380 lines
54 KiB
XML
380 lines
54 KiB
XML
<document id="E2F318501A5D64123A7A403E83912F74" ID-CLB-Dataset="20273" ID-DOI="10.11646/zootaxa.5115.1.9" ID-GBIF-Dataset="183d6d80-cdfb-4ec4-9500-17cb2e89f635" ID-ISSN="1175-5326" ID-Zenodo-Dep="6346178" ID-ZooBank="5FF52BDB-1928-4C4A-ACA2-47B5B869A611" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646989739340" checkinUser="plazi" docAuthor="Starkevich, Pavel, Podėnas, Sigitas & Sivell, Duncan" docDate="2022" docId="03B38A2C5A17265029BAFEF1FAA09999" docLanguage="en" docName="zootaxa.5115.1.9.pdf" docOrigin="Zootaxa 5115 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tipula (Vestiplex) tanycera Alexander 1961" docType="treatment" docVersion="6" lastPageNumber="144" masterDocId="FF8AF2545A1D265D292DFF92FF859F25" masterDocTitle="Tipula (Vestiplex) levisoni sp. nov., a new cranefly (Diptera, Tipulidae) from the Western Himalayas" masterLastPageNumber="145" masterPageNumber="131" pageNumber="141" updateTime="1699317865228" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="05B63C2A473C8230C7800825CF8BF223">Tipula (Vestiplex) levisoni sp. nov., a new cranefly (Diptera, Tipulidae) from the Western Himalayas</mods:title>
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<mods:namePart id="BE9A9C6451FAA451FC9053DEEB375B34">Starkevich, Pavel</mods:namePart>
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<mods:affiliation id="DBB6CC7B51BB0334571036D9C15F3D39">Nature Research Centre, Akademijos str. 2, LT- 08412 Vilnius, Lithuania</mods:affiliation>
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<mods:nameIdentifier id="4B3F7968D6AD1C60F366C84D590089D7" type="email">pavel.starkevic@gmail.com</mods:nameIdentifier>
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<mods:namePart id="AC38041E769BD56E30C770A47C71527E">Podėnas, Sigitas</mods:namePart>
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<mods:affiliation id="DCC0C948CDB668BC05B2B9E693928FB6">Nature Research Centre, Akademijos str. 2, LT- 08412 Vilnius, Lithuania & Life Sciences Centre of Vilnius University, Sauletekio str. 7, LT- 10257 Vilnius, Lithuania</mods:affiliation>
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<mods:namePart id="71B07A8D84E3CD99D23402A99BEEBFA9">Sivell, Duncan</mods:namePart>
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<mods:affiliation id="B5D1CBF2FF3910DA228809EA3CC7B4E0">Natural History Museum, Cromwell Road, London, SW 7 5 BD, United Kingdom</mods:affiliation>
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<mods:date id="866D24CD699D514161FA99EA1F2A1C88">2022</mods:date>
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<treatment id="03B38A2C5A17265029BAFEF1FAA09999" ID-DOI="http://doi.org/10.5281/zenodo.6346176" ID-GBIF-Taxon="193941083" ID-Zenodo-Dep="6346176" LSID="urn:lsid:plazi:treatment:03B38A2C5A17265029BAFEF1FAA09999" httpUri="http://treatment.plazi.org/id/03B38A2C5A17265029BAFEF1FAA09999" lastPageId="13" lastPageNumber="144" pageId="10" pageNumber="141">
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<paragraph id="8BA53B3A5A17265729BAFEF1FDE19E5B" blockId="10.[151,612,355,419]" box="[151,612,355,382]" pageId="10" pageNumber="141">
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<taxonomicName id="4C1A40B95A17265729BAFEF1FDE19E5B" ID-CoL="576QL" authority="Alexander" authorityName="Alexander" authorityYear="1961" box="[151,612,355,382]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="141" phylum="Arthropoda" rank="species" species="tanycera" subGenus="Vestiplex">
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<emphasis id="B96EE7285A17265729BAFEF1FDE19E5B" bold="true" box="[151,612,355,382]" pageId="10" pageNumber="141">
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<emphasis id="B96EE7285A17265729BAFEF1FF639E5B" bold="true" box="[151,230,355,382]" italics="true" pageId="10" pageNumber="141">Tipula</emphasis>
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(
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<emphasis id="B96EE7285A17265729DAFEF1FEE69E5B" bold="true" box="[247,355,355,382]" italics="true" pageId="10" pageNumber="141">Vestiplex</emphasis>
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)
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<emphasis id="B96EE7285A1726572859FEF6FE599E5B" bold="true" box="[372,476,356,382]" italics="true" pageId="10" pageNumber="141">tanycera</emphasis>
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Alexander
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</emphasis>
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</taxonomicName>
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</paragraph>
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<subSubSection id="C30068B15A17265729BAFE1AFEA29E86" box="[151,295,392,419]" pageId="10" pageNumber="141" type="description">
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<paragraph id="8BA53B3A5A17265729BAFE1AFEA29E86" blockId="10.[151,612,355,419]" box="[151,295,392,419]" pageId="10" pageNumber="141">
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(
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<figureCitation id="132127BF5A17265729B2FE1AFEA59E86" box="[159,288,392,419]" captionStart-0="FIGURES 38–40" captionStart-1="FIGURES 41–50" captionStart-2="FIGURES 51–56" captionStartId-0="10.[151,264,1964,1988]" captionStartId-1="11.[151,264,1865,1889]" captionStartId-2="12.[151,264,1887,1911]" captionTargetBox-0="[185,1414,827,1928]" captionTargetBox-1="[210,1377,191,1829]" captionTargetBox-2="[177,1411,190,1849]" captionTargetId-0="figure-228@10.[174,1414,817,1941]" captionTargetId-1="figure-20@11.[198,1389,181,1841]" captionTargetId-2="figure-26@12.[166,1422,181,1864]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="FIGURES 38–40. Tipula (Vestiplex) tanycera. 38, male and female, paratypes; 39, male, thorax, dorsal view; 40, wing. Scale of 38, 40 = 0.5 mm; 39 = 1 mm." captionText-1="FIGURES 41–50. Male terminalia of Tipula (Vestiplex) tanycera. 41, hypopygium, dorsal view; 42, hypopygium, lateral view; 43, epandrium, dorsal view; 44, left gonocoxite, lateral view; 45, left outer gonostylus; 46, left inner gonostylus, lateral view; 47, aedeagal guide, lateral view; 48, gonocoxal fragment, dorsal view; 49, sperm pump, dorsal view; 50, distal part of intromittent organ, lateral view. Abbreviations: mt, mid-dorsal tooth of inner gonostylus; pt, posterolateral tooth of epandrium. Scale a: 41, 42 = 0.5 mm; b: 43–49 = 0.5 mm; c: 50 = 0.25 mm." captionText-2="FIGURES 51–56. Female terminalia of Tipula (Vestiplex) tanycera. 51, ovipositor, left lateral view; 52, sternite 8 with hypovalvae, ventral view; 53, sternite 9, dorsal view; 54, furca, dorsal view; 55, bursa copulatrix, dorsal view; 56, spermatheca. Abbreviation: cds, cul-de-sac of bursa copulatrix. Scale a: 50 = 1 mm; b: 51–54 = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6346192" figureDoi-1="http://doi.org/10.5281/zenodo.6346194" figureDoi-2="http://doi.org/10.5281/zenodo.6346196" httpUri-0="https://zenodo.org/record/6346192/files/figure.png" httpUri-1="https://zenodo.org/record/6346194/files/figure.png" httpUri-2="https://zenodo.org/record/6346196/files/figure.png" pageId="10" pageNumber="141">Figs 38–56</figureCitation>
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)
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</paragraph>
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<subSubSection id="C30068B15A17265729BAFE55FCA69EDB" pageId="10" pageNumber="141" type="reference_group">
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<paragraph id="8BA53B3A5A17265729BAFE55FD419EFA" blockId="10.[151,803,455,510]" box="[151,708,455,479]" pageId="10" pageNumber="141">
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<treatmentCitationGroup id="AB0A1C145A17265729BAFE55FD419EFA" box="[151,708,455,479]" pageId="10" pageNumber="141">
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<treatmentCitation id="0ABB1D2B5A17265729BAFE55FD459EFA" author="Alexander, C. P." box="[151,704,455,479]" page="142" pageId="10" pageNumber="141" year="1961">
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<taxonomicName id="4C1A40B95A17265729BAFE55FD459EFA" ID-CoL="576QL" authority="Alexander, 1961: 140 - 142" authorityName="Alexander" authorityPageNumber="140 - 142" authorityYear="1961" box="[151,704,455,479]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="141" phylum="Arthropoda" rank="species" species="tanycera" subGenus="Vestiplex">
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<emphasis id="B96EE7285A17265729BAFE55FF5D9EFA" box="[151,216,455,479]" italics="true" pageId="10" pageNumber="141">Tipula</emphasis>
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(
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||
<emphasis id="B96EE7285A17265729CAFE55FEC49EFA" box="[231,321,455,479]" italics="true" pageId="10" pageNumber="141">Vestiplex</emphasis>
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)
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<emphasis id="B96EE7285A172657287DFE5AFE2C9EFA" box="[336,425,456,479]" italics="true" pageId="10" pageNumber="141">tanycera</emphasis>
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<bibRefCitation id="EF8B46CB5A1726572883FE55FD459EFA" author="Alexander, C. P." box="[430,704,455,479]" pageId="10" pageNumber="141" pagination="97 - 149" refId="ref7670" refString="Alexander, C. P. (1961). New or little-known Tipulidae from eastern Asia (Diptera). XLVIII. Philippine Journal of Science, 90, 97 - 149." type="journal article" year="1961">Alexander, 1961: 140–142</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
.
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</treatmentCitationGroup>
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</paragraph>
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||
<paragraph id="8BA53B3A5A17265729BAFE74FCA69EDB" blockId="10.[151,803,455,510]" box="[151,803,486,510]" pageId="10" pageNumber="141">
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<treatmentCitationGroup id="AB0A1C145A17265729BAFE74FCA69EDB" box="[151,803,486,510]" pageId="10" pageNumber="141">
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<treatmentCitation id="0ABB1D2B5A17265729BAFE74FC9A9EDB" author="Alexander, C. P. & Alexander, M. M." box="[151,799,486,510]" page="65" pageId="10" pageNumber="141" year="1973">
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||
<taxonomicName id="4C1A40B95A17265729BAFE74FC9A9EDB" ID-CoL="576QL" authority=": Alexander and Alexander, 1973: 65" authorityName=": Alexander and Alexander" authorityPageNumber="65" authorityYear="1973" box="[151,799,486,510]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="141" phylum="Arthropoda" rank="species" species="tanycera" subGenus="Vestiplex">
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<emphasis id="B96EE7285A17265729BAFE74FF5D9EDB" box="[151,216,486,510]" italics="true" pageId="10" pageNumber="141">Tipula</emphasis>
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(
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||
<emphasis id="B96EE7285A17265729CAFE74FEC49EDB" box="[231,321,486,510]" italics="true" pageId="10" pageNumber="141">Vestiplex</emphasis>
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||
)
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<emphasis id="B96EE7285A172657287DFE75FE2C9EDB" box="[336,425,487,510]" italics="true" pageId="10" pageNumber="141">tanycera</emphasis>
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:
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<bibRefCitation id="EF8B46CB5A1726572899FE74FC9A9EDB" author="Alexander, C. P. & Alexander, M. M." box="[436,799,486,510]" pageId="10" pageNumber="141" refId="ref7736" refString="Alexander, C. P. & Alexander, M. M. (1973) Tipulidae. In: Delfinado, M. D. & Hardy, D. E. (Ed.), A Catalog of the Diptera of the Oriental Region. Vol. 1. The University of Hawaii Press, Honolulu, pp. 10 - 224." type="book" year="1973">Alexander and Alexander, 1973: 65</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30068B15A17265729BAFDB5FD279C3C" pageId="10" pageNumber="141" type="materials_examined">
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<paragraph id="8BA53B3A5A17265729BAFDB5FD279C3C" blockId="10.[151,1437,550,793]" pageId="10" pageNumber="141">
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<emphasis id="B96EE7285A17265729BAFDB5FE4E9D65" bold="true" box="[151,459,550,577]" pageId="10" pageNumber="141">Type material examined.</emphasis>
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||
<materialsCitation id="3B7231675A17265728FAFDB4FC249D40" ID-GBIF-Occurrence="3699968308" collectingDate="1953-07-02" collectionCode="USNM" country="Pakistan" county="Schmid" elevation="3353" location="Salf-ul-Maluk" municipality="Northwest Frontier Province" pageId="10" pageNumber="141" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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||
<emphasis id="B96EE7285A17265728FAFDB4FDF39D65" bold="true" box="[471,630,550,576]" pageId="10" pageNumber="141">
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<typeStatus id="54A185985A17265728FAFDB4FDF39D65" box="[471,630,550,576]" pageId="10" pageNumber="141" type="holotype">HOLOTYPE</typeStatus>
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</emphasis>
|
||
<specimenCount id="9D1CF0B35A1726572BAEFDB5FD1F9D64" box="[643,666,551,577]" pageId="10" pageNumber="141" type="male">♂</specimenCount>
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||
, labelled:
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<location id="8EC56DE15A1726572A34FDB5FC3E9D64" LSID="urn:lsid:plazi:treatment:03B38A2C5A17265029BAFEF1FAA09999:8EC56DE15A1726572A34FDB5FC3E9D64" box="[793,955,551,577]" country="Pakistan" county="Schmid" municipality="Northwest Frontier Province" name="Salf-ul-Maluk" pageId="10" pageNumber="141">Salf-ul-Maluk</location>
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,
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||
<collectingMunicipality id="6BC1A1405A1726572AE6FDB5FA929D64" box="[971,1303,551,577]" pageId="10" pageNumber="141">Northwest Frontier Province</collectingMunicipality>
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[
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<collectingCountry id="F30D7BAA5A1726572C07FDB4FA149D64" box="[1322,1425,550,577]" name="Pakistan" pageId="10" pageNumber="141">Pakistan</collectingCountry>
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],
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<quantity id="4CE296DF5A17265729BAFDD9FE869D40" box="[151,259,587,613]" metricMagnitude="3" metricUnit="m" metricValue="3.3528" pageId="10" pageNumber="141" unit="ft" value="11000.0">
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<elevation id="0037DC095A17265729BAFDD9FE869D40" box="[151,259,587,613]" metricMagnitude="3" metricUnit="m" metricValue="3.3528" pageId="10" pageNumber="141" unit="ft" value="11000.0">11,000 ft.</elevation>
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</quantity>
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[
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<quantity id="4CE296DF5A172657283CFDD8FEEB9D40" box="[273,366,586,613]" metricMagnitude="3" metricUnit="m" metricValue="3.35" pageId="10" pageNumber="141" unit="m" value="3350.0">
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<elevation id="0037DC095A172657283CFDD8FEEB9D40" box="[273,366,586,613]" metricMagnitude="3" metricUnit="m" metricValue="3.35" pageId="10" pageNumber="141" unit="m" value="3350.0">3350 m</elevation>
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||
</quantity>
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||
], along lake margin,
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||
<date id="FFA41DFA5A1726572B71FDD8FD559D40" box="[604,720,586,613]" pageId="10" pageNumber="141" value="1953-07-02">
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<collectingDate id="EFE0E4125A1726572B71FDD8FD559D40" box="[604,720,586,613]" pageId="10" pageNumber="141" value="1953-07-02">1953.vii.2</collectingDate>
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</date>
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,
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<collectingCounty id="62C443B65A1726572BF1FDD9FCB79D40" box="[732,818,587,613]" pageId="10" pageNumber="141">Schmid</collectingCounty>
|
||
(
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||
<collectionCode id="ED0BA3FF5A1726572A6CFDD9FC1D9D40" box="[833,920,587,613]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="141" type="Museum">USNM</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="3B7231675A1726572A81FDD8FD189C3C" ID-GBIF-Occurrence="3699968306" collectingDate="1953-07-13" collectionCode="USNM" collectorName="No. PS & C. P. Alexander" country="Pakistan" county="Schmid" elevation="3435" location="Kalarian Baihk Northwest Frontier Province" municipality="Northwest Frontier Province" pageId="10" pageNumber="141" specimenCount="13" specimenCount-female="2" specimenCount-male="11" typeStatus="paratype">
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||
<emphasis id="B96EE7285A1726572A81FDD8FBDC9D40" bold="true" box="[940,1113,586,613]" pageId="10" pageNumber="141">
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<typeStatus id="54A185985A1726572A81FDD8FBD59D41" box="[940,1104,586,612]" pageId="10" pageNumber="141" type="paratype">PARATYPES</typeStatus>
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||
:
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</emphasis>
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same data as holotype (
|
||
<specimenCount id="9D1CF0B35A1726572C44FDD9FA129D40" box="[1385,1431,587,613]" pageId="10" pageNumber="141" type="male">4 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D1CF0B35A17265729BAFDFDFF459DAC" box="[151,192,623,649]" pageId="10" pageNumber="141" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED0BA3FF5A17265729E3FDFDFEA49DAC" box="[206,289,623,649]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="141" type="Museum">USNM</collectionCode>
|
||
) [among them
|
||
<specimenCount id="9D1CF0B35A17265728E5FDFDFE749DAC" box="[456,497,623,649]" pageId="10" pageNumber="141" type="male">1 ♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D1CF0B35A1726572B0BFDFDFDCA9DAC" box="[550,591,623,649]" pageId="10" pageNumber="141" type="female">1 ♀</specimenCount>
|
||
(female labeled as
|
||
<typeStatus id="54A185985A1726572A0BFDFDFC079DAF" box="[806,898,623,650]" pageId="10" pageNumber="141" type="allotype">allotype</typeStatus>
|
||
) pinned together on same pin, both dissected for present study, male genitalia number
|
||
<collectorName id="26EF5EEC5A1726572B14FD01FD0D9D88" box="[569,648,659,685]" pageId="10" pageNumber="141">No. PS</collectorName>
|
||
0377m, female genitalia number
|
||
<collectorName id="26EF5EEC5A1726572AD6FD01FBCC9D88" box="[1019,1097,659,685]" pageId="10" pageNumber="141">No. PS</collectorName>
|
||
0378f and other male with antenna, wing and genitalia on slide mounted by
|
||
<collectorName id="26EF5EEC5A1726572BB4FD25FCC39DF4" box="[665,838,694,721]" pageId="10" pageNumber="141">C.P. Alexander</collectorName>
|
||
];
|
||
<location id="8EC56DE15A1726572A7BFD25FAC39DF4" LSID="urn:lsid:plazi:treatment:03B38A2C5A17265029BAFEF1FAA09999:8EC56DE15A1726572A7BFD25FAC39DF4" box="[854,1350,695,721]" name="Kalarian Baihk Northwest Frontier Province" pageId="10" pageNumber="141">Kalarian Baihk Northwest Frontier Province</location>
|
||
,
|
||
<quantity id="4CE296DF5A1726572C7DFD25FF359DD0" metricMagnitude="3" metricUnit="m" metricValue="3.435096" pageId="10" pageNumber="141" unit="ft" value="11270.0">
|
||
<elevation id="0037DC095A1726572C7DFD25FF359DD0" metricMagnitude="3" metricUnit="m" metricValue="3.435096" pageId="10" pageNumber="141" unit="ft" value="11270.0">11,270 ft.</elevation>
|
||
</quantity>
|
||
[
|
||
<quantity id="4CE296DF5A1726572990FD48FE9C9DD0" box="[189,281,730,757]" metricMagnitude="3" metricUnit="m" metricValue="3.435" pageId="10" pageNumber="141" unit="m" value="3435.0">
|
||
<elevation id="0037DC095A1726572990FD48FE9C9DD0" box="[189,281,730,757]" metricMagnitude="3" metricUnit="m" metricValue="3.435" pageId="10" pageNumber="141" unit="m" value="3435.0">3435 m</elevation>
|
||
</quantity>
|
||
], along lake delta,
|
||
<date id="FFA41DFA5A17265728C3FD48FDF59DD0" box="[494,624,730,757]" pageId="10" pageNumber="141" value="1953-07-13">
|
||
<collectingDate id="EFE0E4125A17265728C3FD48FDF59DD0" box="[494,624,730,757]" pageId="10" pageNumber="141" value="1953-07-13">1953.vii.13</collectingDate>
|
||
</date>
|
||
, Schmid (
|
||
<specimenCount id="9D1CF0B35A1726572BCFFD48FC8B9DD0" box="[738,782,730,757]" pageId="10" pageNumber="141" type="male">5 ♂</specimenCount>
|
||
,
|
||
<collectionCode id="ED0BA3FF5A1726572A36FD49FCEB9DD0" box="[795,878,731,757]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="141" type="Museum">USNM</collectionCode>
|
||
) [among them
|
||
<specimenCount id="9D1CF0B35A1726572D34FD49FBDB9DD0" box="[1049,1118,731,757]" pageId="10" pageNumber="141" type="male">one ♂</specimenCount>
|
||
with antenna, leg, wing and genitalia on slide mounted by
|
||
<collectorName id="26EF5EEC5A17265728C7FD6DFD129C3C" box="[490,663,766,793]" pageId="10" pageNumber="141">C.P. Alexander</collectorName>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF656BB25A17265729BAF83EFE5B98CD" ID-DOI="http://doi.org/10.5281/zenodo.6346192" ID-Zenodo-Dep="6346192" httpUri="https://zenodo.org/record/6346192/files/figure.png" pageId="10" pageNumber="141" startId="10.[151,264,1964,1988]" targetBox="[185,1414,827,1928]" targetPageId="10">
|
||
<paragraph id="8BA53B3A5A17265729BAF83EFE5B98CD" blockId="10.[151,1436,1964,2025]" pageId="10" pageNumber="141">
|
||
<emphasis id="B96EE7285A17265729BAF83EFED098E1" bold="true" box="[151,341,1964,1988]" pageId="10" pageNumber="141">FIGURES 38–40.</emphasis>
|
||
<taxonomicName id="4C1A40B95A1726572876F83FFDEB98E1" authorityName=": Alexander and Alexander" authorityYear="1973" box="[347,622,1965,1989]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="141" phylum="Arthropoda" rank="species" species="tanycera" subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1726572876F83FFE1998E0" box="[347,412,1965,1989]" italics="true" pageId="10" pageNumber="141">Tipula</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1726572881F83FFD8398E0" box="[428,518,1965,1989]" italics="true" pageId="10" pageNumber="141">Vestiplex</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1726572B38F83FFDEB98E1" box="[533,622,1965,1988]" italics="true" pageId="10" pageNumber="141">tanycera</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B96EE7285A1726572B56F83EFD1E98E1" bold="true" box="[635,667,1964,1988]" pageId="10" pageNumber="141">38,</emphasis>
|
||
male and female, paratypes;
|
||
<emphasis id="B96EE7285A1726572AEAF83EFC6298E1" bold="true" box="[967,999,1964,1988]" pageId="10" pageNumber="141">39,</emphasis>
|
||
male, thorax, dorsal view;
|
||
<emphasis id="B96EE7285A1726572DD2F83EFA9B98E1" bold="true" box="[1279,1310,1964,1988]" pageId="10" pageNumber="141">40,</emphasis>
|
||
wing. Scale of 38, 40 = 0.5 mm; 39 = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF656BB25A16265629BAF8DBFD2198D4" ID-DOI="http://doi.org/10.5281/zenodo.6346194" ID-Zenodo-Dep="6346194" httpUri="https://zenodo.org/record/6346194/files/figure.png" pageId="11" pageNumber="142" startId="11.[151,264,1865,1889]" targetBox="[210,1377,191,1829]" targetPageId="11">
|
||
<paragraph id="8BA53B3A5A16265629BAF8DBFD2198D4" blockId="11.[151,1437,1865,2033]" pageId="11" pageNumber="142">
|
||
<emphasis id="B96EE7285A16265629BAF8DBFED19844" bold="true" box="[151,340,1865,1889]" pageId="11" pageNumber="142">FIGURES 41–50.</emphasis>
|
||
Male terminalia of
|
||
<taxonomicName id="4C1A40B95A1626562B36F8DBFCA99844" authorityName=": Alexander and Alexander" authorityYear="1973" box="[539,812,1865,1889]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="142" phylum="Arthropoda" rank="species" species="tanycera" subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1626562B36F8DBFDD99844" box="[539,604,1865,1889]" italics="true" pageId="11" pageNumber="142">Tipula</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1626562B46F8DBFD409844" box="[619,709,1865,1889]" italics="true" pageId="11" pageNumber="142">Vestiplex</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1626562BFEF8D8FCA99844" box="[723,812,1866,1889]" italics="true" pageId="11" pageNumber="142">tanycera</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B96EE7285A1626562A15F8DBFCDD9844" bold="true" box="[824,856,1865,1889]" pageId="11" pageNumber="142">41,</emphasis>
|
||
hypopygium, dorsal view;
|
||
<emphasis id="B96EE7285A1626562D40F8DBFB099844" bold="true" box="[1133,1164,1865,1889]" pageId="11" pageNumber="142">42,</emphasis>
|
||
hypopygium, lateral view;
|
||
<emphasis id="B96EE7285A16265629BAF8FFFF3298A0" bold="true" box="[151,183,1901,1925]" pageId="11" pageNumber="142">43,</emphasis>
|
||
epandrium, dorsal view;
|
||
<emphasis id="B96EE7285A1626562899F8FFFE5198A0" bold="true" box="[436,468,1901,1925]" pageId="11" pageNumber="142">44,</emphasis>
|
||
left gonocoxite, lateral view;
|
||
<emphasis id="B96EE7285A1626562BD3F8FFFC9898A0" bold="true" box="[766,797,1901,1925]" pageId="11" pageNumber="142">45,</emphasis>
|
||
left outer gonostylus;
|
||
<emphasis id="B96EE7285A1626562AD0F8FFFB9998A0" bold="true" box="[1021,1052,1901,1925]" pageId="11" pageNumber="142">46,</emphasis>
|
||
left inner gonostylus, lateral view;
|
||
<emphasis id="B96EE7285A1626562C50F8FFFA1998A0" bold="true" box="[1405,1436,1901,1925]" pageId="11" pageNumber="142">47,</emphasis>
|
||
aedeagal guide, lateral view;
|
||
<emphasis id="B96EE7285A16265628EDF803FE65988C" bold="true" box="[448,480,1937,1961]" pageId="11" pageNumber="142">48,</emphasis>
|
||
gonocoxal fragment, dorsal view;
|
||
<emphasis id="B96EE7285A1626562A6FF803FCE7988C" bold="true" box="[834,866,1937,1961]" pageId="11" pageNumber="142">49,</emphasis>
|
||
sperm pump, dorsal view;
|
||
<emphasis id="B96EE7285A1626562D5AF803FB12988C" bold="true" box="[1143,1175,1937,1961]" pageId="11" pageNumber="142">50,</emphasis>
|
||
distal part of intromittent organ, lateral view. Abbreviations: mt, mid-dorsal tooth of inner gonostylus; pt, posterolateral tooth of epandrium. Scale a: 41, 42 = 0.5 mm; b: 43–49 = 0.5 mm; c: 50 = 0.25 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF656BB25A11265129BAF8CDFBAB989A" ID-DOI="http://doi.org/10.5281/zenodo.6346196" ID-Zenodo-Dep="6346196" httpUri="https://zenodo.org/record/6346196/files/figure.png" pageId="12" pageNumber="143" startId="12.[151,264,1887,1911]" targetBox="[177,1411,190,1849]" targetPageId="12">
|
||
<paragraph id="8BA53B3A5A11265129BAF8CDFBAB989A" blockId="12.[151,1437,1887,1984]" pageId="12" pageNumber="143">
|
||
<emphasis id="B96EE7285A11265129BAF8CDFED39852" bold="true" box="[151,342,1887,1911]" pageId="12" pageNumber="143">FIGURES 51–56.</emphasis>
|
||
Female terminalia of
|
||
<taxonomicName id="4C1A40B95A1126512B12F8F2FCD09852" authorityName=": Alexander and Alexander" authorityYear="1973" box="[575,853,1888,1912]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="143" phylum="Arthropoda" rank="species" species="tanycera" subGenus="Vestiplex">
|
||
<emphasis id="B96EE7285A1126512B12F8F2FD05985D" box="[575,640,1888,1912]" italics="true" pageId="12" pageNumber="143">Tipula</emphasis>
|
||
(
|
||
<emphasis id="B96EE7285A1126512BBCF8F2FD6E985D" box="[657,747,1888,1912]" italics="true" pageId="12" pageNumber="143">Vestiplex</emphasis>
|
||
)
|
||
<emphasis id="B96EE7285A1126512BD1F8F2FCD09852" box="[764,853,1888,1911]" italics="true" pageId="12" pageNumber="143">tanycera</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B96EE7285A1126512A49F8CDFC069852" bold="true" box="[868,899,1887,1911]" pageId="12" pageNumber="143">51,</emphasis>
|
||
ovipositor, left lateral view;
|
||
<emphasis id="B96EE7285A1126512D99F8CDFB569852" bold="true" box="[1204,1235,1887,1911]" pageId="12" pageNumber="143">52,</emphasis>
|
||
sternite 8 with hypovalvae, ventral view;
|
||
<emphasis id="B96EE7285A11265128A0F811FE2898BE" bold="true" box="[397,429,1923,1947]" pageId="12" pageNumber="143">53,</emphasis>
|
||
sternite 9, dorsal view;
|
||
<emphasis id="B96EE7285A1126512B88F811FD4098BE" bold="true" box="[677,709,1923,1947]" pageId="12" pageNumber="143">54,</emphasis>
|
||
furca, dorsal view;
|
||
<emphasis id="B96EE7285A1126512ABFF816FC3798B9" bold="true" box="[914,946,1924,1948]" pageId="12" pageNumber="143">55,</emphasis>
|
||
bursa copulatrix, dorsal view;
|
||
<emphasis id="B96EE7285A1126512DDDF811FA9598BE" bold="true" box="[1264,1296,1923,1947]" pageId="12" pageNumber="143">56,</emphasis>
|
||
spermatheca. Abbreviation: cds, cul-de-sac of bursa copulatrix. Scale a: 50 = 1 mm; b: 51–54 = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C30068B15A10265029EAFF04FAC19974" pageId="13" pageNumber="144" type="description">
|
||
<paragraph id="8BA53B3A5A10265029EAFF04FE469FF1" blockId="13.[151,1437,150,1725]" pageId="13" pageNumber="144">
|
||
<emphasis id="B96EE7285A10265029EAFF04FE419F95" bold="true" box="[199,452,150,176]" pageId="13" pageNumber="144">Redescription. Male.</emphasis>
|
||
Body length 17.3–20.0 mm, wing length 19.4–21.0 mm. General body colouration brownish yellow (
|
||
<figureCitation id="132127BF5A102650284FFF28FE339FF1" box="[354,438,186,212]" captionStart="FIGURES 38–40" captionStartId="10.[151,264,1964,1988]" captionTargetBox="[185,1414,827,1928]" captionTargetId="figure-228@10.[174,1414,817,1941]" captionTargetPageId="10" captionText="FIGURES 38–40. Tipula (Vestiplex) tanycera. 38, male and female, paratypes; 39, male, thorax, dorsal view; 40, wing. Scale of 38, 40 = 0.5 mm; 39 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6346192" httpUri="https://zenodo.org/record/6346192/files/figure.png" pageId="13" pageNumber="144">Fig. 38</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA53B3A5A10265029EAFF4CFE619E89" blockId="13.[151,1437,150,1725]" pageId="13" pageNumber="144">
|
||
<emphasis id="B96EE7285A10265029EAFF4CFE819FDD" box="[199,260,222,248]" italics="true" pageId="13" pageNumber="144">Head</emphasis>
|
||
. Vertex and occiput yellowish grey pruinescent with brown median line. Rostrum brown, dorsally thinly dusted with grey pruinescence. Nasus short and distinct. Antenna 13-segmented, when bent backwards extends beyond wing base. Scape and pedicel yellow, flagellum brown. Each flagellomere, except first, with weak basal enlargement and incision, and additional apical incision; shape of segments slightly sinuous. Apical flagellomere small. Longest verticils shorter than corresponding segments. Palpus with first segment yellowish, second brownish yellow, other brownish black.
|
||
</paragraph>
|
||
<paragraph id="8BA53B3A5A10265029EAFE24FE289D8D" blockId="13.[151,1437,150,1725]" pageId="13" pageNumber="144">
|
||
<emphasis id="B96EE7285A10265029EAFE24FE9D9EF5" box="[199,280,438,464]" italics="true" pageId="13" pageNumber="144">Thorax</emphasis>
|
||
. Whole thorax brown, dusted with grey pruinescence (
|
||
<figureCitation id="132127BF5A1026502AA5FE24FC5B9EF5" box="[904,990,438,464]" captionStart="FIGURES 38–40" captionStartId="10.[151,264,1964,1988]" captionTargetBox="[185,1414,827,1928]" captionTargetId="figure-228@10.[174,1414,817,1941]" captionTargetPageId="10" captionText="FIGURES 38–40. Tipula (Vestiplex) tanycera. 38, male and female, paratypes; 39, male, thorax, dorsal view; 40, wing. Scale of 38, 40 = 0.5 mm; 39 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6346192" httpUri="https://zenodo.org/record/6346192/files/figure.png" pageId="13" pageNumber="144">Fig. 39</figureCitation>
|
||
). Pronotum brown, with median brown line. Prescutum and presutural scutum with four brown, grey dusted stripes, bordered by brown. Medial stripes fused in basal half. Interspaces grey. Postsutural scutum grey, each lobe with two brown spots, bordered by brown. Scutellum and mediotergite brown, dusted with grey, with brown median line. Pleura brown. Coxae yellow, dusted with grey, trochanters yellow, femora and tibiae yellow, darkened at tip; first two tarsal segments yellow, distally brown, rest of segments brown. Tarsal claw without tooth. Wing patterned with brown (
|
||
<figureCitation id="132127BF5A1026502D55FDF8FB489DA1" box="[1144,1229,618,644]" captionStart="FIGURES 38–40" captionStartId="10.[151,264,1964,1988]" captionTargetBox="[185,1414,827,1928]" captionTargetId="figure-228@10.[174,1414,817,1941]" captionTargetPageId="10" captionText="FIGURES 38–40. Tipula (Vestiplex) tanycera. 38, male and female, paratypes; 39, male, thorax, dorsal view; 40, wing. Scale of 38, 40 = 0.5 mm; 39 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6346192" httpUri="https://zenodo.org/record/6346192/files/figure.png" pageId="13" pageNumber="144">Fig. 40</figureCitation>
|
||
). Halter yellowish brown with brown knob.
|
||
</paragraph>
|
||
<paragraph id="8BA53B3A5A10265029EAFD20FB519DE9" blockId="13.[151,1437,150,1725]" box="[199,1236,690,717]" pageId="13" pageNumber="144">
|
||
<emphasis id="B96EE7285A10265029EAFD20FEB49DE9" box="[199,305,690,716]" italics="true" pageId="13" pageNumber="144">Abdomen</emphasis>
|
||
. Brownish yellow, with brown dorsal and lateral stripes, terminal segments brown.
|
||
</paragraph>
|
||
<paragraph id="8BA53B3A5A10265029EAFD45FD869A28" blockId="13.[151,1437,150,1725]" pageId="13" pageNumber="144">
|
||
<emphasis id="B96EE7285A10265029EAFD45FED39DD5" box="[199,342,727,752]" italics="true" pageId="13" pageNumber="144">Hypopygium</emphasis>
|
||
(
|
||
<figureCitation id="132127BF5A102650284AFD44FE699DD4" box="[359,492,726,753]" captionStart="FIGURES 41–50" captionStartId="11.[151,264,1865,1889]" captionTargetBox="[210,1377,191,1829]" captionTargetId="figure-20@11.[198,1389,181,1841]" captionTargetPageId="11" captionText="FIGURES 41–50. Male terminalia of Tipula (Vestiplex) tanycera. 41, hypopygium, dorsal view; 42, hypopygium, lateral view; 43, epandrium, dorsal view; 44, left gonocoxite, lateral view; 45, left outer gonostylus; 46, left inner gonostylus, lateral view; 47, aedeagal guide, lateral view; 48, gonocoxal fragment, dorsal view; 49, sperm pump, dorsal view; 50, distal part of intromittent organ, lateral view. Abbreviations: mt, mid-dorsal tooth of inner gonostylus; pt, posterolateral tooth of epandrium. Scale a: 41, 42 = 0.5 mm; b: 43–49 = 0.5 mm; c: 50 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346194" httpUri="https://zenodo.org/record/6346194/files/figure.png" pageId="13" pageNumber="144">Figs 41–50</figureCitation>
|
||
). Brown, broader than rest of abdomen. Epandrium forming large concave, sclerotized, crescent in outline saucer-shaped plate (
|
||
<figureCitation id="132127BF5A1026502BBEFD68FC909C31" box="[659,789,762,788]" captionStart="FIGURES 41–50" captionStartId="11.[151,264,1865,1889]" captionTargetBox="[210,1377,191,1829]" captionTargetId="figure-20@11.[198,1389,181,1841]" captionTargetPageId="11" captionText="FIGURES 41–50. Male terminalia of Tipula (Vestiplex) tanycera. 41, hypopygium, dorsal view; 42, hypopygium, lateral view; 43, epandrium, dorsal view; 44, left gonocoxite, lateral view; 45, left outer gonostylus; 46, left inner gonostylus, lateral view; 47, aedeagal guide, lateral view; 48, gonocoxal fragment, dorsal view; 49, sperm pump, dorsal view; 50, distal part of intromittent organ, lateral view. Abbreviations: mt, mid-dorsal tooth of inner gonostylus; pt, posterolateral tooth of epandrium. Scale a: 41, 42 = 0.5 mm; b: 43–49 = 0.5 mm; c: 50 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346194" httpUri="https://zenodo.org/record/6346194/files/figure.png" pageId="13" pageNumber="144">Figs 41–43</figureCitation>
|
||
). Main body of plate brownish yellow with blackened rim. Posterior margin of plate broadly emarginated, medially with shallow U-shaped emargination, laterally bordered with obtuse projections or denticles (
|
||
<bibRefCitation id="EF8B46CB5A1026502B1EFCD0FD6E9C79" author="Alexander, C. P." box="[563,747,834,860]" pageId="13" pageNumber="144" pagination="97 - 149" refId="ref7670" refString="Alexander, C. P. (1961). New or little-known Tipulidae from eastern Asia (Diptera). XLVIII. Philippine Journal of Science, 90, 97 - 149." type="journal article" year="1961">Alexander 1961</bibRefCitation>
|
||
, pl. 4, fig. 33;
|
||
<bibRefCitation id="EF8B46CB5A1026502AA3FCD0FBC29C79" author="Starkevich, P." box="[910,1095,834,860]" pageId="13" pageNumber="144" refId="ref8272" refString="Starkevich, P. (2012) Taxonomic and phylogentic review of the subgenus Tipula (Vestiplex) Bezzi, 1924 (Diptera, Tipulidae). Doctoral dissertation, Vilnius University, Biomedical Sciences, Zoology, Vilnius, 219 pp." type="book" year="2012">Starkevich 2012</bibRefCitation>
|
||
, fig. 142) and additional small denticles on either side. Lateral angle of plate broad and obtuse. Anterior and lateral portions of plate raised into sclerotised rim, posterolaterally terminating in black obtuse teeth. Gonocoxite unarmed (
|
||
<figureCitation id="132127BF5A1026502DAAFC18FB589C81" box="[1159,1245,906,932]" captionStart="FIGURES 41–50" captionStartId="11.[151,264,1865,1889]" captionTargetBox="[210,1377,191,1829]" captionTargetId="figure-20@11.[198,1389,181,1841]" captionTargetPageId="11" captionText="FIGURES 41–50. Male terminalia of Tipula (Vestiplex) tanycera. 41, hypopygium, dorsal view; 42, hypopygium, lateral view; 43, epandrium, dorsal view; 44, left gonocoxite, lateral view; 45, left outer gonostylus; 46, left inner gonostylus, lateral view; 47, aedeagal guide, lateral view; 48, gonocoxal fragment, dorsal view; 49, sperm pump, dorsal view; 50, distal part of intromittent organ, lateral view. Abbreviations: mt, mid-dorsal tooth of inner gonostylus; pt, posterolateral tooth of epandrium. Scale a: 41, 42 = 0.5 mm; b: 43–49 = 0.5 mm; c: 50 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346194" httpUri="https://zenodo.org/record/6346194/files/figure.png" pageId="13" pageNumber="144">Fig. 44</figureCitation>
|
||
). Outer gonostylus flattened, nearly oval (
|
||
<figureCitation id="132127BF5A1026502892FC3CFD919CEC" box="[447,532,942,969]" captionStart="FIGURES 41–50" captionStartId="11.[151,264,1865,1889]" captionTargetBox="[210,1377,191,1829]" captionTargetId="figure-20@11.[198,1389,181,1841]" captionTargetPageId="11" captionText="FIGURES 41–50. Male terminalia of Tipula (Vestiplex) tanycera. 41, hypopygium, dorsal view; 42, hypopygium, lateral view; 43, epandrium, dorsal view; 44, left gonocoxite, lateral view; 45, left outer gonostylus; 46, left inner gonostylus, lateral view; 47, aedeagal guide, lateral view; 48, gonocoxal fragment, dorsal view; 49, sperm pump, dorsal view; 50, distal part of intromittent organ, lateral view. Abbreviations: mt, mid-dorsal tooth of inner gonostylus; pt, posterolateral tooth of epandrium. Scale a: 41, 42 = 0.5 mm; b: 43–49 = 0.5 mm; c: 50 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346194" httpUri="https://zenodo.org/record/6346194/files/figure.png" pageId="13" pageNumber="144">Fig. 45</figureCitation>
|
||
). Inner gonostylus large, claw-shaped curved plate with single, large mid-dorsal tooth (
|
||
<figureCitation id="132127BF5A10265029CDFC40FEB19CC8" box="[224,308,978,1005]" captionStart="FIGURES 41–50" captionStartId="11.[151,264,1865,1889]" captionTargetBox="[210,1377,191,1829]" captionTargetId="figure-20@11.[198,1389,181,1841]" captionTargetPageId="11" captionText="FIGURES 41–50. Male terminalia of Tipula (Vestiplex) tanycera. 41, hypopygium, dorsal view; 42, hypopygium, lateral view; 43, epandrium, dorsal view; 44, left gonocoxite, lateral view; 45, left outer gonostylus; 46, left inner gonostylus, lateral view; 47, aedeagal guide, lateral view; 48, gonocoxal fragment, dorsal view; 49, sperm pump, dorsal view; 50, distal part of intromittent organ, lateral view. Abbreviations: mt, mid-dorsal tooth of inner gonostylus; pt, posterolateral tooth of epandrium. Scale a: 41, 42 = 0.5 mm; b: 43–49 = 0.5 mm; c: 50 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346194" httpUri="https://zenodo.org/record/6346194/files/figure.png" pageId="13" pageNumber="144">Fig. 46</figureCitation>
|
||
); beak extends into blackened rostrum. Aedeagal guide in shape of narrow, elongated tube, base with dorsal flange handle-shaped, membranous at base, ventral flange flattened; distal margin bent inward forming two microscopic hooks (
|
||
<figureCitation id="132127BF5A1026502857FB88FE4A9B11" box="[378,463,1050,1076]" captionStart="FIGURES 41–50" captionStartId="11.[151,264,1865,1889]" captionTargetBox="[210,1377,191,1829]" captionTargetId="figure-20@11.[198,1389,181,1841]" captionTargetPageId="11" captionText="FIGURES 41–50. Male terminalia of Tipula (Vestiplex) tanycera. 41, hypopygium, dorsal view; 42, hypopygium, lateral view; 43, epandrium, dorsal view; 44, left gonocoxite, lateral view; 45, left outer gonostylus; 46, left inner gonostylus, lateral view; 47, aedeagal guide, lateral view; 48, gonocoxal fragment, dorsal view; 49, sperm pump, dorsal view; 50, distal part of intromittent organ, lateral view. Abbreviations: mt, mid-dorsal tooth of inner gonostylus; pt, posterolateral tooth of epandrium. Scale a: 41, 42 = 0.5 mm; b: 43–49 = 0.5 mm; c: 50 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346194" httpUri="https://zenodo.org/record/6346194/files/figure.png" pageId="13" pageNumber="144">Fig. 47</figureCitation>
|
||
). Gonocoxal fragment large with lateral and medial sclerites well-developed (
|
||
<figureCitation id="132127BF5A1026502C16FB88FA159B11" box="[1339,1424,1050,1076]" captionStart="FIGURES 41–50" captionStartId="11.[151,264,1865,1889]" captionTargetBox="[210,1377,191,1829]" captionTargetId="figure-20@11.[198,1389,181,1841]" captionTargetPageId="11" captionText="FIGURES 41–50. Male terminalia of Tipula (Vestiplex) tanycera. 41, hypopygium, dorsal view; 42, hypopygium, lateral view; 43, epandrium, dorsal view; 44, left gonocoxite, lateral view; 45, left outer gonostylus; 46, left inner gonostylus, lateral view; 47, aedeagal guide, lateral view; 48, gonocoxal fragment, dorsal view; 49, sperm pump, dorsal view; 50, distal part of intromittent organ, lateral view. Abbreviations: mt, mid-dorsal tooth of inner gonostylus; pt, posterolateral tooth of epandrium. Scale a: 41, 42 = 0.5 mm; b: 43–49 = 0.5 mm; c: 50 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346194" httpUri="https://zenodo.org/record/6346194/files/figure.png" pageId="13" pageNumber="144">Fig. 48</figureCitation>
|
||
). Medial sclerites fused, anterior apodeme distinct, slightly flattened, posterior part broadly flattened and expanded at base and arched distally. Lateral sclerites nearly V-shaped. Sperm pump with small and flattened central vesicle (
|
||
<figureCitation id="132127BF5A102650298DFB14FF7D9B85" box="[160,248,1158,1184]" captionStart="FIGURES 41–50" captionStartId="11.[151,264,1865,1889]" captionTargetBox="[210,1377,191,1829]" captionTargetId="figure-20@11.[198,1389,181,1841]" captionTargetPageId="11" captionText="FIGURES 41–50. Male terminalia of Tipula (Vestiplex) tanycera. 41, hypopygium, dorsal view; 42, hypopygium, lateral view; 43, epandrium, dorsal view; 44, left gonocoxite, lateral view; 45, left outer gonostylus; 46, left inner gonostylus, lateral view; 47, aedeagal guide, lateral view; 48, gonocoxal fragment, dorsal view; 49, sperm pump, dorsal view; 50, distal part of intromittent organ, lateral view. Abbreviations: mt, mid-dorsal tooth of inner gonostylus; pt, posterolateral tooth of epandrium. Scale a: 41, 42 = 0.5 mm; b: 43–49 = 0.5 mm; c: 50 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346194" httpUri="https://zenodo.org/record/6346194/files/figure.png" pageId="13" pageNumber="144">Fig. 49</figureCitation>
|
||
). Compressor apodeme with median shallow emargination. Posterior immovable apodeme longer than compressor apodeme, large and extended laterally into broad plate, medially with darker ridge, anterior margin bent inwards. Anterior immovable apodeme nearly rounded, narrowed at lateral margin. Aedeagus with distal part ventrally membranous (
|
||
<figureCitation id="132127BF5A102650288FFB60FE739A28" box="[418,502,1266,1293]" captionStart="FIGURES 41–50" captionStartId="11.[151,264,1865,1889]" captionTargetBox="[210,1377,191,1829]" captionTargetId="figure-20@11.[198,1389,181,1841]" captionTargetPageId="11" captionText="FIGURES 41–50. Male terminalia of Tipula (Vestiplex) tanycera. 41, hypopygium, dorsal view; 42, hypopygium, lateral view; 43, epandrium, dorsal view; 44, left gonocoxite, lateral view; 45, left outer gonostylus; 46, left inner gonostylus, lateral view; 47, aedeagal guide, lateral view; 48, gonocoxal fragment, dorsal view; 49, sperm pump, dorsal view; 50, distal part of intromittent organ, lateral view. Abbreviations: mt, mid-dorsal tooth of inner gonostylus; pt, posterolateral tooth of epandrium. Scale a: 41, 42 = 0.5 mm; b: 43–49 = 0.5 mm; c: 50 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6346194" httpUri="https://zenodo.org/record/6346194/files/figure.png" pageId="13" pageNumber="144">Fig. 50</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA53B3A5A10265029EAFA84FAE19A15" blockId="13.[151,1437,150,1725]" box="[199,1380,1302,1329]" pageId="13" pageNumber="144">
|
||
<emphasis id="B96EE7285A10265029EAFA84FEA09A15" bold="true" box="[199,293,1302,1328]" pageId="13" pageNumber="144">Female.</emphasis>
|
||
Body length
|
||
<quantity id="4CE296DF5A1026502890FA84FDA69A14" box="[445,547,1302,1329]" metricMagnitude="-2" metricUnit="m" metricValue="2.34" pageId="13" pageNumber="144" unit="mm" value="23.4">23.4 mm</quantity>
|
||
, wing length
|
||
<quantity id="4CE296DF5A1026502B97FA84FCA49A14" box="[698,801,1302,1329]" metricMagnitude="-2" metricUnit="m" metricValue="1.94" pageId="13" pageNumber="144" unit="mm" value="19.4">19.4 mm</quantity>
|
||
. Generally similar to male in body colour (
|
||
<figureCitation id="132127BF5A1026502C2EFA84FAD29A15" box="[1283,1367,1302,1328]" captionStart="FIGURES 38–40" captionStartId="10.[151,264,1964,1988]" captionTargetBox="[185,1414,827,1928]" captionTargetId="figure-228@10.[174,1414,817,1941]" captionTargetPageId="10" captionText="FIGURES 38–40. Tipula (Vestiplex) tanycera. 38, male and female, paratypes; 39, male, thorax, dorsal view; 40, wing. Scale of 38, 40 = 0.5 mm; 39 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6346192" httpUri="https://zenodo.org/record/6346192/files/figure.png" pageId="13" pageNumber="144">Fig. 38</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA53B3A5A10265029EAFAA8FAC19974" blockId="13.[151,1437,150,1725]" pageId="13" pageNumber="144">
|
||
<emphasis id="B96EE7285A10265029EAFAA8FE1D9A71" box="[199,408,1338,1364]" italics="true" pageId="13" pageNumber="144">Female terminalia</emphasis>
|
||
(
|
||
<figureCitation id="132127BF5A102650288AFAA8FDAF9A70" box="[423,554,1338,1365]" captionStart="FIGURES 51–56" captionStartId="12.[151,264,1887,1911]" captionTargetBox="[177,1411,190,1849]" captionTargetId="figure-26@12.[166,1422,181,1864]" captionTargetPageId="12" captionText="FIGURES 51–56. Female terminalia of Tipula (Vestiplex) tanycera. 51, ovipositor, left lateral view; 52, sternite 8 with hypovalvae, ventral view; 53, sternite 9, dorsal view; 54, furca, dorsal view; 55, bursa copulatrix, dorsal view; 56, spermatheca. Abbreviation: cds, cul-de-sac of bursa copulatrix. Scale a: 50 = 1 mm; b: 51–54 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6346196" httpUri="https://zenodo.org/record/6346196/files/figure.png" pageId="13" pageNumber="144">Figs 51–56</figureCitation>
|
||
). Tergite 10 shiny brown. Cercus brownish yellow, about as long as tergite 10, slender, narrowing towards upcurved tip (
|
||
<figureCitation id="132127BF5A1026502B5BFACCFD4E9A5C" box="[630,715,1374,1401]" captionStart="FIGURES 51–56" captionStartId="12.[151,264,1887,1911]" captionTargetBox="[177,1411,190,1849]" captionTargetId="figure-26@12.[166,1422,181,1864]" captionTargetPageId="12" captionText="FIGURES 51–56. Female terminalia of Tipula (Vestiplex) tanycera. 51, ovipositor, left lateral view; 52, sternite 8 with hypovalvae, ventral view; 53, sternite 9, dorsal view; 54, furca, dorsal view; 55, bursa copulatrix, dorsal view; 56, spermatheca. Abbreviation: cds, cul-de-sac of bursa copulatrix. Scale a: 50 = 1 mm; b: 51–54 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6346196" httpUri="https://zenodo.org/record/6346196/files/figure.png" pageId="13" pageNumber="144">Fig. 51</figureCitation>
|
||
). Ventral margin distally with distinct serrations. Hypovalva in shape of pale, elongated slender filament, with short trichia at tip (
|
||
<figureCitation id="132127BF5A1026502AA1FA10FC679AB8" box="[908,994,1410,1437]" captionStart="FIGURES 51–56" captionStartId="12.[151,264,1887,1911]" captionTargetBox="[177,1411,190,1849]" captionTargetId="figure-26@12.[166,1422,181,1864]" captionTargetPageId="12" captionText="FIGURES 51–56. Female terminalia of Tipula (Vestiplex) tanycera. 51, ovipositor, left lateral view; 52, sternite 8 with hypovalvae, ventral view; 53, sternite 9, dorsal view; 54, furca, dorsal view; 55, bursa copulatrix, dorsal view; 56, spermatheca. Abbreviation: cds, cul-de-sac of bursa copulatrix. Scale a: 50 = 1 mm; b: 51–54 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6346196" httpUri="https://zenodo.org/record/6346196/files/figure.png" pageId="13" pageNumber="144">Fig. 52</figureCitation>
|
||
). Median incision between hypovalvae deeper than posterior margin of sternite 8. Lateral incision relatively deep and broad, about 1.5 times as deep as maximal width. Posterolateral angle of sternite 8 rounded. Sternite 9 with posterior part shovel-shaped, margin rounded, medially forming groove; anterior parts flattened at base (
|
||
<figureCitation id="132127BF5A1026502AAEFA7CFC53992C" box="[899,982,1518,1545]" captionStart="FIGURES 51–56" captionStartId="12.[151,264,1887,1911]" captionTargetBox="[177,1411,190,1849]" captionTargetId="figure-26@12.[166,1422,181,1864]" captionTargetPageId="12" captionText="FIGURES 51–56. Female terminalia of Tipula (Vestiplex) tanycera. 51, ovipositor, left lateral view; 52, sternite 8 with hypovalvae, ventral view; 53, sternite 9, dorsal view; 54, furca, dorsal view; 55, bursa copulatrix, dorsal view; 56, spermatheca. Abbreviation: cds, cul-de-sac of bursa copulatrix. Scale a: 50 = 1 mm; b: 51–54 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6346196" httpUri="https://zenodo.org/record/6346196/files/figure.png" pageId="13" pageNumber="144">Fig. 53</figureCitation>
|
||
). Furca anteriorly narrow but posteriorly broad (
|
||
<figureCitation id="132127BF5A10265029CAF980FEB89908" box="[231,317,1554,1581]" captionStart="FIGURES 51–56" captionStartId="12.[151,264,1887,1911]" captionTargetBox="[177,1411,190,1849]" captionTargetId="figure-26@12.[166,1422,181,1864]" captionTargetPageId="12" captionText="FIGURES 51–56. Female terminalia of Tipula (Vestiplex) tanycera. 51, ovipositor, left lateral view; 52, sternite 8 with hypovalvae, ventral view; 53, sternite 9, dorsal view; 54, furca, dorsal view; 55, bursa copulatrix, dorsal view; 56, spermatheca. Abbreviation: cds, cul-de-sac of bursa copulatrix. Scale a: 50 = 1 mm; b: 51–54 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6346196" httpUri="https://zenodo.org/record/6346196/files/figure.png" pageId="13" pageNumber="144">Fig. 54</figureCitation>
|
||
). Bursa copulatrix with spermathecal ducts sclerotised at base, in shape of slightly curved, slender, dark brown sticks (
|
||
<figureCitation id="132127BF5A1026502843F9A4FE479974" box="[366,450,1590,1617]" captionStart="FIGURES 51–56" captionStartId="12.[151,264,1887,1911]" captionTargetBox="[177,1411,190,1849]" captionTargetId="figure-26@12.[166,1422,181,1864]" captionTargetPageId="12" captionText="FIGURES 51–56. Female terminalia of Tipula (Vestiplex) tanycera. 51, ovipositor, left lateral view; 52, sternite 8 with hypovalvae, ventral view; 53, sternite 9, dorsal view; 54, furca, dorsal view; 55, bursa copulatrix, dorsal view; 56, spermatheca. Abbreviation: cds, cul-de-sac of bursa copulatrix. Scale a: 50 = 1 mm; b: 51–54 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6346196" httpUri="https://zenodo.org/record/6346196/files/figure.png" pageId="13" pageNumber="144">Fig. 55</figureCitation>
|
||
). Cul-de-sac of bursa copulatrix curved. Spermatheca nearly spherical (
|
||
<figureCitation id="132127BF5A1026502DCEF9A4FABD9974" box="[1251,1336,1590,1617]" captionStart="FIGURES 51–56" captionStartId="12.[151,264,1887,1911]" captionTargetBox="[177,1411,190,1849]" captionTargetId="figure-26@12.[166,1422,181,1864]" captionTargetPageId="12" captionText="FIGURES 51–56. Female terminalia of Tipula (Vestiplex) tanycera. 51, ovipositor, left lateral view; 52, sternite 8 with hypovalvae, ventral view; 53, sternite 9, dorsal view; 54, furca, dorsal view; 55, bursa copulatrix, dorsal view; 56, spermatheca. Abbreviation: cds, cul-de-sac of bursa copulatrix. Scale a: 50 = 1 mm; b: 51–54 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6346196" httpUri="https://zenodo.org/record/6346196/files/figure.png" pageId="13" pageNumber="144">Fig. 56</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30068B15A10265029EAF9C8FAA39950" box="[199,1318,1626,1653]" pageId="13" pageNumber="144" type="distribution">
|
||
<paragraph id="8BA53B3A5A10265029EAF9C8FAA39950" blockId="13.[151,1437,150,1725]" box="[199,1318,1626,1653]" pageId="13" pageNumber="144">
|
||
<emphasis id="B96EE7285A10265029EAF9C8FEE49951" bold="true" box="[199,353,1626,1652]" pageId="13" pageNumber="144">Distribution.</emphasis>
|
||
<collectingCountry id="F30D7BAA5A102650284AF9C8FE439951" box="[359,454,1626,1652]" name="Pakistan" pageId="13" pageNumber="144">Pakistan</collectingCountry>
|
||
(
|
||
<collectingRegion id="49DEF5D85A10265028F8F9C8FD4F9951" box="[469,714,1626,1652]" country="Pakistan" name="Khyber Pakhtunkhwa" pageId="13" pageNumber="144">Khyber Pakhtunkhwa</collectingRegion>
|
||
[=
|
||
<collectingRegion id="49DEF5D85A1026502BDCF9C8FBC69950" box="[753,1091,1626,1653]" country="Pakistan" name="Khyber Pakhtunkhwa" pageId="13" pageNumber="144">North-West Frontier Province</collectingRegion>
|
||
]) (
|
||
<bibRefCitation id="EF8B46CB5A1026502D4DF9C8FA9C9950" author="Alexander, C. P." box="[1120,1305,1626,1653]" pageId="13" pageNumber="144" pagination="97 - 149" refId="ref7670" refString="Alexander, C. P. (1961). New or little-known Tipulidae from eastern Asia (Diptera). XLVIII. Philippine Journal of Science, 90, 97 - 149." type="journal article" year="1961">Alexander 1961</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30068B15A10265029EAF9ECFAA09999" pageId="13" pageNumber="144" type="discussion">
|
||
<paragraph id="8BA53B3A5A10265029EAF9ECFAA09999" blockId="13.[151,1437,150,1725]" pageId="13" pageNumber="144">
|
||
<emphasis id="B96EE7285A10265029EAF9ECFEBE99BD" bold="true" box="[199,315,1662,1688]" pageId="13" pageNumber="144">Remarks.</emphasis>
|
||
The species is known only from the
|
||
<typeStatus id="54A185985A1026502BCEF9EDFC9699BC" box="[739,787,1663,1689]" pageId="13" pageNumber="144">type</typeStatus>
|
||
series collected near Salf ul Maluk Lake, a mountainous lake located at the northern end of the Kaghan Valley, in Saiful Muluk National Park, northern
|
||
<collectingCountry id="F30D7BAA5A1026502D93F930FAA49999" box="[1214,1313,1698,1724]" name="Pakistan" pageId="13" pageNumber="144">Pakistan</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |