frontal setiferous punctures along medial side of eye; temporal setiferous puncture situated at the same level as posterior margin of eye; basal setiferous puncture closer to nuchal constriction than to posterior frontal setiferous puncture.</paragraph>
<paragraph id="8BF936ECBD10FFED9D81DADFFB527216" blockId="9.[151,1437,1044,2004]" pageId="9" pageNumber="36">Pronotum slightly wider than long, PW/PL = 1.05, nearly parallel-sided, basal margin rounded; two setiferous punctures in each dorsal row; one setiferous puncture in each sublateral row; large lateral setiferous puncture slightly behind level of last puncture in dorsal row and with socket touching lateral marginal groove; posterior ventral plate bearing middle longitudinal carina and not divided into two parts.</paragraph>
<paragraph id="8BF936ECBD10FFED9D81D92AFCD97273" blockId="9.[151,1437,1044,2004]" box="[197,810,1605,1631]" pageId="9" pageNumber="36">Scutellum densely punctuated but lacking any setae.</paragraph>
<paragraph id="8BF936ECBD10FFED9D81D905FAD97286" blockId="9.[151,1437,1044,2004]" pageId="9" pageNumber="36">Elytra wider than long, EW/EL = 1.07, ESL/EL = 0.51; each elytron with two large antero-lateral setae; surface covered with coarse and dense setiferous punctures becoming slightly sparser towards apex.</paragraph>
<paragraph id="8BF936ECBD10FFED9D81D9DAFCDA7313" blockId="9.[151,1437,1044,2004]" pageId="9" pageNumber="36">Setiferous punctures on each abdominal tergite sparser and smaller than those on elytron, becoming sparser towards apex of each tergite, surface among punctures covered with dense and rather vague transverse microsculpture which only becoming more obvious around punctures and near basal ridge; sternite III with middle portion of basal ridge sharply pointed backward.</paragraph>
<paragraph id="8BF936ECBD10FFEE9D81D825FC9F760B" blockId="9.[151,1437,1044,2004]" lastBlockId="10.[151,1437,152,738]" lastPageId="10" lastPageNumber="37" pageId="9" pageNumber="36">
Male sternite VII (
<figureCitation id="137D2A69BD10FFED9CDBD825FE177348" box="[415,484,1866,1892]" captionStart="FIGURE 7" captionStartId="10.[151,258,1712,1736]" captionTargetBox="[152,1421,785,1711]" captionTargetId="figure@10.[151,1436,772,1715]" captionTargetPageId="10" captionText="FIGURE 7. Indoquedius radius sp. nov.: A, male sternite VII; B, male tergite VIII; C, male sternite VIII; D, male sternite IX; E, male tergite X; F, aedeagus in lateral view; G, aedeagus in parameral view; H, underside of paramere. Scales = 0.5 mm." httpUri="https://zenodo.org/record/198067/files/figure.png" pageId="9" pageNumber="36">Fig. 7</figureCitation>
A) with four large apical setae on each side, apical margin with a shallow middle emargination, a small area around the emargination impunctate; tergite VIII (
<figureCitation id="137D2A69BD10FFED9974D81FFB8473A6" box="[1072,1143,1904,1930]" captionStart="FIGURE 7" captionStartId="10.[151,258,1712,1736]" captionTargetBox="[152,1421,785,1711]" captionTargetId="figure@10.[151,1436,772,1715]" captionTargetPageId="10" captionText="FIGURE 7. Indoquedius radius sp. nov.: A, male sternite VII; B, male tergite VIII; C, male sternite VIII; D, male sternite IX; E, male tergite X; F, aedeagus in lateral view; G, aedeagus in parameral view; H, underside of paramere. Scales = 0.5 mm." httpUri="https://zenodo.org/record/198067/files/figure.png" pageId="9" pageNumber="36">Fig. 7</figureCitation>
B) with two large lateral setae on each side, middle portion of basal ridge arcuated backward; sternite VIII (
<figureCitation id="137D2A69BD10FFED99C4D8FAFB3B7383" box="[1152,1224,1941,1967]" captionStart="FIGURE 7" captionStartId="10.[151,258,1712,1736]" captionTargetBox="[152,1421,785,1711]" captionTargetId="figure@10.[151,1436,772,1715]" captionTargetPageId="10" captionText="FIGURE 7. Indoquedius radius sp. nov.: A, male sternite VII; B, male tergite VIII; C, male sternite VIII; D, male sternite IX; E, male tergite X; F, aedeagus in lateral view; G, aedeagus in parameral view; H, underside of paramere. Scales = 0.5 mm." httpUri="https://zenodo.org/record/198067/files/figure.png" pageId="9" pageNumber="36">Fig. 7</figureCitation>
C) with two large setae on each side, middle portion of basal ridge slightly arcuated backward, apical margin with a middle emargination, a small subtriangular area around emargination impunctate; sternite IX (
<figureCitation id="137D2A69BD13FFEE99F4DFF7FB0A749E" box="[1200,1273,152,178]" captionStart="FIGURE 7" captionStartId="10.[151,258,1712,1736]" captionTargetBox="[152,1421,785,1711]" captionTargetId="figure@10.[151,1436,772,1715]" captionTargetPageId="10" captionText="FIGURE 7. Indoquedius radius sp. nov.: A, male sternite VII; B, male tergite VIII; C, male sternite VIII; D, male sternite IX; E, male tergite X; F, aedeagus in lateral view; G, aedeagus in parameral view; H, underside of paramere. Scales = 0.5 mm." httpUri="https://zenodo.org/record/198067/files/figure.png" pageId="10" pageNumber="37">Fig. 7</figureCitation>
D) with basal portion small, apical margin with a deep middle emargination, one large subapical seta on each side; tergite X (
<figureCitation id="137D2A69BD13FFEE9DE4DF8DFF1A74D0" box="[160,233,226,252]" captionStart="FIGURE 7" captionStartId="10.[151,258,1712,1736]" captionTargetBox="[152,1421,785,1711]" captionTargetId="figure@10.[151,1436,772,1715]" captionTargetPageId="10" captionText="FIGURE 7. Indoquedius radius sp. nov.: A, male sternite VII; B, male tergite VIII; C, male sternite VIII; D, male sternite IX; E, male tergite X; F, aedeagus in lateral view; G, aedeagus in parameral view; H, underside of paramere. Scales = 0.5 mm." httpUri="https://zenodo.org/record/198067/files/figure.png" pageId="10" pageNumber="37">Fig. 7</figureCitation>
E) with basal side broadly concaved, apical margin with a deep middle emargination; aedeagus in lateral view (
<figureCitation id="137D2A69BD13FFEE9C7CDE67FE72750E" box="[312,385,264,290]" captionStart="FIGURE 7" captionStartId="10.[151,258,1712,1736]" captionTargetBox="[152,1421,785,1711]" captionTargetId="figure@10.[151,1436,772,1715]" captionTargetPageId="10" captionText="FIGURE 7. Indoquedius radius sp. nov.: A, male sternite VII; B, male tergite VIII; C, male sternite VIII; D, male sternite IX; E, male tergite X; F, aedeagus in lateral view; G, aedeagus in parameral view; H, underside of paramere. Scales = 0.5 mm." httpUri="https://zenodo.org/record/198067/files/figure.png" pageId="10" pageNumber="37">Fig. 7</figureCitation>
F) with apex of paramere not protruding beyond that of median lobe, paramere nearly straight throughout whole length, median lobe with apex slightly bent towards paramere, aedeagus in parameral view (
<figureCitation id="137D2A69BD13FFEE9C1FDE3DFE537540" box="[347,416,338,364]" captionStart="FIGURE 7" captionStartId="10.[151,258,1712,1736]" captionTargetBox="[152,1421,785,1711]" captionTargetId="figure@10.[151,1436,772,1715]" captionTargetPageId="10" captionText="FIGURE 7. Indoquedius radius sp. nov.: A, male sternite VII; B, male tergite VIII; C, male sternite VIII; D, male sternite IX; E, male tergite X; F, aedeagus in lateral view; G, aedeagus in parameral view; H, underside of paramere. Scales = 0.5 mm." httpUri="https://zenodo.org/record/198067/files/figure.png" pageId="10" pageNumber="37">Fig. 7</figureCitation>
G) with paramere from base to apex gradually narrowed and forming rod-shaped apical portion, apex blunt; median lobe from base to apex first nearly parallel-sided and then narrowed at about apical 1/10, apex blunt; paramere with base wider and other portion narrower than median lobe; apical portion of paramere with four small apical setae and two subapical setae on each side, sensory peg setae on underside distributed along apical and lateral sides, forming one apical group (including 7 peg setae) and two lateral groups (including 34 peg setae) (
<figureCitation id="137D2A69BD13FFEE9F63DD62FD9E760B" box="[551,621,525,551]" captionStart="FIGURE 7" captionStartId="10.[151,258,1712,1736]" captionTargetBox="[152,1421,785,1711]" captionTargetId="figure@10.[151,1436,772,1715]" captionTargetPageId="10" captionText="FIGURE 7. Indoquedius radius sp. nov.: A, male sternite VII; B, male tergite VIII; C, male sternite VIII; D, male sternite IX; E, male tergite X; F, aedeagus in lateral view; G, aedeagus in parameral view; H, underside of paramere. Scales = 0.5 mm." httpUri="https://zenodo.org/record/198067/files/figure.png" pageId="10" pageNumber="37">Fig. 7</figureCitation>
H). Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C35C6567BD13FFEE9D81DD5DFDD37660" box="[197,544,562,588]" pageId="10" pageNumber="37" type="distribution">
<paragraph id="8BF936ECBD13FFEE9D81DD5DFDD37660" blockId="10.[151,1437,152,738]" box="[197,544,562,588]" pageId="10" pageNumber="37">
<emphasis id="B932EAFEBD13FFEE9D81DD5DFE967660" bold="true" box="[197,357,562,588]" pageId="10" pageNumber="37">Distribution.</emphasis>
<collectingCountry id="F351767CBD13FFEE9C28DD5DFE417660" box="[364,434,562,588]" name="China" pageId="10" pageNumber="37">China</collectingCountry>
(Fujian).
</paragraph>
</subSubSection>
<subSubSection id="C35C6567BD13FFEE9D81DD38FDF676BB" pageId="10" pageNumber="37" type="discussion">
<paragraph id="8BF936ECBD13FFEE9D81DD38FDF676BB" blockId="10.[151,1437,152,738]" pageId="10" pageNumber="37">
<emphasis id="B932EAFEBD13FFEE9D81DD38FECD765D" bold="true" box="[197,318,599,625]" pageId="10" pageNumber="37">Remarks.</emphasis>
This new species has an entire paramere with the distribution pattern of sensory peg setae being different from other congeners.
</paragraph>
</subSubSection>
<subSubSection id="C35C6567BD13FFEE9D81DDCDFE9876CE" pageId="10" pageNumber="37" type="etymology">
<paragraph id="8BF936ECBD13FFEE9D81DDCDFE9876CE" blockId="10.[151,1437,152,738]" pageId="10" pageNumber="37">
<emphasis id="B932EAFEBD13FFEE9D81DDCDFEA07690" bold="true" box="[197,339,674,700]" pageId="10" pageNumber="37">Etymology.</emphasis>
The species name is derived from the Latin word “
<taxonomicName id="4C464D6FBD13FFEE9EFADDCDFBF97697" box="[958,1034,674,699]" class="Insecta" family="Staphylinidae" genus="Indoquedius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="37" phylum="Arthropoda" rank="species" species="radius">
<emphasis id="B932EAFEBD13FFEE9EFADDCDFBF97697" box="[958,1034,674,699]" italics="true" pageId="10" pageNumber="37">radius</emphasis>
</taxonomicName>
” (arm, beam, ray) referring to its straight paramere.
</paragraph>
</subSubSection>
</treatment>
</document>