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<document ID-DOI="10.11646/zootaxa.5250.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="7705657" ID-ZooBank="4729EAF3-8B78-4AAD-B373-9179D008F239" approvalRequired="130" approvalRequired_for_document="1" approvalRequired_for_originalDoi="1" approvalRequired_for_textStreams="123" approvalRequired_for_treatments="5" checkinTime="1678178160677" checkinUser="plazi" docAuthor="WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI &amp; HUANG, JIA" docDate="2023" docId="0393914C2A1E8744C282FCEE6B79FCD6" docLanguage="en" docName="zootaxa.5250.1.1.pdf" docOrigin="Zootaxa 5250 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5250.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.5:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="5" docTitle="Stegana (Steganina) cornuta Li &amp; Chen, sp. nov." docType="treatment" docVersion="3" lastPageNumber="61" masterDocId="FFAAE9342A228778C215FF896958FB43" masterDocTitle="Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae)" masterLastPageNumber="109" masterPageNumber="1" pageNumber="61" updateTime="1678205790159" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>WANG, YA-LIAN</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>LI, BING-XUE</mods:namePart>
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<mods:namePart>CUI, MENG-DI</mods:namePart>
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<mods:namePart>ZHANG, YUAN</mods:namePart>
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<mods:namePart>WANG, LE</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>ZHANG, CAI-HONG</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>TSAUR, SHUN-CHERN</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>CHEN, HONG-WEI</mods:namePart>
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<mods:namePart>HUANG, JIA</mods:namePart>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-03-07</mods:number>
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<mods:number>5250</mods:number>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7705571" ID-Zenodo-Dep="7705571" LSID="urn:lsid:plazi:treatment:0393914C2A1E8744C282FCEE6B79FCD6" httpUri="http://treatment.plazi.org/id/0393914C2A1E8744C282FCEE6B79FCD6" lastPageNumber="61" pageId="60" pageNumber="61">
<subSubSection box="[151,749,870,897]" pageId="60" pageNumber="61" type="nomenclature">
<paragraph blockId="60.[151,749,870,933]" box="[151,749,870,897]" pageId="60" pageNumber="61">
<heading bold="true" box="[151,749,870,897]" fontSize="11" level="1" pageId="60" pageNumber="61" reason="1">
<emphasis bold="true" box="[151,749,870,897]" pageId="60" pageNumber="61">
<taxonomicName authority="Li &amp; Chen" authorityName="Li &amp; Chen" box="[151,642,870,897]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="cornuta" status="sp. nov." subGenus="Steganina">
<emphasis bold="true" box="[151,249,871,897]" italics="true" pageId="60" pageNumber="61">Stegana</emphasis>
(
<emphasis bold="true" box="[266,388,871,897]" italics="true" pageId="60" pageNumber="61">Steganina</emphasis>
)
<emphasis bold="true" box="[405,500,871,897]" italics="true" pageId="60" pageNumber="61">cornuta</emphasis>
Li &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel box="[656,749,871,897]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,355,907,933]" pageId="60" pageNumber="61" type="description">
<paragraph blockId="60.[151,749,870,933]" box="[151,355,907,933]" pageId="60" pageNumber="61">
(
<figureCitation box="[159,306,907,933]" captionStart="FIGURE 7" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[186,1415,182,1919]" captionTargetId="figure-30@30.[173,1415,181,1919]" captionTargetPageId="30" captionText="FIGURE 7. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) cornuta Li &amp; Chen, sp. nov.; C, D. S. (S.) cucullata Li &amp; Chen, sp. nov.; E, F. S. (S.) cultella Cui &amp; Chen, sp. nov.; G, H. S. (S.) curvitabulata Cui &amp; Chen, sp. nov.; I, J. S. (S.) daiya Cui &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705681" httpUri="https://zenodo.org/record/7705681/files/figure.png" pageId="60" pageNumber="61">Figs. 7A, 7B</figureCitation>
,
<figureCitation box="[318,347,907,933]" captionStart="FIGURE 32" captionStartId="61.[151,250,1052,1076]" captionTargetBox="[161,1426,190,1020]" captionTargetId="figure-19@61.[151,1436,181,1028]" captionTargetPageId="61" captionText="FIGURE 32. Stegana (Steganina) cornuta Li &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705744" httpUri="https://zenodo.org/record/7705744/files/figure.png" pageId="60" pageNumber="61">32</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="60" pageNumber="61" type="diagnosis">
<paragraph blockId="60.[151,1437,978,1941]" pageId="60" pageNumber="61">
<emphasis bold="true" box="[151,274,978,1004]" pageId="60" pageNumber="61">Diagnosis.</emphasis>
This species closely resembles
<emphasis box="[627,641,979,1005]" italics="true" pageId="60" pageNumber="61">S</emphasis>
.
<taxonomicName authority="Wu &amp; Chen, 2010" authorityName="Wu &amp; Chen" authorityYear="2010" box="[655,1047,979,1005]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="angulistrata" subGenus="Steganina">
(
<emphasis box="[664,678,979,1005]" italics="true" pageId="60" pageNumber="61">S</emphasis>
.)
<emphasis box="[701,838,979,1005]" italics="true" pageId="60" pageNumber="61">angulistrata</emphasis>
<bibRefCitation author="Wu, L. &amp; Chen, H. W." box="[844,1047,979,1005]" pageId="60" pageNumber="61" pagination="69 - 75" refId="ref74898" refString="Wu, L. &amp; Chen, H. W. (2010) Three new species of Stegana (Steganina) (Diptera: Drosophilidae) from southern China. Oriental Insects, 44, 69 - 75. https: // doi. org / 10.1080 / 00305316.2010.10417607" type="journal article" year="2010">
Wu &amp; Chen,
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</taxonomicName>
in the shape of male terminalia (as fig.
<quantity box="[195,265,1015,1041]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="15.24" metricValueMin="2.54" pageId="60" pageNumber="61" unit="in" value="3.5" valueMax="6.0" valueMin="1.0">16 in</quantity>
<bibRefCitation author="Wu, L. &amp; Chen, H. W." box="[270,466,1015,1041]" pageId="60" pageNumber="61" pagination="69 - 75" refId="ref74898" refString="Wu, L. &amp; Chen, H. W. (2010) Three new species of Stegana (Steganina) (Diptera: Drosophilidae) from southern China. Oriental Insects, 44, 69 - 75. https: // doi. org / 10.1080 / 00305316.2010.10417607" type="journal article" year="2010">Wu &amp; Chen 2010</bibRefCitation>
,
<figureCitation box="[476,557,1015,1041]" captionStart="FIGURE 32" captionStartId="61.[151,250,1052,1076]" captionTargetBox="[161,1426,190,1020]" captionTargetId="figure-19@61.[151,1436,181,1028]" captionTargetPageId="61" captionText="FIGURE 32. Stegana (Steganina) cornuta Li &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705744" httpUri="https://zenodo.org/record/7705744/files/figure.png" pageId="60" pageNumber="61">Fig. 32</figureCitation>
), but can be distinguished from the latter by having the aedeagus medially lacking serrated process but mediolaterally with a pair of horned processes (
<figureCitation box="[966,1104,1051,1077]" captionStart="FIGURE 32" captionStartId="61.[151,250,1052,1076]" captionTargetBox="[161,1426,190,1020]" captionTargetId="figure-19@61.[151,1436,181,1028]" captionTargetPageId="61" captionText="FIGURE 32. Stegana (Steganina) cornuta Li &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705744" httpUri="https://zenodo.org/record/7705744/files/figure.png" pageId="60" pageNumber="61">Fig. 32C, D</figureCitation>
); the aedeagal sheath distally slender and strongly dorsad curved (
<figureCitation box="[558,689,1087,1113]" captionStart="FIGURE 32" captionStartId="61.[151,250,1052,1076]" captionTargetBox="[161,1426,190,1020]" captionTargetId="figure-19@61.[151,1436,181,1028]" captionTargetPageId="61" captionText="FIGURE 32. Stegana (Steganina) cornuta Li &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705744" httpUri="https://zenodo.org/record/7705744/files/figure.png" pageId="60" pageNumber="61">Fig. 32E, F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="60" pageNumber="61" type="description">
<paragraph blockId="60.[151,1437,978,1941]" pageId="60" pageNumber="61">
<emphasis bold="true" box="[199,345,1122,1148]" pageId="60" pageNumber="61">Description.</emphasis>
Male and female: First flagellomere brown. Clypeus yellow. Mesonotum yellow (
<figureCitation box="[1275,1366,1123,1149]" captionStart="FIGURE 7" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[186,1415,182,1919]" captionTargetId="figure-30@30.[173,1415,181,1919]" captionTargetPageId="30" captionText="FIGURE 7. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) cornuta Li &amp; Chen, sp. nov.; C, D. S. (S.) cucullata Li &amp; Chen, sp. nov.; E, F. S. (S.) cultella Cui &amp; Chen, sp. nov.; G, H. S. (S.) curvitabulata Cui &amp; Chen, sp. nov.; I, J. S. (S.) daiya Cui &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705681" httpUri="https://zenodo.org/record/7705681/files/figure.png" pageId="60" pageNumber="61">Fig. 7A</figureCitation>
). Katepisternum yellowish (
<figureCitation box="[408,498,1159,1185]" captionStart="FIGURE 7" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[186,1415,182,1919]" captionTargetId="figure-30@30.[173,1415,181,1919]" captionTargetPageId="30" captionText="FIGURE 7. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) cornuta Li &amp; Chen, sp. nov.; C, D. S. (S.) cucullata Li &amp; Chen, sp. nov.; E, F. S. (S.) cultella Cui &amp; Chen, sp. nov.; G, H. S. (S.) curvitabulata Cui &amp; Chen, sp. nov.; I, J. S. (S.) daiya Cui &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705681" httpUri="https://zenodo.org/record/7705681/files/figure.png" pageId="60" pageNumber="61">Fig. 7B</figureCitation>
). Scutellum yellow (
<figureCitation box="[732,823,1159,1185]" captionStart="FIGURE 7" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[186,1415,182,1919]" captionTargetId="figure-30@30.[173,1415,181,1919]" captionTargetPageId="30" captionText="FIGURE 7. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) cornuta Li &amp; Chen, sp. nov.; C, D. S. (S.) cucullata Li &amp; Chen, sp. nov.; E, F. S. (S.) cultella Cui &amp; Chen, sp. nov.; G, H. S. (S.) curvitabulata Cui &amp; Chen, sp. nov.; I, J. S. (S.) daiya Cui &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705681" httpUri="https://zenodo.org/record/7705681/files/figure.png" pageId="60" pageNumber="61">Fig. 7A</figureCitation>
). Halter mostly grey (
<figureCitation box="[1071,1161,1159,1185]" captionStart="FIGURE 7" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[186,1415,182,1919]" captionTargetId="figure-30@30.[173,1415,181,1919]" captionTargetPageId="30" captionText="FIGURE 7. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) cornuta Li &amp; Chen, sp. nov.; C, D. S. (S.) cucullata Li &amp; Chen, sp. nov.; E, F. S. (S.) cultella Cui &amp; Chen, sp. nov.; G, H. S. (S.) curvitabulata Cui &amp; Chen, sp. nov.; I, J. S. (S.) daiya Cui &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705681" httpUri="https://zenodo.org/record/7705681/files/figure.png" pageId="60" pageNumber="61">Fig. 7B</figureCitation>
). Fore femur with a row of 3 or 4 long setae on distal part of posteroventral surface. Abdominal tergites brown to dark brown (
<figureCitation box="[1299,1421,1195,1221]" captionStart="FIGURE 7" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[186,1415,182,1919]" captionTargetId="figure-30@30.[173,1415,181,1919]" captionTargetPageId="30" captionText="FIGURE 7. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) cornuta Li &amp; Chen, sp. nov.; C, D. S. (S.) cucullata Li &amp; Chen, sp. nov.; E, F. S. (S.) cultella Cui &amp; Chen, sp. nov.; G, H. S. (S.) curvitabulata Cui &amp; Chen, sp. nov.; I, J. S. (S.) daiya Cui &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705681" httpUri="https://zenodo.org/record/7705681/files/figure.png" pageId="60" pageNumber="61">Fig. 7A, B</figureCitation>
); sternites brown, slightly wider than long. Male terminalia (
<figureCitation box="[820,906,1231,1257]" captionStart="FIGURE 32" captionStartId="61.[151,250,1052,1076]" captionTargetBox="[161,1426,190,1020]" captionTargetId="figure-19@61.[151,1436,181,1028]" captionTargetPageId="61" captionText="FIGURE 32. Stegana (Steganina) cornuta Li &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705744" httpUri="https://zenodo.org/record/7705744/files/figure.png" pageId="60" pageNumber="61">Fig. 32</figureCitation>
): Epandrium with
<emphasis box="[1118,1144,1231,1256]" italics="true" pageId="60" pageNumber="61">ca</emphasis>
. 9 setae on dorsal to posterior portions per side (
<figureCitation box="[427,533,1267,1293]" captionStart="FIGURE 32" captionStartId="61.[151,250,1052,1076]" captionTargetBox="[161,1426,190,1020]" captionTargetId="figure-19@61.[151,1436,181,1028]" captionTargetPageId="61" captionText="FIGURE 32. Stegana (Steganina) cornuta Li &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705744" httpUri="https://zenodo.org/record/7705744/files/figure.png" pageId="60" pageNumber="61">Fig. 32A</figureCitation>
). Surstylus with 1 prensiseta, several setulae, and serrated processes near distal margin (
<figureCitation box="[245,380,1303,1329]" captionStart="FIGURE 32" captionStartId="61.[151,250,1052,1076]" captionTargetBox="[161,1426,190,1020]" captionTargetId="figure-19@61.[151,1436,181,1028]" captionTargetPageId="61" captionText="FIGURE 32. Stegana (Steganina) cornuta Li &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705744" httpUri="https://zenodo.org/record/7705744/files/figure.png" pageId="60" pageNumber="61">Fig. 32A, B</figureCitation>
). Cercus lacking pubescence (
<figureCitation box="[719,823,1303,1329]" captionStart="FIGURE 32" captionStartId="61.[151,250,1052,1076]" captionTargetBox="[161,1426,190,1020]" captionTargetId="figure-19@61.[151,1436,181,1028]" captionTargetPageId="61" captionText="FIGURE 32. Stegana (Steganina) cornuta Li &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705744" httpUri="https://zenodo.org/record/7705744/files/figure.png" pageId="60" pageNumber="61">Fig. 32A</figureCitation>
). Hypandrium expanded anteromedially (
<figureCitation box="[1287,1424,1303,1329]" captionStart="FIGURE 32" captionStartId="61.[151,250,1052,1076]" captionTargetBox="[161,1426,190,1020]" captionTargetId="figure-19@61.[151,1436,181,1028]" captionTargetPageId="61" captionText="FIGURE 32. Stegana (Steganina) cornuta Li &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705744" httpUri="https://zenodo.org/record/7705744/files/figure.png" pageId="60" pageNumber="61">Fig. 32C, D</figureCitation>
). Pregonite small, with 2 sensilla (
<figureCitation box="[512,647,1339,1365]" captionStart="FIGURE 32" captionStartId="61.[151,250,1052,1076]" captionTargetBox="[161,1426,190,1020]" captionTargetId="figure-19@61.[151,1436,181,1028]" captionTargetPageId="61" captionText="FIGURE 32. Stegana (Steganina) cornuta Li &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705744" httpUri="https://zenodo.org/record/7705744/files/figure.png" pageId="60" pageNumber="61">Fig. 32C, D</figureCitation>
). Aedeagus rounded distally, with numerous minute, serrated processes laterally and much pubescence distoventrally and sublaterally (
<figureCitation box="[870,1010,1375,1401]" captionStart="FIGURE 32" captionStartId="61.[151,250,1052,1076]" captionTargetBox="[161,1426,190,1020]" captionTargetId="figure-19@61.[151,1436,181,1028]" captionTargetPageId="61" captionText="FIGURE 32. Stegana (Steganina) cornuta Li &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705744" httpUri="https://zenodo.org/record/7705744/files/figure.png" pageId="60" pageNumber="61">Fig. 32C, D</figureCitation>
). Aedeagal sheath with some minute, serrated processes distally (
<figureCitation box="[459,589,1411,1437]" captionStart="FIGURE 32" captionStartId="61.[151,250,1052,1076]" captionTargetBox="[161,1426,190,1020]" captionTargetId="figure-19@61.[151,1436,181,1028]" captionTargetPageId="61" captionText="FIGURE 32. Stegana (Steganina) cornuta Li &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705744" httpUri="https://zenodo.org/record/7705744/files/figure.png" pageId="60" pageNumber="61">Fig. 32E, F</figureCitation>
). Subepandrial sclerite slightly vaulted, split medially (
<figureCitation box="[1208,1313,1411,1437]" captionStart="FIGURE 32" captionStartId="61.[151,250,1052,1076]" captionTargetBox="[161,1426,190,1020]" captionTargetId="figure-19@61.[151,1436,181,1028]" captionTargetPageId="61" captionText="FIGURE 32. Stegana (Steganina) cornuta Li &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705744" httpUri="https://zenodo.org/record/7705744/files/figure.png" pageId="60" pageNumber="61">Fig. 32G</figureCitation>
).
</paragraph>
<paragraph blockId="60.[151,1437,978,1941]" pageId="60" pageNumber="61">
<emphasis bold="true" box="[199,527,1446,1473]" pageId="60" pageNumber="61">Measurements and indices.</emphasis>
BL = 3.00 mm in
<typeStatus box="[744,842,1447,1473]" pageId="60" pageNumber="61">holotype</typeStatus>
(
<quantity box="[858,1003,1447,1473]" metricMagnitude="-2" metricUnit="m" metricValue="6.8326" metricValueMax="7.7216" metricValueMin="5.9436" pageId="60" pageNumber="61" unit="in" value="2.69" valueMax="3.04" valueMin="2.34">2.343.04 in</quantity>
4♁,
<quantity box="[1062,1205,1447,1473]" metricMagnitude="-2" metricUnit="m" metricValue="8.0264" metricValueMax="8.7376" metricValueMin="7.3152" pageId="60" pageNumber="61" unit="in" value="3.16" valueMax="3.44" valueMin="2.88">2.883.44 in</quantity>
<specimenCount box="[1213,1248,1446,1473]" pageId="60" pageNumber="61" type="female">5♀</specimenCount>
<typeStatus box="[1257,1369,1447,1473]" pageId="60" pageNumber="61">paratypes</typeStatus>
), ThL =
<quantity box="[175,276,1483,1509]" metricMagnitude="-3" metricUnit="m" metricValue="1.48" pageId="60" pageNumber="61" unit="mm" value="1.48">1.48 mm</quantity>
(
<quantity box="[292,436,1483,1509]" metricMagnitude="-2" metricUnit="m" metricValue="3.3909" metricValueMax="3.7084" metricValueMin="3.0734" pageId="60" pageNumber="61" unit="in" value="1.335" valueMax="1.46" valueMin="1.21">1.211.46 in</quantity>
♁,
<quantity box="[480,622,1483,1509]" metricMagnitude="-2" metricUnit="m" metricValue="3.9242999999999997" metricValueMax="4.572" metricValueMin="3.2766" pageId="60" pageNumber="61" unit="in" value="1.545" valueMax="1.8" valueMin="1.29">1.291.80 in</quantity>
<specimenCount box="[631,650,1483,1509]" pageId="60" pageNumber="61" type="female"></specimenCount>
), WL =
<quantity box="[750,852,1483,1509]" metricMagnitude="-3" metricUnit="m" metricValue="2.32" pageId="60" pageNumber="61" unit="mm" value="2.32">2.32 mm</quantity>
(
<quantity box="[867,1011,1482,1509]" metricMagnitude="-2" metricUnit="m" metricValue="6.223" metricValueMax="6.5786" metricValueMin="5.8674" pageId="60" pageNumber="61" unit="in" value="2.45" valueMax="2.59" valueMin="2.31">2.312.59 in</quantity>
♁,
<quantity box="[1055,1198,1483,1509]" metricMagnitude="-2" metricUnit="m" metricValue="6.5405" metricValueMax="6.9088" metricValueMin="6.1722" pageId="60" pageNumber="61" unit="in" value="2.575" valueMax="2.72" valueMin="2.43">2.432.72 in</quantity>
<specimenCount box="[1206,1225,1483,1509]" pageId="60" pageNumber="61" type="female"></specimenCount>
), WW =
<quantity box="[1335,1437,1483,1509]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="60" pageNumber="61" unit="mm" value="1.2">1.20 mm</quantity>
(
<quantity box="[159,302,1519,1545]" metricMagnitude="-2" metricUnit="m" metricValue="2.6162" metricValueMax="2.7686" metricValueMin="2.4638" pageId="60" pageNumber="61" unit="in" value="1.03" valueMax="1.09" valueMin="0.97">0.971.09 in</quantity>
♁,
<quantity box="[346,487,1519,1545]" metricMagnitude="-2" metricUnit="m" metricValue="2.7304999999999997" metricValueMax="3.1496" metricValueMin="2.3114" pageId="60" pageNumber="61" unit="in" value="1.075" valueMax="1.24" valueMin="0.91">0.911.24 in</quantity>
<specimenCount box="[495,514,1519,1545]" pageId="60" pageNumber="61" type="female"></specimenCount>
), arb = 9/5 (69/46), avd = 0.83 (0.760.93), adf = 1.50 (0.911.29), flw = 2.50 (1.742.16), FW/HW = 0.48 (0.270.46), ch/o = 0.17 (0.260.37), prorb = 1.50 (0.531.41), rcorb = 0.70 (0.59 1.16), vb = 0.56 (0.461.22), dcl = 0.33 (0.430.69), presctl = 0.57 (0.390.63), sctl = 2.23 (1.472.27), sterno = 0.64 (0.530.96), orbito = 1.40 (1.671.98), dcp = 0.28 (0.160.39), sctlp = 1.89 (1.552.08), C = 1.74 (1.532.13), 4c = 1.55 (1.061.38), 4v = 2.40 (1.362.10), 5x = 1.50 (1.062.00), ac = 15.50 (8.3617.55), M = 0.75 (0.580.72), C3F = 0.81 (0.560.71).
</paragraph>
</subSubSection>
<subSubSection pageId="60" pageNumber="61" type="materials_examined">
<paragraph blockId="60.[151,1437,978,1941]" pageId="60" pageNumber="61">
<emphasis bold="true" box="[199,389,1734,1761]" pageId="60" pageNumber="61">Type materials.</emphasis>
<materialsCitation collectingDate="2012-05-13" collectionCode="SCAU" collectorName="H. W. Chen" country="China" elevation="1420" latitude="22.738611" location="Pu'er" longLatPrecision="21" longitude="101.11694" municipality="Yixiang" pageId="60" pageNumber="61" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus box="[401,505,1735,1761]" pageId="60" pageNumber="61">Holotype</typeStatus>
♁ (
<collectionCode box="[558,637,1735,1761]" pageId="60" pageNumber="61">SCAU</collectionCode>
, No. 125586, DNA #1414, 2665),
<collectingCountry box="[1055,1148,1735,1761]" name="China" pageId="60" pageNumber="61">CHINA</collectingCountry>
:
<collectingMunicipality box="[1162,1254,1735,1761]" pageId="60" pageNumber="61">Yixiang</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:0393914C2A1E8744C282FCEE6B79FCD6:8EE576812A1E8744C6E3F94E6C68FDA2" box="[1270,1328,1735,1761]" country="China" latitude="22.738611" longLatPrecision="21" longitude="101.11694" municipality="Yixiang" name="Pu'er" pageId="60" pageNumber="61" stateProvince="Yunnan">Pu'er</location>
,
<collectingRegion box="[1344,1433,1735,1761]" country="China" name="Yunnan" pageId="60" pageNumber="61">Yunnan</collectingRegion>
,
<geoCoordinate box="[151,285,1771,1797]" degrees="22" direction="north" minutes="44" orientation="latitude" pageId="60" pageNumber="61" precision="15" seconds="19" value="22.738611">22°44'19''N</geoCoordinate>
,
<geoCoordinate box="[301,444,1771,1797]" degrees="101" direction="east" minutes="07" orientation="longitude" pageId="60" pageNumber="61" precision="15" seconds="01" value="101.11694">101°07'01''E</geoCoordinate>
, alt.
<quantity box="[508,588,1771,1797]" metricMagnitude="3" metricUnit="m" metricValue="1.42" pageId="60" pageNumber="61" unit="m" value="1420.0">
<elevation box="[508,588,1771,1797]" metricMagnitude="3" metricUnit="m" metricValue="1.42" pageId="60" pageNumber="61" unit="m" value="1420.0">1420m</elevation>
</quantity>
,
<date box="[605,718,1771,1797]" pageId="60" pageNumber="61" value="2012-05-13">
<collectingDate box="[605,718,1771,1797]" pageId="60" pageNumber="61" value="2012-05-13">13.v.2012</collectingDate>
</date>
, ex tree trunk,
<collectorName box="[904,1037,1771,1797]" pageId="60" pageNumber="61">H.W. Chen</collectorName>
</materialsCitation>
.
<materialsCitation collectionCode="SCAU" country="China" pageId="60" pageNumber="61" specimenCount="5" specimenCount-female="5" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus box="[1052,1165,1771,1797]" pageId="60" pageNumber="61">Paratypes</typeStatus>
: 2♁,
<specimenCount box="[1234,1269,1770,1797]" pageId="60" pageNumber="61" type="female">5♀</specimenCount>
(
<collectionCode box="[1289,1368,1771,1797]" pageId="60" pageNumber="61">SCAU</collectionCode>
, Nos. 12558793, DNA #1414, 1939, 2665, 2702), 1
</materialsCitation>
♁,
<materialsCitation collectingDate="2013-03-28" collectingDateMax="2016-05-10" collectingDateMin="2013-03-28" collectionCode="SCAU" collectorName="J. J. Gao &amp; J. Huang" country="China" elevation="1320" latitude="23.168888" location="Cangyuan" longLatPrecision="21" longitude="99.23111" municipality="Mengdong" pageId="60" pageNumber="61" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount box="[713,748,1807,1833]" pageId="60" pageNumber="61" type="female">1♀</specimenCount>
on
<date box="[791,957,1807,1833]" pageId="60" pageNumber="61" value="2013-03-28" valueMax="2013-03-30" valueMin="2013-03-28">
<collectingDate box="[791,957,1807,1833]" pageId="60" pageNumber="61" value="2013-03-28" valueMax="2013-03-30" valueMin="2013-03-28">2830.iii.2013</collectingDate>
</date>
,
<collectorName box="[969,1061,1807,1833]" pageId="60" pageNumber="61">J.J. Gao</collectorName>
, other data same as holotype; 1♁ (
<collectionCode box="[159,238,1843,1869]" pageId="60" pageNumber="61">SCAU</collectionCode>
, No. 125594, DNA #2173),
<collectingMunicipality box="[548,672,1843,1869]" pageId="60" pageNumber="61">Mengdong</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:0393914C2A1E8744C282FCEE6B79FCD6:8EE576812A1E8744C0BFF8BA6A46FC0E" box="[682,798,1843,1869]" latitude="23.168888" longLatPrecision="21" longitude="99.23111" municipality="Mengdong" name="Cangyuan" pageId="60" pageNumber="61" stateProvince="Yunnan">Cangyuan</location>
,
<collectingRegion box="[807,896,1843,1869]" country="China" name="Yunnan" pageId="60" pageNumber="61">Yunnan</collectingRegion>
,
<geoCoordinate box="[905,1038,1843,1869]" degrees="23" direction="north" minutes="10" orientation="latitude" pageId="60" pageNumber="61" precision="15" seconds="08" value="23.168888">23°10'08''N</geoCoordinate>
,
<geoCoordinate box="[1048,1178,1842,1869]" degrees="99" direction="east" minutes="13" orientation="longitude" pageId="60" pageNumber="61" precision="15" seconds="52" value="99.23111">99°13'52''E</geoCoordinate>
, alt.
<quantity box="[1229,1309,1843,1869]" metricMagnitude="3" metricUnit="m" metricValue="1.3199999999999998" pageId="60" pageNumber="61" unit="m" value="1320.0">
<elevation box="[1229,1309,1843,1869]" metricMagnitude="3" metricUnit="m" metricValue="1.3199999999999998" pageId="60" pageNumber="61" unit="m" value="1320.0">1320m</elevation>
</quantity>
,
<date box="[1319,1432,1843,1869]" pageId="60" pageNumber="61" value="2016-05-10">
<collectingDate box="[1319,1432,1843,1869]" pageId="60" pageNumber="61" value="2016-05-10">10.v.2016</collectingDate>
</date>
, ex tree trunk,
<collectorName box="[307,410,1879,1905]" pageId="60" pageNumber="61">J. Huang</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[199,545,1914,1941]" pageId="60" pageNumber="61" type="distribution">
<paragraph blockId="60.[151,1437,978,1941]" box="[199,545,1914,1941]" pageId="60" pageNumber="61">
<emphasis bold="true" box="[199,353,1914,1940]" pageId="60" pageNumber="61">Distribution.</emphasis>
<collectingCountry box="[359,426,1915,1941]" name="China" pageId="60" pageNumber="61">China</collectingCountry>
(
<collectingRegion box="[441,533,1915,1941]" country="China" name="Yunnan" pageId="60" pageNumber="61">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>