583 lines
61 KiB
XML
583 lines
61 KiB
XML
<document id="DFC91C3C905A280D5564187C9A06B8B6" ID-DOI="10.11646/zootaxa.4039.4.5" ID-GBIF-Dataset="5d48e0d6-4af9-422e-af66-81503dd42524" ID-ISSN="1175-5326" ID-Zenodo-Dep="241183" ID-ZooBank="C6FD1263-3CAB-47F3-8661-700612D0B59A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461333100469" checkinUser="plazi" docAuthor="Huang, Hao, Bartolozzi, Luca & Chen, Chang-Chin" docDate="2015" docId="0393D10F5A697554D39FAA80F33FFD89" docLanguage="en" docName="zt04039p565.pdf" docOrigin="Zootaxa 4039 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Echinoaesalus borneoensis Huang & Imura 2011, new status" docType="treatment" docVersion="9" lastPageNumber="558" masterDocId="FFAAA9775A687551D308AE67F265FC59" masterDocTitle="Notes on the genus Echinoaesalus Zelenka, 1993 (Coleoptera: Lucanidae), with the description of a new species from the Malay Peninsula" masterLastPageNumber="565" masterPageNumber="553" pageNumber="554" updateTime="1698618809578" updateUser="plazi">
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<mods:title id="80FE2C71013652A6D19465C6DEE3BBF1">Notes on the genus Echinoaesalus Zelenka, 1993 (Coleoptera: Lucanidae), with the description of a new species from the Malay Peninsula</mods:title>
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<mods:namePart id="0536C0C4FAF173153FAE5735CA8A0899">Huang, Hao</mods:namePart>
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<mods:namePart id="94E02882F7AEB13A3E0925C027A54E33">Bartolozzi, Luca</mods:namePart>
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<mods:namePart id="57BB5380C5181670018FDF95C508B8D1">Chen, Chang-Chin</mods:namePart>
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<mods:title id="75DC45C72D098E7E82A5024D0150DEAB">Zootaxa</mods:title>
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<treatment id="0393D10F5A697554D39FAA80F33FFD89" ID-DOI="http://doi.org/10.5281/zenodo.6097384" ID-GBIF-Taxon="119644246" ID-Zenodo-Dep="6097384" LSID="urn:lsid:plazi:treatment:0393D10F5A697554D39FAA80F33FFD89" httpUri="http://treatment.plazi.org/id/0393D10F5A697554D39FAA80F33FFD89" lastPageId="5" lastPageNumber="558" pageId="1" pageNumber="554">
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<subSubSection id="C32033925A697550D39FAA80F140F97A" pageId="1" pageNumber="554" type="nomenclature">
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<paragraph id="8B8560195A697550D39FAA80F1E0F95B" blockId="1.[151,901,1255,1315]" box="[151,901,1255,1282]" pageId="1" pageNumber="554">
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<heading id="D0CDD7755A697550D39FAA80F1E0F95B" bold="true" box="[151,901,1255,1282]" fontSize="11" level="1" pageId="1" pageNumber="554" reason="1">
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<emphasis id="B94EBC0B5A697550D39FAA80F1E0F95B" bold="true" box="[151,901,1255,1282]" pageId="1" pageNumber="554">
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<taxonomicName id="4C3A1B9A5A697550D39FAA80F09EF95B" ID-CoL="6DVSB" authority="Huang & Imura, 2011" authorityName="Huang & Imura" authorityYear="2011" box="[151,763,1255,1282]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="borneoensis" status="stat. nov.">
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<emphasis id="B94EBC0B5A697550D39FAA80F384F958" bold="true" box="[151,481,1255,1281]" italics="true" pageId="1" pageNumber="554">Echinoaesalus borneoensis</emphasis>
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<bibRefCitation id="EFAB1DE85A697550D2E0AA8FF09EF95B" author="Huang" box="[488,763,1256,1282]" pageId="1" pageNumber="554" refString="Huang, H., Imura, Y. & Wu, L. (2011) Description of three new taxa of Echinoaesalus Zelenka from Borneo and Java (Coleoptera, Lucanidae). Kogane, 12, 97 - 107." type="journal article" year="2011">Huang & Imura, 2011</bibRefCitation>
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</taxonomicName>
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<taxonomicNameLabel id="A27D01705A697550D00AAA8FF1E0F95B" box="[770,901,1256,1282]" pageId="1" pageNumber="554">new status</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8560195A697550D39FAB6DF140F97A" blockId="1.[151,901,1255,1315]" box="[151,805,1290,1315]" pageId="1" pageNumber="554">
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(
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||
<figureCitation id="13017C9C5A697550D397AB6DF28BF97A" box="[159,238,1290,1315]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1750,1773]" captionTargetBox="[183,1395,205,1699]" captionTargetId="figure@2.[183,1395,193,1713]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Antenna. 1, Echinoaesalus hidakai, L 12, western Malaysia; 2, E. borneoensis, L 11, Borneo; 3, E. timidus, L 19, western Malaysia; 4, E. sabahensis, L 14, Borneo; 5, E. gedeensis, holotype, Java; 6, E. cechovskyi, holotype, western Malaysia; 7, E. cechovskyi, paratype, Singapore. Arrow directed to end of third antennomere." httpUri="https://zenodo.org/record/241184/files/figure.png" pageId="1" pageNumber="554">Figs. 2</figureCitation>
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||
,
|
||
<figureCitation id="13017C9C5A697550D3F2AB6CF356F97A" box="[250,307,1291,1315]" captionStart="FIGURES 8 – 30" captionStartId="2.[151,264,1842,1865]" captionText="FIGURES 8 – 30. Mentum under same scale (scale bar 0.5 mm). 8, Echinoaesalus hidakai, L 12, western Malaysia; 9, E. borneoensis, L 01, Borneo; 10, E. hidakai, L 10, western Malaysia; 11, E. borneoensis, L 07, Borneo; 12, E. sabahensis, after Krikken (2008), Borneo; 13, E. sabahensis, L 14, Borneo; 14, E. cechovskyi, holotype, western Malaysia; 15, E. cechovskyi, paratype, Singapore; 16 – 18, E. timidus; 16, L 21; 17, L 22; 18, L 18; 19, E. javanus, after Krikken (2008), holotype, Java; 20, E. gedeensis, holotype, Java; 21, E. timidus, after Krikken (2008), holotype, probably male; 22 – 30, E. timidus; 22, L 08; 23, L 02; 24, L 17; 25, L 23; 26, L 09; 27, L 06; 28, L 20; 29, L 13; 30, L 29." pageId="1" pageNumber="554">9, 11</figureCitation>
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||
,
|
||
<figureCitation id="13017C9C5A697550D237AB6CF3EDF97A" box="[319,392,1291,1315]" captionStart="FIGURES 35 – 52" captionStartId="3.[151,264,1853,1876]" captionTargetBox="[202,1395,193,1742]" captionTargetId="figure@3.[202,1396,193,1744]" captionTargetPageId="3" captionText="FIGURES 35 – 52. Protibia (left), mesotibia (middle), and metatibia (right), or just protibia. 35, Echinoaesalus hidakai, L 12; 36, E. hidakai, L 10; 37, E. borneoensis, L 11; 38, E. borneoensis, L 07; 39 – 48, E. timidus; 39, L 17; 40, L 13; 41, L 08; 42, L 02; 43, L 23; 44, L 21; 45, L 22; 46, L 18; 47, L 09; 48, L 06; 49, E. sabahensis, L 14; 50, E. cechovskyi, holotype; 51, E. cechovskyi, paratype; 52, E. gedeensis, holotype." httpUri="https://zenodo.org/record/241185/files/figure.png" pageId="1" pageNumber="554">37–38</figureCitation>
|
||
,
|
||
<figureCitation id="13017C9C5A697550D29BAB6DF3D7F97A" box="[403,434,1290,1315]" captionStart="FIGURES 53 – 56" captionStartId="3.[151,264,1976,1999]" captionText="FIGURES 53 – 56. Hindwing under same scale (scale bar 1 mm). 53, Echinoaesalus timidus, L 08, Sumatra; 54, E. timidus, L 06, Borneo; 55, E. borneoensis, paratype, Borneo; 56, E. cechovskyi, paratype, Singapore." pageId="1" pageNumber="554">55</figureCitation>
|
||
,
|
||
<figureCitation id="13017C9C5A697550D2B5AB6DF063F97A" box="[445,518,1290,1315]" captionStart="FIGURES 57 – 76" captionStartId="4.[151,264,1567,1590]" captionTargetBox="[151,1436,203,1545]" captionTargetId="figure@4.[151,1436,193,1546]" captionTargetPageId="4" captionText="FIGURES 57 – 76. Prosternal process in ventral view. 57, Echinoaesalus hidakai, L 12; 58, E. hidakai, L 10; 59 – 62, E. borneoensis; 59, L 11; 60, L 01; 61, L 07; 62, L 16; 63, E. gedeensis, holotype; 64 – 67, E. timidus; 64, L 08; 65, L 02; 66, L 17; 67, L 21; 68, E. sabahensis, L 14; 69, E. cechovskyi, L 28; 70 – 73, E. timidus; 70, L 09; 71, L 06; 72, L 20; 73, L 13; 74, E. sabahensis, after Krikken 2008; 75, E. cechovskyi, paratype; 76, E. javanus, after Krikken (2008), holotype." httpUri="https://zenodo.org/record/241186/files/figure.png" pageId="1" pageNumber="554">59–62</figureCitation>
|
||
, 78–79, 91–92, 104–106)
|
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</paragraph>
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</subSubSection>
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<subSubSection id="C32033925A697550D39FAB34F087F931" box="[151,738,1362,1384]" pageId="1" pageNumber="554" type="reference_group">
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<paragraph id="8B8560195A697550D39FAB34F087F931" blockId="1.[151,738,1362,1384]" box="[151,738,1362,1384]" pageId="1" pageNumber="554">
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<taxonomicName id="4C3A1B9A5A697550D39FAB34F087F931" ID-CoL="6DVSB" authority="Huang & Imura, 2011" authorityName="Huang & Imura" authorityYear="2011" box="[151,738,1362,1384]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="subSpecies" species="hidakai" subSpecies="borneoensis">
|
||
<emphasis id="B94EBC0B5A697550D39FAB34F39BF931" box="[151,510,1363,1384]" italics="true" pageId="1" pageNumber="554">Echinoaesalus hidakai borneoensis</emphasis>
|
||
<bibRefCitation id="EFAB1DE85A697550D10BAB34F087F931" author="Huang" box="[515,738,1362,1384]" pageId="1" pageNumber="554" refString="Huang, H., Imura, Y. & Wu, L. (2011) Description of three new taxa of Echinoaesalus Zelenka from Borneo and Java (Coleoptera, Lucanidae). Kogane, 12, 97 - 107." type="journal article" year="2011">Huang & Imura, 2011</bibRefCitation>
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</taxonomicName>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C32033925A697550D39FABF2F7EBFA67" pageId="1" pageNumber="554" type="materials_examined">
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<paragraph id="8B8560195A697550D39FABF2F7EBFA67" blockId="1.[151,1437,1429,2033]" pageId="1" pageNumber="554">
|
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<emphasis id="B94EBC0B5A697550D39FABF2F29AF9F7" bold="true" box="[151,255,1429,1454]" pageId="1" pageNumber="554">Material</emphasis>
|
||
.
|
||
<emphasis id="B94EBC0B5A697550D205ABF2F3C9F9F7" bold="true" box="[269,428,1429,1454]" pageId="1" pageNumber="554">
|
||
<collectingCountry id="F32D20895A697550D205ABF2F3C9F9F7" box="[269,428,1429,1454]" name="Indonesia" pageId="1" pageNumber="554">INDONESIA</collectingCountry>
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</emphasis>
|
||
:
|
||
<specimenCount id="9D3CAB905A697550D2B2ABF1F381F9F4" box="[442,484,1430,1454]" pageId="1" pageNumber="554" type="generic">1 ♂</specimenCount>
|
||
(L01), BORNEO, South Kalimantan Province, Kandangan district, Loksado,
|
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<quantity id="4CC2CDFC5A697550D65DABF1F7F9F9F7" box="[1365,1436,1429,1454]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="554" unit="km" value="17.0">17 km</quantity>
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NE,
|
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<quantity id="4CC2CDFC5A697550D3C4ABDDF373F988" box="[204,278,1466,1490]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="554" unit="m" value="900.0">900 m</quantity>
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,
|
||
<date id="FF8446D95A697550D22BABDEF3C6F98B" box="[291,419,1465,1490]" pageId="1" pageNumber="554" value="1997-11-15">15.XI.1997</date>
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–
|
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<date id="FF8446D95A697550D2BBABDEF056F98B" box="[435,563,1465,1490]" pageId="1" pageNumber="554" value="1998-11-15">15.XI.1998</date>
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;
|
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<specimenCount id="9D3CAB905A697550D148ABDDF00EF988" box="[576,619,1466,1490]" pageId="1" pageNumber="554" type="generic">1 ♂</specimenCount>
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(L11),
|
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<emphasis id="B94EBC0B5A697550D1CAABDEF13DF98B" bold="true" box="[706,856,1465,1490]" pageId="1" pageNumber="554">
|
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<collectingCountry id="F32D20895A697550D1CAABDEF13DF98B" box="[706,856,1465,1490]" name="Malaysia" pageId="1" pageNumber="554">MALAYSIA</collectingCountry>
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</emphasis>
|
||
, Sabah, Mt. Kinabalu, near park headquarters,
|
||
<date id="FF8446D95A697550D67AABDDF375F9AF" pageId="1" pageNumber="554" value="2006-10-21" valueMax="2006-10-22" valueMin="2006-10-21">21– 22.X.2006</date>
|
||
, R. Novak; 1 ♀ (L07),
|
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<collectingCountry id="F32D20895A697550D13CABBAF0AFF9AF" box="[564,714,1501,1526]" name="Malaysia" pageId="1" pageNumber="554">
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<emphasis id="B94EBC0B5A697550D13CABBAF0AFF9AF" bold="true" box="[564,714,1501,1526]" pageId="1" pageNumber="554">MALAYSIA</emphasis>
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</collectingCountry>
|
||
, Sabah, Crocker range,
|
||
<date id="FF8446D95A697550D0E5ABBAF605F9AF" box="[1005,1120,1501,1526]" pageId="1" pageNumber="554" value="2007-03-05">5.III.2007</date>
|
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.
|
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<emphasis id="B94EBC0B5A697550D778ABBAF763F9AF" bold="true" box="[1136,1286,1501,1526]" pageId="1" pageNumber="554">
|
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<collectingCountry id="F32D20895A697550D778ABBAF763F9AF" box="[1136,1286,1501,1526]" name="Malaysia" pageId="1" pageNumber="554">MALAYSIA</collectingCountry>
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</emphasis>
|
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: 1 ♀ (L27), Kelantan,
|
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<quantity id="4CC2CDFC5A697550D202A866F337FA43" box="[266,338,1537,1562]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="554" unit="km" value="60.0">60 km</quantity>
|
||
NE of Tanah Rala, Tanah Kerajaan,
|
||
<quantity id="4CC2CDFC5A697550D1F9A865F12CFA40" box="[753,841,1538,1562]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="554" unit="m" value="1000.0">1000 m</quantity>
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,
|
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<date id="FF8446D95A697550D05DA865F198FA43" box="[853,1021,1538,1562]" pageId="1" pageNumber="554" value="2007-04-12" valueMax="2007-04-30" valueMin="2007-04-12">12–30.IV.2007</date>
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, P. Cechovsky; 1 ♀ (L16), Kelantan,
|
||
<quantity id="4CC2CDFC5A697550D39FA841F2BBFA67" box="[151,222,1573,1598]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="554" unit="km" value="30.0">30 km</quantity>
|
||
NW of Gua Musang, Ulu Lalat Mt., Kampong Sungai Om,
|
||
<quantity id="4CC2CDFC5A697550D088A841F66AFA64" box="[896,1039,1574,1598]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="1" pageNumber="554" unit="m" value="900.0" valueMax="1000.0" valueMin="800.0">800–1000 m</quantity>
|
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,
|
||
<date id="FF8446D95A697550D713A841F683FA67" box="[1051,1254,1574,1598]" pageId="1" pageNumber="554" value="2012-05-22" valueMax="2012-06-14" valueMin="2012-05-22">22.V.–14.VI.2012</date>
|
||
, P. Cechovsky.
|
||
</paragraph>
|
||
</subSubSection>
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<subSubSection id="C32033925A697554D3CFA82EF291FCA1" lastPageId="5" lastPageNumber="558" pageId="1" pageNumber="554" type="discussion">
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<paragraph id="8B8560195A697550D3CFA82EF62EFAAB" blockId="1.[151,1437,1429,2033]" pageId="1" pageNumber="554">
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<emphasis id="B94EBC0B5A697550D3CFA82EF351FA3B" bold="true" box="[199,308,1609,1634]" pageId="1" pageNumber="554">Remarks</emphasis>
|
||
. This taxon was originally described as a subspecies of
|
||
<taxonomicName id="4C3A1B9A5A697550D0BAA82DF643FA38" box="[946,1062,1609,1633]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="hidakai">
|
||
<emphasis id="B94EBC0B5A697550D0BAA82DF643FA38" box="[946,1062,1609,1633]" italics="true" pageId="1" pageNumber="554">E. hidakai</emphasis>
|
||
</taxonomicName>
|
||
, based on the similarity between them and overall lack of material. However in the new material we have examined, female specimens of both taxa were collected from the same area in Kelantan,
|
||
<collectingCountry id="F32D20895A697550D1C8A8F6F14FFAF3" box="[704,810,1681,1706]" name="Malaysia" pageId="1" pageNumber="554">Malaysia</collectingCountry>
|
||
. An examination of female genitalia shows significant differences between the two taxa in length of spermathecal duct, size of hemisternite, and length of stylus. This sympatric fact strongly supports
|
||
<taxonomicName id="4C3A1B9A5A697550D10EA8BEF152FAA8" box="[518,823,1753,1777]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="borneoensis">
|
||
<emphasis id="B94EBC0B5A697550D10EA8BEF152FAA8" box="[518,823,1753,1777]" italics="true" pageId="1" pageNumber="554">Echinoaesalus borneoensis</emphasis>
|
||
</taxonomicName>
|
||
to be a separate species.
|
||
</paragraph>
|
||
<paragraph id="8B8560195A697555D3CFA89AF6C7FB56" blockId="1.[151,1437,1429,2033]" lastBlockId="4.[151,1437,1567,1807]" lastPageId="4" lastPageNumber="557" pageId="1" pageNumber="554">
|
||
The following differences are found between
|
||
<taxonomicName id="4C3A1B9A5A697550D1E9A899F1E9FB4C" box="[737,908,1789,1813]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="borneoensis">
|
||
<emphasis id="B94EBC0B5A697550D1E9A899F1E9FB4C" box="[737,908,1789,1813]" italics="true" pageId="1" pageNumber="554">E. borneoensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B94EBC0B5A697550D0C4A899F62FFB4C" box="[972,1098,1789,1813]" italics="true" pageId="1" pageNumber="554">
|
||
<taxonomicName id="4C3A1B9A5A697550D0C4A899F620FB4C" box="[972,1093,1789,1813]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="hidakai">E. hidakai</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
based on an examination of additional specimens: 1) mentum in both sexes more transverse in
|
||
<taxonomicName id="4C3A1B9A5A697550D084A945F651FB60" box="[908,1076,1825,1849]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="borneoensis">
|
||
<emphasis id="B94EBC0B5A697550D084A945F651FB60" box="[908,1076,1825,1849]" italics="true" pageId="1" pageNumber="554">E. borneoensis</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<emphasis id="B94EBC0B5A697550D79AA945F768FB60" box="[1170,1293,1825,1849]" italics="true" pageId="1" pageNumber="554">
|
||
<taxonomicName id="4C3A1B9A5A697550D79AA945F76CFB60" box="[1170,1289,1825,1849]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="hidakai">E. hidakai</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
with clearly defined round pits in
|
||
<taxonomicName id="4C3A1B9A5A697550D284A921F056FB04" box="[396,563,1861,1885]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="borneoensis">
|
||
<emphasis id="B94EBC0B5A697550D284A921F056FB04" box="[396,563,1861,1885]" italics="true" pageId="1" pageNumber="554">E. borneoensis</emphasis>
|
||
</taxonomicName>
|
||
but with elongate and connected pits in
|
||
<taxonomicName id="4C3A1B9A5A697550D70AA921F613FB04" box="[1026,1142,1861,1885]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="hidakai">
|
||
<emphasis id="B94EBC0B5A697550D70AA921F613FB04" box="[1026,1142,1861,1885]" italics="true" pageId="1" pageNumber="554">E. hidakai</emphasis>
|
||
</taxonomicName>
|
||
; 2) pronotal plate of both sexes shorter in
|
||
<taxonomicName id="4C3A1B9A5A697550D250A90AF067FBDD" box="[344,514,1900,1924]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="borneoensis">
|
||
<emphasis id="B94EBC0B5A697550D250A90AF067FBDD" box="[344,514,1900,1924]" italics="true" pageId="1" pageNumber="554">E. borneoensis</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<emphasis id="B94EBC0B5A697550D161A90AF082FBDD" box="[617,743,1900,1924]" italics="true" pageId="1" pageNumber="554">
|
||
<taxonomicName id="4C3A1B9A5A697550D161A90AF086FBDD" box="[617,739,1900,1924]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="hidakai">E. hidakai</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
more evident in female; 3) 9th abdominal segment of male shorter in
|
||
<taxonomicName id="4C3A1B9A5A697550D201A9F6F3CAFBF1" box="[265,431,1936,1960]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="borneoensis">
|
||
<emphasis id="B94EBC0B5A697550D201A9F6F3CAFBF1" box="[265,431,1936,1960]" italics="true" pageId="1" pageNumber="554">E. borneoensis</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<taxonomicName id="4C3A1B9A5A697550D104A9F6F0E5FBF1" box="[524,640,1936,1960]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="hidakai">
|
||
<emphasis id="B94EBC0B5A697550D104A9F6F0E5FBF1" box="[524,640,1936,1960]" italics="true" pageId="1" pageNumber="554">E. hidakai</emphasis>
|
||
</taxonomicName>
|
||
; 4) median lobe of parameres markedly thinner in
|
||
<taxonomicName id="4C3A1B9A5A697550D7B5A9F6F701FBF1" box="[1213,1380,1936,1960]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="borneoensis">
|
||
<emphasis id="B94EBC0B5A697550D7B5A9F6F701FBF1" box="[1213,1380,1936,1960]" italics="true" pageId="1" pageNumber="554">E. borneoensis</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<emphasis id="B94EBC0B5A697550D3BCA9D2F34AFB95" box="[180,303,1972,1996]" italics="true" pageId="1" pageNumber="554">
|
||
<taxonomicName id="4C3A1B9A5A697550D3BCA9D2F34EFB95" box="[180,299,1972,1996]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="hidakai">E. hidakai</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
completely pigmented in
|
||
<taxonomicName id="4C3A1B9A5A697550D15FA9D2F0AEFB95" box="[599,715,1972,1996]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="hidakai">
|
||
<emphasis id="B94EBC0B5A697550D15FA9D2F0AEFB95" box="[599,715,1972,1996]" italics="true" pageId="1" pageNumber="554">E. hidakai</emphasis>
|
||
</taxonomicName>
|
||
but with lateral colorless splits in
|
||
<taxonomicName id="4C3A1B9A5A697550D75BA9D2F69FFB95" box="[1107,1274,1972,1996]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="borneoensis">
|
||
<emphasis id="B94EBC0B5A697550D75BA9D2F69FFB95" box="[1107,1274,1972,1996]" italics="true" pageId="1" pageNumber="554">E. borneoensis</emphasis>
|
||
</taxonomicName>
|
||
; 5) parameres markedly shorter in
|
||
<taxonomicName id="4C3A1B9A5A697550D271A9BEF07AFBA9" box="[377,543,2008,2032]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="borneoensis">
|
||
<emphasis id="B94EBC0B5A697550D271A9BEF07AFBA9" box="[377,543,2008,2032]" italics="true" pageId="1" pageNumber="554">E. borneoensis</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<taxonomicName id="4C3A1B9A5A697550D172A9BEF088FBA9" box="[634,749,2008,2032]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="hidakai">
|
||
<emphasis id="B94EBC0B5A697550D172A9BEF088FBA9" box="[634,749,2008,2032]" italics="true" pageId="1" pageNumber="554">E. hidakai</emphasis>
|
||
</taxonomicName>
|
||
; 6) hemisternite of female genitalia smaller with shorter
|
||
<emphasis id="B94EBC0B5A6C7555D39FA8FBF332FAEB" bold="true" box="[151,343,1692,1715]" pageId="4" pageNumber="557">FIGURES 77–89.</emphasis>
|
||
9th abdominal segment of male in ventral view under same scale (scale bar
|
||
<quantity id="4CC2CDFC5A6C7555D78FA8FBF6AFFAEB" box="[1159,1226,1692,1714]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="557" unit="mm" value="1.0">1 mm</quantity>
|
||
). 77,
|
||
<taxonomicName id="4C3A1B9A5A6C7555D600A8FAF286FA89" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="hidakai">
|
||
<emphasis id="B94EBC0B5A6C7555D600A8FAF286FA89" italics="true" pageId="4" pageNumber="557">Echinoaesalus hidakai</emphasis>
|
||
</taxonomicName>
|
||
, L12, western
|
||
<collectingCountry id="F32D20895A6C7555D270A8DCF3BDFA88" box="[376,472,1723,1745]" name="Malaysia" pageId="4" pageNumber="557">Malaysia</collectingCountry>
|
||
; 78,
|
||
<taxonomicName id="4C3A1B9A5A6C7555D10FA8DBF0F8FA89" box="[519,669,1723,1745]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="borneoensis">
|
||
<emphasis id="B94EBC0B5A6C7555D10FA8DBF0F8FA89" box="[519,669,1723,1745]" italics="true" pageId="4" pageNumber="557">E. borneoensis</emphasis>
|
||
</taxonomicName>
|
||
, L11, Borneo; 79,
|
||
<taxonomicName id="4C3A1B9A5A6C7555D052A8DBF195FA89" box="[858,1008,1723,1745]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="borneoensis">
|
||
<emphasis id="B94EBC0B5A6C7555D052A8DBF195FA89" box="[858,1008,1723,1745]" italics="true" pageId="4" pageNumber="557">E. borneoensis</emphasis>
|
||
</taxonomicName>
|
||
, L01, Borneo; 80–86,
|
||
<taxonomicName id="4C3A1B9A5A6C7555D7DCA8DBF75FFA89" box="[1236,1338,1723,1745]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="timidus">
|
||
<emphasis id="B94EBC0B5A6C7555D7DCA8DBF75FFA89" box="[1236,1338,1723,1745]" italics="true" pageId="4" pageNumber="557">E. timidus</emphasis>
|
||
</taxonomicName>
|
||
; 80, L08; 81, L02; 82, L17; 83, L23; 84, L21; 85, L22; 86, L18; 87,
|
||
<taxonomicName id="4C3A1B9A5A6C7555D1F2A8BCF1E3FAB6" box="[762,902,1754,1776]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="sabahensis">
|
||
<emphasis id="B94EBC0B5A6C7555D1F2A8BCF1E3FAB6" box="[762,902,1754,1776]" italics="true" pageId="4" pageNumber="557">E. sabahensis</emphasis>
|
||
</taxonomicName>
|
||
, L14, Borneo; 88,
|
||
<taxonomicName id="4C3A1B9A5A6C7555D745A8BCF6BDFAB6" box="[1101,1240,1754,1776]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="cechovskyi">
|
||
<emphasis id="B94EBC0B5A6C7555D745A8BCF6BDFAB6" box="[1101,1240,1754,1776]" italics="true" pageId="4" pageNumber="557">E. cechovskyi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="5481DEBB5A6C7555D7EEA8BDF724FAA9" box="[1254,1345,1754,1776]" pageId="4" pageNumber="557" type="holotype">holotype</typeStatus>
|
||
, western
|
||
<collectingCountry id="F32D20895A6C7555D39FA89EF292FB56" box="[151,247,1785,1807]" name="Malaysia" pageId="4" pageNumber="557">Malaysia</collectingCountry>
|
||
; 89,
|
||
<taxonomicName id="4C3A1B9A5A6C7555D22FA89DF3C3FB57" box="[295,422,1785,1807]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="gedeensis">
|
||
<emphasis id="B94EBC0B5A6C7555D22FA89DF3C3FB57" box="[295,422,1785,1807]" italics="true" pageId="4" pageNumber="557">E. gedeensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="5481DEBB5A6C7555D2BAA89EF068FB56" box="[434,525,1785,1807]" pageId="4" pageNumber="557" type="holotype">holotype</typeStatus>
|
||
, Java. Arrow directed to pigmented terminal end of ventral plate.
|
||
</paragraph>
|
||
<caption id="DF4530915A6A7553D39FA8B1F62CFB73" httpUri="https://zenodo.org/record/241184/files/figure.png" pageId="2" pageNumber="555" targetBox="[183,1395,205,1699]" targetPageId="2">
|
||
<paragraph id="8B8560195A6A7553D39FA8B1F62CFB73" blockId="2.[151,1437,1750,2018]" pageId="2" pageNumber="555">
|
||
<emphasis id="B94EBC0B5A6A7553D39FA8B1F359FAB5" bold="true" box="[151,316,1750,1773]" pageId="2" pageNumber="555">FIGURES 1–7.</emphasis>
|
||
Antenna. 1,
|
||
<taxonomicName id="4C3A1B9A5A6A7553D2CEA8B0F0CAFAB5" box="[454,687,1751,1772]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="hidakai">
|
||
<emphasis id="B94EBC0B5A6A7553D2CEA8B0F0CAFAB5" box="[454,687,1751,1772]" italics="true" pageId="2" pageNumber="555">Echinoaesalus hidakai</emphasis>
|
||
</taxonomicName>
|
||
, L12, western Malaysia; 2,
|
||
<taxonomicName id="4C3A1B9A5A6A7553D0DEA8B0F60AFAB5" box="[982,1135,1751,1772]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="borneoensis">
|
||
<emphasis id="B94EBC0B5A6A7553D0DEA8B0F60AFAB5" box="[982,1135,1751,1772]" italics="true" pageId="2" pageNumber="555">E. borneoensis</emphasis>
|
||
</taxonomicName>
|
||
, L11, Borneo; 3,
|
||
<taxonomicName id="4C3A1B9A5A6A7553D624A8B0F7F3FAB5" box="[1324,1430,1751,1772]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="timidus">
|
||
<emphasis id="B94EBC0B5A6A7553D624A8B0F7F3FAB5" box="[1324,1430,1751,1772]" italics="true" pageId="2" pageNumber="555">E. timidus</emphasis>
|
||
</taxonomicName>
|
||
, L19, western Malaysia; 4,
|
||
<taxonomicName id="4C3A1B9A5A6A7553D2A7A891F059FB53" box="[431,572,1781,1803]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="sabahensis">
|
||
<emphasis id="B94EBC0B5A6A7553D2A7A891F059FB53" box="[431,572,1781,1803]" italics="true" pageId="2" pageNumber="555">E. sabahensis</emphasis>
|
||
</taxonomicName>
|
||
, L14, Borneo; 5,
|
||
<taxonomicName id="4C3A1B9A5A6A7553D1F2A891F11FFB53" box="[762,890,1781,1803]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="gedeensis">
|
||
<emphasis id="B94EBC0B5A6A7553D1F2A891F11FFB53" box="[762,890,1781,1803]" italics="true" pageId="2" pageNumber="555">E. gedeensis</emphasis>
|
||
</taxonomicName>
|
||
, holotype, Java; 6,
|
||
<emphasis id="B94EBC0B5A6A7553D742A891F6B8FB53" box="[1098,1245,1781,1803]" italics="true" pageId="2" pageNumber="555">
|
||
<taxonomicName id="4C3A1B9A5A6A7553D742A891F6BCFB53" box="[1098,1241,1781,1803]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="cechovskyi">E. cechovskyi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
holotype, western Malaysia; 7,
|
||
<taxonomicName id="4C3A1B9A5A6A7553D212A972F3C6FB70" box="[282,419,1812,1834]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="cechovskyi">
|
||
<emphasis id="B94EBC0B5A6A7553D212A972F3C6FB70" box="[282,419,1812,1834]" italics="true" pageId="2" pageNumber="555">E. cechovskyi</emphasis>
|
||
</taxonomicName>
|
||
, paratype, Singapore. Arrow directed to end of third antennomere.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4530915A6A7553D39FA955F170FBBB" pageId="2" pageNumber="555">
|
||
<paragraph id="8B8560195A6A7553D39FA955F170FBBB" blockId="2.[151,1437,1750,2018]" pageId="2" pageNumber="555">
|
||
<emphasis id="B94EBC0B5A6A7553D39FA955F32EFB11" bold="true" box="[151,331,1842,1865]" pageId="2" pageNumber="555">FIGURES 8–30.</emphasis>
|
||
Mentum under same scale (scale bar 0.5 mm). 8,
|
||
<taxonomicName id="4C3A1B9A5A6A7553D065A954F632FB11" box="[877,1111,1843,1864]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="hidakai">
|
||
<emphasis id="B94EBC0B5A6A7553D065A954F632FB11" box="[877,1111,1843,1864]" italics="true" pageId="2" pageNumber="555">Echinoaesalus hidakai</emphasis>
|
||
</taxonomicName>
|
||
, L12, western Malaysia; 9,
|
||
<taxonomicName id="4C3A1B9A5A6A7553D68EA953F374FB3E" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="borneoensis">
|
||
<emphasis id="B94EBC0B5A6A7553D68EA953F374FB3E" italics="true" pageId="2" pageNumber="555">E. borneoensis</emphasis>
|
||
</taxonomicName>
|
||
, L01, Borneo; 10,
|
||
<taxonomicName id="4C3A1B9A5A6A7553D2D1A935F027FB3E" box="[473,578,1874,1895]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="hidakai">
|
||
<emphasis id="B94EBC0B5A6A7553D2D1A935F027FB3E" box="[473,578,1874,1895]" italics="true" pageId="2" pageNumber="555">E. hidakai</emphasis>
|
||
</taxonomicName>
|
||
, L10, western Malaysia; 11,
|
||
<taxonomicName id="4C3A1B9A5A6A7553D07AA935F66EFB3E" box="[882,1035,1874,1895]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="borneoensis">
|
||
<emphasis id="B94EBC0B5A6A7553D07AA935F66EFB3E" box="[882,1035,1874,1895]" italics="true" pageId="2" pageNumber="555">E. borneoensis</emphasis>
|
||
</taxonomicName>
|
||
, L07, Borneo; 12,
|
||
<taxonomicName id="4C3A1B9A5A6A7553D7DAA935F73AFB3E" box="[1234,1375,1874,1895]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="sabahensis">
|
||
<emphasis id="B94EBC0B5A6A7553D7DAA935F73AFB3E" box="[1234,1375,1874,1895]" italics="true" pageId="2" pageNumber="555">E. sabahensis</emphasis>
|
||
</taxonomicName>
|
||
, after Krikken (2008), Borneo; 13,
|
||
<taxonomicName id="4C3A1B9A5A6A7553D2CEA916F031FBDC" box="[454,596,1904,1926]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="sabahensis">
|
||
<emphasis id="B94EBC0B5A6A7553D2CEA916F031FBDC" box="[454,596,1904,1926]" italics="true" pageId="2" pageNumber="555">E. sabahensis</emphasis>
|
||
</taxonomicName>
|
||
, L14, Borneo; 14,
|
||
<emphasis id="B94EBC0B5A6A7553D014A916F1CAFBDC" box="[796,943,1904,1926]" italics="true" pageId="2" pageNumber="555">
|
||
<taxonomicName id="4C3A1B9A5A6A7553D014A916F1CEFBDC" box="[796,939,1904,1926]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="cechovskyi">E. cechovskyi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
holotype, western Malaysia; 15,
|
||
<emphasis id="B94EBC0B5A6A7553D602A916F7F9FBDC" box="[1290,1436,1904,1926]" italics="true" pageId="2" pageNumber="555">
|
||
<taxonomicName id="4C3A1B9A5A6A7553D602A916F7FDFBDC" box="[1290,1432,1904,1926]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="cechovskyi">E. cechovskyi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
paratype, Singapore; 16–18,
|
||
<taxonomicName id="4C3A1B9A5A6A7553D2B3A9E8F047FBFD" box="[443,546,1935,1956]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="timidus">
|
||
<emphasis id="B94EBC0B5A6A7553D2B3A9E8F047FBFD" box="[443,546,1935,1956]" italics="true" pageId="2" pageNumber="555">E. timidus</emphasis>
|
||
</taxonomicName>
|
||
; 16, L21; 17, L22; 18, L18; 19,
|
||
<taxonomicName id="4C3A1B9A5A6A7553D062A9E8F1B0FBFD" box="[874,981,1935,1956]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="javanus">
|
||
<emphasis id="B94EBC0B5A6A7553D062A9E8F1B0FBFD" box="[874,981,1935,1956]" italics="true" pageId="2" pageNumber="555">E. javanus</emphasis>
|
||
</taxonomicName>
|
||
, after Krikken (2008), holotype, Java; 20,
|
||
<taxonomicName id="4C3A1B9A5A6A7553D68EA9E8F29FFB9B" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="gedeensis">
|
||
<emphasis id="B94EBC0B5A6A7553D68EA9E8F29FFB9B" italics="true" pageId="2" pageNumber="555">E. gedeensis</emphasis>
|
||
</taxonomicName>
|
||
, holotype, Java; 21,
|
||
<taxonomicName id="4C3A1B9A5A6A7553D2C5A9C9F050FB9B" box="[461,565,1965,1987]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="timidus">
|
||
<emphasis id="B94EBC0B5A6A7553D2C5A9C9F050FB9B" box="[461,565,1965,1987]" italics="true" pageId="2" pageNumber="555">E. timidus</emphasis>
|
||
</taxonomicName>
|
||
, after Krikken (2008), holotype, probably male; 22–30,
|
||
<taxonomicName id="4C3A1B9A5A6A7553D77CA9C9F6BEFB9B" box="[1140,1243,1965,1987]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="timidus">
|
||
<emphasis id="B94EBC0B5A6A7553D77CA9C9F6BEFB9B" box="[1140,1243,1965,1987]" italics="true" pageId="2" pageNumber="555">E. timidus</emphasis>
|
||
</taxonomicName>
|
||
; 22, L08; 23, L02; 24, L17; 25, L23; 26, L09; 27, L06; 28, L20; 29, L13; 30, L29.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4530915A6B7552D39FA886F3D2FB6C" pageId="3" pageNumber="556">
|
||
<paragraph id="8B8560195A6B7552D39FA886F3D2FB6C" blockId="3.[151,1436,1761,2029]" pageId="3" pageNumber="556">
|
||
<emphasis id="B94EBC0B5A6B7552D39FA886F33EFAAE" bold="true" box="[151,347,1761,1784]" pageId="3" pageNumber="556">FIGURES 31–34.</emphasis>
|
||
Labium (left half, with ligula) and maxilla (right half) under same scale (scale bar 0.2 mm). 31,
|
||
<taxonomicName id="4C3A1B9A5A6B7552D39FA966F318FB4F" box="[151,381,1793,1814]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="556" phylum="Arthropoda" rank="species" species="hidakai">
|
||
<emphasis id="B94EBC0B5A6B7552D39FA966F318FB4F" box="[151,381,1793,1814]" italics="true" pageId="3" pageNumber="556">Echinoaesalus hidakai</emphasis>
|
||
</taxonomicName>
|
||
, L12, western Malaysia; 32,
|
||
<taxonomicName id="4C3A1B9A5A6B7552D1AEA965F16AFB4F" box="[678,783,1793,1815]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="556" phylum="Arthropoda" rank="species" species="hidakai">
|
||
<emphasis id="B94EBC0B5A6B7552D1AEA965F16AFB4F" box="[678,783,1793,1815]" italics="true" pageId="3" pageNumber="556">E. hidakai</emphasis>
|
||
</taxonomicName>
|
||
, L10, western Malaysia; 33,
|
||
<taxonomicName id="4C3A1B9A5A6B7552D730A965F6C5FB4F" box="[1080,1184,1793,1815]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="556" phylum="Arthropoda" rank="species" species="timidus">
|
||
<emphasis id="B94EBC0B5A6B7552D730A965F6C5FB4F" box="[1080,1184,1793,1815]" italics="true" pageId="3" pageNumber="556">E. timidus</emphasis>
|
||
</taxonomicName>
|
||
, L17, western Malaysia; 34,
|
||
<taxonomicName id="4C3A1B9A5A6B7552D3B5A947F346FB6C" box="[189,291,1824,1845]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="556" phylum="Arthropoda" rank="species" species="timidus">
|
||
<emphasis id="B94EBC0B5A6B7552D3B5A947F346FB6C" box="[189,291,1824,1845]" italics="true" pageId="3" pageNumber="556">E. timidus</emphasis>
|
||
</taxonomicName>
|
||
, L06, Borneo.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4530915A6B7552D39FA95AF069FBE9" httpUri="https://zenodo.org/record/241185/files/figure.png" pageId="3" pageNumber="556" targetBox="[202,1395,193,1742]" targetPageId="3">
|
||
<paragraph id="8B8560195A6B7552D39FA95AF069FBE9" blockId="3.[151,1436,1761,2029]" pageId="3" pageNumber="556">
|
||
<emphasis id="B94EBC0B5A6B7552D39FA95AF330FB0A" bold="true" box="[151,341,1853,1876]" pageId="3" pageNumber="556">FIGURES 35–52.</emphasis>
|
||
Protibia (left), mesotibia (middle), and metatibia (right), or just protibia. 35,
|
||
<taxonomicName id="4C3A1B9A5A6B7552D77EA959F73BFB0A" box="[1142,1374,1854,1875]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="556" phylum="Arthropoda" rank="species" species="hidakai">
|
||
<emphasis id="B94EBC0B5A6B7552D77EA959F73BFB0A" box="[1142,1374,1854,1875]" italics="true" pageId="3" pageNumber="556">Echinoaesalus hidakai</emphasis>
|
||
</taxonomicName>
|
||
, L12; 36,
|
||
<taxonomicName id="4C3A1B9A5A6B7552D3B5A939F343FB2B" box="[189,294,1885,1907]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="556" phylum="Arthropoda" rank="species" species="hidakai">
|
||
<emphasis id="B94EBC0B5A6B7552D3B5A939F343FB2B" box="[189,294,1885,1907]" italics="true" pageId="3" pageNumber="556">E. hidakai</emphasis>
|
||
</taxonomicName>
|
||
, L10; 37,
|
||
<taxonomicName id="4C3A1B9A5A6B7552D298A939F042FB2B" box="[400,551,1885,1907]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="556" phylum="Arthropoda" rank="species" species="borneoensis">
|
||
<emphasis id="B94EBC0B5A6B7552D298A939F042FB2B" box="[400,551,1885,1907]" italics="true" pageId="3" pageNumber="556">E. borneoensis</emphasis>
|
||
</taxonomicName>
|
||
, L11; 38,
|
||
<taxonomicName id="4C3A1B9A5A6B7552D199A939F14DFB2B" box="[657,808,1885,1907]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="556" phylum="Arthropoda" rank="species" species="borneoensis">
|
||
<emphasis id="B94EBC0B5A6B7552D199A939F14DFB2B" box="[657,808,1885,1907]" italics="true" pageId="3" pageNumber="556">E. borneoensis</emphasis>
|
||
</taxonomicName>
|
||
, L07; 39–48,
|
||
<taxonomicName id="4C3A1B9A5A6B7552D0B1A939F645FB2B" box="[953,1056,1885,1907]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="556" phylum="Arthropoda" rank="species" species="timidus">
|
||
<emphasis id="B94EBC0B5A6B7552D0B1A939F645FB2B" box="[953,1056,1885,1907]" italics="true" pageId="3" pageNumber="556">E. timidus</emphasis>
|
||
</taxonomicName>
|
||
; 39, L17; 40, L13; 41, L08; 42, L02; 43, L23; 44, L21; 45, L22; 46, L18; 47, L09; 48, L06; 49,
|
||
<taxonomicName id="4C3A1B9A5A6B7552D1E7A91BF11FFBC8" box="[751,890,1916,1937]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="556" phylum="Arthropoda" rank="species" species="sabahensis">
|
||
<emphasis id="B94EBC0B5A6B7552D1E7A91BF11FFBC8" box="[751,890,1916,1937]" italics="true" pageId="3" pageNumber="556">E. sabahensis</emphasis>
|
||
</taxonomicName>
|
||
, L14; 50,
|
||
<emphasis id="B94EBC0B5A6B7552D0EEA91BF612FBC8" box="[998,1143,1916,1937]" italics="true" pageId="3" pageNumber="556">
|
||
<taxonomicName id="4C3A1B9A5A6B7552D0EEA91BF616FBC8" box="[998,1139,1916,1937]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="556" phylum="Arthropoda" rank="species" species="cechovskyi">E. cechovskyi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
holotype; 51,
|
||
<emphasis id="B94EBC0B5A6B7552D603A91BF7F9FBC8" box="[1291,1436,1916,1937]" italics="true" pageId="3" pageNumber="556">
|
||
<taxonomicName id="4C3A1B9A5A6B7552D603A91BF7FDFBC8" box="[1291,1432,1916,1937]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="556" phylum="Arthropoda" rank="species" species="cechovskyi">E. cechovskyi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
paratype; 52,
|
||
<taxonomicName id="4C3A1B9A5A6B7552D229A9FCF3FAFBE9" box="[289,415,1947,1968]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="556" phylum="Arthropoda" rank="species" species="gedeensis">
|
||
<emphasis id="B94EBC0B5A6B7552D229A9FCF3FAFBE9" box="[289,415,1947,1968]" italics="true" pageId="3" pageNumber="556">E. gedeensis</emphasis>
|
||
</taxonomicName>
|
||
, holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4530915A6B7552D39FA9DFF667FBB4" pageId="3" pageNumber="556">
|
||
<paragraph id="8B8560195A6B7552D39FA9DFF667FBB4" blockId="3.[151,1436,1761,2029]" pageId="3" pageNumber="556">
|
||
<emphasis id="B94EBC0B5A6B7552D39FA9DFF336FB97" bold="true" box="[151,339,1976,1999]" pageId="3" pageNumber="556">FIGURES 53–56.</emphasis>
|
||
Hindwing under same scale (scale bar 1 mm). 53,
|
||
<taxonomicName id="4C3A1B9A5A6B7552D047A9DEF651FB97" box="[847,1076,1977,1998]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="556" phylum="Arthropoda" rank="species" species="timidus">
|
||
<emphasis id="B94EBC0B5A6B7552D047A9DEF651FB97" box="[847,1076,1977,1998]" italics="true" pageId="3" pageNumber="556">Echinoaesalus timidus</emphasis>
|
||
</taxonomicName>
|
||
, L08, Sumatra; 54,
|
||
<taxonomicName id="4C3A1B9A5A6B7552D7F3A9DDF704FB97" box="[1275,1377,1977,1999]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="556" phylum="Arthropoda" rank="species" species="timidus">
|
||
<emphasis id="B94EBC0B5A6B7552D7F3A9DDF704FB97" box="[1275,1377,1977,1999]" italics="true" pageId="3" pageNumber="556">E. timidus</emphasis>
|
||
</taxonomicName>
|
||
, L06, Borneo; 55,
|
||
<taxonomicName id="4C3A1B9A5A6B7552D21CA9BFF3CEFBB4" box="[276,427,2008,2029]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="556" phylum="Arthropoda" rank="species" species="borneoensis">
|
||
<emphasis id="B94EBC0B5A6B7552D21CA9BFF3CEFBB4" box="[276,427,2008,2029]" italics="true" pageId="3" pageNumber="556">E. borneoensis</emphasis>
|
||
</taxonomicName>
|
||
, paratype, Borneo; 56,
|
||
<emphasis id="B94EBC0B5A6B7552D191A9BFF14FFBB4" box="[665,810,2008,2029]" italics="true" pageId="3" pageNumber="556">
|
||
<taxonomicName id="4C3A1B9A5A6B7552D191A9BFF140FBB4" box="[665,805,2008,2029]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="556" phylum="Arthropoda" rank="species" species="cechovskyi">E. cechovskyi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
paratype, Singapore.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4530915A6C7555D39FA878F604FAC8" httpUri="https://zenodo.org/record/241186/files/figure.png" pageId="4" pageNumber="557" targetBox="[151,1436,203,1545]" targetPageId="4">
|
||
<paragraph id="8B8560195A6C7555D39FA878F604FAC8" blockId="4.[151,1437,1567,1807]" pageId="4" pageNumber="557">
|
||
<emphasis id="B94EBC0B5A6C7555D39FA878F33FFA6D" bold="true" box="[151,346,1567,1590]" pageId="4" pageNumber="557">FIGURES 57–76.</emphasis>
|
||
Prosternal process in ventral view. 57,
|
||
<taxonomicName id="4C3A1B9A5A6C7555D019A847F198FA6C" box="[785,1021,1568,1589]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="hidakai">
|
||
<emphasis id="B94EBC0B5A6C7555D019A847F198FA6C" box="[785,1021,1568,1589]" italics="true" pageId="4" pageNumber="557">Echinoaesalus hidakai</emphasis>
|
||
</taxonomicName>
|
||
, L12; 58,
|
||
<taxonomicName id="4C3A1B9A5A6C7555D77FA847F680FA6C" box="[1143,1253,1568,1589]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="hidakai">
|
||
<emphasis id="B94EBC0B5A6C7555D77FA847F680FA6C" box="[1143,1253,1568,1589]" italics="true" pageId="4" pageNumber="557">E. hidakai</emphasis>
|
||
</taxonomicName>
|
||
, L10; 59–62,
|
||
<taxonomicName id="4C3A1B9A5A6C7555D68EA847F374FA0A" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="borneoensis">
|
||
<emphasis id="B94EBC0B5A6C7555D68EA847F374FA0A" italics="true" pageId="4" pageNumber="557">E. borneoensis</emphasis>
|
||
</taxonomicName>
|
||
; 59, L11; 60, L01; 61, L07; 62, L16; 63,
|
||
<taxonomicName id="4C3A1B9A5A6C7555D1BDA858F151FA0A" box="[693,820,1598,1620]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="gedeensis">
|
||
<emphasis id="B94EBC0B5A6C7555D1BDA858F151FA0A" box="[693,820,1598,1620]" italics="true" pageId="4" pageNumber="557">E. gedeensis</emphasis>
|
||
</taxonomicName>
|
||
, holotype; 64–67,
|
||
<taxonomicName id="4C3A1B9A5A6C7555D0FBA858F63FFA0A" box="[1011,1114,1598,1620]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="timidus">
|
||
<emphasis id="B94EBC0B5A6C7555D0FBA858F63FFA0A" box="[1011,1114,1598,1620]" italics="true" pageId="4" pageNumber="557">E. timidus</emphasis>
|
||
</taxonomicName>
|
||
; 64, L08; 65, L02; 66, L17; 67, L21; 68,
|
||
<taxonomicName id="4C3A1B9A5A6C7555D3FBA83AF31BFA2B" box="[243,382,1629,1650]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="sabahensis">
|
||
<emphasis id="B94EBC0B5A6C7555D3FBA83AF31BFA2B" box="[243,382,1629,1650]" italics="true" pageId="4" pageNumber="557">E. sabahensis</emphasis>
|
||
</taxonomicName>
|
||
, L14; 69,
|
||
<taxonomicName id="4C3A1B9A5A6C7555D2EFA83AF015FA2B" box="[487,624,1629,1650]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="cechovskyi">
|
||
<emphasis id="B94EBC0B5A6C7555D2EFA83AF015FA2B" box="[487,624,1629,1650]" italics="true" pageId="4" pageNumber="557">E. cechovskyi</emphasis>
|
||
</taxonomicName>
|
||
, L28; 70–73,
|
||
<taxonomicName id="4C3A1B9A5A6C7555D008A83AF103FA2B" box="[768,870,1629,1650]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="timidus">
|
||
<emphasis id="B94EBC0B5A6C7555D008A83AF103FA2B" box="[768,870,1629,1650]" italics="true" pageId="4" pageNumber="557">E. timidus</emphasis>
|
||
</taxonomicName>
|
||
; 70, L09; 71, L06; 72, L20; 73, L13; 74,
|
||
<taxonomicName id="4C3A1B9A5A6C7555D602A83AF7F0FA2B" box="[1290,1429,1629,1650]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="sabahensis">
|
||
<emphasis id="B94EBC0B5A6C7555D602A83AF7F0FA2B" box="[1290,1429,1629,1650]" italics="true" pageId="4" pageNumber="557">E. sabahensis</emphasis>
|
||
</taxonomicName>
|
||
, after Krikken 2008; 75,
|
||
<emphasis id="B94EBC0B5A6C7555D283A81BF07EFAC9" box="[395,539,1659,1681]" italics="true" pageId="4" pageNumber="557">
|
||
<taxonomicName id="4C3A1B9A5A6C7555D283A81BF072FAC9" box="[395,535,1659,1681]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="cechovskyi">E. cechovskyi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
paratype; 76,
|
||
<emphasis id="B94EBC0B5A6C7555D1A4A81BF178FAC8" box="[684,797,1660,1681]" italics="true" pageId="4" pageNumber="557">
|
||
<taxonomicName id="4C3A1B9A5A6C7555D1A4A81BF17FFAC8" box="[684,794,1660,1681]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="javanus">E. javanus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
after Krikken (2008), holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8560195A6C7555D39FA95CF297FB2E" blockId="4.[151,1436,1851,2020]" pageId="4" pageNumber="557">
|
||
stylus in
|
||
<taxonomicName id="4C3A1B9A5A6C7555D20AA95BF3CEFB0A" box="[258,427,1851,1875]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="borneoensis">
|
||
<emphasis id="B94EBC0B5A6C7555D20AA95BF3CEFB0A" box="[258,427,1851,1875]" italics="true" pageId="4" pageNumber="557">E. borneoensis</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<taxonomicName id="4C3A1B9A5A6C7555D106A95BF0E1FB0A" box="[526,644,1851,1875]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="hidakai">
|
||
<emphasis id="B94EBC0B5A6C7555D106A95BF0E1FB0A" box="[526,644,1851,1875]" italics="true" pageId="4" pageNumber="557">E. hidakai</emphasis>
|
||
</taxonomicName>
|
||
; 7) spermathecal duct in
|
||
<taxonomicName id="4C3A1B9A5A6C7555D0A6A95BF632FB0A" box="[942,1111,1851,1875]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="borneoensis">
|
||
<emphasis id="B94EBC0B5A6C7555D0A6A95BF632FB0A" box="[942,1111,1851,1875]" italics="true" pageId="4" pageNumber="557">E. borneoensis</emphasis>
|
||
</taxonomicName>
|
||
nearly half as long as in
|
||
<taxonomicName id="4C3A1B9A5A6C7555D68CA95BF28EFB2E" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="hidakai">
|
||
<emphasis id="B94EBC0B5A6C7555D68CA95BF28EFB2E" italics="true" pageId="4" pageNumber="557">E. hidakai</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8560195A6C7554D3CFA9E4F291FCA1" blockId="4.[151,1436,1851,2020]" lastBlockId="5.[151,1436,151,464]" lastPageId="5" lastPageNumber="558" pageId="4" pageNumber="557">
|
||
<bibRef id="529AF4EF5A6C7554D3CFA9E4F291FCA1" lastPageId="5" lastPageNumber="558" pageId="4" pageNumber="557" type="book">
|
||
<author id="538788FC5A6C7555D3CFA9E4F3E4FBC2" box="[199,385,1923,1948]" fns="I" npo="LF" pageId="4" pageNumber="557">
|
||
Nevertheless,
|
||
<emphasis id="B94EBC0B5A6C7555D261A9E3F3E4FBC2" box="[361,385,1924,1947]" italics="true" pageId="4" pageNumber="557">E.</emphasis>
|
||
</author>
|
||
<title id="F93E8B2F5A6C7555D284A9E4F328FBBD" jopScore="1.2138507" pageId="4" pageNumber="557">
|
||
borneoensis and
|
||
<emphasis id="B94EBC0B5A6C7555D158A9E3F0A2FBC2" box="[592,711,1923,1947]" italics="true" pageId="4" pageNumber="557">E. hidakai</emphasis>
|
||
are very similar and share nearly all the remaining characters other than the characters above. We did not find any differences in labium, maxilla, antenna, and hindwing between the two species
|
||
</title>
|
||
. The sexual dimorphism of
|
||
<emphasis id="B94EBC0B5A6C7555D19DA9ABF15AFBBA" box="[661,831,1995,2019]" italics="true" pageId="4" pageNumber="557">E. borneoensis</emphasis>
|
||
is exactly the same as that of
|
||
<emphasis id="B94EBC0B5A6C7555D7A2A9ABF745FBBA" box="[1194,1312,1995,2019]" italics="true" pageId="4" pageNumber="557">E. hidakai</emphasis>
|
||
. It is very possible that all species of the
|
||
<emphasis id="B94EBC0B5A6D7554D2F4AEFEF014FCE9" box="[508,625,152,176]" italics="true" pageId="5" pageNumber="558">E. hidakai</emphasis>
|
||
group have the same sexual dimorphism, different from that of the
|
||
<emphasis id="B94EBC0B5A6D7554D68CAEFEF28FFC8D" italics="true" pageId="5" pageNumber="558">E. matsuii</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAB1DE85A6D7554D3F6AEDBF3F4FC8D" author="Araya" box="[254,401,188,212]" pageId="5" pageNumber="558" refString="Araya, K., Kon, M. & Johki, Y. (1993) A new species of the genus Aesalus (Coleoptera, Lucanidae) from the Malay Peninsula. Elytra, 21, 93 - 107." type="journal article" year="1993">
|
||
Araya,
|
||
<year id="FF92214A5A6D7554D250AEDBF3F4FC8D" box="[344,401,188,212]" pageId="5" pageNumber="558">1993</year>
|
||
</bibRefCitation>
|
||
)
|
||
<journalOrPublisher id="51FB6AD55A6D7554D2AEAEDAF295FCA1" jopScore="1.7536052" pageId="5" pageNumber="558" type="_" typeScore="8">group by having no sexual dimorphism in posterior margin of last visible abdominal ventrite</journalOrPublisher>
|
||
.
|
||
</bibRef>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32033925A6D7554D3CFAF63F03BFDD1" pageId="5" pageNumber="558" type="description">
|
||
<paragraph id="8B8560195A6D7554D3CFAF63F06DFD19" blockId="5.[151,1436,151,464]" pageId="5" pageNumber="558">
|
||
<emphasis id="B94EBC0B5A6D7554D3CFAF63F313FD44" bold="true" box="[199,374,260,285]" pageId="5" pageNumber="558">Male genitalia</emphasis>
|
||
(
|
||
<figureCitation id="13017C9C5A6D7554D281AF63F072FD45" box="[393,535,260,285]" captionStart="FIGURES 90 – 102" captionStartId="5.[151,264,1797,1820]" captionTargetBox="[151,1436,519,1775]" captionTargetId="figure@5.[151,1436,519,1775]" captionTargetPageId="5" captionText="FIGURES 90 – 102. Aedeagus in dorsal, left lateral, ventral, and right lateral views under same scale (scale bar 1 mm). 90, Echinoaesalus hidakai, L 12, western Malaysia; 91, E. borneoensis, L 11, Borneo; 92, E. borneoensis, L 01, Borneo; 93 – 98, E. timidus; 93, L 08; 94, L 02; 95, L 17; 96, L 23; 97, L 21; 98, L 22; 99, E. sabahensis, L 14, Borneo; 100, E. timidus, L 18; 101, E. cechovskyi, holotype, western Malaysia; 102, E. gedeensis, holotype, Java. Arrows directed to ends of parameres." httpUri="https://zenodo.org/record/241187/files/figure.png" pageId="5" pageNumber="558">Figs. 91–92</figureCitation>
|
||
). The male genitalia taken from the new material fit the original description (
|
||
<bibRefCitation id="EFAB1DE85A6D7554D396AF4FF30CFD19" author="Huang" box="[158,361,296,320]" pageId="5" pageNumber="558" refString="Huang, H., Imura, Y. & Wu, L. (2011) Description of three new taxa of Echinoaesalus Zelenka from Borneo and Java (Coleoptera, Lucanidae). Kogane, 12, 97 - 107." type="journal article" year="2011">
|
||
Huang
|
||
<emphasis id="B94EBC0B5A6D7554D3FAAF4EF346FD19" box="[242,291,296,320]" italics="true" pageId="5" pageNumber="558">et al</emphasis>
|
||
. 2011
|
||
</bibRefCitation>
|
||
) in all details.
|
||
</paragraph>
|
||
<paragraph id="8B8560195A6D7554D3CFAF2BF03BFDD1" blockId="5.[151,1436,151,464]" pageId="5" pageNumber="558">
|
||
<emphasis id="B94EBC0B5A6D7554D3CFAF2BF3EBFD3C" bold="true" box="[199,398,332,357]" pageId="5" pageNumber="558">Female genitalia</emphasis>
|
||
(
|
||
<figureCitation id="13017C9C5A6D7554D2A8AF2BF02DFD3C" box="[416,584,332,357]" captionStart="FIGURES 103 – 106" captionStartId="6.[151,264,1501,1524]" captionTargetBox="[151,1436,193,1480]" captionTargetId="figure@6.[151,1436,193,1481]" captionTargetPageId="6" captionText="FIGURES 103 – 106. Female genitalia under same scale (scale bar 1 mm). 103, Echinoaesalus hidakai, L 10; 104 – 106, E. borneoensis; 104, L 16; 105, L 07; 106, L 27. Abbreviations: st, stylus; lv, last abdominal ventrite; mo, median oviduct; bd, bursal duct; bc, bursa copulatrix; sd, spermathecal duct; s, spermatheca; sg, spermathecal gland; ag, accessory gland." httpUri="https://zenodo.org/record/241188/files/figure.png" pageId="5" pageNumber="558">Figs. 104–106</figureCitation>
|
||
). Generally as in
|
||
<taxonomicName id="4C3A1B9A5A6D7554D01EAF2AF1E9FD3D" box="[790,908,332,356]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="558" phylum="Arthropoda" rank="species" species="hidakai">
|
||
<emphasis id="B94EBC0B5A6D7554D01EAF2AF1E9FD3D" box="[790,908,332,356]" italics="true" pageId="5" pageNumber="558">E. hidakai</emphasis>
|
||
</taxonomicName>
|
||
, but with smaller hemisternite, shorter stylus, and markedly shorter spermathecal duct.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32033925A6D7554D3CFAFF3F33FFD89" pageId="5" pageNumber="558" type="distribution">
|
||
<paragraph id="8B8560195A6D7554D3CFAFF3F33FFD89" blockId="5.[151,1436,151,464]" pageId="5" pageNumber="558">
|
||
<emphasis id="B94EBC0B5A6D7554D3CFAFF3F33CFDF4" bold="true" box="[199,345,404,429]" pageId="5" pageNumber="558">Distribution</emphasis>
|
||
. Known from Borneo (
|
||
<typeStatus id="5481DEBB5A6D7554D16CAFF3F0F3FDF5" box="[612,662,404,428]" pageId="5" pageNumber="558">type</typeStatus>
|
||
locality) and the Malay Peninsula. Sympatric with
|
||
<taxonomicName id="4C3A1B9A5A6D7554D7D7AFF2F736FDF5" box="[1247,1363,404,428]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="558" phylum="Arthropoda" rank="species" species="hidakai">
|
||
<emphasis id="B94EBC0B5A6D7554D7D7AFF2F736FDF5" box="[1247,1363,404,428]" italics="true" pageId="5" pageNumber="558">E. hidakai</emphasis>
|
||
</taxonomicName>
|
||
in the Malay Peninsula.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |