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<document id="45479D23CDE9DF91D29EE1EDE09ADBB0" ID-DOI="10.5252/cryptogamie-bryologie2020v41a2" ID-ISSN="1776-0992" ID-Zenodo-Dep="12215441" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719006298856" checkinUser="felipe" docAuthor="Reeb, Catherine &amp; Gradstein, Robbert" docDate="2020" docId="03EF87AA652FFFC4CAF9E6FBFE36FE1A" docLanguage="en" docName="CryptogamieBryologie.41.2.11-34.pdf" docOrigin="Cryptogamie, Bryologie 20 (2)" docSource="http://dx.doi.org/10.5252/cryptogamie-bryologie2020v41a2" docStyle="DocumentStyle:3F4A000F5B047ABA0EAB2943AA9373AE.3:CryptogamieBryologie.2019-.journal_article" docStyleId="3F4A000F5B047ABA0EAB2943AA9373AE" docStyleName="CryptogamieBryologie.2019-.journal_article" docStyleVersion="3" docTitle="Riccardia gasparii Reeb &amp; Gradstein 2020, sp. nov." docType="treatment" docVersion="2" lastPageNumber="24" masterDocId="FFD6FFD26522FFCBC97FE62CFFFBFFEA" masterDocTitle="A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key" masterLastPageNumber="34" masterPageNumber="11" pageNumber="22" updateTime="1719020027017" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="2B3B0D0D4DE8A3A3132C3629FBEF840B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8D778CF3DD55410039428C4BAFD87D57">
<mods:title id="801E6EDF5C6648C5E1BCA9E49CE2D38A">A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key</mods:title>
</mods:titleInfo>
<mods:name id="3660425DE02DE102AAF5DD42D3C6A677" type="personal">
<mods:role id="4403822A81551213A814EE4E3031943C">
<mods:roleTerm id="DAB9C19BBD7D1D35FF73ACF18B420E7A">Author</mods:roleTerm>
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<mods:namePart id="83C6762E8DF0F4E08985382E129962A1">Reeb, Catherine</mods:namePart>
</mods:name>
<mods:name id="B295C1242CB4A8BEA63A94530B214AC9" type="personal">
<mods:role id="B13B8C19DF463869A6D4C9F648F351C1">
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<mods:namePart id="95F6B4C310FEACCF04778B3623069F68">Gradstein, Robbert</mods:namePart>
<mods:affiliation id="3A940C121D70FF1181FEFFFB621D8E0B">Institut de Systématique Évolution Biodiversité, Sorbonne Université, MNHN, CNRS, EPHE, UMR 7205 Museum national d’Histoire naturelle, 57 rue Cuvier 75231 Paris cedex 05 (France)</mods:affiliation>
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<mods:title id="5EFE1F6FA7C3DB5801317D0B4D225E0D">Cryptogamie, Bryologie</mods:title>
</mods:titleInfo>
<mods:part id="1B9B7F69DB943454F93657939D9C527C">
<mods:date id="9DCAC0D6FDE0F5B1D7D1E3E3ACEC4212">2020</mods:date>
<mods:detail id="EFDA99CDD77557105AF847363FBDD8A1" type="pubDate">
<mods:number id="AF582A35C1BF9D16983C3BF420F641CA">2020-01-29</mods:number>
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<mods:number id="CF2CCED58A152EB1781F499294D9B732">20</mods:number>
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<mods:classification id="EDD43FAAE17DF6A5F3ACCBB0D19D2537">journal article</mods:classification>
<mods:identifier id="9DCBEEE556B18C3D716F7902A15B948A" type="DOI">10.5252/cryptogamie-bryologie2020v41a2</mods:identifier>
<mods:identifier id="6893AF5BAF2013D557D0A3AA1D56ED8A" type="ISSN">1776-0992</mods:identifier>
<mods:identifier id="DA9636F16F01F2233092D69B3523ED08" type="Zenodo-Dep">12215441</mods:identifier>
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<treatment id="03EF87AA652FFFC4CAF9E6FBFE36FE1A" ID-DOI="http://doi.org/10.5281/zenodo.12215463" ID-Zenodo-Dep="12215463" LSID="urn:lsid:plazi:treatment:03EF87AA652FFFC4CAF9E6FBFE36FE1A" httpUri="http://treatment.plazi.org/id/03EF87AA652FFFC4CAF9E6FBFE36FE1A" lastPageId="15" lastPageNumber="24" pageId="13" pageNumber="22">
<subSubSection id="C35C6537652FFFC6CAF9E6FBFAADFF18" box="[902,1366,215,242]" pageId="13" pageNumber="22" type="nomenclature">
<paragraph id="8BF936BC652FFFC6CAF9E6FBFAADFF18" blockId="13.[902,1366,215,242]" box="[902,1366,215,242]" pageId="13" pageNumber="22">
<heading id="D0B181D0652FFFC6CAF9E6FBFAADFF18" box="[902,1366,215,242]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="22" reason="4">
<taxonomicName id="4C464D3F652FFFC6CAF9E6FBFB06FF1B" authority="Reeb &amp; Gradst." authorityName="Reeb &amp; Gradstein" authorityYear="2020" box="[902,1277,215,242]" class="Jungermanniopsida" family="Aneuraceae" genus="Riccardia" kingdom="Plantae" order="Metzgeriales" pageId="13" pageNumber="22" phylum="Marchantiophyta" rank="species" species="gasparii" status="sp. nov.">
<emphasis id="B932EAAE652FFFC6CAF9E6FBFBB0FF1B" bold="true" box="[902,1099,215,242]" italics="true" pageId="13" pageNumber="22">Riccardia gasparii</emphasis>
Reeb &amp; Gradst.
</taxonomicName>
<taxonomicNameLabel id="A20157D5652FFFC6CC7BE6F4FAADFF18" box="[1284,1366,216,242]" pageId="13" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35C6537652FFFC6CD4EE6DBFB50FEFB" box="[1073,1195,247,273]" pageId="13" pageNumber="22" type="description">
<paragraph id="8BF936BC652FFFC6CD4EE6DBFB50FEFB" blockId="13.[1073,1195,247,273]" box="[1073,1195,247,273]" pageId="13" pageNumber="22">
<heading id="D0B181D0652FFFC6CD4EE6DBFB50FEFB" box="[1073,1195,247,273]" centered="true" fontSize="11" level="3" pageId="13" pageNumber="22" reason="3">
(
<figureCitation id="137D2A39652FFFC6CD47E6DBFB8FFEFB" box="[1080,1140,247,273]" captionStart="FIG" captionStartId="14.[132,143,1826,1844]" captionTargetBox="[248,1342,219,1805]" captionTargetId="figure-24@14.[736,1279,735,1285]" captionTargetPageId="14" captionText="FIG. 3. — Riccardia gasparii Reeb &amp; Gradst. sp. nov.: A, thalli; B, main axis in cross section; C, ultimate branches in cross section; D, male branch; E, female branches.From Reeb MTV1246 (PC0771050), MTM12134 (PC0763884). Riccardia inconspicua (Steph) Reeb &amp; Bardat:F, thalli;G, portion of ultimate branch with papillae; H, branched papilla (arrow); I, ultimate branch and male branch with papillae; J, main axis in cross section; K, ultimate branch in cross section. From Reeb MTM1295 (PC0763882), Pócs &amp; Mwanjabe 6464/BP, Gardiner s.n. (Type of Aneura exigua), Jungner s.n. (Type of Aneura inconspicua). Riccardia longispica (Steph.) Pearson: L, thalli; M, main axis in cross section; N, ultimate branches in cross section; O, two male branches; P, female branch; Q, male branch. From Konya 9644/D, Bardat PF.2 (PC0763860), Pócs 9612/CD, De Lisle 199 (Type), Jones 1542b. Riccardia longispica (Steph.) Pearson phenotype erosa: R, thalli; S,two calyptra’s; T, main axis in cross section; U, thallus mat showing erect, concave ultimate branches; V, ultimate branch in cross section; W, branch apex with papillae and immature gemmae. From Porley U35b, Pócs 6052/CV, Reeb CRAE193 (PC0763859). Scale bars: A, F, L, R, U, 1 mm; B, C, M-P, Q, S, 500 µm; D, E, G-K, T, 200 µm; H, V, W, 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12215447" httpUri="https://zenodo.org/record/12215447/files/figure.png" pageId="13" pageNumber="22">Fig. 3</figureCitation>
A-E)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35C6537652FFFC6CA52E71DFA54FE88" pageId="13" pageNumber="22" type="diagnosis">
<paragraph id="8BF936BC652FFFC6CA52E71DFA54FE88" blockId="13.[811,1455,304,354]" pageId="13" pageNumber="22">
<smallCapsWord id="8D1FA060652FFFC6CA52E71DFC6EFEAD" baselines="322,323" box="[813,917,305,328]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="13" pageNumber="22">DIAGNOSIS</smallCapsWord>
. — Dioicous. Thallus very fleshy, unwinged. Axis circular to biconvex, branches with rounded margins. Male branches stalked.
</paragraph>
</subSubSection>
<subSubSection id="C35C6537652FFFC6CA52E757FA4BFE2D" pageId="13" pageNumber="22" type="materials_examined">
<paragraph id="8BF936BC652FFFC6CA52E757FA4BFE2D" blockId="13.[811,1457,378,455]" pageId="13" pageNumber="22">
<smallCapsWord id="8D1FA060652FFFC6CA52E757FCA7FE7A" baselines="396,396" box="[813,860,379,402]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="13" pageNumber="22">
<typeStatus id="54FD881E652FFFC6CA52E757FCA7FE7A" box="[813,860,379,402]" pageId="13" pageNumber="22">
<collectionCode id="ED57AE79652FFFC6CA52E757FCA7FE7A" box="[813,860,379,402]" pageId="13" pageNumber="22">TYPE</collectionCode>
</typeStatus>
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. —
<emphasis id="B932EAAE652FFFC6CAFDE756FC0EFE78" bold="true" box="[898,1013,378,402]" pageId="13" pageNumber="22">
<collectingCountry id="F351762C652FFFC6CAFDE756FC0EFE78" box="[898,1013,378,402]" name="Madagascar" pageId="13" pageNumber="22">Madagascar</collectingCountry>
</emphasis>
. Alaotra-Mangoro, Vohimana Reserve, Grande Cascade,
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<geoCoordinate id="EE72507B652FFFC6CD68E7B8FB5AFE46" box="[1047,1185,404,428]" degrees="49" direction="east" minutes="29" orientation="longitude" pageId="13" pageNumber="22" precision="1" seconds="56.6" value="49.499054">49°29’56.6”E</geoCoordinate>
,
<quantity id="4CBE9B59652FFFC6CDD1E7B9FB09FE46" box="[1198,1266,405,429]" metricMagnitude="2" metricUnit="m" metricValue="8.49" pageId="13" pageNumber="22" unit="m" value="849.0">849 m</quantity>
,
<date id="FFF8107C652FFFC6CD80E7B9FA8EFE47" box="[1279,1397,405,429]" pageId="13" pageNumber="22" value="2012-06-30">
30.
<collectionCode id="ED57AE79652FFFC6CC62E7B9FAC1FE47" box="[1309,1338,405,429]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="13" pageNumber="22">VI</collectionCode>
.2012
</date>
,
<emphasis id="B932EAAE652FFFC6CCFAE7B9FC66FE2D" italics="true" pageId="13" pageNumber="22">Reeb MTV1246*</emphasis>
(
<typeStatus id="54FD881E652FFFC6CAD7E783FC25FE2D" box="[936,990,431,455]" pageId="13" pageNumber="22" type="holotype">holo-</typeStatus>
,
<collectionCode id="ED57AE79652FFFC6CA99E783FBFCFE2D" box="[998,1031,431,455]" country="France" lsid="urn:lsid:biocol.org:col:15359" name="Museum National d'Histoire Naturelle, Paris (MNHN) - Non-vascular Plants and Fungi" pageId="13" pageNumber="22" type="Herbarium">PC</collectionCode>
[
<collectionCode id="ED57AE79652FFFC6CD72E783FBD0FE2D" box="[1037,1067,431,455]" country="France" lsid="urn:lsid:biocol.org:col:15359" name="Museum National d'Histoire Naturelle, Paris (MNHN) - Non-vascular Plants and Fungi" pageId="13" pageNumber="22" type="Herbarium">PC</collectionCode>
0771050];
<typeStatus id="54FD881E652FFFC6CDEAE783FB41FE2D" box="[1173,1210,431,455]" pageId="13" pageNumber="22" type="isotype">iso-</typeStatus>
,
<collectionCode id="ED57AE79652FFFC6CDBDE783FB18FE2D" box="[1218,1251,431,455]" country="France" lsid="urn:lsid:biocol.org:col:15359" name="Museum National d'Histoire Naturelle, Paris (MNHN) - Non-vascular Plants and Fungi" pageId="13" pageNumber="22" type="Herbarium">PC</collectionCode>
[
<collectionCode id="ED57AE79652FFFC6CD97E783FAFDFE2D" box="[1256,1286,431,455]" country="France" lsid="urn:lsid:biocol.org:col:15359" name="Museum National d'Histoire Naturelle, Paris (MNHN) - Non-vascular Plants and Fungi" pageId="13" pageNumber="22" type="Herbarium">PC</collectionCode>
0763876],
<collectionCode id="ED57AE79652FFFC6CC10E783FA5EFE2D" box="[1391,1445,431,455]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="13" pageNumber="22" type="Herbarium">TAN</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C35C6537652FFFC6CA52E7F3FB1BFCFF" pageId="13" pageNumber="22" type="etymology">
<paragraph id="8BF936BC652FFFC6CA52E7F3FC51FDFB" blockId="13.[813,1454,479,529]" pageId="13" pageNumber="22">
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. — Dedicated to Gaspard Reeb-Leurent, born
<date id="FFF8107C652FFFC6CA52E7D5FC5DFDFB" box="[813,934,505,529]" pageId="13" pageNumber="22" value="2015-07-21">21/07/2015</date>
.
</paragraph>
<paragraph id="8BF936BC652FFFC6CA52E405FB1BFCFF" blockId="13.[811,1457,553,789]" pageId="13" pageNumber="22">
<smallCapsWord id="8D1FA060652FFFC6CA52E405FC7CFDD5" baselines="570,571" box="[813,903,553,576]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Further" pageId="13" pageNumber="22">FURTHER</smallCapsWord>
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. —
<emphasis id="B932EAAE652FFFC6CDFFE405FB0EFDAB" bold="true" box="[1152,1269,553,577]" pageId="13" pageNumber="22">
<collectingCountry id="F351762C652FFFC6CDFFE405FB0EFDAB" box="[1152,1269,553,577]" name="Madagascar" pageId="13" pageNumber="22">Madagascar</collectingCountry>
</emphasis>
. Alaotra-Mangoro, Vohimana Reserve, Grande Cascade,
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,
<geoCoordinate id="EE72507B652FFFC6CC5DE46FFA52FDB1" box="[1314,1449,579,603]" degrees="49" direction="east" minutes="29" orientation="longitude" pageId="13" pageNumber="22" precision="1" seconds="56.6" value="49.499054">49°29’56.6”E</geoCoordinate>
,
<quantity id="4CBE9B59652FFFC6CA54E472FC96FD9F" box="[811,877,606,630]" metricMagnitude="2" metricUnit="m" metricValue="8.49" pageId="13" pageNumber="22" unit="m" value="849.0">849 m</quantity>
,
<date id="FFF8107C652FFFC6CA09E472FC10FD9C" box="[886,1003,606,630]" pageId="13" pageNumber="22" value="2012-06-30">30.VI.2012</date>
,
<emphasis id="B932EAAE652FFFC6CA8BE472FB76FD9C" box="[1012,1165,606,630]" italics="true" pageId="13" pageNumber="22">Reeb MTV1257</emphasis>
(PC, TAN); Maromizaha Reserve,
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,
<geoCoordinate id="EE72507B652FFFC6CD76E454FB69FD7A" box="[1033,1170,632,656]" degrees="48" direction="east" minutes="27" orientation="longitude" pageId="13" pageNumber="22" precision="1" seconds="50.1" value="48.463917">48°27’50.1”E</geoCoordinate>
,
<quantity id="4CBE9B59652FFFC6CDE2E454FB15FD7A" box="[1181,1262,632,656]" metricMagnitude="3" metricUnit="m" metricValue="1.002" pageId="13" pageNumber="22" unit="m" value="1002.0">1002 m</quantity>
,
<date id="FFF8107C652FFFC6CD85E454FA8CFD7A" box="[1274,1399,632,656]" pageId="13" pageNumber="22" value="2012-07-08">08.VII.2012</date>
,
<emphasis id="B932EAAE652FFFC6CCFAE454FC50FD41" italics="true" pageId="13" pageNumber="22">Reeb MTM12124</emphasis>
* (PC, TAN),
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,
<geoCoordinate id="EE72507B652FFFC6CD9EE4BEFAA2FD40" box="[1249,1369,658,682]" degrees="48" direction="east" minutes="27" orientation="longitude" pageId="13" pageNumber="22" precision="15" seconds="51" value="48.46417">48°27’51”E</geoCoordinate>
,
<quantity id="4CBE9B59652FFFC6CC1AE4BFFA51FD40" box="[1381,1450,659,683]" metricMagnitude="2" metricUnit="m" metricValue="9.209999999999999" pageId="13" pageNumber="22" unit="m" value="921.0">921 m</quantity>
,
<date id="FFF8107C652FFFC6CA51E481FC54FD2F" box="[814,943,685,709]" pageId="13" pageNumber="22" value="2012-07-08">08.VII.2012</date>
,
<emphasis id="B932EAAE652FFFC6CAC6E481FB87FD2F" box="[953,1148,685,709]" italics="true" pageId="13" pageNumber="22">Reeb MTM12127A</emphasis>
*,
<emphasis id="B932EAAE652FFFC6CDEAE481FAE8FD2F" box="[1173,1299,685,709]" italics="true" pageId="13" pageNumber="22">MTM12133</emphasis>
*(PC0763885),
<emphasis id="B932EAAE652FFFC6CA52E4E4FC52FD0A" box="[813,937,712,736]" italics="true" pageId="13" pageNumber="22">MTM12134</emphasis>
* (PC0763884, TAN); Analanjirofo, Mananara Nord, E slope of Mahavoho hills,
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, 49°46.9’47.5”E,
<quantity id="4CBE9B59652FFFC6CC47E4CEFA52FD13" box="[1336,1449,738,762]" metricMagnitude="2" metricUnit="m" metricValue="2.6" metricValueMax="3.0" metricValueMin="2.2" pageId="13" pageNumber="22" unit="m" value="260.0" valueMax="300.0" valueMin="220.0">220-300 m</quantity>
,
<date id="FFF8107C652FFFC6CA52E4D1FC48FCFF" box="[813,947,765,789]" pageId="13" pageNumber="22" value="1998-08-14">14.VIII.1998</date>
,
<emphasis id="B932EAAE652FFFC6CAC2E4D1FB6BFCFF" box="[957,1168,764,789]" italics="true" pageId="13" pageNumber="22">Pócs &amp; Szabo 9878/W</emphasis>
(EGR).
</paragraph>
</subSubSection>
<subSubSection id="C35C6537652FFFC6CA52E501FBE3FC29" pageId="13" pageNumber="22" type="distribution">
<paragraph id="8BF936BC652FFFC6CA52E501FC5CFCB4" blockId="13.[813,1455,812,862]" pageId="13" pageNumber="22">
<smallCapsWord id="8D1FA060652FFFC6CA52E501FC41FCA9" baselines="830,831" box="[813,954,813,836]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="13" pageNumber="22">DISTRIBUTION</smallCapsWord>
. — Alaotra-Mangoro and Analanjirofo regions of
<collectingCountry id="F351762C652FFFC6CA51E56AFC58FCB4" box="[814,931,838,862]" name="Madagascar" pageId="13" pageNumber="22">Madagascar</collectingCountry>
.
</paragraph>
<paragraph id="8BF936BC652FFFC6CA52E55BFBE3FC29" blockId="13.[811,1455,886,963]" pageId="13" pageNumber="22">
<smallCapsWord id="8D1FA060652FFFC6CA52E55BFC78FC67" baselines="904,905" box="[813,899,887,910]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Habitat" pageId="13" pageNumber="22">HABITAT</smallCapsWord>
. — On rocks or boulders sometimes covered with a humus layer, in humid evergreen lowland and montane forests, between about 200 and
<quantity id="4CBE9B59652FFFC6CABBE587FBEFFC29" box="[964,1044,939,963]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="22" unit="m" value="1000.0">1000 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C35C6537652FFFC4CA52E5D9FE36FE1A" lastPageId="15" lastPageNumber="24" pageId="13" pageNumber="22" type="description">
<paragraph id="8BF936BC652FFFC6CA52E5D9FC46FBE7" blockId="13.[813,957,1013,1039]" box="[813,957,1013,1039]" pageId="13" pageNumber="22">
<heading id="D0B181D0652FFFC6CA52E5D9FC46FBE7" box="[813,957,1013,1039]" fontSize="9" level="3" pageId="13" pageNumber="22" reason="5">
<smallCapsWord id="8D1FA060652FFFC6CA52E5D9FC46FBE7" baselines="1032,1032" box="[813,957,1013,1039]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="13" pageNumber="22">DESCRIPTION</smallCapsWord>
</heading>
</paragraph>
<paragraph id="8BF936BC652FFFC6CA52E238FC6DFBC4" blockId="13.[813,918,1044,1070]" box="[813,918,1044,1070]" pageId="13" pageNumber="22">Dioicous.</paragraph>
<paragraph id="8BF936BC652FFFC6CA52E278FC83FB84" blockId="13.[813,888,1108,1134]" box="[813,888,1108,1134]" pageId="13" pageNumber="22">
<heading id="D0B181D0652FFFC6CA52E278FC83FB84" box="[813,888,1108,1134]" fontSize="11" level="3" pageId="13" pageNumber="22" reason="9">
<emphasis id="B932EAAE652FFFC6CA52E278FC83FB84" box="[813,888,1108,1134]" italics="true" pageId="13" pageNumber="22">Thallus</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936BC652FFFC6CA52E258FC67FAC7" blockId="13.[811,1457,1140,1325]" pageId="13" pageNumber="22">
Dull olive green to green when fresh, green to yellow brown in herbarium, very fleshy, rigid but not leathery (easily breaking when manipulated), to
<quantity id="4CBE9B59652FFFC6CD5BE298FBA3FB24" box="[1060,1112,1204,1230]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="22" unit="cm" value="2.0">2 cm</quantity>
long, 1-3-pinnate, not palmate, branches alternate to subopposite, reiterations sometimes present, stolons common, branches mostly wider than the main axis.
</paragraph>
<paragraph id="8BF936BC652FFFC6CA52E378FC6EFA87" blockId="13.[813,917,1364,1389]" box="[813,917,1364,1389]" pageId="13" pageNumber="22">
<heading id="D0B181D0652FFFC6CA52E378FC6EFA87" box="[813,917,1364,1389]" fontSize="11" level="3" pageId="13" pageNumber="22" reason="9">
<emphasis id="B932EAAE652FFFC6CA52E378FC6EFA87" box="[813,917,1364,1389]" italics="true" pageId="13" pageNumber="22">Main axis</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936BC652FFFC6CA52E35FFC22F9E7" blockId="13.[811,1455,1395,1549]" pageId="13" pageNumber="22">Irregularly defined to well-defined in mature thalli, width variable, in cross section circular to ellipsoid at the base, circular to biconvex in the middle, 7-9(-16) cells thick, margins obtuse to rounded, not winged, epidermis cells 3-4× smaller than inner cells.</paragraph>
<paragraph id="8BF936BC652FFFC6CA52E01FFC1AF9A6" blockId="13.[813,993,1586,1612]" box="[813,993,1586,1612]" pageId="13" pageNumber="22">
<heading id="D0B181D0652FFFC6CA52E01FFC1AF9A6" box="[813,993,1586,1612]" fontSize="11" level="3" pageId="13" pageNumber="22" reason="9">
<emphasis id="B932EAAE652FFFC6CA52E01FFC1AF9A6" box="[813,993,1586,1612]" italics="true" pageId="13" pageNumber="22">Primary branches</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936BC652FFFC6CA52E07FFB67F986" blockId="13.[813,1180,1618,1645]" box="[813,1180,1618,1645]" pageId="13" pageNumber="22">
To
<quantity id="4CBE9B59652FFFC6CA2FE07FFC6AF987" box="[848,913,1619,1645]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="22" unit="mm" value="7.0">7 mm</quantity>
long, often overlapping.
</paragraph>
<paragraph id="8BF936BC652FFFC6CA52E0BEFC12F946" blockId="13.[813,1001,1682,1708]" box="[813,1001,1682,1708]" pageId="13" pageNumber="22">
<heading id="D0B181D0652FFFC6CA52E0BEFC12F946" box="[813,1001,1682,1708]" fontSize="11" level="3" pageId="13" pageNumber="22" reason="9">
<emphasis id="B932EAAE652FFFC6CA52E0BEFC12F946" box="[813,1001,1682,1708]" italics="true" pageId="13" pageNumber="22">Ultimate branches</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936BC652FFFC6CA52E09EFAA3F8C6" blockId="13.[811,1455,1714,1836]" pageId="13" pageNumber="22">
Tongue-shaped to oblong, often sinuose (horizontally curved), to
<quantity id="4CBE9B59652FFFC6CA3AE0FEFC78F906" box="[837,899,1746,1772]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="13" pageNumber="22" unit="mm" value="6.0">6 mm</quantity>
long and
<quantity id="4CBE9B59652FFFC6CA98E0FEFBC2F906" box="[999,1081,1746,1772]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="13" pageNumber="22" unit="mm" value="1.8">1.8 mm</quantity>
wide, in cross section plano-convex, 5-6(-7) cells thick, margins obtuse, not winged, epidermis cells 2-3× smaller than inner cells, branch apex rounded.
</paragraph>
<paragraph id="8BF936BC652FFFC6CA52E17DFC18F881" blockId="13.[813,995,1873,1899]" box="[813,995,1873,1899]" pageId="13" pageNumber="22">
<heading id="D0B181D0652FFFC6CA52E17DFC18F881" box="[813,995,1873,1899]" fontSize="11" level="3" pageId="13" pageNumber="22" reason="9">
<emphasis id="B932EAAE652FFFC6CA52E17DFC18F881" box="[813,995,1873,1899]" italics="true" pageId="13" pageNumber="22">Mucilage papillae</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936BC652FFFC6CA52E15DFBE2F861" blockId="13.[813,1049,1905,1932]" box="[813,1049,1905,1932]" pageId="13" pageNumber="22">Apical, not persistent.</paragraph>
<paragraph id="8BF936BC652FFFC6CA52E19DFC6EF821" blockId="13.[813,917,1969,1995]" box="[813,917,1969,1995]" pageId="13" pageNumber="22">
<heading id="D0B181D0652FFFC6CA52E19DFC6EF821" box="[813,917,1969,1995]" fontSize="11" level="3" pageId="13" pageNumber="22" reason="9">
<emphasis id="B932EAAE652FFFC6CA52E19DFC6EF821" box="[813,917,1969,1995]" italics="true" pageId="13" pageNumber="22">Oil bodies</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936BC652FFFC6CA52E1FEFC6EF806" blockId="13.[813,917,2002,2028]" box="[813,917,2002,2028]" pageId="13" pageNumber="22">
<collectingRegion id="4982F85E652FFFC6CA52E1FEFCA3F806" box="[813,856,2002,2028]" country="Cameroon" name="North" pageId="13" pageNumber="22">Not</collectingRegion>
seen.
</paragraph>
<caption id="DF396634652CFFC5C9FBE10EFE3EF800" ID-DOI="http://doi.org/10.5281/zenodo.12215447" ID-Zenodo-Dep="12215447" httpUri="https://zenodo.org/record/12215447/files/figure.png" pageId="14" pageNumber="23" startId="14.[132,143,1826,1844]" targetBox="[248,1342,219,1805]" targetPageId="14" targetType="figure">
<paragraph id="8BF936BC652CFFC5C9FBE10EFE3EF800" blockId="14.[132,1457,1826,2026]" pageId="14" pageNumber="23">
<smallCapsWord id="8D1FA060652CFFC5C9FBE10EFF66F8D9" baselines="1839,1840" box="[132,157,1826,1844]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="14" pageNumber="23">FIG</smallCapsWord>
. 3. —
<taxonomicName id="4C464D3F652CFFC5C9A9E10EFE0EF8DE" authority="Reeb &amp; Gradst." authorityName="Reeb &amp; Gradstein" authorityYear="2020" box="[214,501,1826,1844]" class="Jungermanniopsida" family="Aneuraceae" genus="Riccardia" kingdom="Plantae" order="Metzgeriales" pageId="14" pageNumber="23" phylum="Marchantiophyta" rank="species" species="gasparii" status="sp. nov.">
<emphasis id="B932EAAE652CFFC5C9A9E10EFE90F8DE" box="[214,363,1826,1844]" italics="true" pageId="14" pageNumber="23">Riccardia gasparii</emphasis>
Reeb &amp; Gradst.
</taxonomicName>
<taxonomicNameLabel id="A20157D5652CFFC5C886E10EFDBBF8DE" box="[505,576,1826,1844]" pageId="14" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
:
<emphasis id="B932EAAE652CFFC5CB39E10EFDA8F8DE" bold="true" box="[582,595,1826,1844]" pageId="14" pageNumber="23">A</emphasis>
, thalli;
<emphasis id="B932EAAE652CFFC5CBEFE10EFD66F8DE" bold="true" box="[656,669,1826,1844]" pageId="14" pageNumber="23">B</emphasis>
, main axis in cross section;
<emphasis id="B932EAAE652CFFC5CAF1E10EFC67F8DE" bold="true" box="[910,924,1826,1844]" pageId="14" pageNumber="23">C</emphasis>
, ultimate branches in cross section;
<emphasis id="B932EAAE652CFFC5CDACE10EFB1AF8DE" bold="true" box="[1235,1249,1826,1844]" pageId="14" pageNumber="23">D</emphasis>
, male branch;
<emphasis id="B932EAAE652CFFC5CC1FE10EFA97F8DE" bold="true" box="[1376,1388,1826,1844]" pageId="14" pageNumber="23">E</emphasis>
, female branches.From
<emphasis id="B932EAAE652CFFC5C874E115FE6FF8A1" box="[267,404,1849,1867]" italics="true" pageId="14" pageNumber="23">Reeb MTV1246 (</emphasis>
PC0771050
<emphasis id="B932EAAE652CFFC5C889E115FD98F8A1" box="[502,611,1849,1867]" italics="true" pageId="14" pageNumber="23">), MTM12134</emphasis>
(PC0763884).
<taxonomicName id="4C464D3F652CFFC5CBA2E115FBAAF8A1" authority="(Steph) Reeb &amp; Bardat" authorityName="Reeb &amp; Bardat" baseAuthorityName="Steph" box="[733,1105,1849,1867]" class="Jungermanniopsida" family="Aneuraceae" genus="Riccardia" kingdom="Plantae" order="Metzgeriales" pageId="14" pageNumber="23" phylum="Marchantiophyta" rank="species" species="inconspicua">
<emphasis id="B932EAAE652CFFC5CBA2E115FC68F8A1" box="[733,915,1849,1867]" italics="true" pageId="14" pageNumber="23">Riccardia inconspicua</emphasis>
(Steph) Reeb &amp; Bardat
</taxonomicName>
:
<emphasis id="B932EAAE652CFFC5CD28E115FB99F8A1" bold="true" box="[1111,1122,1849,1867]" pageId="14" pageNumber="23">F</emphasis>
, thalli;
<emphasis id="B932EAAE652CFFC5CDE3E115FB51F8A1" bold="true" box="[1180,1194,1849,1867]" pageId="14" pageNumber="23">G</emphasis>
, portion of ultimate branch with papillae;
<emphasis id="B932EAAE652CFFC5C9AFE17CFF25F888" bold="true" box="[208,222,1872,1890]" pageId="14" pageNumber="23">H</emphasis>
, branched papilla (arrow);
<emphasis id="B932EAAE652CFFC5C8C2E17CFE38F888" bold="true" box="[445,451,1872,1890]" pageId="14" pageNumber="23">I</emphasis>
, ultimate branch and male branch with papillae;
<emphasis id="B932EAAE652CFFC5CA23E17CFC9DF888" bold="true" box="[860,870,1872,1890]" pageId="14" pageNumber="23">J</emphasis>
, main axis in cross section;
<emphasis id="B932EAAE652CFFC5CD2AE17CFB99F888" bold="true" box="[1109,1122,1872,1890]" pageId="14" pageNumber="23">K</emphasis>
, ultimate branch in cross section. From
<emphasis id="B932EAAE652CFFC5C9FBE14BFEF2F893" box="[132,265,1895,1913]" italics="true" pageId="14" pageNumber="23">Reeb MTM1295</emphasis>
(PC0763882),
<emphasis id="B932EAAE652CFFC5C8FBE14AFD2CF893" box="[388,727,1894,1913]" italics="true" pageId="14" pageNumber="23">Pócs &amp; Mwanjabe 6464/BP, Gardiner s.n.</emphasis>
(Type of
<taxonomicName id="4C464D3F652CFFC5CA5EE14BFC61F893" box="[801,922,1895,1913]" class="Jungermanniopsida" family="Aneuraceae" genus="Aneura" kingdom="Plantae" order="Metzgeriales" pageId="14" pageNumber="23" phylum="Marchantiophyta" rank="species" species="exigua">Aneura exigua</taxonomicName>
),
<emphasis id="B932EAAE652CFFC5CAD9E14BFBF6F893" box="[934,1037,1895,1913]" italics="true" pageId="14" pageNumber="23">Jungner s.n.</emphasis>
(Type of
<taxonomicName id="4C464D3F652CFFC5CD29E14BFB01F893" box="[1110,1274,1895,1913]" class="Jungermanniopsida" family="Aneuraceae" genus="Aneura" kingdom="Plantae" order="Metzgeriales" pageId="14" pageNumber="23" phylum="Marchantiophyta" rank="species" species="inconspicua">
<emphasis id="B932EAAE652CFFC5CD29E14BFB01F893" box="[1110,1274,1895,1913]" italics="true" pageId="14" pageNumber="23">Aneura inconspicua</emphasis>
</taxonomicName>
).
<taxonomicName id="4C464D3F652CFFC5CC76E14BFEEAF865" authority="(Steph.) Pearson" authorityName="Pearson" baseAuthorityName="Steph." class="Jungermanniopsida" family="Aneuraceae" genus="Riccardia" kingdom="Plantae" order="Metzgeriales" pageId="14" pageNumber="23" phylum="Marchantiophyta" rank="species" species="longispica">
<emphasis id="B932EAAE652CFFC5CC76E14BFA54F893" box="[1289,1455,1895,1913]" italics="true" pageId="14" pageNumber="23">Riccardia longispica</emphasis>
(Steph.) Pearson
</taxonomicName>
:
<emphasis id="B932EAAE652CFFC5C866E151FEDFF865" bold="true" box="[281,292,1917,1935]" pageId="14" pageNumber="23">L</emphasis>
, thalli;
<emphasis id="B932EAAE652CFFC5C81FE151FE8AF865" bold="true" box="[352,369,1917,1935]" pageId="14" pageNumber="23">M</emphasis>
, main axis in cross section;
<emphasis id="B932EAAE652CFFC5CB22E151FD90F865" bold="true" box="[605,619,1917,1935]" pageId="14" pageNumber="23">N</emphasis>
, ultimate branches in cross section;
<emphasis id="B932EAAE652CFFC5CAE1E151FC56F865" bold="true" box="[926,941,1917,1935]" pageId="14" pageNumber="23">O</emphasis>
, two male branches;
<emphasis id="B932EAAE652CFFC5CD1FE151FB97F865" bold="true" box="[1120,1132,1917,1935]" pageId="14" pageNumber="23">P</emphasis>
, female branch;
<emphasis id="B932EAAE652CFFC5CD86E151FAF3F865" bold="true" box="[1273,1288,1917,1935]" pageId="14" pageNumber="23">Q</emphasis>
, male branch. From
<emphasis id="B932EAAE652CFFC5C9FBE1B8FF00F84C" box="[132,251,1940,1958]" italics="true" pageId="14" pageNumber="23">Konya 9644/D</emphasis>
,
<emphasis id="B932EAAE652CFFC5C877E1B8FE93F84C" box="[264,360,1940,1958]" italics="true" pageId="14" pageNumber="23">Bardat PF.2</emphasis>
(PC0763860),
<emphasis id="B932EAAE652CFFC5C898E1B8FD2DF84C" box="[487,726,1940,1958]" italics="true" pageId="14" pageNumber="23">Pócs 9612/CD, De Lisle 199</emphasis>
(Type),
<emphasis id="B932EAAE652CFFC5CA66E1B8FC7DF84C" box="[793,902,1940,1958]" italics="true" pageId="14" pageNumber="23">Jones 1542b</emphasis>
.
<taxonomicName id="4C464D3F652CFFC5CAEEE1B8FB36F84C" authority="(Steph.) Pearson" authorityName="Pearson" baseAuthorityName="Steph." box="[913,1229,1940,1958]" class="Jungermanniopsida" family="Aneuraceae" genus="Riccardia" kingdom="Plantae" order="Metzgeriales" pageId="14" pageNumber="23" phylum="Marchantiophyta" rank="species" species="longispica">
<emphasis id="B932EAAE652CFFC5CAEEE1B8FBC2F84C" box="[913,1081,1940,1958]" italics="true" pageId="14" pageNumber="23">Riccardia longispica</emphasis>
(Steph.) Pearson
</taxonomicName>
phenotype
<taxonomicName id="4C464D3F652CFFC5CC4EE1B8FAA5F84C" box="[1329,1374,1940,1958]" class="Jungermanniopsida" family="Aneuraceae" genus="Riccardia" kingdom="Plantae" order="Metzgeriales" pageId="14" pageNumber="23" phylum="Marchantiophyta" rank="species" species="erosa">
<emphasis id="B932EAAE652CFFC5CC4EE1B8FAA5F84C" box="[1329,1374,1940,1958]" italics="true" pageId="14" pageNumber="23">erosa</emphasis>
</taxonomicName>
:
<emphasis id="B932EAAE652CFFC5CC15E1B8FA8CF84C" bold="true" box="[1386,1399,1940,1958]" pageId="14" pageNumber="23">R</emphasis>
, thalli;
<emphasis id="B932EAAE652CFFC5C9FBE186FF6BF856" bold="true" box="[132,144,1962,1980]" pageId="14" pageNumber="23">S</emphasis>
,two calyptra’s;
<emphasis id="B932EAAE652CFFC5C86AE187FEDBF857" bold="true" box="[277,288,1963,1981]" pageId="14" pageNumber="23">T</emphasis>
, main axis in cross section;
<emphasis id="B932EAAE652CFFC5CB6EE187FDE4F857" bold="true" box="[529,543,1963,1981]" pageId="14" pageNumber="23">U</emphasis>
, thallus mat showing erect, concave ultimate branches;
<emphasis id="B932EAAE652CFFC5CA84E187FBFCF857" bold="true" box="[1019,1031,1963,1981]" pageId="14" pageNumber="23">V</emphasis>
, ultimate branch in cross section;
<emphasis id="B932EAAE652CFFC5CC56E187FAC0F857" bold="true" box="[1321,1339,1963,1981]" pageId="14" pageNumber="23">W</emphasis>
, branch apex with papillae and immature gemmae. From
<emphasis id="B932EAAE652CFFC5C892E1EDFDA8F839" box="[493,595,1985,2003]" italics="true" pageId="14" pageNumber="23">Porley U35b</emphasis>
,
<emphasis id="B932EAAE652CFFC5CB21E1EDFC9BF839" box="[606,864,1985,2003]" italics="true" pageId="14" pageNumber="23">Pócs 6052/CV, Reeb CRAE193</emphasis>
(PC0763859). Scale bars: A, F, L, R, U, 1 mm; B, C, M-P, Q, S, 500 µm; D, E, G-K, T, 200 µm; H, V, W, 100 µm.
</paragraph>
</caption>
<paragraph id="8BF936BC652DFFC4C9FBE6FBFEE2FF1B" blockId="15.[132,281,215,241]" box="[132,281,215,241]" pageId="15" pageNumber="24">
<heading id="D0B181D0652DFFC4C9FBE6FBFEE2FF1B" box="[132,281,215,241]" fontSize="11" level="3" pageId="15" pageNumber="24" reason="9">
<emphasis id="B932EAAE652DFFC4C9FBE6FBFEE2FF1B" box="[132,281,215,241]" italics="true" pageId="15" pageNumber="24">Male branches</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936BC652DFFC4C9FBE6DBFD17FEDB" blockId="15.[132,775,247,306]" pageId="15" pageNumber="24">On the main axis, solitary, stalked, with a scale-like wing narrower than the axis, the wing crenulate by larger cells.</paragraph>
<paragraph id="8BF936BC652DFFC4C9FBE77BFED7FE9B" blockId="15.[132,300,343,369]" box="[132,300,343,369]" pageId="15" pageNumber="24">
<heading id="D0B181D0652DFFC4C9FBE77BFED7FE9B" box="[132,300,343,369]" fontSize="11" level="3" pageId="15" pageNumber="24" reason="9">
<emphasis id="B932EAAE652DFFC4C9FBE77BFED7FE9B" box="[132,300,343,369]" italics="true" pageId="15" pageNumber="24">Female branches</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936BC652DFFC4C9FBE75BFCF2FE7B" blockId="15.[132,777,375,401]" box="[132,777,375,401]" pageId="15" pageNumber="24">Laterally inserted, straight, bordered by a ciliate, scale-like wing.</paragraph>
<paragraph id="8BF936BC652DFFC4C9FDE79BFF25FE3B" blockId="15.[130,222,439,465]" box="[130,222,439,465]" pageId="15" pageNumber="24">
<heading id="D0B181D0652DFFC4C9FDE79BFF25FE3B" box="[130,222,439,465]" fontSize="11" level="3" pageId="15" pageNumber="24" reason="9">
<emphasis id="B932EAAE652DFFC4C9FDE79BFF25FE3B" box="[130,222,439,465]" italics="true" pageId="15" pageNumber="24">Gemmae</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936BC652DFFC4C9FBE7FAFE36FE1A" blockId="15.[132,461,470,497]" box="[132,461,470,497]" pageId="15" pageNumber="24">Occasionally present, 2-celled.</paragraph>
</subSubSection>
</treatment>
</document>
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