563 lines
71 KiB
XML
563 lines
71 KiB
XML
<document id="3FEDFFA5381EE5D5FAA2EFECFFE3B912" ID-CLB-Dataset="60679" ID-DOI="10.1080/00222933.2023.2218574" ID-GBIF-Dataset="eeb76dde-544a-4e23-8597-979f0540d5ab" ID-ISSN="1464-5262" ID-Zenodo-Dep="8221626" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1690876754604" checkinUser="plazi" docAuthor="Weijola, Valter & Kraus, Fred" docDate="2023" docId="03EF87F6961AFFBE5BB5FE09FCB5F969" docLanguage="en" docName="JNATHIST.57.9-12.947-975.pdf" docOrigin="Journal of Natural History 57 (13 - 16)" docSource="http://dx.doi.org/10.1080/00222933.2023.2218574" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Varanus tanimbar Weijola & Kraus 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="964" masterDocId="FFD6FF8E9616FFAF5A44FFFCFFF4FFA9" masterDocTitle="Two new species of monitor lizards (Squamata: Varanus) endemic to the Louisiade and Tanimbar Archipelagos with a key to the subgenus Euprepiosaurus" masterLastPageNumber="975" masterPageNumber="947" pageNumber="959" updateTime="1691426842078" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="6B1233F7B0585348851931A1467596A3" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="2F3DCA47570726C24E698398999A9694">
|
||
<mods:title id="18C7994CE3A18066D586440EDBAC37F3">Two new species of monitor lizards (Squamata: Varanus) endemic to the Louisiade and Tanimbar Archipelagos with a key to the subgenus Euprepiosaurus</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="25A4F9E2316BEC17F162BDE8AEB6DFC5" type="personal">
|
||
<mods:role id="86EA36FED982E072F380D04C4BA80F88">
|
||
<mods:roleTerm id="38FC9AF01D013CBD7867A699539E7CC9">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="3797968EEE90EB2175D50348C641F23A">Weijola, Valter</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="B260CCEDBC0264B269EE92AEED72BF09" type="personal">
|
||
<mods:role id="EDA9A4C851CF352509BEB4A51BC59D5A">
|
||
<mods:roleTerm id="6A7B5EB8D396C44070C62EA490736520">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="884295D94DE99B599E7A41BBB0D294C3">Kraus, Fred</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="11847E92D716157201D0B2830C7FB549">text</mods:typeOfResource>
|
||
<mods:relatedItem id="0C8A2D7E8DB5E8C5EE606AFC4EBBE3EB" type="host">
|
||
<mods:titleInfo id="B7F5AEC5051E40D619A680D654AA5757">
|
||
<mods:title id="D293BC5DD91C3058AA6194BE1D858AFD">Journal of Natural History</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="2CF36FF1337A1A7AF737B1862EDBD72F">
|
||
<mods:date id="0A8763840747536BCCFE0C675476A0F3">2023</mods:date>
|
||
<mods:detail id="076951F5A6DFA02FF85E4998588BF8C6" type="pubDate">
|
||
<mods:number id="0EF56F0623276E08D9B533016E3973B7">2023-06-21</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="6EDB5C86937D9216FD6120D18D91469E" type="volume">
|
||
<mods:number id="3961D8C1FF5021F30DE0AE2BED982F46">57</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="F8E8006023657E68F86DB63D1E0783D4" type="issue">
|
||
<mods:number id="2DFE4444B62BF97EC8060073FFC3D4D5">13 - 16</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="308202241FC0998C8E037499003AEA9F" unit="page">
|
||
<mods:start id="D3A504DFD3725B13AAA72A1789348B01">947</mods:start>
|
||
<mods:end id="9BDC0FD799DEFB66EB5B27A58BBA6068">975</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="6CF9FECC240CC6923F74331F6A228FFD">
|
||
<mods:url id="560BF1FE31151AF337B7A3534A710855">http://dx.doi.org/10.1080/00222933.2023.2218574</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="87BF748C693855681FFDBFEA078756C2">journal article</mods:classification>
|
||
<mods:identifier id="9156896AD275A12245ABDDE18CDDDD5E" type="CLB-Dataset">60679</mods:identifier>
|
||
<mods:identifier id="B77011576C08B66DE8D9111165BE984D" type="DOI">10.1080/00222933.2023.2218574</mods:identifier>
|
||
<mods:identifier id="97430AA10F55EBFB6961FDD3AE4E3CEB" type="GBIF-Dataset">eeb76dde-544a-4e23-8597-979f0540d5ab</mods:identifier>
|
||
<mods:identifier id="4FFAC1BD2E361E8B7C390FB761B261D4" type="ISSN">1464-5262</mods:identifier>
|
||
<mods:identifier id="1B5227E967AADAFD98C11CD236458AF1" type="Zenodo-Dep">8221626</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03EF87F6961AFFBE5BB5FE09FCB5F969" ID-DOI="http://doi.org/10.5281/zenodo.8221622" ID-Zenodo-Dep="8221622" LSID="urn:lsid:plazi:treatment:03EF87F6961AFFBE5BB5FE09FCB5F969" httpUri="http://treatment.plazi.org/id/03EF87F6961AFFBE5BB5FE09FCB5F969" lastPageId="17" lastPageNumber="964" pageId="12" pageNumber="959">
|
||
<subSubSection id="C35C656B961AFFA35BB5FE09FCC7FDA6" box="[497,819,501,527]" pageId="12" pageNumber="959" type="nomenclature">
|
||
<paragraph id="8BF936E0961AFFA35BB5FE09FCC7FDA6" blockId="12.[497,819,501,562]" box="[497,819,501,527]" pageId="12" pageNumber="959">
|
||
<heading id="D0B1818C961AFFA35BB5FE09FCC7FDA6" box="[497,819,501,527]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="959" reason="5">
|
||
<taxonomicName id="4C464D63961AFFA35BB5FE09FD3BFDA6" authority="Weijola & Kraus, 2023" authorityName="Weijola & Kraus" authorityYear="2023" box="[497,719,501,527]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="12" pageNumber="959" phylum="Chordata" rank="species" species="tanimbar" status="sp. nov.">
|
||
<emphasis id="B932EAF2961AFFA35BB5FE09FD3BFDA6" bold="true" box="[497,719,501,527]" italics="true" pageId="12" pageNumber="959">Varanus tanimbar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2015789961AFFA35893FE09FCC7FDA6" box="[727,819,501,527]" pageId="12" pageNumber="959" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35C656B961AFFA3587AFDE4FD11FD9B" box="[574,741,536,562]" pageId="12" pageNumber="959" type="description">
|
||
<paragraph id="8BF936E0961AFFA3587AFDE4FD11FD9B" blockId="12.[497,819,501,562]" box="[574,741,536,562]" pageId="12" pageNumber="959">
|
||
(
|
||
<figureCitation id="137D2A65961AFFA35802FDE4FD29FD9B" box="[582,733,536,562]" captionStart-0="Figure 8" captionStart-1="Figure 9" captionStart-2="Figure 10" captionStart-3="Figure 11" captionStart-4="Figure 12" captionStartId-0="13.[160,225,1531,1554]" captionStartId-1="14.[160,225,851,874]" captionStartId-2="14.[160,225,1496,1519]" captionStartId-3="15.[160,225,896,919]" captionStartId-4="15.[160,225,1700,1723]" captionTargetBox-0="[163,1153,142,1499]" captionTargetBox-1="[163,1153,142,820]" captionTargetBox-2="[163,1153,934,1465]" captionTargetBox-3="[163,1153,142,864]" captionTargetBox-4="[163,1153,1004,1668]" captionTargetId-0="figure-71@13.[163,1153,142,1499]" captionTargetId-1="figure-40@14.[163,1153,142,820]" captionTargetId-2="figure-61@14.[163,1153,934,1465]" captionTargetId-3="figure-24@15.[163,1153,142,865]" captionTargetId-4="figure-3@15.[163,1153,1004,1668]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="14" captionTargetPageId-3="15" captionTargetPageId-4="15" captionText-0="Figure 8. Dorsal, ventral and head profile of the holotype (WAM112255) of Varanus tanimbar sp. nov." captionText-1="Figure 9. Dorsal and ventral aspect of paratype WAM 109940 of Varanus tanimbar sp. nov." captionText-2="Figure 10. Adult Varanus tanimbar sp. nov. male killed by local hunters near Saumlaki, Yamdena (photo VW)." captionText-3="Figure 11. Juvenile Varanus tanimbar sp. nov. caught by local hunters at Saumlaki,Yamdena (photo VW)." captionText-4="Figure 12. Adult Varanus tanimbar sp.nov. (WAM 109896) Lorulun, Yamdena Island (photo Richard How)." figureDoi-0="http://doi.org/10.5281/zenodo.8221644" figureDoi-1="http://doi.org/10.5281/zenodo.8221646" figureDoi-2="http://doi.org/10.5281/zenodo.8221648" figureDoi-3="http://doi.org/10.5281/zenodo.8221650" figureDoi-4="http://doi.org/10.5281/zenodo.8221652" httpUri-0="https://zenodo.org/record/8221644/files/figure.png" httpUri-1="https://zenodo.org/record/8221646/files/figure.png" httpUri-2="https://zenodo.org/record/8221648/files/figure.png" httpUri-3="https://zenodo.org/record/8221650/files/figure.png" httpUri-4="https://zenodo.org/record/8221652/files/figure.png" pageId="12" pageNumber="959">Figures 8–12</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35C656B961AFFA35AE4FD93FE9CFC09" pageId="12" pageNumber="959" type="materials_examined">
|
||
<materialsCitation id="3B2E3CBD961AFFA35AE4FD93FB75FD66" collectingDate="1993-04-24" collectionCode="RH, RJ, DK" collectorName="Richard How & Ronald Johnstone & Darrell Kitchener" country="Indonesia" county="Tanimbar Islands" location="Latdalam" municipality="Yamdena Island" pageId="12" pageNumber="959" specimenCode="WAM112255" specimenCount="1" typeStatus="holotype">
|
||
<paragraph id="8BF936E0961AFFA35AE4FD93FEFFFD20" blockId="12.[160,1155,623,719]" box="[160,267,623,649]" pageId="12" pageNumber="959">
|
||
<emphasis id="B932EAF2961AFFA35AE4FD93FEFFFD20" bold="true" box="[160,267,623,649]" italics="true" pageId="12" pageNumber="959">
|
||
<typeStatus id="54FD8842961AFFA35AE4FD93FEFFFD20" box="[160,267,623,649]" pageId="12" pageNumber="959">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF936E0961AFFA35AE4FD6FFB75FD66" blockId="12.[160,1155,623,719]" pageId="12" pageNumber="959">
|
||
<specimenCode id="DBE09E9B961AFFA35AE4FD6FFEDAFD05" box="[160,302,659,684]" collectionCode="WAM" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="12" pageNumber="959" type="Museum">WAM112255</specimenCode>
|
||
(
|
||
<figureCitation id="137D2A65961AFFA35B79FD6FFE6DFD05" box="[317,409,659,684]" captionStart="Figure 8" captionStartId="13.[160,225,1531,1554]" captionTargetBox="[163,1153,142,1499]" captionTargetId="figure-71@13.[163,1153,142,1499]" captionTargetPageId="13" captionText="Figure 8. Dorsal, ventral and head profile of the holotype (WAM112255) of Varanus tanimbar sp. nov." figureDoi="http://doi.org/10.5281/zenodo.8221644" httpUri="https://zenodo.org/record/8221644/files/figure.png" pageId="12" pageNumber="959">Figure 8</figureCitation>
|
||
) collected by
|
||
<collectorName id="26B35336961AFFA35873FD6FFD32FD05" box="[567,710,659,684]" pageId="12" pageNumber="959">Richard How</collectorName>
|
||
(
|
||
<collectionCode id="ED57AE25961AFFA35897FD6FFD0CFD05" box="[723,760,659,684]" pageId="12" pageNumber="959">RH</collectionCode>
|
||
),
|
||
<collectorName id="26B35336961AFFA3594CFD6FFC25FD05" box="[776,977,659,684]" pageId="12" pageNumber="959">Ronald Johnstone</collectorName>
|
||
(
|
||
<collectionCode id="ED57AE25961AFFA3599BFD6FFC0FFD05" box="[991,1019,659,684]" pageId="12" pageNumber="959">RJ</collectionCode>
|
||
) and
|
||
<collectorName id="26B35336961AFFA35E7DFD6FFEFFFD66" pageId="12" pageNumber="959">Darrell Kitchener</collectorName>
|
||
(
|
||
<collectionCode id="ED57AE25961AFFA35B5CFD4AFEC9FD66" box="[280,317,694,719]" pageId="12" pageNumber="959">DK</collectionCode>
|
||
),
|
||
<date id="FFF81020961AFFA35B0AFD4AFE12FD66" box="[334,486,694,719]" pageId="12" pageNumber="959" value="1993-04-24">
|
||
<collectingDate id="EFBCE9C8961AFFA35B0AFD4AFE12FD66" box="[334,486,694,719]" pageId="12" pageNumber="959" value="1993-04-24">24 April 1993</collectingDate>
|
||
</date>
|
||
, at
|
||
<location id="8E99603B961AFFA3584AFD4AFD8CFD66" LSID="urn:lsid:plazi:treatment:03EF87F6961AFFBE5BB5FE09FCB5F969:8E99603B961AFFA3584AFD4AFD8CFD66" box="[526,632,694,719]" country="Indonesia" county="Tanimbar Islands" municipality="Yamdena Island" name="Latdalam" pageId="12" pageNumber="959">Latdalam</location>
|
||
,
|
||
<collectingMunicipality id="6B9DAC9A961AFFA358C7FD4AFCCDFD66" box="[643,825,694,719]" pageId="12" pageNumber="959">Yamdena Island</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62984E6C961AFFA35907FD4AFBF7FD66" box="[835,1027,694,719]" pageId="12" pageNumber="959">Tanimbar Islands</collectingCounty>
|
||
,
|
||
<collectingCountry id="F3517670961AFFA35E4AFD4AFB89FD66" box="[1038,1149,694,719]" name="Indonesia" pageId="12" pageNumber="959">Indonesia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B2E3CBD961AFFA35AE4FD07FDCAFCD4" collectingDate="1993-04-18" collectingDateMax="1993-04-20" collectingDateMin="1993-04-18" collectionCode="WAM" county="Yamdena Island" location="Yamdena Island" municipality="Lorulun" pageId="12" pageNumber="959" specimenCode="WAM109940, WAM109896, WAM112323" specimenCount="1" typeStatus="paratype">
|
||
<paragraph id="8BF936E0961AFFA35AE4FD07FEE1FCBC" blockId="12.[160,1156,763,928]" box="[160,277,763,789]" pageId="12" pageNumber="959">
|
||
<emphasis id="B932EAF2961AFFA35AE4FD07FEE1FCBC" bold="true" box="[160,277,763,789]" italics="true" pageId="12" pageNumber="959">
|
||
<typeStatus id="54FD8842961AFFA35AE4FD07FEE1FCBC" box="[160,277,763,789]" pageId="12" pageNumber="959">Paratypes</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF936E0961AFFA35AE4FCE3FE9CFC09" blockId="12.[160,1156,763,928]" pageId="12" pageNumber="959">
|
||
<specimenCode id="DBE09E9B961AFFA35AE4FCE3FED9FC91" box="[160,301,799,824]" collectionCode="WAM" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="12" pageNumber="959" type="Museum">WAM109940</specimenCode>
|
||
(
|
||
<figureCitation id="137D2A65961AFFA35B78FCE3FE6CFC91" box="[316,408,799,824]" captionStart="Figure 9" captionStartId="14.[160,225,851,874]" captionTargetBox="[163,1153,142,820]" captionTargetId="figure-40@14.[163,1153,142,820]" captionTargetPageId="14" captionText="Figure 9. Dorsal and ventral aspect of paratype WAM 109940 of Varanus tanimbar sp. nov." figureDoi="http://doi.org/10.5281/zenodo.8221646" httpUri="https://zenodo.org/record/8221646/files/figure.png" pageId="12" pageNumber="959">Figure 9</figureCitation>
|
||
) collected by RH, RJ and DK,
|
||
<date id="FFF81020961AFFA358A2FCE3FC8AFC91" box="[742,894,799,824]" pageId="12" pageNumber="959" value="1993-04-20">
|
||
<collectingDate id="EFBCE9C8961AFFA358A2FCE3FC8AFC91" box="[742,894,799,824]" pageId="12" pageNumber="959" value="1993-04-20">20 April 1993</collectingDate>
|
||
</date>
|
||
, at Latdalam,
|
||
<location id="8E99603B961AFFA35E5FFCE3FF17FCF3" LSID="urn:lsid:plazi:treatment:03EF87F6961AFFBE5BB5FE09FCB5F969:8E99603B961AFFA35E5FFCE3FF17FCF3" county="Yamdena Island" municipality="Lorulun" name="Yamdena Island" pageId="12" pageNumber="959">Yamdena Island</location>
|
||
,
|
||
<specimenCode id="DBE09E9B961AFFA35AA8FCBDFE8DFCF3" box="[236,377,833,858]" collectionCode="WAM" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="12" pageNumber="959" type="Museum">WAM109896</specimenCode>
|
||
collected by RH, RJ and DK,
|
||
<date id="FFF81020961AFFA358F7FCBDFCB3FCF3" box="[691,839,833,858]" pageId="12" pageNumber="959" value="1993-04-18">
|
||
<collectingDate id="EFBCE9C8961AFFA358F7FCBDFCB3FCF3" box="[691,839,833,858]" pageId="12" pageNumber="959" value="1993-04-18">18 April 1993</collectingDate>
|
||
</date>
|
||
, at
|
||
<collectingMunicipality id="6B9DAC9A961AFFA35929FCBDFC30FCF3" box="[877,964,833,858]" pageId="12" pageNumber="959">Lorulun</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62984E6C961AFFA3598AFCBDFB74FCF3" box="[974,1152,833,858]" pageId="12" pageNumber="959">Yamdena Island</collectingCounty>
|
||
, and
|
||
<specimenCode id="DBE09E9B961AFFA35A8BFC98FEA8FCD4" box="[207,348,868,893]" collectionCode="WAM" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="12" pageNumber="959" type="Museum">WAM112323</specimenCode>
|
||
collected by RJ and I.
|
||
<materialsCitation id="3B2E3CBD961AFFA3580EFC98FE90FC09" country="Indonesia" location="Selaru Island" municipality="Tanimbar Islands" pageId="12" pageNumber="959" specimenCount="1" typeStatus="paratype">
|
||
Maryanto, 27 April, at Adaut,
|
||
<location id="8E99603B961AFFA359C2FC98FBE6FCD4" LSID="urn:lsid:plazi:treatment:03EF87F6961AFFBE5BB5FE09FCB5F969:8E99603B961AFFA359C2FC98FBE6FCD4" box="[902,1042,868,893]" country="Indonesia" municipality="Tanimbar Islands" name="Selaru Island" pageId="12" pageNumber="959">Selaru Island</location>
|
||
,
|
||
<collectingMunicipality id="6B9DAC9A961AFFA35E5FFC98FF19FC09" pageId="12" pageNumber="959">Tanimbar Islands</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="F3517670961AFFA35AB3FC7BFE90FC09" box="[247,356,903,928]" name="Indonesia" pageId="12" pageNumber="959">Indonesia</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
</subSubSection>
|
||
<subSubSection id="C35C656B961AFFA35AE4FC30FC6CFB82" pageId="12" pageNumber="959" type="etymology">
|
||
<paragraph id="8BF936E0961AFFA35AE4FC30FEE9FC4F" blockId="12.[160,1156,972,1067]" box="[160,285,972,998]" pageId="12" pageNumber="959">
|
||
<emphasis id="B932EAF2961AFFA35AE4FC30FEE9FC4F" bold="true" box="[160,285,972,998]" italics="true" pageId="12" pageNumber="959">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF936E0961AFFA35AE4FC0CFC6CFB82" blockId="12.[160,1156,972,1067]" pageId="12" pageNumber="959">
|
||
The specific epithet
|
||
<taxonomicName id="4C464D63961AFFA35BDEFC0CFDF5FBA0" authorityName="Weijola & Kraus" authorityYear="2023" box="[410,513,1008,1033]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="12" pageNumber="959" phylum="Chordata" rank="species" species="tanimbar">
|
||
<emphasis id="B932EAF2961AFFA35BDEFC0CFDF5FBA0" box="[410,513,1008,1033]" italics="true" pageId="12" pageNumber="959">tanimbar</emphasis>
|
||
</taxonomicName>
|
||
is a noun in apposition and refers to the Tanimbar Archipelago of
|
||
<collectingRegion id="4982F802961AFFA35B09FBEEFE56FB82" box="[333,418,1042,1067]" country="Indonesia" name="Maluku Utara" pageId="12" pageNumber="959">Maluku</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3517670961AFFA35BE8FBEEFDEFFB82" box="[428,539,1042,1067]" name="Indonesia" pageId="12" pageNumber="959">Indonesia</collectingCountry>
|
||
, to which this species is endemic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35C656B961AFFA15AE4FBA4FCC5F93E" lastPageId="14" lastPageNumber="961" pageId="12" pageNumber="959" type="diagnosis">
|
||
<paragraph id="8BF936E0961AFFA35AE4FBA4FEE0FBDB" blockId="12.[160,1156,1112,1554]" box="[160,276,1112,1138]" pageId="12" pageNumber="959">
|
||
<emphasis id="B932EAF2961AFFA35AE4FBA4FEE0FBDB" bold="true" box="[160,276,1112,1138]" italics="true" pageId="12" pageNumber="959">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF936E0961AFFA35AE4FB87FD55F9BB" blockId="12.[160,1156,1112,1554]" pageId="12" pageNumber="959">
|
||
<taxonomicName id="4C464D63961AFFA35AE4FB87FE91FB3D" authority="Weijola & Kraus, 2023" authorityName="Weijola & Kraus" authorityYear="2023" box="[160,357,1147,1172]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="12" pageNumber="959" phylum="Chordata" rank="species" species="tanimbar" status="sp. nov.">
|
||
<emphasis id="B932EAF2961AFFA35AE4FB87FE91FB3D" box="[160,357,1147,1172]" italics="true" pageId="12" pageNumber="959">Varanus tanimbar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2015789961AFFA35B28FB87FE37FB3D" box="[364,451,1147,1172]" pageId="12" pageNumber="959" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is a member of the subgenus
|
||
<taxonomicName id="4C464D63961AFFA3595DFB87FC4AFB3D" authorityName="Fitzinger" authorityYear="1843" box="[793,958,1147,1172]" pageId="12" pageNumber="959" rank="subGenus" subGenus="Euprepiosaurus">
|
||
<emphasis id="B932EAF2961AFFA3595DFB87FC4AFB3D" box="[793,958,1147,1172]" italics="true" pageId="12" pageNumber="959">Euprepiosaurus</emphasis>
|
||
</taxonomicName>
|
||
, which is defined by having the unique combination of an asymmetric sperm groove and laterally compressed tail (
|
||
<bibRefCitation id="EFD74B11961AFFA35B76FB3DFDF6FB73" author="Ziegler T & Schmitz A & Koch A & Bohme W" box="[306,514,1217,1242]" pageId="12" pageNumber="959" pagination="1 - 28" refId="ref13085" refString="Ziegler T, Schmitz A, Koch A, Bohme W. 2007. A review of the subgenus Euprepiosaurus of Varanus (Squamata: Varanidae): morphological and molecular phylogeny, distribution and zoogeography, with an identification key for the members of the V. indicus and the V. prasinus species groups. Zootaxa. 1472: 1 - 28." type="journal article" year="2007">
|
||
Ziegler
|
||
<emphasis id="B932EAF2961AFFA35BCEFB3DFE48FB73" box="[394,444,1217,1242]" italics="true" pageId="12" pageNumber="959">et al</emphasis>
|
||
. 2007
|
||
</bibRefCitation>
|
||
). Molecular phylogenetic analyses place it within the
|
||
<taxonomicName id="4C464D63961AFFA35E2BFB3DFF19FB55" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="12" pageNumber="959" phylum="Chordata" rank="species" species="indicus">
|
||
<emphasis id="B932EAF2961AFFA35E2BFB3DFF19FB55" italics="true" pageId="12" pageNumber="959">V. indicus</emphasis>
|
||
</taxonomicName>
|
||
species group defined by the unilateral paryphasmata ornamentation of the hemipenis and lack of blue pigmentation on the tail (
|
||
<bibRefCitation id="EFD74B11961AFFA3594AFAFAFC14FAB6" author="Ziegler T & Schmitz A & Koch A & Bohme W" box="[782,992,1286,1311]" pageId="12" pageNumber="959" pagination="1 - 28" refId="ref13085" refString="Ziegler T, Schmitz A, Koch A, Bohme W. 2007. A review of the subgenus Euprepiosaurus of Varanus (Squamata: Varanidae): morphological and molecular phylogeny, distribution and zoogeography, with an identification key for the members of the V. indicus and the V. prasinus species groups. Zootaxa. 1472: 1 - 28." type="journal article" year="2007">
|
||
Ziegler
|
||
<emphasis id="B932EAF2961AFFA3592CFAFAFC6EFAB6" box="[872,922,1286,1311]" italics="true" pageId="12" pageNumber="959">et al</emphasis>
|
||
. 2007
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD74B11961AFFA359ABFAFAFF23FAEB" author="Weijola V & Vahtera V & Lindqvist C & Kraus F" pageId="12" pageNumber="959" pagination="1053 - 1066" refId="ref12987" refString="Weijola V, Vahtera V, Lindqvist C, Kraus F. 2019. A molecular phylogeny for the Pacific monitor lizards (Varanus subgenus Euprepiosaurus) reveals a recent and rapid radiation with high levels of cryptic diversity. Zool J Linn Soc. 186 (4): 1053 - 1066. doi: 10.1093 / zoolinnean / zlz 002." type="journal article" year="2019">
|
||
Weijola
|
||
<emphasis id="B932EAF2961AFFA35E0FFAFAFB8AFAB6" box="[1099,1150,1286,1311]" italics="true" pageId="12" pageNumber="959">et al</emphasis>
|
||
. 2019
|
||
</bibRefCitation>
|
||
). It can be distinguished from all other species of
|
||
<taxonomicName id="4C464D63961AFFA35972FAD5FC28FAEB" authorityName="Fitzinger" authorityYear="1843" box="[822,988,1321,1346]" pageId="12" pageNumber="975" rank="subGenus" subGenus="Euprepiosaurus">
|
||
<emphasis id="B932EAF2961AFFA35972FAD5FC28FAEB" box="[822,988,1321,1346]" italics="true" pageId="12" pageNumber="959">Euprepiosaurus</emphasis>
|
||
</taxonomicName>
|
||
by its unique combination of: (1) dorsum black with more or less well-defined crossbands composed of lemon-yellow ocelli and/or dots; (2) tail black with distinct yellow bands; (3) dorsal aspect of tongue blue-grey; (4) temporal region typically ornamented with a dark temporal band and a lemon-yellow postocular stripe; (5) venter pale yellow; (6) mid-body scale rows (S) 145–161; (7) mid-dorsal scale rows (XY) 159–170; (8) ventral scale rows (T) 107–109; (9) scales around neck (m) 109–119.
|
||
</paragraph>
|
||
<paragraph id="8BF936E0961AFFA15AE4F9C2FCC5F93E" blockId="12.[160,1156,1598,1728]" lastBlockId="14.[160,1156,1593,1687]" lastPageId="14" lastPageNumber="961" pageId="12" pageNumber="959">
|
||
<emphasis id="B932EAF2961AFFA35AE4F9C2FEB4F9F1" bold="true" box="[160,320,1598,1624]" italics="true" pageId="12" pageNumber="959">Comparisons.</emphasis>
|
||
<taxonomicName id="4C464D63961AFFA35B15F9C3FDECF9F1" authority="Weijola & Kraus, 2023" authorityName="Weijola & Kraus" authorityYear="2023" box="[337,536,1599,1624]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="12" pageNumber="959" phylum="Chordata" rank="species" species="tanimbar" status="sp. nov.">
|
||
<emphasis id="B932EAF2961AFFA35B15F9C3FDECF9F1" box="[337,536,1599,1624]" italics="true" pageId="12" pageNumber="959">Varanus tanimbar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2015789961AFFA3585BF9C3FD83F9F1" box="[543,631,1599,1624]" pageId="12" pageNumber="959" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is the only species of
|
||
<taxonomicName id="4C464D63961AFFA35934F9C3FC3FF9F1" authorityName="Merrem" authorityYear="1820" box="[880,971,1599,1624]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="12" pageNumber="959" phylum="Chordata" rank="genus">
|
||
<emphasis id="B932EAF2961AFFA35934F9C3FC3FF9F1" box="[880,971,1599,1624]" italics="true" pageId="12" pageNumber="959">Varanus</emphasis>
|
||
</taxonomicName>
|
||
known to occur in the Tanimbar Islands. It can be distinguished from other allopatric species of
|
||
<taxonomicName id="4C464D63961AFFA35AE5F979FEBEF937" authorityName="Fitzinger" authorityYear="1843" box="[161,330,1669,1694]" pageId="12" pageNumber="975" rank="subGenus" subGenus="Euprepiosaurus">
|
||
<emphasis id="B932EAF2961AFFA35AE5F979FEBEF937" box="[161,330,1669,1694]" italics="true" pageId="12" pageNumber="959">Euprepiosaurus</emphasis>
|
||
</taxonomicName>
|
||
present in the
|
||
<collectingCountry id="F3517670961AFFA35BB8F979FD92F937" box="[508,614,1669,1694]" name="Indonesia" pageId="12" pageNumber="959">Moluccas</collectingCountry>
|
||
by differences in scalation and colour pattern: from
|
||
<taxonomicName id="4C464D63961AFFA35AA5F95BFE7DF969" authorityName="Ziegler, Bohme and Philipp" authorityYear="1999" box="[225,393,1703,1728]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="12" pageNumber="959" phylum="Chordata" rank="species" species="caerulivirens">
|
||
<emphasis id="B932EAF2961AFFA35AA5F95BFE7DF969" box="[225,393,1703,1728]" italics="true" pageId="12" pageNumber="959">V. caerulivirens</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C464D63961AFFA35B8DF95BFDB1F969" authorityName="Harvey and Barker UTA R" authorityYear="1998" box="[457,581,1703,1728]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="12" pageNumber="959" phylum="Chordata" rank="species" species="yuwonoi">
|
||
<emphasis id="B932EAF2961AFFA35B8DF95BFDB1F969" box="[457,581,1703,1728]" italics="true" pageId="12" pageNumber="959">V. yuwonoi</emphasis>
|
||
</taxonomicName>
|
||
by the lack of blue pigmentation on the tail (vs present in
|
||
<taxonomicName id="4C464D63961BFFA25B65F9C0FE3DF9FC" authorityName="Ziegler, Bohme and Philipp" authorityYear="1999" box="[289,457,1596,1621]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="13" pageNumber="960" phylum="Chordata" rank="species" species="caerulivirens">
|
||
<emphasis id="B932EAF2961BFFA25B65F9C0FE3DF9FC" box="[289,457,1596,1621]" italics="true" pageId="13" pageNumber="960">V. caerulivirens</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C464D63961BFFA2584EF9C0FD72F9FC" authorityName="Harvey and Barker UTA R" authorityYear="1998" box="[522,646,1596,1621]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="13" pageNumber="960" phylum="Chordata" rank="species" species="yuwonoi">
|
||
<emphasis id="B932EAF2961BFFA2584EF9C0FD72F9FC" box="[522,646,1596,1621]" italics="true" pageId="13" pageNumber="960">V. yuwonoi</emphasis>
|
||
</taxonomicName>
|
||
), from
|
||
<taxonomicName id="4C464D63961BFFA2589BF9C0FCC5F9FC" authorityName="Weijola and Sweet, RMNH" authorityYear="2010" box="[735,817,1596,1621]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="13" pageNumber="960" phylum="Chordata" rank="species" species="obor">
|
||
<emphasis id="B932EAF2961BFFA2589BF9C0FCC5F9FC" box="[735,817,1596,1621]" italics="true" pageId="13" pageNumber="960">V. obor</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C464D63961BFFA25935F9C0FC07F9FC" authorityName="Bohme and Ziegler" authorityYear="2005" box="[881,1011,1596,1621]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="13" pageNumber="960" phylum="Chordata" rank="species" species="zugorum">
|
||
<emphasis id="B932EAF2961BFFA25935F9C0FC07F9FC" box="[881,1011,1596,1621]" italics="true" pageId="13" pageNumber="960">V. zugorum</emphasis>
|
||
</taxonomicName>
|
||
by its black dorsum with rows of yellow ocelli (vs solid black in
|
||
<taxonomicName id="4C464D63961BFFA2595AF9A3FC85F9D1" authorityName="Weijola and Sweet, RMNH" authorityYear="2010" box="[798,881,1631,1656]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="13" pageNumber="960" phylum="Chordata" rank="species" species="obor">
|
||
<emphasis id="B932EAF2961BFFA2595AF9A3FC85F9D1" box="[798,881,1631,1656]" italics="true" pageId="13" pageNumber="960">V. obor</emphasis>
|
||
</taxonomicName>
|
||
and solid brown in
|
||
<taxonomicName id="4C464D63961BFFA25E2BF9A3FEF7F932" authorityName="Bohme and Ziegler" authorityYear="2005" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="13" pageNumber="960" phylum="Chordata" rank="species" species="zugorum">
|
||
<emphasis id="B932EAF2961BFFA25E2BF9A3FEF7F932" italics="true" pageId="13" pageNumber="960">V. zugorum</emphasis>
|
||
</taxonomicName>
|
||
), from
|
||
<taxonomicName id="4C464D63961BFFA25B18F97EFE25F932" authorityName="Bohme and Ziegler" authorityYear="1997" box="[348,465,1666,1691]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="13" pageNumber="960" phylum="Chordata" rank="species" species="melinus">
|
||
<emphasis id="B932EAF2961BFFA25B18F97EFE25F932" box="[348,465,1666,1691]" italics="true" pageId="13" pageNumber="960">V. melinus</emphasis>
|
||
</taxonomicName>
|
||
by its black (vs yellow in
|
||
<taxonomicName id="4C464D63961BFFA2594FF97EFC74F932" authorityName="Bohme and Ziegler" authorityYear="1997" box="[779,896,1666,1691]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="13" pageNumber="960" phylum="Chordata" rank="species" species="melinus">
|
||
<emphasis id="B932EAF2961BFFA2594FF97EFC74F932" box="[779,896,1666,1691]" italics="true" pageId="13" pageNumber="960">V. melinus</emphasis>
|
||
</taxonomicName>
|
||
) head and dark grey/ blue tongue (vs pink in
|
||
<taxonomicName id="4C464D63961BFFA25BF9F958FDC5F914" authorityName="Bohme and Ziegler" authorityYear="1997" box="[445,561,1700,1725]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="13" pageNumber="960" phylum="Chordata" rank="species" species="melinus">
|
||
<emphasis id="B932EAF2961BFFA25BF9F958FDC5F914" box="[445,561,1700,1725]" italics="true" pageId="13" pageNumber="960">V. melinus</emphasis>
|
||
</taxonomicName>
|
||
), and from
|
||
<taxonomicName id="4C464D63961BFFA258FAF958FC73F914" authorityName="Ziegler, Bohme and Schmitz" authorityYear="2009" box="[702,903,1700,1725]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="13" pageNumber="960" phylum="Chordata" rank="species" species="rainerguentheri">
|
||
<emphasis id="B932EAF2961BFFA258FAF958FC73F914" box="[702,903,1700,1725]" italics="true" pageId="13" pageNumber="960">V. rainerguentheri</emphasis>
|
||
</taxonomicName>
|
||
by having the dorsal pattern of yellow ocelli arrayed more or less into crossbands (vs more irregularly scattered yellow spots in
|
||
<taxonomicName id="4C464D639618FFA15B8EF9A7FD66F9DD" authorityName="Ziegler, Bohme and Schmitz" authorityYear="2009" box="[458,658,1627,1652]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="14" pageNumber="961" phylum="Chordata" rank="species" species="rainerguentheri">
|
||
<emphasis id="B932EAF29618FFA15B8EF9A7FD66F9DD" box="[458,658,1627,1652]" italics="true" pageId="14" pageNumber="961">V. rainerguentheri</emphasis>
|
||
</taxonomicName>
|
||
) and the higher mid-body (S) scale counts (
|
||
<quantity id="4CBE9B059618FFA15AE3F982FED2F93E" box="[167,294,1662,1687]" metricMagnitude="0" metricUnit="m" metricValue="3.8862" metricValueMax="4.0893999999999995" metricValueMin="3.683" pageId="14" pageNumber="961" unit="in" value="153.0" valueMax="161.0" valueMin="145.0">145–161 in</quantity>
|
||
<taxonomicName id="4C464D639618FFA15B6AF982FE44F93E" authorityName="Weijola & Kraus" authorityYear="2023" box="[302,432,1662,1687]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="14" pageNumber="961" phylum="Chordata" rank="species" species="tanimbar">
|
||
<emphasis id="B932EAF29618FFA15B6AF982FE44F93E" box="[302,432,1662,1687]" italics="true" pageId="14" pageNumber="961">V. tanimbar</emphasis>
|
||
</taxonomicName>
|
||
vs
|
||
<quantity id="4CBE9B059618FFA15B93F982FDA1F93E" box="[471,597,1662,1687]" metricMagnitude="0" metricUnit="m" metricValue="3.2893" metricValueMax="3.5305999999999997" metricValueMin="3.048" pageId="14" pageNumber="961" unit="in" value="129.5" valueMax="139.0" valueMin="120.0">120–139 in</quantity>
|
||
<taxonomicName id="4C464D639618FFA15819F982FCD0F93E" authorityName="Ziegler, Bohme and Schmitz" authorityYear="2009" box="[605,804,1662,1687]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="14" pageNumber="961" phylum="Chordata" rank="species" species="rainerguentheri">
|
||
<emphasis id="B932EAF29618FFA15819F982FCD0F93E" box="[605,804,1662,1687]" italics="true" pageId="14" pageNumber="961">V. rainerguentheri</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF396668961BFFA25AE4FA07FB70F9BB" ID-DOI="http://doi.org/10.5281/zenodo.8221644" ID-Zenodo-Dep="8221644" box="[160,1156,1531,1554]" httpUri="https://zenodo.org/record/8221644/files/figure.png" pageId="13" pageNumber="960" startId="13.[160,225,1531,1554]" targetBox="[163,1153,142,1499]" targetPageId="13" targetType="figure">
|
||
<paragraph id="8BF936E0961BFFA25AE4FA07FB70F9BB" blockId="13.[160,1156,1531,1554]" box="[160,1156,1531,1554]" pageId="13" pageNumber="960">
|
||
<emphasis id="B932EAF2961BFFA25AE4FA07FF0EF9BB" bold="true" box="[160,250,1531,1554]" pageId="13" pageNumber="960">Figure 8.</emphasis>
|
||
Dorsal, ventral and head profile of the holotype (WAM112255) of
|
||
<taxonomicName id="4C464D63961BFFA259C1FA07FBC5F9BB" authority="Weijola & Kraus, 2023" authorityName="Weijola & Kraus" authorityYear="2023" box="[901,1073,1531,1554]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="13" pageNumber="960" phylum="Chordata" rank="species" species="tanimbar" status="sp. nov.">
|
||
<emphasis id="B932EAF2961BFFA259C1FA07FBC5F9BB" box="[901,1073,1531,1554]" italics="true" pageId="13" pageNumber="960">Varanus tanimbar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2015789961BFFA25E7CFA07FB70F9BB" box="[1080,1156,1531,1554]" pageId="13" pageNumber="960" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3966689618FFA15AE4FCAFFBD1FCC3" ID-DOI="http://doi.org/10.5281/zenodo.8221646" ID-Zenodo-Dep="8221646" box="[160,1061,851,874]" httpUri="https://zenodo.org/record/8221646/files/figure.png" pageId="14" pageNumber="961" startId="14.[160,225,851,874]" targetBox="[163,1153,142,820]" targetPageId="14" targetType="figure">
|
||
<paragraph id="8BF936E09618FFA15AE4FCAFFBD1FCC3" blockId="14.[160,1061,851,874]" box="[160,1061,851,874]" pageId="14" pageNumber="961">
|
||
<emphasis id="B932EAF29618FFA15AE4FCAFFF0EFCC3" bold="true" box="[160,250,851,874]" pageId="14" pageNumber="961">Figure 9.</emphasis>
|
||
Dorsal and ventral aspect of paratype WAM 109940 of
|
||
<taxonomicName id="4C464D639618FFA15967FCAFFC24FCC3" authority="Weijola & Kraus, 2023" authorityName="Weijola & Kraus" authorityYear="2023" box="[803,976,851,874]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="14" pageNumber="961" phylum="Chordata" rank="species" species="tanimbar" status="sp. nov.">
|
||
<emphasis id="B932EAF29618FFA15967FCAFFC24FCC3" box="[803,976,851,874]" italics="true" pageId="14" pageNumber="961">Varanus tanimbar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20157899618FFA1599CFCAFFBD1FCC3" box="[984,1061,851,874]" pageId="14" pageNumber="961" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3966689618FFA15AE4FA24FEECF9A5" ID-DOI="http://doi.org/10.5281/zenodo.8221648" ID-Zenodo-Dep="8221648" httpUri="https://zenodo.org/record/8221648/files/figure.png" pageId="14" pageNumber="961" startId="14.[160,225,1496,1519]" targetBox="[163,1153,934,1465]" targetPageId="14" targetType="figure">
|
||
<paragraph id="8BF936E09618FFA15AE4FA24FEECF9A5" blockId="14.[160,1155,1496,1548]" pageId="14" pageNumber="961">
|
||
<emphasis id="B932EAF29618FFA15AE4FA24FEFEFA46" bold="true" box="[160,266,1496,1519]" pageId="14" pageNumber="961">Figure 10.</emphasis>
|
||
Adult
|
||
<taxonomicName id="4C464D639618FFA15B17FA24FDF6FA46" authority="Weijola & Kraus, 2023" authorityName="Weijola & Kraus" authorityYear="2023" box="[339,514,1496,1519]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="14" pageNumber="961" phylum="Chordata" rank="species" species="tanimbar" status="sp. nov.">
|
||
<emphasis id="B932EAF29618FFA15B17FA24FDF6FA46" box="[339,514,1496,1519]" italics="true" pageId="14" pageNumber="961">Varanus tanimbar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20157899618FFA15848FA24FDA8FA46" box="[524,604,1496,1519]" pageId="14" pageNumber="961" rank="species">sp. nov.</taxonomicNameLabel>
|
||
male killed by local hunters near Saumlaki, Yamdena (photo VW).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3966689619FFA05AE4FC7CFB70FC3E" ID-DOI="http://doi.org/10.5281/zenodo.8221650" ID-Zenodo-Dep="8221650" box="[160,1156,896,919]" httpUri="https://zenodo.org/record/8221650/files/figure.png" pageId="15" pageNumber="962" startId="15.[160,225,896,919]" targetBox="[163,1153,142,864]" targetPageId="15" targetType="figure">
|
||
<paragraph id="8BF936E09619FFA05AE4FC7CFB70FC3E" blockId="15.[160,1156,896,919]" box="[160,1156,896,919]" pageId="15" pageNumber="962">
|
||
<emphasis id="B932EAF29619FFA05AE4FC7CFEF2FC3E" bold="true" box="[160,262,896,919]" pageId="15" pageNumber="962">Figure 11.</emphasis>
|
||
Juvenile
|
||
<taxonomicName id="4C464D639619FFA05B19FC7CFDF0FC3E" authority="Weijola & Kraus, 2023" authorityName="Weijola & Kraus" authorityYear="2023" box="[349,516,896,919]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="15" pageNumber="962" phylum="Chordata" rank="species" species="tanimbar" status="sp. nov.">
|
||
<emphasis id="B932EAF29619FFA05B19FC7CFDF0FC3E" box="[349,516,896,919]" italics="true" pageId="15" pageNumber="962">Varanus tanimbar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20157899619FFA0584DFC7CFDA7FC3E" box="[521,595,896,919]" pageId="15" pageNumber="962" rank="species">sp. nov.</taxonomicNameLabel>
|
||
caught by local hunters at Saumlaki, Yamdena (photo VW).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3966689619FFA05AE4F958FB70F912" ID-DOI="http://doi.org/10.5281/zenodo.8221652" ID-Zenodo-Dep="8221652" box="[160,1156,1700,1723]" httpUri="https://zenodo.org/record/8221652/files/figure.png" pageId="15" pageNumber="962" startId="15.[160,225,1700,1723]" targetBox="[163,1153,1004,1668]" targetPageId="15" targetType="figure">
|
||
<paragraph id="8BF936E09619FFA05AE4F958FB70F912" blockId="15.[160,1156,1700,1723]" box="[160,1156,1700,1723]" pageId="15" pageNumber="962">
|
||
<emphasis id="B932EAF29619FFA05AE4F958FEF1F912" bold="true" box="[160,261,1700,1723]" pageId="15" pageNumber="962">Figure 12.</emphasis>
|
||
Adult
|
||
<taxonomicName id="4C464D639619FFA05B07F958FE1DF912" authority="Weijola & Kraus, 2023" authorityName="Weijola & Kraus" authorityYear="2023" box="[323,489,1700,1723]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="15" pageNumber="962" phylum="Chordata" rank="species" species="tanimbar" status="sp. nov.">
|
||
<emphasis id="B932EAF29619FFA05B07F958FE1DF912" box="[323,489,1700,1723]" italics="true" pageId="15" pageNumber="962">Varanus tanimbar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20157899619FFA05BAAF958FDC3F912" box="[494,567,1700,1723]" pageId="15" pageNumber="962" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(WAM 109896) Lorulun, Yamdena Island (photo Richard How).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C35C656B9606FFBE5A84FF72FE21FB11" lastPageId="17" lastPageNumber="964" pageId="16" pageNumber="963" type="description">
|
||
<paragraph id="8BF936E09606FFBF5A84FF72FE01FE33" blockId="16.[159,1156,142,410]" pageId="16" pageNumber="963">
|
||
Within the
|
||
<taxonomicName id="4C464D639606FFBF5B71FF72FE62FF0E" box="[309,406,142,167]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="16" pageNumber="963" phylum="Chordata" rank="species" species="indicus">
|
||
<emphasis id="B932EAF29606FFBF5B71FF72FE62FF0E" box="[309,406,142,167]" italics="true" pageId="16" pageNumber="963">V. indicus</emphasis>
|
||
</taxonomicName>
|
||
group,
|
||
<taxonomicName id="4C464D639606FFBF5BA3FF72FD57FF0E" authority="Weijola & Kraus, 2023" authorityName="Weijola & Kraus" authorityYear="2023" box="[487,675,142,167]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="16" pageNumber="963" phylum="Chordata" rank="species" species="tanimbar" status="sp. nov.">
|
||
<emphasis id="B932EAF29606FFBF5BA3FF72FD57FF0E" box="[487,675,142,167]" italics="true" pageId="16" pageNumber="963">Varanus tanimbar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20157899606FFBF58ECFF72FD0FFF0E" box="[680,763,142,167]" pageId="16" pageNumber="963" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is only likely to be confused with the allopatric
|
||
<taxonomicName id="4C464D639606FFBF5B4CFF4DFEA5FF63" authorityName="Bohme, Jacobs, Koppetsch and Schmitz" authorityYear="2019" box="[264,337,177,202]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="16" pageNumber="963" phylum="Chordata" rank="species" species="colei">
|
||
<emphasis id="B932EAF29606FFBF5B4CFF4DFEA5FF63" box="[264,337,177,202]" italics="true" pageId="16" pageNumber="963">V. colei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C464D639606FFBF5B1EFF4DFE21FF63" authorityName="Lesson" authorityYear="1830" box="[346,469,177,202]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="16" pageNumber="963" phylum="Chordata" rank="species" species="douarrha">
|
||
<emphasis id="B932EAF29606FFBF5B1EFF4DFE21FF63" box="[346,469,177,202]" italics="true" pageId="16" pageNumber="963">V. douarrha</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C464D639606FFBF5B9AFF4DFDCEFF63" authorityName="Bohme, Horn and Ziegler" authorityYear="1994" box="[478,570,177,202]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="16" pageNumber="963" phylum="Chordata" rank="species" species="finschi">
|
||
<emphasis id="B932EAF29606FFBF5B9AFF4DFDCEFF63" box="[478,570,177,202]" italics="true" pageId="16" pageNumber="963">V. finschi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C464D639606FFBF5800FF4DFD51FF63" box="[580,677,177,202]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="16" pageNumber="963" phylum="Chordata" rank="species" species="indicus">
|
||
<emphasis id="B932EAF29606FFBF5800FF4DFD51FF63" box="[580,677,177,202]" italics="true" pageId="16" pageNumber="963">V. indicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C464D639606FFBF58EBFF4DFCC1FF63" authorityName="Koch, Arida, Schmitz, Bohme and Ziegler" authorityYear="2009" box="[687,821,177,202]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="16" pageNumber="963" phylum="Chordata" rank="species" species="lirungensis">
|
||
<emphasis id="B932EAF29606FFBF58EBFF4DFCC1FF63" box="[687,821,177,202]" italics="true" pageId="16" pageNumber="963">V. lirungensis</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C464D639606FFBF5913FF4DFC06FF63" authority="Weijola & Kraus, 2023" authorityName="Weijola & Kraus" authorityYear="2023" box="[855,1010,177,202]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="16" pageNumber="963" phylum="Chordata" rank="species" species="louisiadensis" status="sp. nov.">
|
||
<emphasis id="B932EAF29606FFBF5913FF4DFC06FF63" box="[855,1010,177,202]" italics="true" pageId="16" pageNumber="963">V. louisiadensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20157899606FFBF59B3FF4DFBBFFF63" box="[1015,1099,177,202]" pageId="16" pageNumber="963" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, with which it shares a similar dorsal pattern composed of transverse bands of yellow ocelli. It can be distinguished from
|
||
<taxonomicName id="4C464D639606FFBF5BD5FF0AFE2EFEA6" authorityName="Bohme, Jacobs, Koppetsch and Schmitz" authorityYear="2019" box="[401,474,246,271]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="16" pageNumber="963" phylum="Chordata" rank="species" species="colei">
|
||
<emphasis id="B932EAF29606FFBF5BD5FF0AFE2EFEA6" box="[401,474,246,271]" italics="true" pageId="16" pageNumber="963">V. colei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C464D639606FFBF5BA0FF0AFDABFEA6" authorityName="Lesson" authorityYear="1830" box="[484,607,246,271]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="16" pageNumber="963" phylum="Chordata" rank="species" species="douarrha">
|
||
<emphasis id="B932EAF29606FFBF5BA0FF0AFDABFEA6" box="[484,607,246,271]" italics="true" pageId="16" pageNumber="963">V. douarrha</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C464D639606FFBF582DFF0AFD31FEA6" authorityName="Bohme, Horn and Ziegler" authorityYear="1994" box="[617,709,246,271]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="16" pageNumber="963" phylum="Chordata" rank="species" species="finschi">
|
||
<emphasis id="B932EAF29606FFBF582DFF0AFD31FEA6" box="[617,709,246,271]" italics="true" pageId="16" pageNumber="963">V. finschi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C464D639606FFBF588BFF0AFCA2FEA6" authorityName="Koch, Arida, Schmitz, Bohme and Ziegler" authorityYear="2009" box="[719,854,246,271]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="16" pageNumber="963" phylum="Chordata" rank="species" species="lirungensis">
|
||
<emphasis id="B932EAF29606FFBF588BFF0AFCA2FEA6" box="[719,854,246,271]" italics="true" pageId="16" pageNumber="963">V. lirungensis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C464D639606FFBF59D4FF0AFBD8FEA6" authority="Weijola & Kraus, 2023" authorityName="Weijola & Kraus" authorityYear="2023" box="[912,1068,246,271]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="16" pageNumber="963" phylum="Chordata" rank="species" species="louisiadensis" status="sp. nov.">
|
||
<emphasis id="B932EAF29606FFBF59D4FF0AFBD8FEA6" box="[912,1068,246,271]" italics="true" pageId="16" pageNumber="963">V. louisiadensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20157899606FFBF5E75FF0AFB70FEA6" box="[1073,1156,246,271]" pageId="16" pageNumber="963" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the typical presence of a distinct yellow postocular stripe (vs absent in the other species). It can be distinguished from
|
||
<taxonomicName id="4C464D639606FFBF5B80FEC0FDD3FEFC" box="[452,551,316,341]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="16" pageNumber="963" phylum="Chordata" rank="species" species="indicus">
|
||
<emphasis id="B932EAF29606FFBF5B80FEC0FDD3FEFC" box="[452,551,316,341]" italics="true" pageId="16" pageNumber="963">V. indicus</emphasis>
|
||
</taxonomicName>
|
||
by its completely blue-grey tongue (vs bicoloured pink and grey in
|
||
<taxonomicName id="4C464D639606FFBF5B5BFEA2FE75FEDE" box="[287,385,350,375]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="16" pageNumber="963" phylum="Chordata" rank="species" species="indicus">
|
||
<emphasis id="B932EAF29606FFBF5B5BFEA2FE75FEDE" box="[287,385,350,375]" italics="true" pageId="16" pageNumber="963">V. indicus</emphasis>
|
||
</taxonomicName>
|
||
) and higher average scale counts in characters S (154 vs 141.4), T (107.5 vs 96.3) and m (112.5 vs 103.3).
|
||
</paragraph>
|
||
<paragraph id="8BF936E09606FFBF5AE4FE3AFD4EFC29" blockId="16.[160,1156,454,1728]" pageId="16" pageNumber="963">
|
||
<emphasis id="B932EAF29606FFBF5AE4FE3AFE1AFE49" bold="true" box="[160,494,454,480]" italics="true" pageId="16" pageNumber="963">
|
||
Description of the
|
||
<typeStatus id="54FD88429606FFBF5B3BFE3AFE1DFE49" box="[383,489,454,480]" pageId="16" pageNumber="963">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Juvenile, abdomen opened with both single vertical and longitudinal slits; basal 1/3 of tail with longitudinal cut along ventral side. Habitus slender, total length
|
||
<quantity id="4CBE9B059606FFBF5B63FDF1FE71FD8F" box="[295,389,525,550]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="963" unit="mm" value="500.0">500 mm</quantity>
|
||
, SVL
|
||
<quantity id="4CBE9B059606FFBF5BFFFDF1FDEDFD8F" box="[443,537,525,550]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="963" unit="mm" value="200.0">200 mm</quantity>
|
||
, tail length
|
||
<quantity id="4CBE9B059606FFBF58D3FDF1FD00FD8F" box="[663,756,525,550]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="16" pageNumber="963" unit="mm" value="300.0">300 mm</quantity>
|
||
; head elongate. Ground colouration of dorsum, neck and head dark brown to black; densely scattered with pale grey/blue (yellow in life) scales; small blotches and rings/rosettes indistinctly arranged as crossbands over dorsum. These rings and spots are composed of grey/blue (yellow in life) scales; the skin surrounding the scales is grey/black. Tail black, banded on distal 2/3 of length by 17 indistinct bands, each 5–7 scale rows wide. Tail 1.5 times as long as body, relatively elongate (36.14 times as long as high at midlength). Dorsal keel formed by 1–4 enlarged mid-dorsal caudal scales starting at ~
|
||
<quantity id="4CBE9B059606FFBF5809FD03FD55FCB1" box="[589,673,767,792]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="16" pageNumber="963" unit="mm" value="35.0">35 mm</quantity>
|
||
posterior to tail base. Tail round at base, becoming triangular where keel is formed and increasingly laterally compressed to around midlength. Ventral ground colour pale cream, intersected by numerous (23) grey crossbands between cloaca and gular fold.
|
||
</paragraph>
|
||
<paragraph id="8BF936E09606FFBF5A84FC76FBB4FB87" blockId="16.[160,1156,454,1728]" pageId="16" pageNumber="963">
|
||
Ventral sides of limbs cream coloured and intersected by mosaic of narrow grey bands. Pale cream stripe
|
||
<quantity id="4CBE9B059606FFBF5B2DFC51FE14FC6F" box="[361,480,941,966]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="16" pageNumber="963" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
|
||
(~18 scale rows) wide between snout and gular fold; throat rapidly turning marmorated lateral to this. Tail cream ventrally, densely covered by indistinct grey crossbands most clearly visible on basal and distal 1/3 of tail. Limbs black dorsally with white/grey (yellow in life) spots composed of about 1–7 scales.
|
||
</paragraph>
|
||
<paragraph id="8BF936E09606FFBF5A84FBCBFBCFFB72" blockId="16.[160,1156,454,1728]" pageId="16" pageNumber="963">Head 1.96 times as long as wide, dorsal aspect black with small amount of white on most scales, yellow parietal scale, dark temporal band ~5 scales wide, and white postocular stripe ~2 scales wide. Supralabials and infralabials paler than rostrum and dorsal aspect of head. Nostrils slightly elongate, pointed at anterior corner, closer to snout than to eye. Nasal capsules slightly expanded, with shallow sagittal groove on rostrum.</paragraph>
|
||
<paragraph id="8BF936E09606FFBF5A84FB19FBB3F969" blockId="16.[160,1156,454,1728]" pageId="16" pageNumber="963">Nuchal scales of irregular shape on upper neck, round at mid-neck, and increasingly elongate towards shoulders; nuchals domed, with one to several scale pits, surrounded by one or two rows of granules. Dorsal scales oval, slightly irregular in size, domed or keeled (towards abdominal region), most with single pit posteriorly and surrounded by single row of granules. Lateral caudal scales rectangular, elongate, with a single pit centrally or posteriorly. Ventral caudal scales rectangular, elongate, strongly keeled, twice length of mid-dorsal caudals, occurring in row of 10 at midlength of tail. Ventral scales of pubic region polygonal to round, bordered posteriorly by row of granules. Abdominal scales rectangular with rounded corners, bordered posteriorly by row of granules. Ventral scales become increasingly less elongate and round towards chest and finally quadrate or polygonal and irregular in shape near gular fold; ventrals polished, often equipped with single pit and with few granules posteriorly. Gular scales rounded with few granules along posterior edge but become oval to rectangular and elongate towards mental region; mental scales larger laterally, rectangular or polygonal and densely covered in pits.</paragraph>
|
||
<paragraph id="8BF936E09607FFBE5A84FF72FC55FE9B" blockId="17.[160,1156,142,549]" pageId="17" pageNumber="964">Infrafemorals and infratibials round or polygonal, often with a few granules along posterior margin. Infracarpals and infratarsals round, of irregular size, highly domed. Subdigital scales irregular in size and shape, highly domed to almost flat. Twelve rows of enlarged scales along outer margin of fourth toe; four slightly enlarged scales along outer margin of third toe. Claws brown/translucent, sharp, recurved.</paragraph>
|
||
<paragraph id="8BF936E09607FFBE5A84FEC0FE24FD8C" blockId="17.[160,1156,142,549]" pageId="17" pageNumber="964">Occipital scales relatively small, irregular, polygonal. Five enlarged, irregularly shaped supraocular scales, elongate rectangular to pentagonal, densely covered with pits. Scales on forehead and rostrum larger than occipital scales, polygonal, flattened. Supralabials rectangular or pentagonal, covered with pits. Infralabials of irregular size, triangular or polygonal. Tines and dorsal aspect of distal half of tongue grey. Lateral sides of tongue white with only a small amount of grey pigment; distal half of ventral aspect pale blue with a grey central groove.</paragraph>
|
||
<paragraph id="8BF936E09607FFBE5AE4FDACFBA7FDC3" blockId="17.[160,1107,592,618]" box="[160,1107,592,618]" pageId="17" pageNumber="964">
|
||
<emphasis id="B932EAF29607FFBE5AE4FDACFECEFDC3" bold="true" box="[160,314,592,618]" italics="true" pageId="17" pageNumber="964">Scale counts.</emphasis>
|
||
S: 158, XY: 159, DOR: 170, T: 107, VEN: 128, X: 44, m: 109, P: 43, Q: 91.
|
||
</paragraph>
|
||
<paragraph id="8BF936E09607FFBE5AE4FD6AFCDAFD7B" blockId="17.[160,1156,662,722]" pageId="17" pageNumber="964">
|
||
<emphasis id="B932EAF29607FFBE5AE4FD6AFEA0FD19" bold="true" box="[160,340,662,688]" italics="true" pageId="17" pageNumber="964">Measurements.</emphasis>
|
||
SVL:
|
||
<quantity id="4CBE9B059607FFBE5BDCFD6AFE02FD06" box="[408,502,662,687]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="964" unit="mm" value="200.0">200 mm</quantity>
|
||
, F:
|
||
<quantity id="4CBE9B059607FFBE5853FD6AFD81FD06" box="[535,629,662,687]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="964" unit="mm" value="300.0">300 mm</quantity>
|
||
, TL:
|
||
<quantity id="4CBE9B059607FFBE58E7FD6AFCF5FD06" box="[675,769,662,687]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="964" unit="mm" value="500.0">500 mm</quantity>
|
||
, E:
|
||
<quantity id="4CBE9B059607FFBE5967FD6AFC75FD06" box="[803,897,662,687]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" pageId="17" pageNumber="964" unit="mm" value="125.0">125 mm</quantity>
|
||
, D:
|
||
<quantity id="4CBE9B059607FFBE59E3FD6AFC0CFD06" box="[935,1016,662,687]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="17" pageNumber="964" unit="mm" value="75.0">75 mm</quantity>
|
||
, A:
|
||
<quantity id="4CBE9B059607FFBE5E59FD6AFB74FD06" box="[1053,1152,662,687]" metricMagnitude="-2" metricUnit="m" metricValue="3.53" pageId="17" pageNumber="964" unit="mm" value="35.3">35.3 mm</quantity>
|
||
, B:
|
||
<quantity id="4CBE9B059607FFBE5AFFFD45FEF9FD7B" box="[187,269,697,722]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="17" pageNumber="964" unit="mm" value="18.0">18 mm</quantity>
|
||
, C:
|
||
<quantity id="4CBE9B059607FFBE5B70FD45FE6DFD7B" box="[308,409,697,722]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="17" pageNumber="964" unit="mm" value="12.5">12.5 mm</quantity>
|
||
, G:
|
||
<quantity id="4CBE9B059607FFBE5B86FD45FDD3FD7B" box="[450,551,697,722]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="17" pageNumber="964" unit="mm" value="10.8">10.8 mm</quantity>
|
||
, H: 7.0 mm, I: 18.0 mm.
|
||
</paragraph>
|
||
<paragraph id="8BF936E09607FFBE5AE4FD02FB81FCB1" blockId="17.[160,1141,766,792]" box="[160,1141,766,792]" pageId="17" pageNumber="964">
|
||
<emphasis id="B932EAF29607FFBE5AE4FD02FC29FCB1" bold="true" box="[160,989,766,792]" italics="true" pageId="17" pageNumber="964">
|
||
Scale counts, measurements and proportion indices of the
|
||
<typeStatus id="54FD88429607FFBE591EFD02FC79FCB1" box="[858,909,766,792]" pageId="17" pageNumber="964">type</typeStatus>
|
||
series.
|
||
</emphasis>
|
||
See
|
||
<tableCitation id="C6C4035B9607FFBE5E59FD03FB84FCB1" box="[1053,1136,767,792]" captionStart="Table 3" captionStartId="10.[133,189,696,719]" captionTargetPageId="10" captionText="Table 3. Scale counts, measurements (in mm), and mensural ratios for the type series of V. louisiadensis sp. nov. and V. tanimbar sp. nov." httpUri="http://table.plazi.org/id/DF396668961CFFA55AC1FD44FA22FD66" pageId="17" pageNumber="964" tableUuid="DF396668961CFFA55AC1FD44FA22FD66">Table 3</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF936E09607FFBE5AE4FCBFFBD3FBA3" blockId="17.[160,1155,835,1034]" pageId="17" pageNumber="964">
|
||
<emphasis id="B932EAF29607FFBE5AE4FCBFFDDFFCF4" bold="true" box="[160,555,835,861]" italics="true" pageId="17" pageNumber="964">Variation and colouration in life.</emphasis>
|
||
The
|
||
<typeStatus id="54FD88429607FFBE5829FCB8FD2FFCF4" box="[621,731,836,861]" pageId="17" pageNumber="964">paratypes</typeStatus>
|
||
are similar to the
|
||
<typeStatus id="54FD88429607FFBE59F4FCB8FBE0FCF4" box="[944,1044,836,861]" pageId="17" pageNumber="964">holotype</typeStatus>
|
||
although there is some variation in the amount of dark pigmentation on the gular region, venter and ventral surfaces of limbs (
|
||
<figureCitation id="137D2A659607FFBE5BB8FC75FD92FC0B" box="[508,614,905,930]" captionStart="Figure 8" captionStartId="13.[160,225,1531,1554]" captionTargetBox="[163,1153,142,1499]" captionTargetId="figure-71@13.[163,1153,142,1499]" captionTargetPageId="13" captionText="Figure 8. Dorsal, ventral and head profile of the holotype (WAM112255) of Varanus tanimbar sp. nov." figureDoi="http://doi.org/10.5281/zenodo.8221644" httpUri="https://zenodo.org/record/8221644/files/figure.png" pageId="17" pageNumber="964">Figures 8</figureCitation>
|
||
,
|
||
<figureCitation id="137D2A659607FFBE5837FC75FD75FC0B" box="[627,641,905,930]" captionStart="Figure 9" captionStartId="14.[160,225,851,874]" captionTargetBox="[163,1153,142,820]" captionTargetId="figure-40@14.[163,1153,142,820]" captionTargetPageId="14" captionText="Figure 9. Dorsal and ventral aspect of paratype WAM 109940 of Varanus tanimbar sp. nov." figureDoi="http://doi.org/10.5281/zenodo.8221646" httpUri="https://zenodo.org/record/8221646/files/figure.png" pageId="17" pageNumber="964">9</figureCitation>
|
||
). While most individuals have a well-defined yellow temporal band, this was absent in one of seven individuals seen by us (
|
||
<figureCitation id="137D2A659607FFBE5E49FC50FB82FC6C" box="[1037,1142,940,965]" captionStart="Figure 12" captionStartId="15.[160,225,1700,1723]" captionTargetBox="[163,1153,1004,1668]" captionTargetId="figure-3@15.[163,1153,1004,1668]" captionTargetPageId="15" captionText="Figure 12. Adult Varanus tanimbar sp.nov. (WAM 109896) Lorulun, Yamdena Island (photo Richard How)." figureDoi="http://doi.org/10.5281/zenodo.8221652" httpUri="https://zenodo.org/record/8221652/files/figure.png" pageId="17" pageNumber="964">Figure 12</figureCitation>
|
||
). In life, dorsal ground colour black; throat, dorsal spots, and ocelli greenish lemon-yellow (
|
||
<figureCitation id="137D2A659607FFBE5AECFC0DFEC3FBA3" box="[168,311,1009,1034]" captionStart-0="Figure 8" captionStart-1="Figure 9" captionStart-2="Figure 10" captionStart-3="Figure 11" captionStart-4="Figure 12" captionStartId-0="13.[160,225,1531,1554]" captionStartId-1="14.[160,225,851,874]" captionStartId-2="14.[160,225,1496,1519]" captionStartId-3="15.[160,225,896,919]" captionStartId-4="15.[160,225,1700,1723]" captionTargetBox-0="[163,1153,142,1499]" captionTargetBox-1="[163,1153,142,820]" captionTargetBox-2="[163,1153,934,1465]" captionTargetBox-3="[163,1153,142,864]" captionTargetBox-4="[163,1153,1004,1668]" captionTargetId-0="figure-71@13.[163,1153,142,1499]" captionTargetId-1="figure-40@14.[163,1153,142,820]" captionTargetId-2="figure-61@14.[163,1153,934,1465]" captionTargetId-3="figure-24@15.[163,1153,142,865]" captionTargetId-4="figure-3@15.[163,1153,1004,1668]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="14" captionTargetPageId-3="15" captionTargetPageId-4="15" captionText-0="Figure 8. Dorsal, ventral and head profile of the holotype (WAM112255) of Varanus tanimbar sp. nov." captionText-1="Figure 9. Dorsal and ventral aspect of paratype WAM 109940 of Varanus tanimbar sp. nov." captionText-2="Figure 10. Adult Varanus tanimbar sp. nov. male killed by local hunters near Saumlaki, Yamdena (photo VW)." captionText-3="Figure 11. Juvenile Varanus tanimbar sp. nov. caught by local hunters at Saumlaki,Yamdena (photo VW)." captionText-4="Figure 12. Adult Varanus tanimbar sp.nov. (WAM 109896) Lorulun, Yamdena Island (photo Richard How)." figureDoi-0="http://doi.org/10.5281/zenodo.8221644" figureDoi-1="http://doi.org/10.5281/zenodo.8221646" figureDoi-2="http://doi.org/10.5281/zenodo.8221648" figureDoi-3="http://doi.org/10.5281/zenodo.8221650" figureDoi-4="http://doi.org/10.5281/zenodo.8221652" httpUri-0="https://zenodo.org/record/8221644/files/figure.png" httpUri-1="https://zenodo.org/record/8221646/files/figure.png" httpUri-2="https://zenodo.org/record/8221648/files/figure.png" httpUri-3="https://zenodo.org/record/8221650/files/figure.png" httpUri-4="https://zenodo.org/record/8221652/files/figure.png" pageId="17" pageNumber="964">Figures 8–12</figureCitation>
|
||
). Tongue pink basally with grey blue on tines and dorsodistal half.
|
||
</paragraph>
|
||
<paragraph id="8BF936E09607FFBE5AE4FBCAFE21FB11" blockId="17.[160,1156,1078,1208]" pageId="17" pageNumber="964">
|
||
<emphasis id="B932EAF29607FFBE5AE4FBCAFF06FBF9" bold="true" box="[160,242,1078,1104]" italics="true" pageId="17" pageNumber="964">Range.</emphasis>
|
||
<taxonomicName id="4C464D639607FFBE5B45FBCBFE33FBF9" authority="Weijola & Kraus, 2023" authorityName="Weijola & Kraus" authorityYear="2023" box="[257,455,1079,1104]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="17" pageNumber="964" phylum="Chordata" rank="species" species="tanimbar" status="sp. nov.">
|
||
<emphasis id="B932EAF29607FFBE5B45FBCBFE33FBF9" box="[257,455,1079,1104]" italics="true" pageId="17" pageNumber="964">Varanus tanimbar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20157899607FFBE5B8AFBCBFDD1FBF9" box="[462,549,1079,1104]" pageId="17" pageNumber="964" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is known from Yamdena and Selaru – the two largest islands in the Tanimbar Archipelago. The species is also likely to occur on surrounding islands such as Larat, Fordate, Wotap, Wuliaru, Selu and Sera, but its presence on those islands needs confirmation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35C656B9607FFBE5AE4FB18FE02FA02" pageId="17" pageNumber="964" type="biology_ecology">
|
||
<paragraph id="8BF936E09607FFBE5AE4FB18FE02FA02" blockId="17.[160,1156,1252,1451]" pageId="17" pageNumber="964">
|
||
<emphasis id="B932EAF29607FFBE5AE4FB18FEADFB57" bold="true" box="[160,345,1252,1278]" italics="true" pageId="17" pageNumber="964">Natural history.</emphasis>
|
||
There are no specific details about the biology of this species, but it is likely similar in diet and habitat preferences to other closely related species of the
|
||
<taxonomicName id="4C464D639607FFBE5E2BFAFBFF18FAEA" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="17" pageNumber="964" phylum="Chordata" rank="species" species="indicus">
|
||
<emphasis id="B932EAF29607FFBE5E2BFAFBFF18FAEA" italics="true" pageId="17" pageNumber="964">V. indicus</emphasis>
|
||
</taxonomicName>
|
||
group, which are habitat generalists of mangroves, coastal, and lowland forests (
|
||
<bibRefCitation id="EFD74B119607FFBE5AE2FAB0FE41FACC" author="Weijola V & Sweet S" box="[166,437,1356,1381]" pageId="17" pageNumber="964" pagination="14 - 23" refId="ref12838" refString="Weijola V, Sweet S. 2015. A single species of mangrove monitor (Varanus) occupies Ambon, Seram, Buru and Saparua, Moluccas, Indonesia. Amphib Reptile Conserv. 9: 14 - 23." type="journal article" year="2015">Weijola and Sweet 2015</bibRefCitation>
|
||
). During a brief visit to Yamdena in 2009, VW found the species being hunted for food by local residents, and this might well suppress populations in the vicinity of human settlements.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35C656B9607FFBE5AE4FA2AFCB5F969" pageId="17" pageNumber="964" type="discussion">
|
||
<paragraph id="8BF936E09607FFBE5AE4FA2AFEF2FA59" blockId="17.[160,1156,1494,1728]" box="[160,262,1494,1520]" pageId="17" pageNumber="964">
|
||
<emphasis id="B932EAF29607FFBE5AE4FA2AFEF2FA59" bold="true" box="[160,262,1494,1520]" italics="true" pageId="17" pageNumber="964">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF936E09607FFBE5AE4FA06FCB5F969" blockId="17.[160,1156,1494,1728]" pageId="17" pageNumber="964">
|
||
<taxonomicName id="4C464D639607FFBE5AE4FA06FE9CF9BA" authority="Weijola & Kraus, 2023" authorityName="Weijola & Kraus" authorityYear="2023" box="[160,360,1530,1555]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="17" pageNumber="964" phylum="Chordata" rank="species" species="tanimbar" status="sp. nov.">
|
||
<emphasis id="B932EAF29607FFBE5AE4FA06FE9CF9BA" box="[160,360,1530,1555]" italics="true" pageId="17" pageNumber="964">Varanus tanimbar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20157899607FFBE5B35FA06FE3FF9BA" box="[369,459,1530,1555]" pageId="17" pageNumber="964" rank="species">sp. nov.</taxonomicNameLabel>
|
||
formed a well-supported clade together with the Moluccan species
|
||
<taxonomicName id="4C464D639607FFBE5ABCF9E1FEAAF99F" box="[248,350,1565,1590]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="17" pageNumber="964" phylum="Chordata" rank="species" species="indicus">
|
||
<emphasis id="B932EAF29607FFBE5ABCF9E1FEAAF99F" box="[248,350,1565,1590]" italics="true" pageId="17" pageNumber="964">V. indicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C464D639607FFBE5BD1F9E1FDF7F99F" authorityName="Bohme and Ziegler" authorityYear="1997" box="[405,515,1565,1590]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="17" pageNumber="964" phylum="Chordata" rank="species" species="melinus">
|
||
<emphasis id="B932EAF29607FFBE5BD1F9E1FDF7F99F" box="[405,515,1565,1590]" italics="true" pageId="17" pageNumber="964">V. melinus</emphasis>
|
||
</taxonomicName>
|
||
in the BEAST analysis based on the 16S and ND4 markers used by
|
||
<bibRefCitation id="EFD74B119607FFBE5B47F9C3FE10F9F1" author="Weijola V & Vahtera V & Lindqvist C & Kraus F" box="[259,484,1599,1624]" pageId="17" pageNumber="964" pagination="1053 - 1066" refId="ref12987" refString="Weijola V, Vahtera V, Lindqvist C, Kraus F. 2019. A molecular phylogeny for the Pacific monitor lizards (Varanus subgenus Euprepiosaurus) reveals a recent and rapid radiation with high levels of cryptic diversity. Zool J Linn Soc. 186 (4): 1053 - 1066. doi: 10.1093 / zoolinnean / zlz 002." type="journal article" year="2019">
|
||
Weijola
|
||
<emphasis id="B932EAF29607FFBE5B1AF9C3FE7AF9F1" box="[350,398,1599,1624]" italics="true" pageId="17" pageNumber="964">et al</emphasis>
|
||
. (2019)
|
||
</bibRefCitation>
|
||
. Within this Moluccan clade,
|
||
<taxonomicName id="4C464D639607FFBE5976F9C3FC40F9F1" authority="Weijola & Kraus, 2023" authorityName="Weijola & Kraus" authorityYear="2023" box="[818,948,1599,1624]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="17" pageNumber="964" phylum="Chordata" rank="species" species="tanimbar" status="sp. nov.">
|
||
<emphasis id="B932EAF29607FFBE5976F9C3FC40F9F1" box="[818,948,1599,1624]" italics="true" pageId="17" pageNumber="964">V. tanimbar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20157899607FFBE59F8F9C3FBE0F9F1" box="[956,1044,1599,1624]" pageId="17" pageNumber="964" rank="species">sp. nov.</taxonomicNameLabel>
|
||
formed a basal sister lineage to the closely related
|
||
<taxonomicName id="4C464D639607FFBE582DF99EFD3BF9D2" box="[617,719,1634,1659]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="17" pageNumber="964" phylum="Chordata" rank="species" species="indicus">
|
||
<emphasis id="B932EAF29607FFBE582DF99EFD3BF9D2" box="[617,719,1634,1659]" italics="true" pageId="17" pageNumber="964">V. indicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C464D639607FFBE5942F99EFC81F9D2" authorityName="Bohme and Ziegler" authorityYear="1997" box="[774,885,1634,1659]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="17" pageNumber="964" phylum="Chordata" rank="species" species="melinus">
|
||
<emphasis id="B932EAF29607FFBE5942F99EFC81F9D2" box="[774,885,1634,1659]" italics="true" pageId="17" pageNumber="964">V. melinus</emphasis>
|
||
</taxonomicName>
|
||
. ND4 pairwise distances between
|
||
<taxonomicName id="4C464D639607FFBE5B4DF979FE7DF937" authority="Weijola & Kraus, 2023" authorityName="Weijola & Kraus" authorityYear="2023" box="[265,393,1669,1694]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="17" pageNumber="964" phylum="Chordata" rank="species" species="tanimbar" status="sp. nov.">
|
||
<emphasis id="B932EAF29607FFBE5B4DF979FE7DF937" box="[265,393,1669,1694]" italics="true" pageId="17" pageNumber="964">V. tanimbar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20157899607FFBE5BD4F979FE13F937" box="[400,487,1669,1694]" pageId="17" pageNumber="964" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and these two species are 3.4% and 3.9%, respectively, and 2.3% between
|
||
<taxonomicName id="4C464D639607FFBE5B03F95BFE5BF969" box="[327,431,1703,1728]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="17" pageNumber="964" phylum="Chordata" rank="species" species="indicus">
|
||
<emphasis id="B932EAF29607FFBE5B03F95BFE5BF969" box="[327,431,1703,1728]" italics="true" pageId="17" pageNumber="964">V. indicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C464D639607FFBE5BACF95BFCC9F969" authority="(Weijola et al. 2019)" baseAuthorityName="Weijola" baseAuthorityYear="2019" box="[488,829,1703,1728]" class="Squamata" family="Varanidae" genus="Varanus" kingdom="Animalia" pageId="17" pageNumber="964" phylum="Chordata" rank="species" species="melinus">
|
||
<emphasis id="B932EAF29607FFBE5BACF95BFDACF969" box="[488,600,1703,1728]" italics="true" pageId="17" pageNumber="964">V. melinus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD74B119607FFBE5821F95BFCC1F969" author="Weijola V & Vahtera V & Lindqvist C & Kraus F" box="[613,821,1703,1728]" pageId="17" pageNumber="964" pagination="1053 - 1066" refId="ref12987" refString="Weijola V, Vahtera V, Lindqvist C, Kraus F. 2019. A molecular phylogeny for the Pacific monitor lizards (Varanus subgenus Euprepiosaurus) reveals a recent and rapid radiation with high levels of cryptic diversity. Zool J Linn Soc. 186 (4): 1053 - 1066. doi: 10.1093 / zoolinnean / zlz 002." type="journal article" year="2019">
|
||
Weijola
|
||
<emphasis id="B932EAF29607FFBE5885F95BFD05F969" box="[705,753,1703,1728]" italics="true" pageId="17" pageNumber="964">et al</emphasis>
|
||
. 2019
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |