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<document id="13E36A319C5D5525D835079EF1748934" ID-CLB-Dataset="276840" ID-DOI="10.11646/zootaxa.5228.2.4" ID-GBIF-Dataset="c2b7eabc-5c5c-4b8c-8f5d-fdb96d1df158" ID-ISSN="1175-5326" ID-Zenodo-Dep="7532281" ID-ZooBank="0B15B8F9-142C-4785-8CD8-D24409FA733E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1673571446165" checkinUser="plazi" docAuthor="Zarkani, Agustin, Ercan, Cansu, Apriyanto, Dwinardi &amp; Kaydan, Mehmet Bora" docDate="2023" docId="03EB87E0136B7A35B59247A1B679FAC0" docLanguage="en" docName="zootaxa.5228.2.4.pdf" docOrigin="Zootaxa 5228 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dysmicoccus zeynepae Zarkani &amp; Kaydan 2023, sp. n." docType="treatment" docVersion="4" lastPageNumber="164" masterDocId="FFD2FF98136F7A32B5054222B529FFB9" masterDocTitle="Studies on mealybugs (Hemiptera: Pseudococcidae) in Indonesia, with description of a new species and three new country records" masterLastPageNumber="172" masterPageNumber="157" pageNumber="161" updateTime="1699477565528" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="F0FA357BAEE2E4146899CEF1682BEE7C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6D0D937F6F15A440BD83C06AAF7E09F2">
<mods:title id="02C61FB321040E679E480459AE3669FD">Studies on mealybugs (Hemiptera: Pseudococcidae) in Indonesia, with description of a new species and three new country records</mods:title>
</mods:titleInfo>
<mods:name id="B83D430476552391FF3F9D7A7F9D132E" type="personal">
<mods:role id="D0FB8048C534D8A1367F61F1C10CEF09">
<mods:roleTerm id="F55EA2C167E6A9AFCBDDE9B7083148FC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FFAAA6C4A422FBD5968C7232803180D3">Zarkani, Agustin</mods:namePart>
<mods:nameIdentifier id="A5465DE189875F4F050533F08029F932" type="ORCID">0000-0001-9837-5019</mods:nameIdentifier>
<mods:affiliation id="C0DEE1210EAA57199E0E8CBF09B4CFDB">agustinzarkani @ unib. ac. id; https: // orcid. org / 0000 - 0001 - 9837 - 5019</mods:affiliation>
<mods:nameIdentifier id="187BDD3754AF0683040AADA382651CCB" type="email">agustinzarkani@unib.ac.id</mods:nameIdentifier>
</mods:name>
<mods:name id="18C8A5C4615B085105F73159A7754C82" type="personal">
<mods:role id="250BC5FCD013BE6293B31D68AF0DA8EE">
<mods:roleTerm id="C992764FF23E9BFABD6E54F5C6DD4BD4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5494F7EE97EF4828407171E1A3F984B8">Ercan, Cansu</mods:namePart>
<mods:nameIdentifier id="17D3584441E2F8E085934DD6A429338A" type="ORCID">0000-0003-3516-9183</mods:nameIdentifier>
<mods:affiliation id="DB6C87BECE1812C85FFE424E151BB8B3">Biotechnology Development and Research Centre, Çukurova University, 01250, Adana, Turkey. &amp; cansuercann 01 @ gmail. com; https: // orcid. org / 0000 - 0003 - 3516 - 9183</mods:affiliation>
<mods:nameIdentifier id="D8AD007FC8B1ECCC4D314BC89234E236" type="email">cansuercann01@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="E9EA72D36ACC91F57153C9A60DE6AD02" type="personal">
<mods:role id="7A880E84F97F7F250D45896807C3C8B0">
<mods:roleTerm id="28DB0962A747A59DB60D294DA4BF8B43">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F975F49E52666E862779A380DB5469AC">Apriyanto, Dwinardi</mods:namePart>
<mods:nameIdentifier id="72124BD47B2316D72630005EA9721921" type="ORCID">0000-0002-5286-9010</mods:nameIdentifier>
<mods:affiliation id="F03469D394E17E498774C37E567A348A">dwinardi 2018 @ gmail. com; https: // orcid. org / 0000 - 0002 - 5286 - 9010</mods:affiliation>
<mods:nameIdentifier id="1622B88B65726F471F54A48417A9A09B" type="email">dwinardi2018@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="36DE82B07E2C78FF081A89F309B1F056" type="personal">
<mods:role id="403946F2FAA0AFF8702B896D5666BB07">
<mods:roleTerm id="C031CDE298ED0CE7DDA8E63389BC2C03">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="49228CE2F01FACA1A17BA3C88D8313B9">Kaydan, Mehmet Bora</mods:namePart>
<mods:nameIdentifier id="A68911AC0115E75205B947FE7F8B68D1" type="ORCID">0000-0002-0677-255X</mods:nameIdentifier>
<mods:affiliation id="0A664B619ECA106D9D6AD4E39A387770">Biotechnology Development and Research Centre, Çukurova University, 01250, Adana, Turkey. &amp; bkaydan @ cu. edu. tr; https: // orcid. org / 0000 - 0002 - 0677 - 255 X</mods:affiliation>
<mods:nameIdentifier id="996C87253313B3F9FE2BD846B39CF3EC" type="email">bkaydan@cu.edu.tr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="8296DFAD351B5125163EABD46BA0C56A">text</mods:typeOfResource>
<mods:relatedItem id="1CCDB0573F42E54D43EFA601FDE4A442" type="host">
<mods:titleInfo id="5BB855A1121915DE4D760939D046972D">
<mods:title id="4AE52627FA9F6AB95AB0F4154343F182">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="24D5186B020ED6121C4E728CFDB2B410">
<mods:date id="09B5FB1A05BEB9D75E6D1556E96E1960">2023</mods:date>
<mods:detail id="4FAE7E965605B62F5EA2220A5B01A568" type="pubDate">
<mods:number id="E52F8FE7C30CCF975FEE83986A6947FA">2023-01-12</mods:number>
</mods:detail>
<mods:detail id="4394AE6B0F06E69C2136203257780073" type="volume">
<mods:number id="FF00F6E195490E5D0D1D1ED1F07EF87F">5228</mods:number>
</mods:detail>
<mods:detail id="166497A3961B0A2E3467F3CF56233AF3" type="issue">
<mods:number id="C4F94333FF531E176648C15823BD0307">2</mods:number>
</mods:detail>
<mods:extent id="BCA8FF6BBA1269140439B5A2F7731F82" unit="page">
<mods:start id="5F2860F17E2A333A8142963091FE0C98">157</mods:start>
<mods:end id="C1D8DBC4EC0CF537D6CD949549E13690">172</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="068852FF364161C49B6CDE6C5A614E6A">journal article</mods:classification>
<mods:identifier id="D9C67B2DF3CC650ED239178AAA96B923" type="CLB-Dataset">276840</mods:identifier>
<mods:identifier id="B560F3C5CD07E122FB9E12A0055FE954" type="DOI">10.11646/zootaxa.5228.2.4</mods:identifier>
<mods:identifier id="DC2935A994C3E9E36AC6A9BA5F19E23C" type="GBIF-Dataset">c2b7eabc-5c5c-4b8c-8f5d-fdb96d1df158</mods:identifier>
<mods:identifier id="E23D95C107C5661764F68D71B2543A78" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="FB4AA414DCF046901A923EA830FAB3FD" type="Zenodo-Dep">7532281</mods:identifier>
<mods:identifier id="369820E508AFF78C1B8E2C2E038EE617" type="ZooBank">0B15B8F9-142C-4785-8CD8-D24409FA733E</mods:identifier>
</mods:mods>
<treatment id="03EB87E0136B7A35B59247A1B679FAC0" ID-DOI="http://doi.org/10.5281/zenodo.7532289" ID-GBIF-Taxon="217348829" ID-Zenodo-Dep="7532289" LSID="urn:lsid:plazi:treatment:03EB87E0136B7A35B59247A1B679FAC0" httpUri="http://treatment.plazi.org/id/03EB87E0136B7A35B59247A1B679FAC0" lastPageId="7" lastPageNumber="164" pageId="4" pageNumber="161">
<subSubSection id="C358657D136B7A36B59247A1B7C5FA24" box="[151,748,1410,1437]" pageId="4" pageNumber="161" type="nomenclature">
<paragraph id="8BFD36F6136B7A36B59247A1B7C5FA24" blockId="4.[151,748,1410,1437]" box="[151,748,1410,1437]" pageId="4" pageNumber="161">
<heading id="D0B5819A136B7A36B59247A1B7C5FA24" bold="true" box="[151,748,1410,1437]" fontSize="11" level="1" pageId="4" pageNumber="161" reason="1">
<emphasis id="B936EAE4136B7A36B59247A1B7C5FA24" bold="true" box="[151,748,1410,1437]" pageId="4" pageNumber="161">
<taxonomicName id="4C424D75136B7A36B59247A1B78BFA24" authority="Zarkani &amp; Kaydan" authorityName="Zarkani &amp; Kaydan" authorityYear="2023" box="[151,674,1410,1437]" class="Insecta" family="Pseudococcidae" genus="Dysmicoccus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="161" phylum="Arthropoda" rank="species" species="zeynepae" status="sp. nov.">
<emphasis id="B936EAE4136B7A36B59247A1B483FA24" bold="true" box="[151,426,1411,1437]" italics="true" pageId="4" pageNumber="161">Dysmicoccus zeynepae</emphasis>
Zarkani &amp; Kaydan
</taxonomicName>
<taxonomicNameLabel id="A205579F136B7A36B7AF47A1B7C5FA24" box="[682,748,1411,1437]" pageId="4" pageNumber="161" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C358657D136B7A36B59247E8B77AF8BC" pageId="4" pageNumber="161" type="materials_examined">
<paragraph id="8BFD36F6136B7A36B59247E8B7B1FA5C" blockId="4.[151,1437,1482,1977]" box="[151,664,1482,1509]" pageId="4" pageNumber="161">
<emphasis id="B936EAE4136B7A36B59247E8B450FA5D" bold="true" box="[151,377,1482,1508]" pageId="4" pageNumber="161">Material examined</emphasis>
, all deposited at
<collectionCode id="ED53AE33136B7A36B73D47E9B7BDFA5C" box="[568,660,1483,1509]" pageId="4" pageNumber="161">MMUB</collectionCode>
.
</paragraph>
<paragraph id="8BFD36F6136B7A36B5C247CDB199F9E8" blockId="4.[151,1437,1482,1977]" pageId="4" pageNumber="161">
<emphasis id="B936EAE4136B7A36B5C247CDB41DF9B0" bold="true" box="[199,308,1519,1545]" pageId="4" pageNumber="161">
<typeStatus id="54F98854136B7A36B5C247CDB41DF9B0" box="[199,308,1519,1545]" pageId="4" pageNumber="161" type="holotype">Holotype</typeStatus>
</emphasis>
adult female, left label: AZ205 /
<date id="FFFC1036136B7A36B7C047CDB601F9B0" box="[709,808,1519,1545]" pageId="4" pageNumber="161" value="2008-02-04">4.ii.2008</date>
/
<collectingCountry id="F3557666136B7A36B64347CDB69AF9B0" box="[838,947,1519,1545]" name="Indonesia" pageId="4" pageNumber="161">Indonesia</collectingCountry>
/
<collectingRegion id="4986F814136B7A36B6D547CDB119F9B0" box="[976,1072,1519,1545]" country="Indonesia" name="Sumatera Utara" pageId="4" pageNumber="161">Sumatra</collectingRegion>
,
<collectingRegion id="4986F814136B7A36B14547CDB184F9B0" box="[1088,1197,1519,1545]" country="Indonesia" name="Bengkulu" pageId="4" pageNumber="161">Bengkulu</collectingRegion>
/
<taxonomicName id="4C424D75136B7A36B1CF47CDB0A3F9B0" box="[1226,1418,1519,1545]" class="Magnoliopsida" family="Malvaceae" genus="Durio" kingdom="Plantae" order="Malvales" pageId="4" pageNumber="161" phylum="Tracheophyta" rank="species" species="zibethinus">
<emphasis id="B936EAE4136B7A36B1CF47CDB0A3F9B0" box="[1226,1418,1519,1545]" italics="true" pageId="4" pageNumber="161">Durio zibethinus</emphasis>
</taxonomicName>
/ 10238′33″ E / 0334′54.4″ S /
<quantity id="4CBA9B13136B7A36B4DF4430B70AF994" box="[474,547,1554,1581]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="4" pageNumber="161" unit="m" value="520.0">520 m</quantity>
; right label:
<emphasis id="B936EAE4136B7A36B7A94431B667F994" box="[684,846,1555,1581]" italics="true" pageId="4" pageNumber="161">
<taxonomicName id="4C424D75136B7A36B7A94431B669F995" authorityName="Ferris" authorityYear="1950" box="[684,832,1555,1580]" class="Insecta" family="Pseudococcidae" genus="Dysmicoccus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="161" phylum="Arthropoda" rank="genus">Dysmicoccus</taxonomicName>
/
</emphasis>
<taxonomicName id="4C424D75136B7A36B6514431B1BAF994" authority="Zarkani &amp; Kaydan" authorityName="Zarkani &amp; Kaydan" authorityYear="2023" box="[852,1171,1555,1581]" class="Insecta" family="Pseudococcidae" genus="Dysmicoccus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="161" phylum="Arthropoda" rank="species" species="zeynepae">
<emphasis id="B936EAE4136B7A36B6514431B692F995" box="[852,955,1555,1580]" italics="true" pageId="4" pageNumber="161">zeynepae</emphasis>
Zarkani &amp; Kaydan
</taxonomicName>
,
<specimenCount id="9D44FD7F136B7A36B1984431B1F2F994" box="[1181,1243,1555,1581]" pageId="4" pageNumber="161" type="female">3 ♀♀</specimenCount>
/ coll. A. Zarkani / det.
<collectionCode id="ED53AE33136B7A36B5D34415B42BF9E8" box="[214,258,1591,1617]" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="4" pageNumber="161">MB</collectionCode>
Kaydan. The
<typeStatus id="54F98854136B7A36B4A54415B72BF9E8" box="[416,514,1591,1617]" pageId="4" pageNumber="161" type="holotype">holotype</typeStatus>
specimen is ringed with red ink on the coverslip. In
<collectionCode id="ED53AE33136B7A36B14A4415B182F9E8" box="[1103,1195,1591,1617]" pageId="4" pageNumber="161">MMUB</collectionCode>
.
</paragraph>
<paragraph id="8BFD36F6136B7A36B5C24479B77AF8BC" blockId="4.[151,1437,1482,1977]" pageId="4" pageNumber="161">
<emphasis id="B936EAE4136B7A36B5C24479B46FF9CC" bold="true" box="[199,326,1627,1653]" pageId="4" pageNumber="161">
<typeStatus id="54F98854136B7A36B5C24479B469F9CC" box="[199,320,1627,1653]" pageId="4" pageNumber="161" type="paratype">Paratypes</typeStatus>
,
</emphasis>
<specimenCount id="9D44FD7F136B7A36B4574478B4BEF9CC" box="[338,407,1626,1653]" pageId="4" pageNumber="161" type="female">5 ♀♀</specimenCount>
,
<collectingCountry id="F3557666136B7A36B4AF4479B76EF9CC" box="[426,583,1627,1653]" name="Indonesia" pageId="4" pageNumber="161">INDONESIA</collectingCountry>
: (AZ205) same data as
<typeStatus id="54F98854136B7A36B66D4479B6E7F9CC" box="[872,974,1627,1653]" pageId="4" pageNumber="161" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D44FD7F136B7A36B6DB4479B10AF9CC" box="[990,1059,1627,1653]" pageId="4" pageNumber="161" type="female">3 ♀♀</specimenCount>
, AZ206,
<collectingRegion id="4986F814136B7A36B1924479B1DEF9CC" box="[1175,1271,1627,1653]" country="Indonesia" name="Sumatera Utara" pageId="4" pageNumber="161">Sumatra</collectingRegion>
,
<collectingRegion id="4986F814136B7A36B0024479B05DF9CC" box="[1287,1396,1627,1653]" country="Indonesia" name="Bengkulu" pageId="4" pageNumber="161">Bengkulu</collectingRegion>
on
<taxonomicName id="4C424D75136B7A36B592445DB771F920" authority="Corr." authorityName="Corr." box="[151,600,1663,1689]" class="Magnoliopsida" family="Meliaceae" genus="Lansium" kingdom="Plantae" order="Sapindales" pageId="4" pageNumber="161" phylum="Tracheophyta" rank="species" species="parasiticum">
<emphasis id="B936EAE4136B7A36B592445DB4AFF921" box="[151,390,1663,1688]" italics="true" pageId="4" pageNumber="161">Lansium parasiticum</emphasis>
Corr. (Meliaceae)
</taxonomicName>
, 0359′28.0″ S, 10225′50.4″ E,
<date id="FFFC1036136B7A36B6B3445DB100F920" box="[950,1065,1663,1689]" pageId="4" pageNumber="161" value="2018-02-11">11.ii.2018</date>
, coll. A. Zarkani;
<specimenCount id="9D44FD7F136B7A36B1FD445DB010F920" box="[1272,1337,1663,1689]" pageId="4" pageNumber="161" type="female">3 ♀♀</specimenCount>
, AZ207,
<collectingRegion id="4986F814136B7A36B5924481B5DEF904" box="[151,247,1699,1725]" country="Indonesia" name="Sumatera Utara" pageId="4" pageNumber="161">Sumatra</collectingRegion>
,
<collectingRegion id="4986F814136B7A36B4044481B447F904" box="[257,366,1699,1725]" country="Indonesia" name="Bengkulu" pageId="4" pageNumber="161">Bengkulu</collectingRegion>
on
<taxonomicName id="4C424D75136B7A36B4934481B774F905" box="[406,605,1699,1725]" class="Magnoliopsida" family="Sapotaceae" genus="Manilkara" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="161" phylum="Tracheophyta" rank="species" species="zapota">
<emphasis id="B936EAE4136B7A36B4934481B774F905" box="[406,605,1699,1725]" italics="true" pageId="4" pageNumber="161">Manilkara zapota</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C424D75136B7A36B78C4481B624F904" box="[649,781,1699,1725]" class="Magnoliopsida" family="Sapotaceae" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="161" phylum="Tracheophyta" rank="family">Sapotaceae</taxonomicName>
), 0400′05.7″ S, 10226′52.1″ E,
<date id="FFFC1036136B7A36B16D4481B1F5F904" box="[1128,1244,1699,1725]" pageId="4" pageNumber="161" value="2018-02-12">12.ii.2018</date>
, coll. A. Zarkani;
<specimenCount id="9D44FD7F136B7A36B59244E5B5F0F958" box="[151,217,1735,1761]" pageId="4" pageNumber="161" type="female">3 ♀♀</specimenCount>
, AZ208,
<collectingRegion id="4986F814136B7A36B44044E5B48CF958" box="[325,421,1735,1761]" country="Indonesia" name="Sumatera Utara" pageId="4" pageNumber="161">Sumatra</collectingRegion>
,
<collectingRegion id="4986F814136B7A36B4B744E5B70BF958" box="[434,546,1735,1761]" country="Indonesia" name="Bengkulu" pageId="4" pageNumber="161">Bengkulu</collectingRegion>
,
<taxonomicName id="4C424D75136B7A36B72A44E5B16AF958" authority="Lindl. ex De Will." authorityName="De Will." box="[559,1091,1735,1761]" class="Magnoliopsida" family="Rubiaceae" genus="Coffea" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="161" phylum="Tracheophyta" rank="species" species="robusta">
<emphasis id="B936EAE4136B7A36B72A44E5B7FEF958" box="[559,727,1735,1761]" italics="true" pageId="4" pageNumber="161">Coffea robusta</emphasis>
Lindl. ex De Will. (Rubiaceae)
</taxonomicName>
, 0336′15.4″ S, 10236′30.8″ E,
<date id="FFFC1036136B7A36B59244C9B422F8BC" box="[151,267,1771,1797]" pageId="4" pageNumber="161" value="2018-02-19">19.ii.2018</date>
, coll. A. Zarkani. In
<collectionCode id="ED53AE33136B7A36B4F644C9B766F8BC" box="[499,591,1771,1797]" pageId="4" pageNumber="161">MMUB</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C358657D136B7A35B5C2452CB456FCE4" lastPageId="7" lastPageNumber="164" pageId="4" pageNumber="161" type="description">
<paragraph id="8BFD36F6136B7A36B5C2452CB722F891" blockId="4.[151,1437,1482,1977]" box="[199,523,1806,1833]" pageId="4" pageNumber="161">
<emphasis id="B936EAE4136B7A36B5C2452CB722F891" bold="true" box="[199,523,1806,1833]" pageId="4" pageNumber="161">Description of adult female</emphasis>
</paragraph>
<paragraph id="8BFD36F6136B7A36B5C24510B1B6F8C8" blockId="4.[151,1437,1482,1977]" pageId="4" pageNumber="161">
<emphasis id="B936EAE4136B7A36B5C24510B48FF8F5" bold="true" box="[199,422,1842,1868]" pageId="4" pageNumber="161">Appearance in life</emphasis>
(
<figureCitation id="13792A73136B7A36B4BD4511B4D2F8F4" box="[440,507,1843,1869]" captionStart="FIGURE 1" captionStartId="5.[151,250,1930,1954]" captionTargetBox="[151,1436,181,1907]" captionTargetId="figure-17@5.[151,1436,181,1907]" captionTargetPageId="5" captionText="FIGURE 1. Dysmicoccus zeynepae Zarkani &amp; Kaydan sp. n. on: (A) Lansium parasiticum Corr.; (B) Coffea robusta Lindl. ex De Will; (C) Manilkara zapota L.; and (D) Durio zibethinus Murr." figureDoi="http://doi.org/10.5281/zenodo.7532285" httpUri="https://zenodo.org/record/7532285/files/figure.png" pageId="4" pageNumber="161">Fig. 1</figureCitation>
). Adult females secrete a thin powdery white wax covering the dorsal surface of their bodies. Living on leaves, flowers and fruits of host plants, commonly attended by ants.
</paragraph>
<paragraph id="8BFD36F6136B7A35B5C24558B1B7FDA0" blockId="4.[151,1437,1482,1977]" lastBlockId="7.[151,1437,150,1401]" lastPageId="7" lastPageNumber="164" pageId="4" pageNumber="161">
<emphasis id="B936EAE4136B7A36B5C24558B73CF82D" bold="true" box="[199,533,1914,1941]" pageId="4" pageNumber="161">Slide-mounted adult female</emphasis>
(based on
<typeStatus id="54F98854136B7A36B7934559B7D1F82C" box="[662,760,1915,1941]" pageId="4" pageNumber="161" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D44FD7F136B7A36B6364558B69EF82C" box="[819,951,1914,1941]" pageId="4" pageNumber="161" type="generic" typeStatus="paratypes">5 paratypes</specimenCount>
) (
<figureCitation id="13792A73136B7A36B6D54559B131F82C" box="[976,1048,1915,1941]" captionStart="FIGURE 2" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[184,1396,195,1910]" captionTargetId="figure-19@6.[171,1416,181,1919]" captionTargetPageId="6" captionText="FIGURE 2. Adult female Dysmicoccus zeynepae Zarkani &amp; Kaydan sp. n., holotype." figureDoi="http://doi.org/10.5281/zenodo.7532291" httpUri="https://zenodo.org/record/7532291/files/figure.png" pageId="4" pageNumber="161">Fig. 2</figureCitation>
): Body oval,
<quantity id="4CBA9B13136B7A36B1B04558B072F82C" box="[1205,1371,1914,1941]" metricMagnitude="-3" metricUnit="m" metricValue="2.245" metricValueMax="2.54" metricValueMin="1.95" pageId="4" pageNumber="161" unit="mm" value="2.245" valueMax="2.54" valueMin="1.95">1.95–2.54 mm</quantity>
long,
<quantity id="4CBA9B13136B7A36B59245BCB414F800" box="[151,317,1950,1977]" metricMagnitude="-3" metricUnit="m" metricValue="1.9449999999999998" metricValueMax="2.25" metricValueMin="1.64" pageId="4" pageNumber="161" unit="mm" value="1.9449999999999998" valueMax="2.25" valueMin="1.64">1.64–2.25 mm</quantity>
wide. Eyes situated on margins, each 42–45 μm wide. Antenna 8 segmented, 340–380 μm long, with 4 fleshy setae each 22.5–25.0 μm long; apical segment 75–80 μm long, 27.5–30.0 μm wide, with apical seta 30–35 μm long. Clypeolabral shield 200–220 μm long, 175–185 μm wide. Labium 3 segmented, 110–120 μm long, 80–85 μm wide. Anterior spiracles each 85–90 μm long, 45–50 μm wide across atrium; posterior spiracles each 95.–115. μm long, 55–60 μm wide across atrium. Circulus rounded-quadrate 80–110 μm wide. Legs well developed; segment lengths for each posterior leg: coxa 160–175 μm, trochanter + femur 145–155 μm, tibia + tarsus 175–180 μm, claw 35.0–37.5 μm. Ratio of length of tibia + tarsus to trochanter + femur 1.16–1.21: 1; ratio of length of tibia to tarsus, 1.2–1.4: 1; ratio of length of trochanter + femur to greatest width of femur, 2.41–2.60: 1; coxa with 30–50 translucent pores; tibia with numerous translucent pores. Tarsal digitules capitate, each 35–40 μm long. Claw digitules capitate, each about 27.5–30.0 μm long. Both pairs of ostioles present, anterior ostioles each with a total for both lips of 28–40 trilocular pores and 4–6 setae; posterior ostioles each with a total for both lips of 58–60 trilocular pores and 6–8 setae. Anal ring about 70 μm wide, bearing 6 setae, each seta 80–90 μm long.
</paragraph>
<caption id="DF3D667E136A7A37B59245A8B610F87F" ID-DOI="http://doi.org/10.5281/zenodo.7532285" ID-Zenodo-Dep="7532285" httpUri="https://zenodo.org/record/7532285/files/figure.png" pageId="5" pageNumber="162" startId="5.[151,250,1930,1954]" targetBox="[151,1436,181,1907]" targetPageId="5">
<paragraph id="8BFD36F6136A7A37B59245A8B610F87F" blockId="5.[151,1436,1930,1991]" pageId="5" pageNumber="162">
<emphasis id="B936EAE4136A7A37B59245A8B424F81B" bold="true" box="[151,269,1930,1954]" pageId="5" pageNumber="162">FIGURE 1</emphasis>
.
<taxonomicName id="4C424D75136A7A37B41F45A9B7EEF81A" authority="Zarkani &amp; Kaydan" authorityName="Zarkani &amp; Kaydan" authorityYear="2023" box="[282,711,1931,1955]" class="Insecta" family="Pseudococcidae" genus="Dysmicoccus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="162" phylum="Arthropoda" rank="species" species="zeynepae" status="sp. nov.">
<emphasis id="B936EAE4136A7A37B41F45A9B72AF81B" box="[282,515,1931,1954]" italics="true" pageId="5" pageNumber="162">Dysmicoccus zeynepae</emphasis>
Zarkani &amp; Kaydan
</taxonomicName>
<emphasis id="B936EAE4136A7A37B7CB45A9B62EF81A" bold="true" box="[718,775,1931,1955]" pageId="5" pageNumber="162">
<taxonomicNameLabel id="A205579F136A7A37B7CB45A9B62EF81A" box="[718,775,1931,1955]" pageId="5" pageNumber="162" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
on: (A)
<taxonomicName id="4C424D75136A7A37B65B45A9B15BF81A" authority="Corr." authorityName="Corr." box="[862,1138,1931,1955]" class="Magnoliopsida" family="Meliaceae" genus="Lansium" kingdom="Plantae" order="Sapindales" pageId="5" pageNumber="162" phylum="Tracheophyta" rank="species" species="parasiticum">
<emphasis id="B936EAE4136A7A37B65B45A9B11DF81B" box="[862,1076,1931,1954]" italics="true" pageId="5" pageNumber="162">Lansium parasiticum</emphasis>
Corr.
</taxonomicName>
; (B)
<taxonomicName id="4C424D75136A7A37B1A145A9B5C3F87E" authority="Lindl. ex De Will" authorityName="De Will" class="Magnoliopsida" family="Rubiaceae" genus="Coffea" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="162" phylum="Tracheophyta" rank="species" species="robusta">
<emphasis id="B936EAE4136A7A37B1A145A9B013F81A" box="[1188,1338,1931,1955]" italics="true" pageId="5" pageNumber="162">Coffea robusta</emphasis>
Lindl. ex De Will
</taxonomicName>
; (C)
<taxonomicName id="4C424D75136A7A37B419458DB4F8F87F" box="[284,465,1967,1991]" class="Magnoliopsida" family="Sapotaceae" genus="Manilkara" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="162" phylum="Tracheophyta" rank="species" species="zapota">
<emphasis id="B936EAE4136A7A37B419458DB4F8F87F" box="[284,465,1967,1991]" italics="true" pageId="5" pageNumber="162">Manilkara zapota</emphasis>
</taxonomicName>
L.; and (D)
<taxonomicName id="4C424D75136A7A37B74A458DB610F87F" authority="Murr." authorityName="Murr." box="[591,825,1967,1991]" class="Magnoliopsida" family="Malvaceae" genus="Durio" kingdom="Plantae" order="Malvales" pageId="5" pageNumber="162" phylum="Tracheophyta" rank="species" species="zibethinus">
<emphasis id="B936EAE4136A7A37B74A458DB7D0F87E" box="[591,761,1967,1991]" italics="true" pageId="5" pageNumber="162">Durio zibethinus</emphasis>
Murr.
</taxonomicName>
</paragraph>
</caption>
<caption id="DF3D667E13697A34B59245B5B6D5F809" ID-DOI="http://doi.org/10.5281/zenodo.7532291" ID-Zenodo-Dep="7532291" box="[151,1020,1943,1968]" httpUri="https://zenodo.org/record/7532291/files/figure.png" pageId="6" pageNumber="163" startId="6.[151,250,1943,1967]" targetBox="[184,1396,195,1910]" targetPageId="6">
<paragraph id="8BFD36F613697A34B59245B5B6D5F809" blockId="6.[151,1020,1943,1968]" box="[151,1020,1943,1968]" pageId="6" pageNumber="163">
<emphasis id="B936EAE413697A34B59245B5B424F816" bold="true" box="[151,269,1943,1967]" pageId="6" pageNumber="163">FIGURE 2</emphasis>
. Adult female
<taxonomicName id="4C424D7513697A34B4A645BAB678F809" authority="Zarkani &amp; Kaydan" authorityName="Zarkani &amp; Kaydan" authorityYear="2023" box="[419,849,1944,1968]" class="Insecta" family="Pseudococcidae" genus="Dysmicoccus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="163" phylum="Arthropoda" rank="species" species="zeynepae" status="sp. nov.">
<emphasis id="B936EAE413697A34B4A645BAB7A5F816" box="[419,652,1944,1967]" italics="true" pageId="6" pageNumber="163">Dysmicoccus zeynepae</emphasis>
Zarkani &amp; Kaydan
</taxonomicName>
<emphasis id="B936EAE413697A34B65245BAB6B9F809" bold="true" box="[855,912,1944,1968]" pageId="6" pageNumber="163">
<taxonomicNameLabel id="A205579F13697A34B65245BAB6B9F809" box="[855,912,1944,1968]" pageId="6" pageNumber="163" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype.
</paragraph>
</caption>
<paragraph id="8BFD36F613687A35B5C24001B790FD74" blockId="7.[151,1437,150,1401]" pageId="7" pageNumber="164">
<emphasis id="B936EAE413687A35B5C24001B400FD85" box="[199,297,547,572]" italics="true" pageId="7" pageNumber="164">Dorsum.</emphasis>
Derm membranous, with 16 pairs of cerarii around body margin, each cerarius with 4–7 enlarged conical setae and 3–5 auxiliary setae. Each anal lobe cerarius set on membranous cuticle and containing 5–7 enlarged conical setae 25–35 μm long, plus 40–45 trilocular pores and 3–5 hair-like auxiliary setae. Dorsal setae flagellate, each 15–75 μm long, scattered throughout dorsum. Trilocular pores, each 3–4 μm in diameter, scattered. Multilocular disc pores and tubular ducts absent.
</paragraph>
<paragraph id="8BFD36F613687A35B5C240F5B456FCE4" blockId="7.[151,1437,150,1401]" pageId="7" pageNumber="164">
<emphasis id="B936EAE413687A35B5C240F5B43AFD49" box="[199,275,727,752]" italics="true" pageId="7" pageNumber="164">Venter.</emphasis>
Setae flagellate, each 30–110 μm long, longest setae located medially on head.Apical setae on anal lobes unusually short, each 80–90 μm long. Multilocular disc pores, each 7–8 μm in diameter, present only immediately around vulva, numbering 8–10. Trilocular pores, each 2.5–3.0 μm across, scattered throughout venter. Oral collar tubular ducts absent.
</paragraph>
</subSubSection>
<subSubSection id="C358657D13687A35B5C24144B757FB7C" pageId="7" pageNumber="164" type="discussion">
<paragraph id="8BFD36F613687A35B5C24144B757FB7C" blockId="7.[151,1437,150,1401]" pageId="7" pageNumber="164">
<emphasis id="B936EAE413687A35B5C24144B466FC39" bold="true" box="[199,335,870,896]" pageId="7" pageNumber="164">Comments.</emphasis>
<taxonomicName id="4C424D7513687A35B45C4145B774FC39" authorityName="Zarkani &amp; Kaydan" authorityYear="2023" box="[345,605,871,896]" class="Insecta" family="Pseudococcidae" genus="Dysmicoccus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="zeynepae">
<emphasis id="B936EAE413687A35B45C4145B774FC39" box="[345,605,871,896]" italics="true" pageId="7" pageNumber="164">Dysmicoccus zeynepae</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C424D7513687A35B63C4145B6E9FC38" authorityName="Williams" authorityYear="1994" box="[825,960,871,897]" class="Insecta" family="Pseudococcidae" genus="Dysmicoccus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="finitimus">
<emphasis id="B936EAE413687A35B63C4145B6E9FC38" box="[825,960,871,897]" italics="true" pageId="7" pageNumber="164">D. finitimus</emphasis>
</taxonomicName>
in having anal lobe and other abdominal cerarii each containing a group of about 2‒7 conical setae. However,
<taxonomicName id="4C424D7513687A35B6C641A9B166FC1D" authorityName="Zarkani &amp; Kaydan" authorityYear="2023" box="[963,1103,907,932]" class="Insecta" family="Pseudococcidae" genus="Dysmicoccus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="zeynepae">
<emphasis id="B936EAE413687A35B6C641A9B166FC1D" box="[963,1103,907,932]" italics="true" pageId="7" pageNumber="164">D. zeynepae</emphasis>
</taxonomicName>
can be readily distinguished from
<taxonomicName id="4C424D7513687A35B5D3418DB475FC70" authorityName="Williams" authorityYear="1994" box="[214,348,943,969]" class="Insecta" family="Pseudococcidae" genus="Dysmicoccus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="finitimus">
<emphasis id="B936EAE413687A35B5D3418DB475FC70" box="[214,348,943,969]" italics="true" pageId="7" pageNumber="164">D. finitimus</emphasis>
</taxonomicName>
in having: (i) no multilocular disc pores and oral collar tubular ducts on dorsum; and (ii) a few multilocular disc pores without oral collar tubular ducts on venter. It is also close to
<taxonomicName id="4C424D7513687A35B16541F1B1CAFC54" box="[1120,1251,979,1005]" class="Insecta" family="Pseudococcidae" genus="Dysmicoccus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="lepelleyi">
<emphasis id="B936EAE413687A35B16541F1B1CAFC54" box="[1120,1251,979,1005]" italics="true" pageId="7" pageNumber="164">D. lepelleyi</emphasis>
</taxonomicName>
in having small legs and translucent pores on hind coxa and femur, but
<taxonomicName id="4C424D7513687A35B60341D5B6B9FBA9" authorityName="Zarkani &amp; Kaydan" authorityYear="2023" box="[774,912,1015,1040]" class="Insecta" family="Pseudococcidae" genus="Dysmicoccus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="zeynepae">
<emphasis id="B936EAE413687A35B60341D5B6B9FBA9" box="[774,912,1015,1040]" italics="true" pageId="7" pageNumber="164">D. zeynepae</emphasis>
</taxonomicName>
lacks oral collar tubular ducts on both dorsum and venter.
<taxonomicName id="4C424D7513687A35B41C4639B732FB8D" authorityName="Zarkani &amp; Kaydan" authorityYear="2023" box="[281,539,1051,1076]" class="Insecta" family="Pseudococcidae" genus="Dysmicoccus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="zeynepae">
<emphasis id="B936EAE413687A35B41C4639B732FB8D" box="[281,539,1051,1076]" italics="true" pageId="7" pageNumber="164">Dysmicoccus zeynepae</emphasis>
</taxonomicName>
is also close to
<taxonomicName id="4C424D7513687A35B7C84639B6D8FB8C" authority="Williams" authorityName="Williams" box="[717,1009,1051,1077]" class="Insecta" family="Pseudococcidae" genus="Dysmicoccus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="castanopseus">
<emphasis id="B936EAE413687A35B7C84639B6ADFB8D" box="[717,900,1051,1076]" italics="true" pageId="7" pageNumber="164">D. castanopseus</emphasis>
Williams
</taxonomicName>
in lacking oral collar tubular ducts on dorsum and venter; however, it can be readily distinguished by having (character state for
<taxonomicName id="4C424D7513687A35B198461DB07CFBE1" box="[1181,1365,1087,1112]" class="Insecta" family="Pseudococcidae" genus="Dysmicoccus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="castanopseus">
<emphasis id="B936EAE413687A35B198461DB07CFBE1" box="[1181,1365,1087,1112]" italics="true" pageId="7" pageNumber="164">D. castanopseus</emphasis>
</taxonomicName>
given in parentheses): (i) small, stout legs with tibia + tarsus obviously shorter than trochanter + femur (large legs with tibia + tarsus very slightly longer than trochanter + femur); and (ii) translucent pores on the hind coxa and femur (translucent pores on hind femur and tibia).
</paragraph>
</subSubSection>
<subSubSection id="C358657D13687A35B5C246EDB623FAB4" pageId="7" pageNumber="164" type="etymology">
<paragraph id="8BFD36F613687A35B5C246EDB623FAB4" blockId="7.[151,1437,150,1401]" pageId="7" pageNumber="164">
<emphasis id="B936EAE413687A35B5C246EDB467FB50" bold="true" box="[199,334,1231,1257]" pageId="7" pageNumber="164">Etymology.</emphasis>
This species is named after Zeynep Kaydan (“mother” of Kaydan’s Laboratory), Zeynep Ģleç and Zeynep Kaya who are good friends of the Kaydan’s lab.
</paragraph>
</subSubSection>
<subSubSection id="C358657D13687A35B5C24735B774FAEC" pageId="7" pageNumber="164" type="biology_ecology">
<paragraph id="8BFD36F613687A35B5C24735B774FAEC" blockId="7.[151,1437,150,1401]" pageId="7" pageNumber="164">
<emphasis id="B936EAE413687A35B5C24735B47FFA88" bold="true" box="[199,342,1303,1329]" pageId="7" pageNumber="164">Host plants.</emphasis>
<taxonomicName id="4C424D7513687A35B4594735B784FA88" box="[348,685,1303,1329]" class="Magnoliopsida" family="Malvaceae" genus="Durio" kingdom="Plantae" order="Malvales" pageId="7" pageNumber="164" phylum="Tracheophyta" rank="species" species="zibethinus">
<emphasis id="B936EAE413687A35B4594735B731FA88" box="[348,536,1303,1329]" italics="true" pageId="7" pageNumber="164">Durio zibethinus</emphasis>
(Malvaceae)
</taxonomicName>
,
<taxonomicName id="4C424D7513687A35B7B24735B11AFA88" box="[695,1075,1303,1329]" class="Magnoliopsida" family="Meliaceae" genus="Lansium" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="164" phylum="Tracheophyta" rank="species" species="parasiticum">
<emphasis id="B936EAE413687A35B7B24735B68DFA89" box="[695,932,1303,1328]" italics="true" pageId="7" pageNumber="164">Lansium parasiticum</emphasis>
(Meliaceae)
</taxonomicName>
,
<taxonomicName id="4C424D7513687A35B13B4735B0B4FA88" box="[1086,1437,1303,1329]" class="Magnoliopsida" family="Sapotaceae" genus="Manilkara" kingdom="Plantae" order="Ericales" pageId="7" pageNumber="164" phylum="Tracheophyta" rank="species" species="zapota">
<emphasis id="B936EAE413687A35B13B4735B02CFA89" box="[1086,1285,1303,1329]" italics="true" pageId="7" pageNumber="164">Manilkara zapota</emphasis>
(Sapotaceae)
</taxonomicName>
and
<taxonomicName id="4C424D7513687A35B5C24719B4D2FAEC" box="[199,507,1339,1365]" class="Magnoliopsida" family="Rubiaceae" genus="Coffea" kingdom="Plantae" order="Gentianales" pageId="7" pageNumber="164" phylum="Tracheophyta" rank="species" species="robusta">
<emphasis id="B936EAE413687A35B5C24719B444FAEC" box="[199,365,1339,1365]" italics="true" pageId="7" pageNumber="164">Coffea robusta</emphasis>
(Rubiaceae)
</taxonomicName>
(
<figureCitation id="13792A7313687A35B70F4719B779FAEC" box="[522,592,1339,1365]" captionStart="FIGURE 2" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[184,1396,195,1910]" captionTargetId="figure-19@6.[171,1416,181,1919]" captionTargetPageId="6" captionText="FIGURE 2. Adult female Dysmicoccus zeynepae Zarkani &amp; Kaydan sp. n., holotype." figureDoi="http://doi.org/10.5281/zenodo.7532291" httpUri="https://zenodo.org/record/7532291/files/figure.png" pageId="7" pageNumber="164">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C358657D13687A35B5C2477CB679FAC0" box="[199,848,1374,1401]" pageId="7" pageNumber="164" type="distribution">
<paragraph id="8BFD36F613687A35B5C2477CB679FAC0" blockId="7.[151,1437,150,1401]" box="[199,848,1374,1401]" pageId="7" pageNumber="164">
<emphasis id="B936EAE413687A35B5C2477CB449FAC0" bold="true" box="[199,352,1374,1401]" pageId="7" pageNumber="164">Distribution.</emphasis>
<collectingCountry id="F355766613687A35B462477DB4FDFAC0" box="[359,468,1375,1401]" name="Indonesia" pageId="7" pageNumber="164">Indonesia</collectingCountry>
(
<collectingRegion id="4986F81413687A35B4E6477DB76BFAC0" box="[483,578,1375,1401]" country="Indonesia" name="Sumatera Utara" pageId="7" pageNumber="164">Sumatra</collectingRegion>
I.,
<collectingRegion id="4986F81413687A35B762477DB66DFAC0" box="[615,836,1375,1401]" country="Indonesia" name="Bengkulu" pageId="7" pageNumber="164">Bengkulu Province</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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