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<document id="7CC29D45B6FA96406DD48B2C0BFCB0FD" ID-DOI="10.11646/zootaxa.3904.4.1" ID-GBIF-Dataset="a1c49726-9afa-4be5-85ef-9d0caabac1fe" ID-ISSN="1175-5326" ID-Zenodo-Dep="239899" ID-ZooBank="38457A27-A15B-4E87-BE72-9A3AB007CAA5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460890290884" checkinUser="plazi" docAuthor="Yoshitomi, Hiroyuki" docDate="2015" docId="03AC6D78C825216549B2EB293196FA58" docLanguage="en" docName="zt03904p500.pdf" docOrigin="Zootaxa 3904 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ypsiloncyphon teruhisai Yoshitomi, 2015, sp. n." docType="treatment" docUuid="A10D515B-99B6-4D36-9D3B-FC22C30D2BA8" docUuidSource="ZooBank" docVersion="9" lastPageNumber="489" masterDocId="FF951500C80121434925E96E356EFFBD" masterDocTitle="New species of the genus Ypsiloncyphon (Insecta: Coleoptera: Scirtidae: Scirtinae)" masterLastPageNumber="500" masterPageNumber="451" pageNumber="487" updateTime="1698359547105" updateUser="plazi">
<mods:mods id="8D2738FDFA2415E8FF6060534B77E717" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7DC4DC9F1CBFEBA4923077886E07EEFF">
<mods:title id="B3DD906411CC99D54D2E99ECA6BB5559">New species of the genus Ypsiloncyphon (Insecta: Coleoptera: Scirtidae: Scirtinae)</mods:title>
</mods:titleInfo>
<mods:name id="28D39DD5865D2B208903808882F2AD82" type="personal">
<mods:role id="3B10AC9FDAB7E39A5DE0B652CB7F3FC9">
<mods:roleTerm id="7FC896B30633766F797805D371BA6652">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9CF2ADFF36EB5B64A2AEE936A76B999E">Yoshitomi, Hiroyuki</mods:namePart>
</mods:name>
<mods:typeOfResource id="38BC3160DE18063F51C6A381D4F9CCCB">text</mods:typeOfResource>
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<mods:title id="E240DA2675F8FEC95464CAFD928F3884">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3DB33D33C715B41414C6691CAC088EF5">
<mods:date id="55882BD85FBF5C8F6C67C88FA252E8E6">2015</mods:date>
<mods:detail id="84D4FD45E2629E4B2108407C476C344D" type="volume">
<mods:number id="E16E6E3B52136DACB88C044B17EB9C2F">3904</mods:number>
</mods:detail>
<mods:detail id="393A4AFE568B5A4C1229E9F3B43885A7" type="issue">
<mods:number id="D036B31C1222811EEA1D2A1818FE4320">4</mods:number>
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<mods:start id="5EC9A4A0489F777C026802173BE9BC4F">451</mods:start>
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<mods:classification id="4E53DB317A7134360C8B4FB9BCE1213C">journal article</mods:classification>
<mods:identifier id="30EBDB01CC4CBA9A905C7D317B328DCA" type="DOI">10.11646/zootaxa.3904.4.1</mods:identifier>
<mods:identifier id="5D79C56EC5101A772416DF147E09B541" type="GBIF-Dataset">a1c49726-9afa-4be5-85ef-9d0caabac1fe</mods:identifier>
<mods:identifier id="109A259972DFB1E2DF60A1AA5DC271A1" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B61DAB1DB3A0C6ABA580D983777AB76B" type="Zenodo-Dep">239899</mods:identifier>
<mods:identifier id="F9823C6C6D0554583BFCD562DCD07113" type="ZooBank">38457A27-A15B-4E87-BE72-9A3AB007CAA5</mods:identifier>
</mods:mods>
<treatment id="03AC6D78C825216549B2EB293196FA58" ID-DOI="http://doi.org/10.5281/zenodo.6111900" ID-GBIF-Taxon="119586896" ID-Zenodo-Dep="6111900" LSID="urn:lsid:zoobank.org:act:A10D515B-99B6-4D36-9D3B-FC22C30D2BA8" httpUri="http://treatment.plazi.org/id/03AC6D78C825216549B2EB293196FA58" lastPageId="38" lastPageNumber="489" pageId="36" pageNumber="487">
<subSubSection id="C31F8FE5C825216749B2EB2934C9FD3E" pageId="36" pageNumber="487" type="nomenclature">
<paragraph id="8BBADC6EC825216749B2EB29376AFDDF" blockId="36.[151,516,583,643]" box="[151,516,583,610]" pageId="36" pageNumber="487">
<heading id="D0F26B02C825216749B2EB29376AFDDF" bold="true" box="[151,516,583,610]" fontSize="11" level="1" pageId="36" pageNumber="487" reason="1">
<emphasis id="B971007CC825216749B2EB29376AFDDF" bold="true" box="[151,516,583,610]" pageId="36" pageNumber="487">
<taxonomicName id="4C05A7EDC825216749B2EB2934D5FDDC" box="[151,443,583,609]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="teruhisai" status="sp. nov.">
<emphasis id="B971007CC825216749B2EB2934D5FDDC" bold="true" box="[151,443,583,609]" italics="true" pageId="36" pageNumber="487">Ypsiloncyphon teruhisai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A242BD07C825216748E7EB26376AFDDF" box="[450,516,584,610]" pageId="36" pageNumber="487" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBADC6EC825216749B2EB0434C9FD3E" blockId="36.[151,516,583,643]" box="[151,423,618,643]" pageId="36" pageNumber="487">
(
<figureCitation id="133EC0EBC825216749BAEB043594FD3E" box="[159,250,618,643]" captionStart="FIGURE 26" captionStartId="32.[151,250,1735,1757]" captionTargetBox="[150,1432,471,1702]" captionTargetId="figure@32.[137,1445,461,1716]" captionTargetPageId="32" captionText="FIGURE 26. Habitus of Ypsiloncyphon spp., holotype, male (except for C, F, H, I). A) Y. wallacei sp. n., B) Y. chiangmaiensis sp. n., C) Y. luteoapicalis (Ruta, 2007), non-type specimen, male, D) Y. flexus sp. n., E) Y. gedensis sp. n., F) Y. gedensis sp. n., paratype, female, G) Y. teruhisai sp. n., H) Y. teruhisai sp. n., paratype, female, I) Y. micans (Klausnitzer, 1973), non-type specimen, male, J) Y. lanigerus sp. n., K) Y. cervus sp. n. Scales: 1.0 mm." httpUri="https://zenodo.org/record/239925/files/figure.png" pageId="36" pageNumber="487">Figs. 26</figureCitation>
G, 26H, 30, 31)
</paragraph>
</subSubSection>
<subSubSection id="C31F8FE5C825216749B2EBDC36EEFB22" pageId="36" pageNumber="487" type="materials_examined">
<paragraph id="8BBADC6EC825216749B2EBDC36EEFB22" blockId="36.[151,1437,690,2011]" pageId="36" pageNumber="487">
<emphasis id="B971007CC825216749B2EBDC3428FD76" bold="true" box="[151,326,690,715]" pageId="36" pageNumber="487">
<typeStatus id="54BE62CCC825216749B2EBDC35BCFD76" box="[151,210,690,715]" pageId="36" pageNumber="487">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54BE62CCC82521674868EBDC34DBFD76" box="[333,437,690,715]" pageId="36" pageNumber="487" type="holotype">Holotype</typeStatus>
(
<collectionCode id="ED1444ABC825216748E3EBDD3765FD76" box="[454,523,691,715]" pageId="36" pageNumber="487">ITBC</collectionCode>
), male, “Poring Park, Ranau, Sabah, Borneo
<collectingCountry id="F3129CFEC82521674D2FEBDC311FFD76" box="[1034,1137,690,715]" name="Malaysia" pageId="36" pageNumber="487">Malaysia</collectingCountry>
<date id="FFBBFAAEC82521674D5DEBDD3068FD76" box="[1144,1286,691,715]" pageId="36" pageNumber="487" value="2008-03-24">24. III. 2008</date>
H. Takizawa leg.”.
<typeStatus id="54BE62CCC825216749FFEBB93429FD52" box="[218,327,727,751]" pageId="36" pageNumber="487" type="paratype">Paratypes</typeStatus>
(
<collectionCode id="ED1444ABC82521674870EBB934F7FD52" box="[341,409,727,751]" pageId="36" pageNumber="487">ITBC</collectionCode>
,
<collectionCode id="ED1444ABC82521674881EBB9349EFD53" box="[420,496,727,750]" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="36" pageNumber="487">EUMJ</collectionCode>
,
<collectionCode id="ED1444ABC825216748DEEBB93724FD52" box="[507,586,727,751]" pageId="36" pageNumber="487">SEHU</collectionCode>
):
<specimenCount id="9D0317E7C82521674B79EBB937C9FD52" box="[604,679,726,751]" pageId="36" pageNumber="487" type="male">1 male</specimenCount>
&amp;
<specimenCount id="9D0317E7C82521674BEFEBB93640FD52" box="[714,814,726,751]" pageId="36" pageNumber="487" type="female">1 female</specimenCount>
, dame data and collector as for the
<typeStatus id="54BE62CCC82521674D92EBB83072FD52" box="[1207,1308,726,751]" pageId="36" pageNumber="487" type="holotype">holotype</typeStatus>
, but “9,
<date id="FFBBFAAEC82521674C5CEBB93585FCAE" pageId="36" pageNumber="487" value="2009-01-10">10. I. 2009</date>
”;
<specimenCount id="9D0317E7C82521674827EB953408FCAE" box="[258,358,762,787]" pageId="36" pageNumber="487" type="female">1 female</specimenCount>
, “Borneo, Sabah Mt. Kinabalu ParkHead Quarter
<date id="FFBBFAAEC82521674ABCEB95314BFCAE" box="[921,1061,763,787]" pageId="36" pageNumber="487" value="1989-03-22">22. III. 1989</date>
K. Matsumoto leg.”, genit. s. no. HY 1032;
<specimenCount id="9D0317E7C8252167482BEA713432FC8A" box="[270,348,798,823]" pageId="36" pageNumber="487" type="male">1 male</specimenCount>
, “[E.
<collectingCountry id="F3129CFEC825216748B8EA703769FC8A" box="[413,519,798,823]" name="Malaysia" pageId="36" pageNumber="487">Malaysia</collectingCountry>
] Poring Hot Spring (alt.
<quantity id="4CFD718BC82521674A3BEA703608FC8A" box="[798,870,798,823]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="36" pageNumber="487" unit="m" value="600.0">600 m</quantity>
) Sabah, Is. Borneo
<date id="FFBBFAAEC82521674D60EA703199FC8A" box="[1093,1271,798,823]" pageId="36" pageNumber="487" value="1993-03-20" valueMax="1993-03-26" valueMin="1993-03-20">20–26. iii. 1993</date>
T. Ueno leg.”, genit. s. no. HY 768;
<specimenCount id="9D0317E7C825216748B3EA2D3493FCE6" box="[406,509,834,859]" pageId="36" pageNumber="487" type="female">1 female</specimenCount>
, “
<collectionCode id="ED1444ABC82521674B30EA2D373AFCE6" box="[533,596,835,859]" httpUri="http://grbio.org/cool/8va9-jhre" name="Universiti Malaysia Sabah" pageId="36" pageNumber="487">UMS</collectionCode>
hill, Kota Kinabalu, Sabah, BORNEO
<date id="FFBBFAAEC82521674D38EA2D31CAFCE6" box="[1053,1188,835,859]" pageId="36" pageNumber="487" value="2007-05-12">12. V. 2007</date>
H. Takizawa leg.”;
<specimenCount id="9D0317E7C82521674CABEA2D35BEFCC2" pageId="36" pageNumber="487" type="male">1 male</specimenCount>
, “Mahua Waterfall, Tambunan, Sabah BORNEO
<date id="FFBBFAAEC82521674A0CEA093684FCC2" box="[809,1002,871,895]" pageId="36" pageNumber="487" value="2007-06-22" valueMax="2007-06-24" valueMin="2007-06-22">22-24. VI. 2007</date>
H. Takizawa leg.”;
<specimenCount id="9D0317E7C82521674DCDEA09303AFCC2" box="[1256,1364,870,895]" pageId="36" pageNumber="487" type="female">1 female</specimenCount>
, “Kg. Madsiang, Penampang, Sabah, BORNEO
<date id="FFBBFAAEC82521674B56EAE53665FC1E" box="[627,779,907,931]" pageId="36" pageNumber="487" value="2007-07-10">10. VII. 2007</date>
H. Takizawa leg.”;
<specimenCount id="9D0317E7C82521674ACBEAE5313CFC1E" box="[1006,1106,906,931]" pageId="36" pageNumber="487" type="female">1 female</specimenCount>
, “Kg. Mong Kuslad, Moyog. Penampang Sabah, BORNEO
<date id="FFBBFAAEC825216748D5EAC137E7FC7A" box="[496,649,943,967]" pageId="36" pageNumber="487" value="2007-07-22">22. VII. 2007</date>
H. Takizawa leg.”;
<specimenCount id="9D0317E7C82521674A4BEAC136B0FC7A" box="[878,990,942,967]" pageId="36" pageNumber="487" type="female">2 females</specimenCount>
, same locality and collector, but “
<date id="FFBBFAAEC82521674C44EAC135BAFC56" pageId="36" pageNumber="487" value="2008-01-13">13. I. 2008</date>
”;
<specimenCount id="9D0317E7C825216749CEEABD3454FC56" box="[235,314,978,1003]" pageId="36" pageNumber="487" type="male">1 male</specimenCount>
, same locality and collector, but “
<date id="FFBBFAAEC82521674B9BEABD363DFC56" box="[702,851,979,1003]" pageId="36" pageNumber="487" value="2008-09-13">13. IX. 2008</date>
”;
<specimenCount id="9D0317E7C82521674A4EEABD36BEFC56" box="[875,976,978,1003]" pageId="36" pageNumber="487" type="female">1 female</specimenCount>
, “Aru Dalan, Bario, Kelambit, highland,
<collectingCountry id="F3129CFEC825216749B2EA983594FBB2" box="[151,250,1014,1039]" name="Malaysia" pageId="36" pageNumber="487">Sarawak</collectingCountry>
,
<collectingCountry id="F3129CFEC82521674820EA983405FBB2" box="[261,363,1014,1039]" name="Malaysia" pageId="36" pageNumber="487">Malaysia</collectingCountry>
<date id="FFBBFAAEC82521674857EA98349FFBB2" box="[370,497,1014,1039]" pageId="36" pageNumber="487" value="2007-09-06">6. IX. 2007</date>
H. Takizawa leg.”;
<specimenCount id="9D0317E7C82521674BEBEA993649FBB2" box="[718,807,1014,1039]" pageId="36" pageNumber="487" type="male">3 males</specimenCount>
, “Bundu Tuhan, Ranau, Sabah BORNEO
<date id="FFBBFAAEC82521674DDBEA993014FBB2" box="[1278,1402,1015,1039]" pageId="36" pageNumber="487" value="2008-03-01">1. III. 2008</date>
H. Takizawa leg.”;
<specimenCount id="9D0317E7C82521674877ED7534F1FB8E" box="[338,415,1050,1075]" pageId="36" pageNumber="487" type="male">1 male</specimenCount>
&amp;
<specimenCount id="9D0317E7C825216748EDED753740FB8E" box="[456,558,1050,1075]" pageId="36" pageNumber="487" type="female">1 female</specimenCount>
, “Mamut, Ranau, Sabah Borneo,
<collectingCountry id="F3129CFEC82521674A97ED743177FB8E" box="[946,1049,1050,1075]" name="Malaysia" pageId="36" pageNumber="487">Malaysia</collectingCountry>
<date id="FFBBFAAEC82521674D07ED7531CBFB8E" box="[1058,1189,1051,1075]" pageId="36" pageNumber="487" value="2008-03-02">2. III. 2008</date>
H. Takizawa leg.”;
<specimenCount id="9D0317E7C82521674CABED7535BEFBEA" pageId="36" pageNumber="487" type="male">1 male</specimenCount>
, “Kg. Guramboi,
<quantity id="4CFD718BC825216748B3ED5034B8FBEA" box="[406,470,1086,1111]" metricMagnitude="4" metricUnit="m" metricValue="4.3" pageId="36" pageNumber="487" unit="km" value="43.0">43km</quantity>
Penampang, Sabah, Borneo
<date id="FFBBFAAEC82521674A3DED5036CDFBEA" box="[792,931,1086,1111]" pageId="36" pageNumber="487" value="2008-03-15">15. III. 2008</date>
H. Takizawa leg.”;
<specimenCount id="9D0317E7C82521674DA7ED51318BFBEA" box="[1154,1253,1086,1111]" pageId="36" pageNumber="487" type="female">1 female</specimenCount>
, “Ridon, Lanjak Entimau wildlife Sanct., Kapit,
<collectingCountry id="F3129CFEC82521674B22ED0C3709FBC6" box="[519,615,1122,1147]" name="Malaysia" pageId="36" pageNumber="487">Sarawak</collectingCountry>
<date id="FFBBFAAEC82521674B54ED0D366BFBC6" box="[625,773,1123,1147]" pageId="36" pageNumber="487" value="2008-06-18">18. VI. 2008</date>
H. Takizawa leg.”;
<specimenCount id="9D0317E7C82521674AD4ED0D3136FBC6" box="[1009,1112,1122,1147]" pageId="36" pageNumber="487" type="female">1 female</specimenCount>
, “Manggis Subst., Kinabalu Park, Ranau, Sabah, BORNEO
<date id="FFBBFAAEC825216748DFEDE937C6FB22" box="[506,680,1159,1183]" pageId="36" pageNumber="487" value="2008-12-09" valueMax="2008-12-10" valueMin="2008-12-09">9-10. XII. 2008</date>
H. Takizawa leg.”.
</paragraph>
</subSubSection>
<subSubSection id="C31F8FE5C825216649E2EDC431E2FE3A" lastPageId="37" lastPageNumber="488" pageId="36" pageNumber="487" type="description">
<paragraph id="8BBADC6EC825216749E2EDC431C7FB5A" blockId="36.[151,1437,690,2011]" pageId="36" pageNumber="487">
<emphasis id="B971007CC825216749E2EDC43437FB7E" bold="true" box="[199,345,1194,1219]" pageId="36" pageNumber="487">Description.</emphasis>
Male. Body (
<figureCitation id="133EC0EBC825216748DDEDC43722FB7E" box="[504,588,1194,1219]" captionStart="FIGURE 26" captionStartId="32.[151,250,1735,1757]" captionTargetBox="[150,1432,471,1702]" captionTargetId="figure@32.[137,1445,461,1716]" captionTargetPageId="32" captionText="FIGURE 26. Habitus of Ypsiloncyphon spp., holotype, male (except for C, F, H, I). A) Y. wallacei sp. n., B) Y. chiangmaiensis sp. n., C) Y. luteoapicalis (Ruta, 2007), non-type specimen, male, D) Y. flexus sp. n., E) Y. gedensis sp. n., F) Y. gedensis sp. n., paratype, female, G) Y. teruhisai sp. n., H) Y. teruhisai sp. n., paratype, female, I) Y. micans (Klausnitzer, 1973), non-type specimen, male, J) Y. lanigerus sp. n., K) Y. cervus sp. n. Scales: 1.0 mm." httpUri="https://zenodo.org/record/239925/files/figure.png" pageId="36" pageNumber="487">Fig. 26</figureCitation>
G) oblong, convex dorsally, shining, closely covered with yellowish-white setae throughout. Coloration of body almost blackish-brown, but antennomeres II–III brown.
</paragraph>
<paragraph id="8BBADC6EC825216749E2ED9C3711FA5E" blockId="36.[151,1437,690,2011]" pageId="36" pageNumber="487">Head moderate in size, slightly convex dorsally, sparsely punctate, almost straight in front margin. Eyes large, prominent; the distance between eyes about 1.7 times as long as the maximum diameter of an eye. Antennae relatively long, reaching about proximal 1/3 of elytra. Pronotum densely punctate; front and lateral margins almost straight; antero-lateral angles obtuse, slightly produced anteriorly; postero-lateral angles rectangular; PW/PL 1.92–2.20 (2.07). Scutellar shield subtriangular, finely punctate. Elytra oblong, widest near the humeral parts, thence gently tapered apically; apices rectangular; EL/EW 1.58–1.69 (1.64); EL/PL 4.75–5.50 (5.15); EW/PW 1.39–1.61 (1.51); TL/EW 1.91–2.02 (1.96).</paragraph>
<paragraph id="8BBADC6EC825216749E2EC80347AF8F7" blockId="36.[151,1437,690,2011]" pageId="36" pageNumber="487">
Caudal margin of sternite VII (
<figureCitation id="133EC0EBC82521674B13EC8037E2F9BA" box="[566,652,1518,1543]" captionStart="FIGURE 30" captionStartId="37.[151,250,1963,1985]" captionTargetBox="[253,1331,453,1935]" captionTargetId="figure@37.[248,1339,445,1942]" captionTargetPageId="37" captionText="FIGURE 30. Ypsiloncyphon teruhisai sp. n., holotype, male. A) sternites V – VII, B) tergite VIII, C) sternite VIII, D) tergite IX, E) sternite IX, F) tegmen, G) penis in ventral view." httpUri="https://zenodo.org/record/239929/files/figure.png" pageId="36" pageNumber="487">Fig. 30</figureCitation>
A) arcuate. Tergite VIII (
<figureCitation id="133EC0EBC82521674A90EC803165F9BA" box="[949,1035,1518,1543]" captionStart="FIGURE 30" captionStartId="37.[151,250,1963,1985]" captionTargetBox="[253,1331,453,1935]" captionTargetId="figure@37.[248,1339,445,1942]" captionTargetPageId="37" captionText="FIGURE 30. Ypsiloncyphon teruhisai sp. n., holotype, male. A) sternites V – VII, B) tergite VIII, C) sternite VIII, D) tergite IX, E) sternite IX, F) tegmen, G) penis in ventral view." httpUri="https://zenodo.org/record/239929/files/figure.png" pageId="36" pageNumber="487">Fig. 30</figureCitation>
B) well sclerotized, wide, sparsely punctate in posterior part, projecting laterally in proximal part, with a pair of long and slender apodemes protruding from near antero-lateral corners; sternite VIII (
<figureCitation id="133EC0EBC82521674B94EF58366AF9F2" box="[689,772,1590,1615]" captionStart="FIGURE 30" captionStartId="37.[151,250,1963,1985]" captionTargetBox="[253,1331,453,1935]" captionTargetId="figure@37.[248,1339,445,1942]" captionTargetPageId="37" captionText="FIGURE 30. Ypsiloncyphon teruhisai sp. n., holotype, male. A) sternites V – VII, B) tergite VIII, C) sternite VIII, D) tergite IX, E) sternite IX, F) tegmen, G) penis in ventral view." httpUri="https://zenodo.org/record/239929/files/figure.png" pageId="36" pageNumber="487">Fig. 30</figureCitation>
C) well sclerotized, short, T-shaped; tergite IX (
<figureCitation id="133EC0EBC82521674C09EF583011F9F2" box="[1324,1407,1590,1615]" captionStart="FIGURE 30" captionStartId="37.[151,250,1963,1985]" captionTargetBox="[253,1331,453,1935]" captionTargetId="figure@37.[248,1339,445,1942]" captionTargetPageId="37" captionText="FIGURE 30. Ypsiloncyphon teruhisai sp. n., holotype, male. A) sternites V – VII, B) tergite VIII, C) sternite VIII, D) tergite IX, E) sternite IX, F) tegmen, G) penis in ventral view." httpUri="https://zenodo.org/record/239929/files/figure.png" pageId="36" pageNumber="487">Fig. 30</figureCitation>
D) lightly sclerotized, trapezoidal, densely covered with short spines in posterior part, with a pair of very long apodemes protruding from near antero-lateral corners; sternite IX (
<figureCitation id="133EC0EBC82521674AB3EF103684F92A" box="[918,1002,1662,1687]" captionStart="FIGURE 30" captionStartId="37.[151,250,1963,1985]" captionTargetBox="[253,1331,453,1935]" captionTargetId="figure@37.[248,1339,445,1942]" captionTargetPageId="37" captionText="FIGURE 30. Ypsiloncyphon teruhisai sp. n., holotype, male. A) sternites V – VII, B) tergite VIII, C) sternite VIII, D) tergite IX, E) sternite IX, F) tegmen, G) penis in ventral view." httpUri="https://zenodo.org/record/239929/files/figure.png" pageId="36" pageNumber="487">Fig. 30</figureCitation>
E) well sclerotized, Y-shaped, bifid in apical 5/6, serrate in apical part, with pointed apex and long subterminal spine. Tegmen (
<figureCitation id="133EC0EBC82521674D95EFCC3068F906" box="[1200,1286,1698,1723]" captionStart="FIGURE 30" captionStartId="37.[151,250,1963,1985]" captionTargetBox="[253,1331,453,1935]" captionTargetId="figure@37.[248,1339,445,1942]" captionTargetPageId="37" captionText="FIGURE 30. Ypsiloncyphon teruhisai sp. n., holotype, male. A) sternites V – VII, B) tergite VIII, C) sternite VIII, D) tergite IX, E) sternite IX, F) tegmen, G) penis in ventral view." httpUri="https://zenodo.org/record/239929/files/figure.png" pageId="36" pageNumber="487">Fig. 30</figureCitation>
F) long, well sclerotized, Y-shaped, shorter than sternite IX, bifid in apical 2/3, with pointed apices. Penis (
<figureCitation id="133EC0EBC82521674D90EFA83066F962" box="[1205,1288,1734,1759]" captionStart="FIGURE 30" captionStartId="37.[151,250,1963,1985]" captionTargetBox="[253,1331,453,1935]" captionTargetId="figure@37.[248,1339,445,1942]" captionTargetPageId="37" captionText="FIGURE 30. Ypsiloncyphon teruhisai sp. n., holotype, male. A) sternites V – VII, B) tergite VIII, C) sternite VIII, D) tergite IX, E) sternite IX, F) tegmen, G) penis in ventral view." httpUri="https://zenodo.org/record/239929/files/figure.png" pageId="36" pageNumber="487">Fig. 30</figureCitation>
G) long, well sclerotized, about 1.2 times as long as tegmen, widest at about proximal 2/3; pala gently tapered; parameroids simple, finely punctate; trigonium short, projecting from proximal 3/7 to 2/3, closely covered with fine serrations in apical part.
</paragraph>
<paragraph id="8BBADC6EC825216649E2EE3834A8FEA0" blockId="36.[151,1437,690,2011]" lastBlockId="37.[151,1437,151,392]" lastPageId="37" lastPageNumber="488" pageId="36" pageNumber="487">
Female. Sexual dimorphism (
<figureCitation id="133EC0EBC82521674B31EE383708F8D2" box="[532,614,1878,1903]" captionStart="FIGURE 26" captionStartId="32.[151,250,1735,1757]" captionTargetBox="[150,1432,471,1702]" captionTargetId="figure@32.[137,1445,461,1716]" captionTargetPageId="32" captionText="FIGURE 26. Habitus of Ypsiloncyphon spp., holotype, male (except for C, F, H, I). A) Y. wallacei sp. n., B) Y. chiangmaiensis sp. n., C) Y. luteoapicalis (Ruta, 2007), non-type specimen, male, D) Y. flexus sp. n., E) Y. gedensis sp. n., F) Y. gedensis sp. n., paratype, female, G) Y. teruhisai sp. n., H) Y. teruhisai sp. n., paratype, female, I) Y. micans (Klausnitzer, 1973), non-type specimen, male, J) Y. lanigerus sp. n., K) Y. cervus sp. n. Scales: 1.0 mm." httpUri="https://zenodo.org/record/239925/files/figure.png" pageId="36" pageNumber="487">Fig. 26</figureCitation>
H) distinct: coloration of elytra brown. PW/PL 2.00–2.26 (2.11); EL/EW 1.68–1.86 (1.74); EL/PL 5.20–6.40 (5.59); EW/PW 1.43–1.63 (1.52); TL/EW 2.00–2.15 (2.06). Caudal margin of sternite VII (
<figureCitation id="133EC0EBC82521674802EEF03419F80A" box="[295,375,1950,1975]" captionStart="FIGURE 31" captionStartId="38.[151,250,1285,1307]" captionTargetBox="[298,1287,188,1255]" captionTargetId="figure@38.[293,1294,185,1262]" captionTargetPageId="38" captionText="FIGURE 31. Ypsiloncyphon teruhisai sp. n., paratype, female. A) sternites V – VII, B) tergite VIII, C) sternite VIII, D) ovipositor." httpUri="https://zenodo.org/record/239930/files/figure.png" pageId="36" pageNumber="487">Fig. 31</figureCitation>
A) arcuate. Tergite VIII (
<figureCitation id="133EC0EBC82521674BB4EEF0378FF80A" box="[657,737,1950,1975]" captionStart="FIGURE 31" captionStartId="38.[151,250,1285,1307]" captionTargetBox="[298,1287,188,1255]" captionTargetId="figure@38.[293,1294,185,1262]" captionTargetPageId="38" captionText="FIGURE 31. Ypsiloncyphon teruhisai sp. n., paratype, female. A) sternites V – VII, B) tergite VIII, C) sternite VIII, D) ovipositor." httpUri="https://zenodo.org/record/239930/files/figure.png" pageId="36" pageNumber="487">Fig. 31</figureCitation>
B) moderately sclerotized, elongate subtriangular, excised and bearing short spines in apical margin, sparsely punctate in apical part, with a pair of long and slender apodemes protruding from antero-lateral corners. Sternite VIII (
<figureCitation id="133EC0EBC82421664BCBE9F93651FF0D" box="[750,831,151,176]" captionStart="FIGURE 31" captionStartId="38.[151,250,1285,1307]" captionTargetBox="[298,1287,188,1255]" captionTargetId="figure@38.[293,1294,185,1262]" captionTargetPageId="38" captionText="FIGURE 31. Ypsiloncyphon teruhisai sp. n., paratype, female. A) sternites V – VII, B) tergite VIII, C) sternite VIII, D) ovipositor." httpUri="https://zenodo.org/record/239930/files/figure.png" pageId="37" pageNumber="488">Fig. 31</figureCitation>
C) moderately sclerotized, oblong, shallowly concave and bearing short spines in apical margin, punctate in apical part, with a long median apodeme. Ovipositor (
<figureCitation id="133EC0EBC82421664C4AE9D235DDFF45" captionStart="FIGURE 31" captionStartId="38.[151,250,1285,1307]" captionTargetBox="[298,1287,188,1255]" captionTargetId="figure@38.[293,1294,185,1262]" captionTargetPageId="38" captionText="FIGURE 31. Ypsiloncyphon teruhisai sp. n., paratype, female. A) sternites V – VII, B) tergite VIII, C) sternite VIII, D) ovipositor." httpUri="https://zenodo.org/record/239930/files/figure.png" pageId="37" pageNumber="488">Fig. 31</figureCitation>
D) very long, with a branch in apical 1/6 of baculus; approximate ratio of stylus, coxite and baculus (n = 1) as -: 20: 169; prehensor absent.
</paragraph>
<paragraph id="8BBADC6EC824216649E2E84931E2FE3A" blockId="37.[151,1437,151,392]" pageId="37" pageNumber="488">
<emphasis id="B971007CC824216649E2E8493413FEFD" bold="true" box="[199,381,295,320]" pageId="37" pageNumber="488">Measurements.</emphasis>
Male (n = 10): TL 2.96–3.58 (3.23) mm; PW 1.00–1.20 (1.09) mm; PL 0.48–0.60 (0.53) mm; EL 2.48–2.98 (2.71) mm; EW 1.55–1.80 (1.65) mm. Female (n = 6): TL 3.07–3.22 (3.16) mm; PW 0.97–1.08 (1.01) mm; PL 0.43–0.50 (0.48) mm; EL 2.59–2.75 (2.68) mm; EW 1.48–1.60 (1.54) mm.
</paragraph>
</subSubSection>
<caption id="DF7A8CE6C824216649B2EEC537F5F85D" httpUri="https://zenodo.org/record/239929/files/figure.png" pageId="37" pageNumber="488" targetBox="[253,1331,453,1935]" targetPageId="37">
<paragraph id="8BBADC6EC824216649B2EEC537F5F85D" blockId="37.[151,1436,1963,2016]" pageId="37" pageNumber="488">
<emphasis id="B971007CC824216649B2EEC53476F87D" bold="true" box="[151,280,1963,1985]" pageId="37" pageNumber="488">FIGURE 30</emphasis>
.
<taxonomicName id="4C05A7EDC82421664800EEC53779F87C" box="[293,535,1963,1985]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="488" phylum="Arthropoda" rank="species" species="teruhisai" status="sp. nov.">
<emphasis id="B971007CC82421664800EEC53779F87C" box="[293,535,1963,1985]" italics="true" pageId="37" pageNumber="488">Ypsiloncyphon teruhisai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A242BD07C82421664B3BEEC23738F87C" box="[542,598,1964,1985]" pageId="37" pageNumber="488" rank="species">
<emphasis id="B971007CC82421664B3BEEC23738F87C" bold="true" box="[542,598,1964,1985]" pageId="37" pageNumber="488">sp. n.</emphasis>
</taxonomicNameLabel>
, holotype, male. A) sternites V–VII, B) tergite VIII, C) sternite VIII, D) tergite IX, E) sternite IX, F) tegmen, G) penis in ventral view.
</paragraph>
</caption>
<caption id="DF7A8CE6C827216549B2EC6B346AFA84" httpUri="https://zenodo.org/record/239930/files/figure.png" pageId="38" pageNumber="489" targetBox="[298,1287,188,1255]" targetPageId="38">
<paragraph id="8BBADC6EC827216549B2EC6B346AFA84" blockId="38.[151,1435,1284,1337]" pageId="38" pageNumber="489">
<emphasis id="B971007CC827216549B2EC6B3473FAA7" bold="true" box="[151,285,1285,1307]" pageId="38" pageNumber="489">FIGURE 31</emphasis>
.
<taxonomicName id="4C05A7EDC82721654815EC6A3746FAA7" box="[304,552,1284,1306]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="489" phylum="Arthropoda" rank="species" species="teruhisai" status="sp. nov.">
<emphasis id="B971007CC82721654815EC6A3746FAA7" box="[304,552,1284,1306]" italics="true" pageId="38" pageNumber="489">Ypsiloncyphon teruhisai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A242BD07C82721654B16EC6B371FFAA7" box="[563,625,1285,1306]" pageId="38" pageNumber="489" rank="species">
<emphasis id="B971007CC82721654B16EC6B371FFAA7" bold="true" box="[563,625,1285,1306]" pageId="38" pageNumber="489">sp. n.</emphasis>
</taxonomicNameLabel>
, paratype, female. A) sternites V–VII, B) tergite VIII, C) sternite VIII, D) ovipositor.
</paragraph>
</caption>
<subSubSection id="C31F8FE5C827216549E2EC0E372FFAC4" box="[199,577,1376,1401]" pageId="38" pageNumber="489" type="distribution">
<paragraph id="8BBADC6EC827216549E2EC0E372FFAC4" blockId="38.[151,1436,1376,1509]" box="[199,577,1376,1401]" pageId="38" pageNumber="489">
<emphasis id="B971007CC827216549E2EC0E340EFAC4" bold="true" box="[199,352,1376,1401]" pageId="38" pageNumber="489">Distribution.</emphasis>
<collectingCountry id="F3129CFEC82721654842EC0E34A0FAC4" box="[359,462,1376,1401]" name="Malaysia" pageId="38" pageNumber="489">Malaysia</collectingCountry>
(Borneo).
</paragraph>
</subSubSection>
<subSubSection id="C31F8FE5C827216549E2ECEA3196FA58" pageId="38" pageNumber="489" type="discussion">
<paragraph id="8BBADC6EC827216549E2ECEA3196FA58" blockId="38.[151,1436,1376,1509]" pageId="38" pageNumber="489">
<emphasis id="B971007CC827216549E2ECEA3455FA20" bold="true" box="[199,315,1412,1437]" pageId="38" pageNumber="489">Remarks.</emphasis>
This species and
<taxonomicName id="4C05A7EDC82721654B21ECEB3663FA20" box="[516,781,1413,1437]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="489" phylum="Arthropoda" rank="species" species="gedensis" status="sp. nov.">
<emphasis id="B971007CC82721654B21ECEB3663FA20" box="[516,781,1413,1437]" italics="true" pageId="38" pageNumber="489">Ypsiloncyphon gedensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A242BD07C82721654A30ECEA363BFA20" box="[789,853,1412,1437]" pageId="38" pageNumber="489" rank="species">
<emphasis id="B971007CC82721654A30ECEA363BFA20" bold="true" box="[789,853,1412,1437]" pageId="38" pageNumber="489">sp. n.</emphasis>
</taxonomicNameLabel>
are remarkable species in the genus in having large (TL about 3.0 mm) and oblong bodies. The differences between them are given in the remarks for
<taxonomicName id="4C05A7EDC82721654DF8ECC73038FA7C" box="[1245,1366,1449,1473]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="489" phylum="Arthropoda" rank="species" species="gedensis" status="sp. nov.">
<emphasis id="B971007CC82721654DF8ECC73187FA7D" box="[1245,1257,1449,1472]" italics="true" pageId="38" pageNumber="489">Y</emphasis>
.
<emphasis id="B971007CC82721654DD3ECC73038FA7C" box="[1270,1366,1449,1473]" italics="true" pageId="38" pageNumber="489">gedensis</emphasis>
</taxonomicName>
<emphasis id="B971007CC82721654C78ECC63420FA58" bold="true" pageId="38" pageNumber="489">
<taxonomicNameLabel id="A242BD07C82721654C78ECC630F2FA7C" box="[1373,1436,1448,1473]" pageId="38" pageNumber="489" rank="species">sp. n.</taxonomicNameLabel>
Etymology.
</emphasis>
This species is named after Mr. Teruhisa Uéno, who collected one of the
<typeStatus id="54BE62CCC82721654DA0ECA3319AFA58" box="[1157,1268,1485,1509]" pageId="38" pageNumber="489" type="paratype">paratypes</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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