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<mods:title id="1603D5A497DB040A9B2E21E15C1F78DA">Review of the Prabhasa Moore, 1878 genus-group (Lepidoptera, Erebidae, Arctiinae)</mods:title>
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<subSubSection id="C35C6527FF9CFFF79EA93B86FDC1FBD4" pageId="1" pageNumber="384" type="nomenclature">
<paragraph id="8BF936ACFF9CFFF79EA93B86FE43FC54" blockId="1.[151,430,940,966]" box="[151,430,940,966]" pageId="1" pageNumber="384">
<heading id="D0B181C0FF9CFFF79EA93B86FE43FC54" bold="true" box="[151,430,940,966]" fontSize="11" level="1" pageId="1" pageNumber="384" reason="1">
<taxonomicName id="4C464D2FFF9CFFF79EA93B86FE43FC54" ID-CoL="94P5P" authority="Moore, 1878" authorityName="Moore" authorityYear="1878" box="[151,430,940,966]" class="Insecta" family="Erebidae" genus="Prabhasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B932EABEFF9CFFF79EA93B86FE43FC54" bold="true" box="[151,430,940,966]" pageId="1" pageNumber="384">
<emphasis id="B932EABEFF9CFFF79EA93B86FEE6FC54" bold="true" box="[151,267,940,966]" italics="true" pageId="1" pageNumber="384">Prabhasa</emphasis>
<bibRefCitation id="EFD74B5DFF9CFFF79F2C3B86FE43FC54" box="[274,430,940,966]" pageId="1" pageNumber="384" refId="ref10349" refString="Moore, F. (1878) A revision of certain genera of European and Asiatic Lithosiinae, With characters of neW genera and species. Proceedings of the Scientific Meetings of the Zoological Society of London, 1878, 3 - 37, pls. 1 - 3." type="proceedings paper">Moore, 1878</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="8BF936ACFF9CFFF79EA93BD8FDC1FBD4" blockId="1.[151,1075,1010,1124]" pageId="1" pageNumber="384">
Proceedings of the Scientific Meetings of the Zoological Society of London
<emphasis id="B932EABEFF9CFFF79DA33BD8FC3DFB9A" bold="true" box="[925,976,1010,1032]" pageId="1" pageNumber="384">1878</emphasis>
: 25–26.
<emphasis id="B932EABEFF9CFFF79EA93C3BFEC8FBB5" bold="true" box="[151,293,1041,1063]" pageId="1" pageNumber="384">Type species:</emphasis>
<taxonomicName id="4C464D2FFF9CFFF79F153C3BFD8BFBB5" authority="Moore, 1878" authorityName="Moore" authorityYear="1878" box="[299,614,1041,1063]" class="Insecta" family="Erebidae" genus="Prabhasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="venosa">
<emphasis id="B932EABEFF9CFFF79F153C3BFE37FBB5" box="[299,474,1041,1063]" italics="true" pageId="1" pageNumber="384">Prabhasa venosa</emphasis>
<bibRefCitation id="EFD74B5DFF9CFFF79FDE3C38FD8BFBB5" box="[480,614,1041,1063]" pageId="1" pageNumber="384" refId="ref10349" refString="Moore, F. (1878) A revision of certain genera of European and Asiatic Lithosiinae, With characters of neW genera and species. Proceedings of the Scientific Meetings of the Zoological Society of London, 1878, 3 - 37, pls. 1 - 3." type="proceedings paper">Moore, 1878</bibRefCitation>
</taxonomicName>
, by subsequent designation (
<bibRefCitation id="EFD74B5DFF9CFFF79DB63C3BFBC5FBB5" box="[904,1064,1041,1063]" pageId="1" pageNumber="384" refId="ref9744" refString="Hampson, G. F. (1894) The fauna of British India, including Ceylon and Burma. Vol. 2. Moths 2. Arctiidae, Agaristidae, Noctuidae. Taylor &amp; Francis, London, XXII + 609 pp." type="book">Hampson 1894</bibRefCitation>
).
<taxonomicName id="4C464D2FFF9CFFF79EA93C1AFEA7FBD4" authority="(Prabhasa)" authorityName="Hampson" authorityYear="1900" baseAuthorityName="Prabhasa" box="[151,330,1072,1094]" class="Insecta" family="Erebidae" genus="Ilema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B932EABEFF9CFFF79EA93C1AFEA7FBD4" box="[151,330,1072,1094]" italics="true" pageId="1" pageNumber="384">Ilema (Prabhasa)</emphasis>
</taxonomicName>
:
<bibRefCitation id="EFD74B5DFF9CFFF79F693C1BFE18FBD4" box="[343,501,1072,1094]" pageId="1" pageNumber="384" refId="ref9788" refString="Hampson, G. F. (1900) Catalogue of the Arctiadae (Nolinae, Lithosianae) in the British Museum. Catalogue of the Lepidoptera Phalaenae in the British Museum, 2, 1 - 590." type="journal article">Hampson 1900</bibRefCitation>
: 138.
</paragraph>
</subSubSection>
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<paragraph id="8BF936ACFF9CFFF79EA93C64FE4CFBF6" blockId="1.[151,1075,1010,1124]" box="[151,417,1102,1124]" pageId="1" pageNumber="384">
<treatmentCitationGroup id="AB561182FF9CFFF79EA93C64FE4CFBF6" box="[151,417,1102,1124]" pageId="1" pageNumber="384">
<taxonomicName id="4C464D2FFF9CFFF79EA93C64FE70FBF6" ID-CoL="927T7" authority=": Strand 1922: 592" authorityName=": Strand" authorityPageNumber="592" authorityYear="1922" box="[151,413,1102,1124]" class="Insecta" family="Erebidae" genus="Eilema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B932EABEFF9CFFF79EA93C64FF32FBF6" box="[151,223,1102,1124]" italics="true" pageId="1" pageNumber="384">Eilema</emphasis>
:
<treatmentCitation id="0AE710BDFF9CFFF79ED23C64FE70FBF6" author="" box="[236,413,1102,1124]" page="592" pageId="1" pageNumber="384">
<bibRefCitation id="EFD74B5DFF9CFFF79ED23C64FE87FBF6" box="[236,362,1102,1124]" pageId="1" pageNumber="384" refId="ref10540" refString="Strand, E. (1922) Arctiidae: Subfam. Lithosiinae. In: Wagner, H. (Ed.), Lepidopterorum Catalogus. Vol. 26. W. Yunk, Berlin, pp. 1 - 899." type="book">Strand 1922</bibRefCitation>
: 592
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
</subSubSection>
<subSubSection id="C35C6527FF9CFFF79EA93CBBFCBCFAA8" pageId="1" pageNumber="384" type="description">
<paragraph id="8BF936ACFF9CFFF79EA93CBBFCBCFAA8" blockId="1.[151,1436,1169,2022]" pageId="1" pageNumber="384">
<emphasis id="B932EABEFF9CFFF79EA93CBBFEC4FB38" bold="true" box="[151,297,1169,1194]" pageId="1" pageNumber="384">Description.</emphasis>
Wing shape of the typical
<taxonomicName id="4C464D2FFF9CFFF79C5D3CBBFD5FFB38" authorityName=": Strand" authorityYear="1922" box="[611,690,1169,1194]" class="Insecta" family="Erebidae" genus="Eilema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B932EABEFF9CFFF79C5D3CBBFD5FFB38" box="[611,690,1169,1194]" italics="true" pageId="1" pageNumber="384">Eilema</emphasis>
</taxonomicName>
habitus, male has tufted androconial fold along the basal half of costal Vein.
<emphasis id="B932EABEFF9CFFF79F213C9FFE39FB5C" bold="true" box="[287,468,1205,1230]" pageId="1" pageNumber="384">Male genitalia:</emphasis>
uncus bifid, cucullus and sacculus distally with narrow processes; ValVe Ventral edge conVeX, with a setose fold in its basal half; aedeagus thin, Vesica membranous, short, thin.
<emphasis id="B932EABEFF9CFFF79AF43CF3FA71FB60" bold="true" box="[1226,1436,1241,1266]" pageId="1" pageNumber="384">Female genitalia:</emphasis>
ductus bursae weakly sclerotized, dorso-Ventrally flattened, its anterior section membranous; corpus bursae elliptical, membranous, with a large round signum posteriorly.
</paragraph>
</subSubSection>
<subSubSection id="C35C6527FF9CFFF49EF93D6FFD2AFEAD" lastPageId="2" lastPageNumber="385" pageId="1" pageNumber="384" type="discussion">
<paragraph id="8BF936ACFF9CFFF49EF93D6FFD2AFEAD" blockId="1.[151,1436,1169,2022]" lastBlockId="2.[151,1436,151,320]" lastPageId="2" lastPageNumber="385" pageId="1" pageNumber="384">
<emphasis id="B932EABEFF9CFFF79EF93D6FFEB5FACC" bold="true" box="[199,344,1349,1374]" pageId="1" pageNumber="384">Remarks. 1.</emphasis>
Černý &amp; Pinratana (2009) combined
<taxonomicName id="4C464D2FFF9CFFF79D3A3D6FFC9CFACC" authorityName="Moore" authorityYear="1878" box="[772,881,1349,1374]" class="Insecta" family="Erebidae" genus="Prabhasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B932EABEFF9CFFF79D3A3D6FFC9CFACC" box="[772,881,1349,1374]" italics="true" pageId="1" pageNumber="384">Prabhasa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C464D2FFF9CFFF79D943D6FFB49FACC" authority="Moore, 1878" authorityName="Moore" authorityYear="1878" box="[938,1188,1349,1374]" class="Insecta" genus="Zadadra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B932EABEFF9CFFF79D943D6FFBE7FACC" box="[938,1034,1349,1374]" italics="true" pageId="1" pageNumber="384">Zadadra</emphasis>
<bibRefCitation id="EFD74B5DFF9CFFF79A2C3D6CFB49FACC" box="[1042,1188,1350,1374]" pageId="1" pageNumber="384" refId="ref10349" refString="Moore, F. (1878) A revision of certain genera of European and Asiatic Lithosiinae, With characters of neW genera and species. Proceedings of the Scientific Meetings of the Zoological Society of London, 1878, 3 - 37, pls. 1 - 3." type="proceedings paper">Moore, 1878</bibRefCitation>
</taxonomicName>
species in one genus; this is not correct: uncus is bifid in
<taxonomicName id="4C464D2FFF9CFFF79C163D43FD78FA10" authorityName="Moore" authorityYear="1878" box="[552,661,1385,1410]" class="Insecta" family="Erebidae" genus="Prabhasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B932EABEFF9CFFF79C163D43FD78FA10" box="[552,661,1385,1410]" italics="true" pageId="1" pageNumber="384">Prabhasa</emphasis>
</taxonomicName>
, but simple and narrow in
<taxonomicName id="4C464D2FFF9CFFF79DF93D43FBC5FA10" authorityName="Moore" authorityYear="1878" box="[967,1064,1385,1410]" class="Insecta" genus="Zadadra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B932EABEFF9CFFF79DF93D43FBC5FA10" box="[967,1064,1385,1410]" italics="true" pageId="1" pageNumber="384">Zadadra</emphasis>
</taxonomicName>
(
<figureCitation id="137D2A29FF9CFFF79A093D43FB61FA10" box="[1079,1164,1385,1410]" captionStart="FIGURES 30–31" captionStartId="13.[151,264,933,955]" captionTargetBox="[170,1433,193,909]" captionTargetId="figure@13.[151,1436,193,912]" captionTargetPageId="13" captionText="FIGURES 30–31. Lithosiina spp.: male genitalia (genital capsules in ventral vieW). 30, Zadadra distorta, Nepal, slide MWM 31500 Volynkin; 31, Macotasa biplagella, holotype, slide BMNH(E) Arct-2840 (©NHMUK)." httpUri="https://zenodo.org/record/1216630/files/figure.png" pageId="1" pageNumber="384">Fig. 30</figureCitation>
);
<taxonomicName id="4C464D2FFF9CFFF79A9F3D43FAE3FA10" authorityName="Moore" authorityYear="1878" box="[1185,1294,1385,1410]" class="Insecta" family="Erebidae" genus="Prabhasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B932EABEFF9CFFF79A9F3D43FAE3FA10" box="[1185,1294,1385,1410]" italics="true" pageId="1" pageNumber="384">Prabhasa</emphasis>
</taxonomicName>
has a setose Ventral fold of sacculus (on the figure 24 it is pressed aboVe costa), whereas no fold is Visible in
<taxonomicName id="4C464D2FFF9CFFF79ADD3DA7FAAEFA34" authorityName="Moore" authorityYear="1878" box="[1251,1347,1421,1446]" class="Insecta" genus="Zadadra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B932EABEFF9CFFF79ADD3DA7FAAEFA34" box="[1251,1347,1421,1446]" italics="true" pageId="1" pageNumber="384">Zadadra</emphasis>
</taxonomicName>
species (see:
<bibRefCitation id="EFD74B5DFF9CFFF79EE93D98FEB5FA58" box="[215,344,1458,1482]" pageId="1" pageNumber="384" refId="ref9702" refString="Fang, C. (2000) Lepidoptera. Arctiidae. In: Fauna Sinica (Insecta). Vol. 19. Science Press, Beijing, pp. 1 - 590. [in Chinese, With English summary]" type="book">Fang 2000</bibRefCitation>
: 240–244; Kirti &amp; Singh 2015: 129–131); howeVer, some other characters, like cucullus and sacculus apices are similar; in any case, these genera are closely related. ReVision of the genus
<taxonomicName id="4C464D2FFF9CFFF79ADD3DFFFAAEFA7C" authorityName="Moore" authorityYear="1878" box="[1251,1347,1493,1518]" class="Insecta" genus="Zadadra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B932EABEFF9CFFF79ADD3DFFFAAEFA7C" box="[1251,1347,1493,1518]" italics="true" pageId="1" pageNumber="384">Zadadra</emphasis>
</taxonomicName>
will be published later by the authors. Here we treat
<taxonomicName id="4C464D2FFF9CFFF79CA73DD3FCEBF980" authorityName="Moore" authorityYear="1878" box="[665,774,1529,1554]" class="Insecta" family="Erebidae" genus="Prabhasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B932EABEFF9CFFF79CA73DD3FCEBF980" box="[665,774,1529,1554]" italics="true" pageId="1" pageNumber="384">Prabhasa</emphasis>
</taxonomicName>
as a probably monotypic genus. All other taXa preViously included to
<taxonomicName id="4C464D2FFF9CFFF79F233E37FE67F9A4" authorityName="Moore" authorityYear="1878" box="[285,394,1565,1590]" class="Insecta" family="Erebidae" genus="Prabhasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B932EABEFF9CFFF79F233E37FE67F9A4" box="[285,394,1565,1590]" italics="true" pageId="1" pageNumber="384">Prabhasa</emphasis>
</taxonomicName>
belong to different genera.
<emphasis id="B932EABEFF9CFFF79CFB3E37FD34F9A4" bold="true" box="[709,729,1565,1590]" pageId="1" pageNumber="384">2.</emphasis>
Specimens from
<collectingCountry id="F351763CFF9CFFF79D9D3E37FC0BF9A4" box="[931,998,1565,1590]" name="China" pageId="1" pageNumber="384">China</collectingCountry>
treated by authors (Daniel 1954;
<bibRefCitation id="EFD74B5DFF9CFFF79B5A3E34FF3EF9C8" pageId="1" pageNumber="384" refId="ref9702" refString="Fang, C. (2000) Lepidoptera. Arctiidae. In: Fauna Sinica (Insecta). Vol. 19. Science Press, Beijing, pp. 1 - 590. [in Chinese, With English summary]" type="book">Fang 2000</bibRefCitation>
;
<bibRefCitation id="EFD74B5DFF9CFFF79EDE3E6BFDDEF9C8" box="[224,563,1601,1626]" pageId="1" pageNumber="384" refId="ref9545" refString="Dubatolov, V. V. &amp; Zolotuhin, V. V. (2011) Does Eilema Hubner, [1819] (Lepidoptera, Arctiidae, Lithosiinae) present one or several genera? Euroasian Entomological Journal, 10 (3), 367 - 379, 380, color pl. VII." type="journal article">DubatoloV &amp; Zolotuhin 2011</bibRefCitation>
; DubatoloV
<emphasis id="B932EABEFF9CFFF79CFF3E68FD11F9C8" box="[705,764,1601,1626]" italics="true" pageId="1" pageNumber="384">et al.</emphasis>
2012) as
<taxonomicName id="4C464D2FFF9CFFF79D4D3E6BFBDAF9CB" authorityName="Moore" authorityYear="1878" box="[883,1079,1601,1626]" class="Insecta" family="Erebidae" genus="Prabhasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="venosa">
<emphasis id="B932EABEFF9CFFF79D4D3E6BFBDAF9CB" box="[883,1079,1601,1626]" italics="true" pageId="1" pageNumber="384">Prabhasa venosa</emphasis>
</taxonomicName>
belong to “
<taxonomicName id="4C464D2FFF9CFFF79AFD3E6BFADDF9C8" authorityName="Moore" authorityYear="1878" box="[1219,1328,1601,1626]" class="Insecta" family="Erebidae" genus="Prabhasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B932EABEFF9CFFF79AFD3E6BFADDF9C8" box="[1219,1328,1601,1626]" italics="true" pageId="1" pageNumber="384">Prabhasa</emphasis>
</taxonomicName>
”
<taxonomicName id="4C464D2FFF9CFFF79B783E6BFEC7F9EC" authority="Moore, 1878" authorityName="Moore" authorityYear="1878" class="Insecta" family="Erebidae" genus="Prabhasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B932EABEFF9CFFF79B783E6BFA76F9C8" box="[1350,1435,1601,1626]" italics="true" pageId="1" pageNumber="384">costalis</emphasis>
<bibRefCitation id="EFD74B5DFF9CFFF79EA93E4CFEC7F9EC" box="[151,298,1638,1662]" pageId="1" pageNumber="384" refId="ref10349" refString="Moore, F. (1878) A revision of certain genera of European and Asiatic Lithosiinae, With characters of neW genera and species. Proceedings of the Scientific Meetings of the Zoological Society of London, 1878, 3 - 37, pls. 1 - 3." type="proceedings paper">
Moore,
<quantity id="4CBE9B49FF9CFFF79ECC3E4CFEC7F9EC" box="[242,298,1638,1662]" metricMagnitude="1" metricUnit="m" metricValue="4.77012" pageId="1" pageNumber="384" unit="in" value="1878.0">1878</quantity>
</bibRefCitation>
</taxonomicName>
in fact. It is not congeneric with
<taxonomicName id="4C464D2FFF9CFFF79C8D3E4FFC9AF9EF" authorityName="Moore" authorityYear="1878" box="[691,887,1637,1662]" class="Insecta" family="Erebidae" genus="Prabhasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="venosa">
<emphasis id="B932EABEFF9CFFF79C8D3E4FFC9AF9EF" box="[691,887,1637,1662]" italics="true" pageId="1" pageNumber="384">Prabhasa venosa</emphasis>
</taxonomicName>
and belongs to another, yet undescribed genus, which is described below as
<taxonomicName id="4C464D2FFF9CFFF79FD33EA3FC27F930" authority="Dubatolov, Volynkin &amp; Kishida" authorityName="Dubatolov &amp; Volynkin &amp; Kishida" authorityYear="2018" box="[493,970,1673,1698]" class="Insecta" family="Erebidae" genus="Chinasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B932EABEFF9CFFF79FD33EA3FDA6F930" box="[493,587,1673,1698]" italics="true" pageId="1" pageNumber="384">Chinasa</emphasis>
DubatoloV, Volynkin &amp; Kishida
</taxonomicName>
,
<emphasis id="B932EABEFF9CFFF79DE73EA3FB8AF930" bold="true" box="[985,1127,1673,1698]" pageId="1" pageNumber="384">
gen.
<date id="FFF8106CFF9CFFF79A2B3EA3FB8AF930" box="[1045,1127,1673,1698]" pageId="1" pageNumber="384">nov. 3.</date>
</emphasis>
Specimens from
<collectingCountry id="F351763CFF9CFFF79B063EA3FA71F930" box="[1336,1436,1673,1698]" name="Thailand" pageId="1" pageNumber="384">Thailand</collectingCountry>
treated by Černý &amp; Pinratana (2009) as
<taxonomicName id="4C464D2FFF9CFFF79C663E87FCF7F957" authorityName="Moore" authorityYear="1878" box="[600,794,1709,1734]" class="Insecta" family="Erebidae" genus="Prabhasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="venosa">
<emphasis id="B932EABEFF9CFFF79C663E87FCF7F957" box="[600,794,1709,1734]" italics="true" pageId="1" pageNumber="384">Prabhasa venosa</emphasis>
</taxonomicName>
belong to the new species described below as
<taxonomicName id="4C464D2FFF9CFFF79B193E87FDA1F978" authority="Dubatolov, Volynkin &amp; Kishida" authorityName="Dubatolov &amp; Volynkin &amp; Kishida" authorityYear="2018" class="Insecta" family="Erebidae" genus="Macohasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="388" phylum="Arthropoda" rank="species" species="cernyi" status="sp. nov.">
<emphasis id="B932EABEFF9CFFF79B193E87FF30F97B" italics="true" pageId="1" pageNumber="384">Macohasa cernyi</emphasis>
DubatoloV, Volynkin &amp; Kishida
</taxonomicName>
,
<emphasis id="B932EABEFF9CFFF79C663EFBFD5FF978" bold="true" box="[600,690,1745,1770]" pageId="1" pageNumber="384">sp. nov.</emphasis>
, which is closely related to
<taxonomicName id="4C464D2FFF9CFFF79DD03EF8FA71F978" authority="(Hampson, 1905)" baseAuthorityName="Hampson" baseAuthorityYear="1905" box="[1006,1436,1745,1770]" class="Insecta" family="Erebidae" genus="Macotasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B932EABEFF9CFFF79DD03EF8FB22F978" box="[1006,1231,1745,1770]" italics="true" pageId="1" pageNumber="384">Macotasa orientalis</emphasis>
(
<bibRefCitation id="EFD74B5DFF9CFFF79AE13EF8FA79F978" box="[1247,1428,1745,1770]" pageId="1" pageNumber="384" refId="ref9827" refString="Hampson, G. F. (1905) Descriptions of neW Genera and Species of Syntomidae, Arctiidae, Agaristidae, and Noctuidae. The Annals and magazine of natural history, Series 7, 15, 425 - 453." type="journal article">Hampson, 1905</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C464D2FFF9CFFF79EF33EDCFD7CF89C" authority="(Hampson, 1918)" baseAuthorityName="Hampson" baseAuthorityYear="1918" box="[205,657,1781,1806]" class="Insecta" family="Erebidae" genus="Macotasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="dimorpha">
<emphasis id="B932EABEFF9CFFF79EF33EDCFE5BF89C" box="[205,438,1781,1806]" italics="true" pageId="1" pageNumber="384">Macotasa dimorpha</emphasis>
(
<bibRefCitation id="EFD74B5DFF9CFFF79FF53EDCFD64F89C" box="[459,649,1782,1806]" pageId="1" pageNumber="384" refId="ref9958" refString="Hampson, G. F. (1918), Descriptions of neW genera and species of Amatidae, Lithosidae, and Noctuidae. Novitates Zoologicae, 25, 93 - 217. https: // doi. org / 10.5962 / bhl. part. 29763" type="journal article">Hampson, 1918</bibRefCitation>
)
</taxonomicName>
. All three species haVe the male genitalia clearly different from
<taxonomicName id="4C464D2FFF9CFFF79EA93F30FE41F8A0" authority="Moore, 1878" authorityName="Moore" authorityYear="1878" box="[151,428,1818,1842]" class="Insecta" family="Erebidae" genus="Macotasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B932EABEFF9CFFF79EA93F30FEEBF8A3" box="[151,262,1818,1841]" italics="true" pageId="1" pageNumber="384">Macotasa</emphasis>
<bibRefCitation id="EFD74B5DFF9CFFF79F2C3F30FE41F8A0" box="[274,428,1818,1842]" pageId="1" pageNumber="384" refId="ref10349" refString="Moore, F. (1878) A revision of certain genera of European and Asiatic Lithosiinae, With characters of neW genera and species. Proceedings of the Scientific Meetings of the Zoological Society of London, 1878, 3 - 37, pls. 1 - 3." type="proceedings paper">Moore, 1878</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C464D2FFF9CFFF79F823F33FDC4F8A0" authorityName="Moore" authorityYear="1878" box="[444,553,1817,1842]" class="Insecta" family="Erebidae" genus="Prabhasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B932EABEFF9CFFF79F823F33FDC4F8A0" box="[444,553,1817,1842]" italics="true" pageId="1" pageNumber="384">Prabhasa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C464D2FFF9CFFF79C523F33FD21F8A0" authorityName="Moore" authorityYear="1878" box="[620,716,1817,1842]" class="Insecta" genus="Zadadra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B932EABEFF9CFFF79C523F33FD21F8A0" box="[620,716,1817,1842]" italics="true" pageId="1" pageNumber="384">Zadadra</emphasis>
</taxonomicName>
and belong to the new genus, which is described below as
<taxonomicName id="4C464D2FFF9CFFF79EA93F17FD4CF8C4" authority="Dubatolov, Volynkin &amp; Kishida" authorityName="Dubatolov &amp; Volynkin &amp; Kishida" authorityYear="2018" box="[151,673,1853,1878]" class="Insecta" family="Erebidae" genus="Macohasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B932EABEFF9CFFF79EA93F17FEE1F8C4" box="[151,268,1853,1878]" italics="true" pageId="1" pageNumber="384">Macohasa</emphasis>
DubatoloV, Volynkin &amp; Kishida
</taxonomicName>
,
<emphasis id="B932EABEFF9CFFF79C8B3F17FCA3F8C4" bold="true" box="[693,846,1853,1878]" pageId="1" pageNumber="384">
gen.
<date id="FFF8106CFF9CFFF79CC93F17FCA3F8C4" box="[759,846,1853,1878]" pageId="1" pageNumber="384">nov. 4.</date>
</emphasis>
Judging the male genitalia structure,
<taxonomicName id="4C464D2FFF9CFFF79B113F17FDFDF8E8" authority="(Hampson, 1894)" baseAuthorityName="Hampson" baseAuthorityYear="1894" class="Insecta" family="Erebidae" genus="Prabhasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="plumbeomicans">
<emphasis id="B932EABEFF9CFFF79B113F17FEAAF8E8" italics="true" pageId="1" pageNumber="384">Prabhasa plumbeomicans</emphasis>
(
<bibRefCitation id="EFD74B5DFF9CFFF79F6A3F48FDE5F8E8" box="[340,520,1890,1914]" pageId="1" pageNumber="384" refId="ref9744" refString="Hampson, G. F. (1894) The fauna of British India, including Ceylon and Burma. Vol. 2. Moths 2. Arctiidae, Agaristidae, Noctuidae. Taylor &amp; Francis, London, XXII + 609 pp." type="book">Hampson, 1894</bibRefCitation>
)
</taxonomicName>
(illustrated by
<bibRefCitation id="EFD74B5DFF9CFFF79C863F4BFBFDF8E8" box="[696,1040,1889,1914]" pageId="1" pageNumber="384" refId="ref9545" refString="Dubatolov, V. V. &amp; Zolotuhin, V. V. (2011) Does Eilema Hubner, [1819] (Lepidoptera, Arctiidae, Lithosiinae) present one or several genera? Euroasian Entomological Journal, 10 (3), 367 - 379, 380, color pl. VII." type="journal article">DubatoloV &amp; Zolotuhin (2011)</bibRefCitation>
and Kirti &amp; Singh (2015)), belongs to the genus
<taxonomicName id="4C464D2FFF9CFFF79F0F3FAFFE7CF80C" authorityName="Moore" authorityYear="1878" box="[305,401,1925,1950]" class="Insecta" genus="Zadadra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B932EABEFF9CFFF79F0F3FAFFE7CF80C" box="[305,401,1925,1950]" italics="true" pageId="1" pageNumber="384">Zadadra</emphasis>
</taxonomicName>
, therefore the new combination is established here:
<taxonomicName id="4C464D2FFF9CFFF79DC43FAFFF37F850" authority="(Hampson, 1894)" authorityName="Dubatolov &amp; Volynkin &amp; Kishida" authorityYear="2018" baseAuthorityName="Hampson" baseAuthorityYear="1894" class="Insecta" genus="Zadadra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="plumbeomicans" status="comb. nov.">
<emphasis id="B932EABEFF9CFFF79DC43FAFFAF8F80C" box="[1018,1301,1925,1950]" italics="true" pageId="1" pageNumber="384">Zadadra plumbeomicans</emphasis>
(
<bibRefCitation id="EFD74B5DFF9CFFF79B163FACFF3EF850" pageId="1" pageNumber="384" refId="ref9744" refString="Hampson, G. F. (1894) The fauna of British India, including Ceylon and Burma. Vol. 2. Moths 2. Arctiidae, Agaristidae, Noctuidae. Taylor &amp; Francis, London, XXII + 609 pp." type="book">Hampson, 1894</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B932EABEFF9CFFF79ED43F83FE62F850" bold="true" box="[234,399,1961,1986]" pageId="1" pageNumber="384">
<taxonomicNameLabel id="A20157C5FF9CFFF79ED43F83FE82F850" box="[234,367,1961,1986]" pageId="1" pageNumber="384">
comb.
<date id="FFF8106CFF9CFFF79F003F83FE82F850" box="[318,367,1961,1986]" pageId="1" pageNumber="384">nov.</date>
</taxonomicNameLabel>
5.
</emphasis>
The male genitalia structure of
<taxonomicName id="4C464D2FFF9CFFF79D333F83FBF4F850" box="[781,1049,1961,1986]" class="Insecta" family="Erebidae" genus="Prabhasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="monastyrskyi">
<emphasis id="B932EABEFF9CFFF79D333F83FBF4F850" box="[781,1049,1961,1986]" italics="true" pageId="1" pageNumber="384">Prabhasa monastyrskyi</emphasis>
</taxonomicName>
DubatoloV, 2012 (see DubatoloV 2012: fig. 5) is more similar to that of
<taxonomicName id="4C464D2FFF9CFFF79C603FE7FD53F874" authorityName="Moore" authorityYear="1878" box="[606,702,1997,2022]" class="Insecta" genus="Zadadra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B932EABEFF9CFFF79C603FE7FD53F874" box="[606,702,1997,2022]" italics="true" pageId="1" pageNumber="384">Zadadra</emphasis>
</taxonomicName>
species, but, howeVer, differs by the robust, bulb-like, strongly setose distal saccular process, and presence of a robust cornutus in Vesica. Here we proVisionally place this species in the genus
<taxonomicName id="4C464D2FFF9FFFF49F1A3896FE69FF47" authorityName="Moore" authorityYear="1878" box="[292,388,188,213]" class="Insecta" genus="Zadadra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B932EABEFF9FFFF49F1A3896FE69FF47" box="[292,388,188,213]" italics="true" pageId="2" pageNumber="385">Zadadra</emphasis>
</taxonomicName>
:
<taxonomicName id="4C464D2FFF9FFFF49FAD3896FC89FF46" authority="(DubatoloV, 2012)" authorityName="Dubatolov &amp; Volynkin &amp; Kishida" authorityYear="2018" baseAuthorityName="Dubatolov" baseAuthorityYear="2012" box="[403,868,188,213]" class="Insecta" genus="Zadadra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="383" phylum="Arthropoda" rank="species" species="monastyrskyi" status="comb. nov.">
<emphasis id="B932EABEFF9FFFF49FAD3896FD62FF47" box="[403,655,188,213]" italics="true" pageId="2" pageNumber="385">Zadadra monastyrskyi</emphasis>
(DubatoloV, 2012)
</taxonomicName>
,
<emphasis id="B932EABEFF9FFFF49D513896FBE1FF47" bold="true" box="[879,1036,188,213]" pageId="2" pageNumber="385">
comb.
<date id="FFF8106CFF9FFFF49D803896FBE1FF47" box="[958,1036,188,213]" pageId="2" pageNumber="385">nov. 6.</date>
</emphasis>
There are seVeral species described or included in
<taxonomicName id="4C464D2FFF9FFFF49F0738F5FE4BFF6A" authorityName="Moore" authorityYear="1878" box="[313,422,223,248]" class="Insecta" family="Erebidae" genus="Prabhasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B932EABEFF9FFFF49F0738F5FE4BFF6A" box="[313,422,223,248]" italics="true" pageId="2" pageNumber="385">Prabhasa</emphasis>
</taxonomicName>
, but their male genitalia are quite different either from
<taxonomicName id="4C464D2FFF9FFFF49A3138F5FB91FF6A" authorityName="Moore" authorityYear="1878" box="[1039,1148,223,248]" class="Insecta" family="Erebidae" genus="Prabhasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B932EABEFF9FFFF49A3138F5FB91FF6A" box="[1039,1148,223,248]" italics="true" pageId="2" pageNumber="385">Prabhasa</emphasis>
</taxonomicName>
or from
<emphasis id="B932EABEFF9FFFF49AE038F5FADDFF6A" box="[1246,1328,223,248]" italics="true" pageId="2" pageNumber="385">Zadara</emphasis>
,
<taxonomicName id="4C464D2FFF9FFFF49B0038F5FA71FF6A" authority="Dubatolov &amp; Volynkin &amp; Kishida, 2018" authorityName="Dubatolov &amp; Volynkin &amp; Kishida" authorityYear="2018" box="[1342,1436,223,248]" class="Insecta" family="Erebidae" genus="Chinasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="384" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B932EABEFF9FFFF49B0038F5FA71FF6A" box="[1342,1436,223,248]" italics="true" pageId="2" pageNumber="385">Chinasa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C464D2FFF9FFFF49EF8392EFED6FE8F" authority="Dubatolov &amp; Volynkin &amp; Kishida, 2018" authorityName="Dubatolov &amp; Volynkin &amp; Kishida" authorityYear="2018" box="[198,315,260,285]" class="Insecta" family="Erebidae" genus="Macohasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="384" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B932EABEFF9FFFF49EF8392EFED6FE8F" box="[198,315,260,285]" italics="true" pageId="2" pageNumber="385">Macohasa</emphasis>
</taxonomicName>
. Their generic placement will be more carefully scrutinized later by the authors of the present paper in the forthcoming reVision of the genus
<taxonomicName id="4C464D2FFF9FFFF49C61390DFD52FED2" authorityName="Moore" authorityYear="1878" box="[607,703,295,320]" class="Insecta" genus="Zadadra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B932EABEFF9FFFF49C61390DFD52FED2" box="[607,703,295,320]" italics="true" pageId="2" pageNumber="385">Zadadra</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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