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<document id="9C629CDB10E7BEF64F7169E408EAB5E1" ID-DOI="10.5281/zenodo.3710074" ID-GBIF-Dataset="f1bbeff3-8e8b-408c-a46c-11595dcf1560" ID-ISSN="1942-1354" ID-Zenodo-Dep="3710074" ID-ZooBank="3EB9D858-BE71-4F1F-BEBD-136898474B51" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584124742059" checkinUser="felipe" docAuthor="Paulsen, M. J." docDate="2018" docId="039A87813E45F05D9BC5FD03C0E04B53" docLanguage="en" docName="InsectaMundi.0666.1-10.pdf.imf" docOrigin="Insecta Mundi 666" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Zelenkaesalus Krikken 2008" docType="treatment" docVersion="7" lastPageNumber="5" masterDocId="FFA3FFF93E43F05B9B05F91BC4084D26" masterDocTitle="Generic changes in the stag beetle tribe Aesalini (Coleoptera: Lucanidae: Aesalinae) with the description of two new species" masterLastPageNumber="10" masterPageNumber="1" pageNumber="5" updateTime="1698728097052" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="BA34D2CF745306A04471F830C3B03DDC">Generic changes in the stag beetle tribe Aesalini (Coleoptera: Lucanidae: Aesalinae) with the description of two new species</mods:title>
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<mods:namePart id="64CAF65603A93DE386FE1F60F79B8BCB">Paulsen, M. J.</mods:namePart>
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<mods:date id="2396F5E9DA8441665C9EA62B073AAF42">2018</mods:date>
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<treatment id="039A87813E45F05D9BC5FD03C0E04B53" ID-DOI="http://doi.org/10.5281/zenodo.3716621" ID-GBIF-Taxon="162488152" ID-Zenodo-Dep="3716621" LSID="urn:lsid:plazi:treatment:039A87813E45F05D9BC5FD03C0E04B53" httpUri="http://treatment.plazi.org/id/039A87813E45F05D9BC5FD03C0E04B53" lastPageNumber="5" pageId="6" pageNumber="5">
<subSubSection id="C329651C3E45F05D9BC5FD03C6414916" box="[192,585,1048,1072]" pageId="6" pageNumber="5" type="nomenclature">
<paragraph id="8B8C36973E45F05D9BC5FD03C6414916" blockId="6.[192,585,1048,1072]" box="[192,585,1048,1072]" pageId="6" pageNumber="5">
<heading id="D0C481FB3E45F05D9BC5FD03C6414916" bold="true" box="[192,585,1048,1072]" fontSize="10" level="2" pageId="6" pageNumber="5" reason="3">
<taxonomicName id="4C334D143E45F05D9BC5FD03C6414916" ID-CoL="648Y4" authority="Krikken, 2008" authorityName="Krikken" authorityYear="2008" box="[192,585,1048,1072]" class="Insecta" family="Lucanidae" genus="Zelenkaesalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B947EA853E45F05D9BC5FD03C6414916" bold="true" box="[192,585,1048,1072]" pageId="6" pageNumber="5">
<emphasis id="B947EA853E45F05D9BC5FD03C58B4916" bold="true" box="[192,387,1048,1072]" italics="true" pageId="6" pageNumber="5">Zelenkaesalus</emphasis>
<bibRefCitation id="EFA24B663E45F05D9A8EFD03C6414916" author="Krikken, J." box="[395,585,1048,1072]" pageId="6" pageNumber="5" pagination="275 - 284" refId="ref5562" refString="Krikken, J. 2008. Zelenkaesalus subgen. n. for three Southeast Asian species of Echinoaesalus Zelenka, including E. javanus sp. n. (Coleoptera: Lucanidae: Aesalinae). Koleopterologische Rundschau 78: 275 - 284." type="journal article" year="2008">Krikken, 2008</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C329651C3E45F05D9BC5FD52C6BF4947" box="[192,695,1096,1121]" pageId="6" pageNumber="5" type="type_taxon">
<paragraph id="8B8C36973E45F05D9BC5FD52C6BF4947" blockId="6.[192,695,1096,1121]" box="[192,695,1096,1121]" pageId="6" pageNumber="5">
<emphasis id="B947EA853E45F05D9BC5FD52C57F4946" bold="true" box="[192,375,1096,1121]" pageId="6" pageNumber="5">
<typeStatus id="548888353E45F05D9BC5FD52C50C4947" box="[192,260,1097,1121]" pageId="6" pageNumber="5">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C334D143E45F05D9A7BFD52C6BF4947" ID-CoL="64ZPV" authority="Krikken." authorityName="Krikken. The" authorityYear="2018" box="[382,695,1097,1121]" class="Insecta" family="Lucanidae" genus="Aesalus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="timidus">
<emphasis id="B947EA853E45F05D9A7BFD52C64A4947" box="[382,578,1097,1121]" italics="true" pageId="6" pageNumber="5">Aesalus timidus</emphasis>
Krikken.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C329651C3E45F05D9BEDFD62C01548E4" pageId="6" pageNumber="5" type="discussion">
<paragraph id="8B8C36973E45F05D9BEDFD62C0344994" blockId="6.[192,1440,1145,1202]" pageId="6" pageNumber="5">
<taxonomicName id="4C334D143E45F05D9BEDFD62C60B49B4" authority="Krikken" authorityName="Krikken" authorityYear="2008" box="[232,515,1145,1170]" class="Insecta" family="Lucanidae" genus="Zelenkaesalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B947EA853E45F05D9BEDFD62C59C49B7" box="[232,404,1145,1169]" italics="true" pageId="6" pageNumber="5">Zelenkaesalus</emphasis>
Krikken
</taxonomicName>
is elevated to generic status and contains the following six species, five of which represent new combinations, and one of which is described herein:
</paragraph>
<paragraph id="8B8C36973E45F05D9BC5FDD0C0C349C5" blockId="6.[192,1227,1226,1251]" box="[192,1227,1226,1251]" pageId="6" pageNumber="5">
<taxonomicName id="4C334D143E45F05D9BC5FDD0C61A49C5" authority="(Huang et al.)" authorityName="Paulsen" authorityYear="2018" baseAuthorityName="Huang et al." baseAuthorityYear="2018" box="[192,530,1226,1251]" class="Insecta" family="Lucanidae" genus="Zelenkaesalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="cechovskyi" status="comb. nov.">
<emphasis id="B947EA853E45F05D9BC5FDD0C55749C4" box="[192,351,1226,1250]" italics="true" pageId="6" pageNumber="5">Z. cechovskyi</emphasis>
(Huang et al.)
</taxonomicName>
,
<taxonomicNameLabel id="A27457FE3E45F05D991AFDD0C68D49C5" box="[543,645,1227,1251]" pageId="6" pageNumber="5" rank="species">n. comb.</taxonomicNameLabel>
<collectingCountry id="F32476073E45F05D9835FDD0C7CF49C5" box="[816,967,1227,1251]" name="Malaysia" pageId="6" pageNumber="5">MALAYSIA</collectingCountry>
:
<collectingRegion id="49F7F8753E45F05D98D4FDD0C01149C5" box="[977,1049,1227,1251]" country="Malaysia" name="Johor" pageId="6" pageNumber="5">Johor</collectingRegion>
;
<collectingCountry id="F32476073E45F05D9F20FDD0C0C349C5" box="[1061,1227,1227,1251]" name="Singapore" pageId="6" pageNumber="5">SINGAPORE</collectingCountry>
</paragraph>
<paragraph id="8B8C36973E45F05D9BC5FDEAC012482E" blockId="6.[192,1050,1264,1289]" box="[192,1050,1264,1289]" pageId="6" pageNumber="5">
<taxonomicName id="4C334D143E45F05D9BC5FDEAC621482F" authority="(Huang and Wu)" authorityName="Paulsen" authorityYear="2018" baseAuthorityName="Huang and Wu" baseAuthorityYear="2018" box="[192,553,1264,1289]" class="Insecta" family="Lucanidae" genus="Zelenkaesalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="gedeensis" status="comb. nov.">
<emphasis id="B947EA853E45F05D9BC5FDEAC55A482E" box="[192,338,1264,1288]" italics="true" pageId="6" pageNumber="5">Z. gedeensis</emphasis>
(Huang and Wu)
</taxonomicName>
,
<taxonomicNameLabel id="A27457FE3E45F05D9930FDEAC693482F" box="[565,667,1265,1289]" pageId="6" pageNumber="5" rank="species">n. comb.</taxonomicNameLabel>
<collectingCountry id="F32476073E45F05D9835FDEAC7DD482F" box="[816,981,1265,1289]" name="Indonesia" pageId="6" pageNumber="5">INDONESIA</collectingCountry>
:
<collectingRegion id="49F7F8753E45F05D98E5FDEAC012482E" box="[992,1050,1265,1288]" country="Indonesia" name="Jawa Timur" pageId="6" pageNumber="5">Java</collectingRegion>
</paragraph>
<paragraph id="8B8C36973E45F05D9BC5FC0CC0124808" blockId="6.[192,1050,1303,1327]" box="[192,1050,1303,1327]" pageId="6" pageNumber="5">
<taxonomicName id="4C334D143E45F05D9BC5FC0CC5CA4809" authority="(Krikken)" authorityName="Paulsen" authorityYear="2018" baseAuthorityName="Krikken" baseAuthorityYear="2018" box="[192,450,1303,1327]" class="Insecta" family="Lucanidae" genus="Zelenkaesalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="javanus" status="comb. nov.">
<emphasis id="B947EA853E45F05D9BC5FC0CC5494808" box="[192,321,1303,1326]" italics="true" pageId="6" pageNumber="5">Z. javanus</emphasis>
(Krikken)
</taxonomicName>
,
<taxonomicNameLabel id="A27457FE3E45F05D9ACBFC0CC63D4809" box="[462,565,1303,1327]" pageId="6" pageNumber="5" rank="species">n. comb.</taxonomicNameLabel>
<collectingCountry id="F32476073E45F05D9835FC0CC7DD4809" box="[816,981,1303,1327]" name="Indonesia" pageId="6" pageNumber="5">INDONESIA</collectingCountry>
:
<collectingRegion id="49F7F8753E45F05D98E5FC0CC0124808" box="[992,1050,1303,1326]" country="Indonesia" name="Jawa Timur" pageId="6" pageNumber="5">Java</collectingRegion>
</paragraph>
<paragraph id="8B8C36973E45F05D9BC5FC26C04A4872" blockId="6.[192,1090,1341,1365]" box="[192,1090,1341,1365]" pageId="6" pageNumber="5">
<taxonomicName id="4C334D143E45F05D9BC5FC26C5B84873" authority="Paulsen" authorityName="Paulsen" authorityYear="2018" box="[192,432,1341,1365]" class="Insecta" family="Lucanidae" genus="Zelenkaesalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="pazuzus" status="sp. nov.">
<emphasis id="B947EA853E45F05D9BC5FC26C54B4872" box="[192,323,1341,1364]" italics="true" pageId="6" pageNumber="5">Z. pazuzus</emphasis>
Paulsen
</taxonomicName>
,
<taxonomicNameLabel id="A27457FE3E45F05D9AB9FC26C5F64872" box="[444,510,1341,1364]" pageId="6" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
<collectingCountry id="F32476073E45F05D9835FC26C7E34873" box="[816,1003,1341,1365]" name="Philippines" pageId="6" pageNumber="5">PHILIPPINES</collectingCountry>
: Luzon
</paragraph>
<paragraph id="8B8C36973E45F05D9BC5FC78C086485D" blockId="6.[192,1166,1378,1403]" box="[192,1166,1378,1403]" pageId="6" pageNumber="5">
<taxonomicName id="4C334D143E45F05D9BC5FC78C5EB485D" authority="(Zelenka)" authorityName="Paulsen" authorityYear="2018" baseAuthorityName="Zelenka" baseAuthorityYear="2018" box="[192,483,1378,1403]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="sabahensis" status="comb. nov.">
<emphasis id="B947EA853E45F05D9BC5FC78C56D485C" box="[192,357,1378,1402]" italics="true" pageId="6" pageNumber="5">Z. sabahensis</emphasis>
(Zelenka)
</taxonomicName>
,
<taxonomicNameLabel id="A27457FE3E45F05D9AEAFC78C65D485D" box="[495,597,1379,1403]" pageId="6" pageNumber="5" rank="species">n. comb.</taxonomicNameLabel>
<collectingCountry id="F32476073E45F05D9835FC78C7CF485D" box="[816,967,1379,1403]" name="Malaysia" pageId="6" pageNumber="5">MALAYSIA</collectingCountry>
:
<collectingRegion id="49F7F8753E45F05D98D4FC78C016485D" box="[977,1054,1379,1403]" country="Malaysia" name="Sabah" pageId="6" pageNumber="5">Sabah</collectingRegion>
(Borneo)
</paragraph>
<paragraph id="8B8C36973E45F05D9BC5FC92C01548E4" blockId="6.[192,1440,1416,1474]" pageId="6" pageNumber="5">
<taxonomicName id="4C334D143E45F05D9BC5FC92C5C84887" authority="(Krikken)" authorityName="Paulsen" authorityYear="2018" baseAuthorityName="Krikken. The" baseAuthorityYear="2018" box="[192,448,1416,1441]" class="Insecta" family="Lucanidae" genus="Aesalus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="timidus" status="comb. nov.">
<emphasis id="B947EA853E45F05D9BC5FC92C5374886" box="[192,319,1416,1440]" italics="true" pageId="6" pageNumber="5">Z. timidus</emphasis>
(Krikken)
</taxonomicName>
,
<taxonomicNameLabel id="A27457FE3E45F05D9AC9FC92C63B4887" box="[460,563,1417,1441]" pageId="6" pageNumber="5" rank="species">n. comb.</taxonomicNameLabel>
<collectingCountry id="F32476073E45F05D9835FC92C7C74887" box="[816,975,1417,1441]" name="Indonesia" pageId="6" pageNumber="5">INDONESIA</collectingCountry>
: Borneo,
<collectingRegion id="49F7F8753E45F05D9F31FC92C0964887" box="[1076,1182,1417,1441]" country="Indonesia" name="Sumatera Utara" pageId="6" pageNumber="5">Sumatra</collectingRegion>
;
<collectingCountry id="F32476073E45F05D9FA3FC92C13F4887" box="[1190,1335,1417,1441]" name="Malaysia" pageId="6" pageNumber="5">MALAYSIA</collectingCountry>
:
<collectingRegion id="49F7F8753E45F05D9E3BFC92C1934887" box="[1342,1435,1417,1441]" country="Malaysia" name="Pahang" pageId="6" pageNumber="5">Pahang</collectingRegion>
,
<collectingRegion id="49F7F8753E45F05D9835FCB2C77448E7" box="[816,892,1449,1473]" country="Malaysia" name="Perak" pageId="6" pageNumber="5">Perak</collectingRegion>
;
<collectingCountry id="F32476073E45F05D988DFCB1C01548E4" box="[904,1053,1450,1474]" name="Thailand" pageId="6" pageNumber="5">THAILAND</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="C329651C3E45F05D9BEDFCC1C0E04B53" pageId="6" pageNumber="5" type="diagnosis">
<paragraph id="8B8C36973E45F05D9BEDFCC1C0E04B53" blockId="6.[192,1441,1498,1653]" pageId="6" pageNumber="5">
The genus is defined by the presence of well-developed sulci on the abdomen (
<figureCitation id="13082A123E45F05D9F97FCC1C0EE48D4" box="[1170,1254,1498,1522]" captionStart="Figures 12" captionStartId="4.[192,287,1819,1841]" captionTargetBox="[192,1407,276,1739]" captionTargetId="figure@4.[192,1440,261,1767]" captionTargetPageId="4" captionText="Figures 12. Zelenka holotypes. 1) Holotype of Echinoaesalus jaechi Zelenka. a) Dorsal habitus. b) Abdomen, ventral view. c) Labels. 2) Holotype female of Echinoaesalus sabahensis Zelenka. a) Dorsal habitus. b) Abdomen, ventral view, showing complete leg-shaped sulci. c) Labels. Photos courtesy H. Schillhammer (Naturhistorisches Museum, Vienna)." figureDoi="http://doi.org/10.5281/zenodo.3710076" httpUri="https://zenodo.org/record/3710076/files/figure.png" pageId="6" pageNumber="5">Fig. 2b</figureCitation>
) and metaster- num for receiving the entire tibia and tarsus. The canthus is short (
<figureCitation id="13082A123E45F05D9F00FCE0C0434B34" box="[1029,1099,1530,1555]" captionStart="Figures 35" captionStartId="5.[192,287,605,627]" captionTargetBox="[192,1439,213,570]" captionTargetId="figure@5.[192,1440,213,571]" captionTargetPageId="5" captionText="Figures 35. Generic differences of ocular canthus/eye and clypeal apex. 3) Echinoaesalus spp. 4) Strabaesalus spp. 5) Zelenkaesalus spp." figureDoi="http://doi.org/10.5281/zenodo.3710078" httpUri="https://zenodo.org/record/3710078/files/figure.png" pageId="6" pageNumber="5">Fig. 5</figureCitation>
) and the eye in dorsal view is almost entirely covered by the pronotum when the head is retracted. The dorsal vestiture includes clumped scales, scattered bristles, and a low covering of minutely branched scales as in
<taxonomicName id="4C334D143E45F05D9FD7FF27C14C4B72" authorityName="Paulsen" authorityYear="2013" box="[1234,1348,1596,1620]" class="Insecta" family="Lucanidae" genus="Trogellus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B947EA853E45F05D9FD7FF27C14C4B72" box="[1234,1348,1596,1620]" italics="true" pageId="6" pageNumber="5">Trogellus</emphasis>
</taxonomicName>
species (
<bibRefCitation id="EFA24B663E45F05D9BCDFF46C57A4B53" author="Paulsen, M. J." box="[200,370,1629,1653]" pageId="6" pageNumber="5" pagination="1 - 25" refId="ref5606" refString="Paulsen, M. J. 2013. A new genus for the Neotropical species of Aesalus Fabricius, with descriptions of eight new species (Coleoptera: Lucanidae: Aesalinae). Insecta Mundi 0325: 1 - 25." type="book chapter" year="2013">Paulsen 2013</bibRefCitation>
). The branched scales generally anchor a waxy exudate, unless abraded.
</paragraph>
</subSubSection>
</treatment>
</document>