275 lines
34 KiB
XML
275 lines
34 KiB
XML
<document id="9E1EE846600EBF828E4B82D860B3214C" ID-CLB-Dataset="29106" ID-DOI="10.11646/zootaxa.4455.2.2" ID-GBIF-Dataset="431a0907-2b69-4c52-86c6-6a0eb0c04d2e" ID-ISSN="1175-5326" ID-Zenodo-Dep="1457167" ID-ZooBank="595A9780-C604-465C-BDE2-589B87A4DB52" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539261406172" checkinUser="plazi" docAuthor="Antić, Dragan Ž., Dražina, Tvrtko, Rađa, Tonći, Lučić, Luka R. & Makarov, Slobodan E." docDate="2018" docId="0383AF242004FFE314DFFB98FBD1FDDC" docLanguage="en" docName="zootaxa.4455.2.2.pdf" docOrigin="Zootaxa 4455 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Typhloiulus clavatus Antić & Dražina & Rađa & Lučić & Makarov 2018, new species" docType="treatment" docVersion="7" lastPageNumber="271" masterDocId="FFBAD75C2008FFEE1448FFEDFFC3FFBC" masterDocTitle="Review of the genus Typhloiulus Latzel, 1884 in the Dinaric region, with a description of four new species and the first description of the male of Typhloiulus insularis Strasser, 1938 (Diplopoda: Julida: Julidae)" masterLastPageNumber="294" masterPageNumber="258" pageNumber="270" updateTime="1698575520044" updateUser="ExternalLinkService">
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<mods:mods id="9EC3D5B95D23F3149F5909A9ABC35B27" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="AB288E7867775CAB1B6DA774D5BC8873">
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<mods:title id="A394F04446722845BE85E0D0E26E377D">Review of the genus Typhloiulus Latzel, 1884 in the Dinaric region, with a description of four new species and the first description of the male of Typhloiulus insularis Strasser, 1938 (Diplopoda: Julida: Julidae)</mods:title>
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<mods:name id="EAE5E078A52E8D3044809E44873550C2" type="personal">
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<mods:roleTerm id="4091B9E334A3B14A6B5AAFC02806D2B7">Author</mods:roleTerm>
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<mods:namePart id="6A4225066A955D19D55D95A24A045F88">Antić, Dragan Ž.</mods:namePart>
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</mods:name>
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<mods:name id="5EEF4B6FA18ADEA08EF59233FBE741DE" type="personal">
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<mods:roleTerm id="277817581EA6ABA6E4CB65F45959E854">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="943ADC0D2897A2321F1F593656FCCFDC">Dražina, Tvrtko</mods:namePart>
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<mods:roleTerm id="CE13801AB6DFF1DA28A324605FB01A7B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="94CC2CBE01AEE371C956578631ACF813">Rađa, Tonći</mods:namePart>
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<mods:roleTerm id="8E858145022D131BBAAE775CEBB932EF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E1E6EB5C32C5AC09A3D4B722C239555A">Lučić, Luka R.</mods:namePart>
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<mods:name id="31C658D685196068AF1FF0C8A3F7A369" type="personal">
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<mods:roleTerm id="2888E53703ABB4EB2BEE9AD2A52D025B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="38FE3CA91C7607ED8A3984F05E34B8FF">Makarov, Slobodan E.</mods:namePart>
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<mods:typeOfResource id="D17CC2099374F204273EB0D6ECA666B1">text</mods:typeOfResource>
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<mods:title id="A0FCA288300906475A7FA8EC4B314004">Zootaxa</mods:title>
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<mods:part id="27CF9372E3D96219D2160FD30F7ED089">
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<mods:date id="50BA39F4BB9EF378AD23F77753B0AB3A">2018</mods:date>
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<mods:number id="A92301A1E1685CE4F88163EE96627BB4">2018-08-02</mods:number>
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<mods:detail id="2C068CA781B0F5388500C9B2E0AF5C0F" type="volume">
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<mods:number id="0EE241F748F563297FD7F00BBB8F0C17">4455</mods:number>
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<mods:identifier id="B4D56238F4EC13B114025347E260EB2E" type="DOI">10.11646/zootaxa.4455.2.2</mods:identifier>
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<mods:identifier id="EDC46092BCB054819A28292ADF0B97B8" type="ISSN">1175-5326</mods:identifier>
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<treatment id="0383AF242004FFE314DFFB98FBD1FDDC" ID-DOI="http://doi.org/10.5281/zenodo.5960762" ID-GBIF-Taxon="148691219" ID-Zenodo-Dep="5960762" LSID="urn:lsid:plazi:treatment:0383AF242004FFE314DFFB98FBD1FDDC" httpUri="http://treatment.plazi.org/id/0383AF242004FFE314DFFB98FBD1FDDC" lastPageId="13" lastPageNumber="271" pageId="12" pageNumber="270">
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<subSubSection id="C3304DB92004FFE214DFFB98FE87FB0C" pageId="12" pageNumber="270" type="nomenclature">
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<paragraph id="8B951E322004FFE214DFFB98FDB8FB33" blockId="12.[151,635,1141,1200]" box="[151,635,1141,1167]" pageId="12" pageNumber="270">
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<heading id="D0DDA95E2004FFE214DFFB98FDB8FB33" bold="true" box="[151,635,1141,1167]" fontSize="11" level="1" pageId="12" pageNumber="270" reason="1">
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<emphasis id="B95EC2202004FFE214DFFB98FDB8FB33" bold="true" box="[151,635,1141,1167]" pageId="12" pageNumber="270">
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<taxonomicName id="4C2A65B12004FFE214DFFB98FE1CFB33" ID-CoL="84DYH" authority="Antic" authorityName="Antić & Dražina & Rađa & Lučić & Makarov" authorityYear="2018" box="[151,479,1141,1167]" class="Diplopoda" family="Julidae" genus="Typhloiulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="12" pageNumber="270" phylum="Arthropoda" rank="species" species="clavatus" status="sp. nov.">
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||
<emphasis id="B95EC2202004FFE214DFFB98FE52FB33" bold="true" box="[151,401,1141,1167]" italics="true" pageId="12" pageNumber="270">Typhloiulus clavatus</emphasis>
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Antić
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</taxonomicName>
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||
,
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<taxonomicNameLabel id="A26D7F5B2004FFE215A4FB98FDB8FB33" box="[492,635,1141,1167]" pageId="12" pageNumber="270" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B951E322004FFE214DFFB7AFE87FB0C" blockId="12.[151,635,1141,1200]" box="[151,324,1175,1200]" pageId="12" pageNumber="270">
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<figureCitation id="131102B72004FFE214DFFB7AFF1DFB0C" box="[151,222,1175,1200]" captionStart="FIGURE 9" captionStartId="15.[151,250,1921,1943]" captionTargetBox="[151,1436,193,1899]" captionTargetId="figure@15.[151,1436,193,1899]" captionTargetPageId="15" captionText="FIGURE 9. Typhloiulus clavatus sp. nov. (NHMSC). A. Paratype female, habitus, lateral view. B. Paratype female, anterior part of the body, lateral view. C. Paratype female, telson, lateral view. D. Paratype female, midbody rings, lateral view. E. Holotype male, body rings 6 and 7, lateral view. Arrow indicates ventral margin of body ring 7. Scale lines: 1 mm." httpUri="https://zenodo.org/record/1457187/files/figure.png" pageId="12" pageNumber="270">Figs 9</figureCitation>
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,
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<figureCitation id="131102B72004FFE214A2FB75FECBFB0C" box="[234,264,1176,1200]" captionStart="FIGURE 10" captionStartId="16.[151,250,1894,1916]" captionTargetBox="[164,1383,238,1796]" captionTargetId="figure@16.[151,1436,193,1872]" captionTargetPageId="16" captionText="FIGURE 10. Typhloiulus clavatus sp. nov., holotype male (NHMSC). A. First pair of legs, anterior view. B. Penis, posterior view. C. Promere, posterior view. D. Gonopods, mesal view. E. Mesomere and opisthomere, mesal view. F. Gonopods, lateral view. Abbreviations: l: lamella; m: mesomere; o: opisthomere; p: promere; s: solenomere; t: telopodite; v: velum; ml: mesal lobe. Scale line: 0.3 mm." httpUri="https://zenodo.org/record/1457189/files/figure.png" pageId="12" pageNumber="270">10</figureCitation>
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,
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<figureCitation id="131102B72004FFE2155CFB7AFE87FB0C" box="[276,324,1175,1200]" captionStart="FIGURE 15" captionStartId="23.[151,250,1539,1561]" captionTargetBox="[199,1385,193,1520]" captionTargetId="figure@23.[199,1385,193,1520]" captionTargetPageId="23" captionText="FIGURE 15. Gonopods of representatives of the subgenus Inversotyphlus Strasser, 1962, mesal view. A. Typhloiulus clavatus sp. nov., holotype male (NHMSC). B. Typhloiulus gracilis sp. nov., paratype male (IZB). C. Typhloiulus opisthonodus sp. nov., holotype male (NHMSC). D. Typhloiulus lobifer Attems, 1951, male from Bitumen mine Minjera, island of Brač. E. Typhloiulus gellianae Makarov & Rađa, 2006, holotype male (ZDR). Scale lines: 0.1 mm." httpUri="https://zenodo.org/record/1457199/files/figure.png" pageId="12" pageNumber="270">15A</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3304DB92004FFE214DFFB32FA55FAA1" pageId="12" pageNumber="270" type="materials_examined">
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<paragraph id="8B951E322004FFE214DFFB32FA55FAA1" blockId="12.[151,1437,1247,2029]" pageId="12" pageNumber="270">
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<materialsCitation id="3B42146F2004FFE214DFFB32FC61FAA0" ID-GBIF-Occurrence="1917075799" collectingDate="2016-04-30" collectionCode="NHMSC" collectorName="T. Rada" country="Croatia" county="Klenovac Pit" latitude="43.7081" location="Material" longLatPrecision="1" longitude="16.776684" municipality="Strazbenica" pageId="12" pageNumber="270" specimenCount="2" typeStatus="holotype">
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<emphasis id="B95EC2202004FFE214DFFB32FEBCFB44" bold="true" box="[151,383,1247,1272]" pageId="12" pageNumber="270">
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<location id="8EF548E92004FFE214DFFB32FF3CFB44" LSID="urn:lsid:plazi:treatment:0383AF242004FFE314DFFB98FBD1FDDC:8EF548E92004FFE214DFFB32FF3CFB44" box="[151,255,1247,1272]" country="Croatia" county="Klenovac Pit" latitude="43.7081" longLatPrecision="1" longitude="16.776684" municipality="Strazbenica" name="Material" pageId="12" pageNumber="270">Material</location>
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examined
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||
</emphasis>
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:
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<location id="8EF548E92004FFE215DBFB0DFE08FB44" LSID="urn:lsid:plazi:treatment:0383AF242004FFE314DFFB98FBD1FDDC:8EF548E92004FFE215DBFB0DFE08FB44" box="[403,459,1248,1272]" country="Croatia" county="Klenovac Pit" latitude="43.7081" longLatPrecision="1" longitude="16.776684" municipality="Strazbenica" name="Type" pageId="12" pageNumber="270">
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<typeStatus id="5491A0902004FFE215DBFB0DFE08FB44" box="[403,459,1248,1272]" pageId="12" pageNumber="270">Type</typeStatus>
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</location>
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specimens.
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||
<typeStatus id="5491A0902004FFE21617FB32FD04FB44" box="[607,711,1247,1272]" pageId="12" pageNumber="270" type="holotype">Holotype</typeStatus>
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||
male,
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<collectingCountry id="F33D5EA22004FFE21755FB0DFC5AFB44" box="[797,921,1248,1272]" name="Croatia" pageId="12" pageNumber="270">CROATIA</collectingCountry>
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,
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<location id="8EF548E92004FFE217E2FB32FBD7FB44" LSID="urn:lsid:plazi:treatment:0383AF242004FFE314DFFB98FBD1FDDC:8EF548E92004FFE217E2FB32FBD7FB44" box="[938,1044,1247,1272]" country="Croatia" county="Klenovac Pit" latitude="43.7081" longLatPrecision="1" longitude="16.776684" municipality="Strazbenica" name="Dalmatia" pageId="12" pageNumber="270">Dalmatia</location>
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,
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<location id="8EF548E92004FFE2106CFB32FB9CFB44" LSID="urn:lsid:plazi:treatment:0383AF242004FFE314DFFB98FBD1FDDC:8EF548E92004FFE2106CFB32FB9CFB44" box="[1060,1119,1247,1272]" country="Croatia" county="Klenovac Pit" latitude="43.7081" longLatPrecision="1" longitude="16.776684" municipality="Strazbenica" name="Otok" pageId="12" pageNumber="270">Otok</location>
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,
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<collectingMunicipality id="6BF184482004FFE21038FB32FB36FB44" box="[1136,1269,1247,1272]" pageId="12" pageNumber="270">Stražbenica</collectingMunicipality>
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,
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<collectingCounty id="62F466BE2004FFE2114DFB32FA5FFB44" box="[1285,1436,1247,1272]" pageId="12" pageNumber="270">Klenovac Pit</collectingCounty>
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(
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<geoCoordinate id="EE1E78F52004FFE214D6FAE9FEF9FAA0" box="[158,314,1284,1308]" direction="north" orientation="latitude" pageId="12" pageNumber="270" precision="1" value="43.7081">43.708100°N</geoCoordinate>
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,
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<geoCoordinate id="EE1E78F52004FFE2150DFAE9FE1FFAA1" box="[325,476,1284,1309]" direction="east" orientation="longitude" pageId="12" pageNumber="270" precision="1" value="16.776684">16.776683°E</geoCoordinate>
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),
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<collectingDate id="EFD0C11A2004FFE215A7FAE9FD4CFAA1" box="[495,655,1284,1309]" pageId="12" pageNumber="270" value="2016-04-30">
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<date id="FF9438F22004FFE215A7FAE9FD4CFAA1" box="[495,655,1284,1309]" pageId="12" pageNumber="270" value="2016-04-30">30 April 2016</date>
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</collectingDate>
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,
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<collectorName id="26DF7BE42004FFE216D2FAE8FD33FAA1" box="[666,752,1284,1309]" pageId="12" pageNumber="270">T. Rađa</collectorName>
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leg. (
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||
<collectionCode id="ED3B86F72004FFE21778FAE9FC59FAA0" box="[816,922,1284,1308]" pageId="12" pageNumber="270">NHMSC</collectionCode>
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)
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</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B42146F2004FFE217E4FAE8FA51FAA1" ID-GBIF-Occurrence="1917075817" box="[940,1426,1284,1309]" collectingDate="2016-04-30" collectionCode="NHMSC" collectorName="T. Rada" country="Croatia" county="Klenovac Pit" latitude="43.7081" location="Material" longLatPrecision="1" longitude="16.776684" municipality="Strazbenica" pageId="12" pageNumber="270" specimenCount="2" typeStatus="paratype">
|
||
<typeStatus id="5491A0902004FFE217E4FAE8FBCDFAA1" box="[940,1038,1285,1309]" pageId="12" pageNumber="270" type="paratype">Paratype</typeStatus>
|
||
female, same data as for
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||
<typeStatus id="5491A0902004FFE21164FAE9FA51FAA1" box="[1324,1426,1284,1309]" pageId="12" pageNumber="270" type="holotype">holotype</typeStatus>
|
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</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3304DB92004FFE2148FFACAFCFDFA48" pageId="12" pageNumber="270" type="diagnosis">
|
||
<paragraph id="8B951E322004FFE2148FFACAFCFDFA48" blockId="12.[151,1437,1247,2029]" pageId="12" pageNumber="270">
|
||
<emphasis id="B95EC2202004FFE2148FFACAFE82FAFC" bold="true" box="[199,321,1319,1344]" pageId="12" pageNumber="270">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C2A65B12004FFE21501FACAFDF3FAFC" authorityName="Antić & Dražina & Rađa & Lučić & Makarov" authorityYear="2018" box="[329,560,1319,1344]" class="Diplopoda" family="Julidae" genus="Typhloiulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="12" pageNumber="270" phylum="Arthropoda" rank="species" species="clavatus">
|
||
<emphasis id="B95EC2202004FFE21501FACAFDF3FAFC" box="[329,560,1319,1344]" italics="true" pageId="12" pageNumber="270">Typhloiulus clavatus</emphasis>
|
||
</taxonomicName>
|
||
is most similar to members of the subgenus
|
||
<emphasis id="B95EC2202004FFE21065FACAFB1AFAFC" box="[1069,1241,1319,1344]" italics="true" pageId="12" pageNumber="270">
|
||
<taxonomicName id="4C2A65B12004FFE21065FACAFB16FAFC" authorityName="Strasser" authorityYear="1962" box="[1069,1237,1319,1344]" class="Diplopoda" family="Julidae" genus="Inversotyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="12" pageNumber="270" phylum="Arthropoda" rank="genus">Inversotyphlus</taxonomicName>
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||
,
|
||
</emphasis>
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||
viz.
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<taxonomicName id="4C2A65B12004FFE2115AFAC4FABBFAFC" authorityName="Attems" authorityYear="1951" box="[1298,1400,1319,1344]" class="Diplopoda" family="Julidae" genus="Typhloiulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="12" pageNumber="270" phylum="Arthropoda" rank="species" species="lobifer">
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||
<emphasis id="B95EC2202004FFE2115AFAC4FABBFAFC" box="[1298,1400,1319,1344]" italics="true" pageId="12" pageNumber="270">T. lobifer</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C2A65B12004FFE211CFFAC4FF3CFAD9" authorityName="Makarov & Rada" authorityYear="2006" class="Diplopoda" family="Julidae" genus="Typhloiulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="12" pageNumber="270" phylum="Arthropoda" rank="species" species="gellianae">
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||
<emphasis id="B95EC2202004FFE211CFFAC4FF3CFAD9" italics="true" pageId="12" pageNumber="270">T. gellianae</emphasis>
|
||
</taxonomicName>
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||
, and herein described
|
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<taxonomicName id="4C2A65B12004FFE215B5FAA0FDADFAD9" authorityName="Antić & Dražina & Rađa & Lučić & Makarov" authorityYear="2018" box="[509,622,1356,1381]" class="Diplopoda" family="Julidae" genus="Typhloiulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="12" pageNumber="270" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B95EC2202004FFE215B5FAA0FDADFAD9" box="[509,622,1356,1381]" italics="true" pageId="12" pageNumber="270">T. gracilis</emphasis>
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||
</taxonomicName>
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and
|
||
<taxonomicName id="4C2A65B12004FFE216EEFAA0FC94FAD9" authorityName="Antić & Dražina & Rađa & Lučić & Makarov" authorityYear="2018" box="[678,855,1356,1381]" class="Diplopoda" family="Julidae" genus="Typhloiulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="12" pageNumber="270" phylum="Arthropoda" rank="species" species="opisthonodus">
|
||
<emphasis id="B95EC2202004FFE216EEFAA0FC94FAD9" box="[678,855,1356,1381]" italics="true" pageId="12" pageNumber="270">T. opisthonodus</emphasis>
|
||
</taxonomicName>
|
||
. These species are characterized by the presence of a very long opisthomere with a similarly shaped velum; the presence of a poorly developed, but visible opisthomeral lamella; and similar male sexual and habitus structures (see Discussion). The new species differs clearly in gonopod details, such as the presence of a robust promere with the apical half slightly curved anteriad and the presence of a very prominent and clavate mesomere.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3304DB92004FFE3148FFA12FE73FE49" lastPageId="13" lastPageNumber="271" pageId="12" pageNumber="270" type="description">
|
||
<paragraph id="8B951E322004FFE3148FFA12FCDAFE11" blockId="12.[151,1437,1247,2029]" lastBlockId="13.[151,1436,151,608]" lastPageId="13" lastPageNumber="271" pageId="12" pageNumber="270">
|
||
<emphasis id="B95EC2202004FFE2148FFA12FE92F9A4" bold="true" box="[199,337,1535,1560]" pageId="12" pageNumber="270">Description</emphasis>
|
||
. Based on
|
||
<typeStatus id="5491A0902004FFE21587FA12FDF2F9A4" box="[463,561,1535,1560]" pageId="12" pageNumber="270" type="holotype">holotype</typeStatus>
|
||
(male)
|
||
<quantity id="4CD2B3D72004FFE216C1F9EDFD1BF9AB" box="[649,728,1536,1560]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="270" unit="mm" value="20.0">20 mm</quantity>
|
||
long, vertical diameter of the largest body ring
|
||
<quantity id="4CD2B3D72004FFE210BAFA12FA9BF9AB" box="[1266,1368,1535,1560]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="12" pageNumber="270" unit="mm" value="1.35">1.35 mm</quantity>
|
||
, body with 41 podous rings + 3 apodous rings + telson.
|
||
<emphasis id="B95EC2202004FFE216F3F9C9FC8FF981" bold="true" box="[699,844,1572,1597]" pageId="12" pageNumber="270">Colouration</emphasis>
|
||
: (
|
||
<figureCitation id="131102B72004FFE2172AF9C9FC7FF980" box="[866,956,1572,1597]" captionStart="FIGURE 9" captionStartId="15.[151,250,1921,1943]" captionTargetBox="[151,1436,193,1899]" captionTargetId="figure@15.[151,1436,193,1899]" captionTargetPageId="15" captionText="FIGURE 9. Typhloiulus clavatus sp. nov. (NHMSC). A. Paratype female, habitus, lateral view. B. Paratype female, anterior part of the body, lateral view. C. Paratype female, telson, lateral view. D. Paratype female, midbody rings, lateral view. E. Holotype male, body rings 6 and 7, lateral view. Arrow indicates ventral margin of body ring 7. Scale lines: 1 mm." httpUri="https://zenodo.org/record/1457187/files/figure.png" pageId="12" pageNumber="270">Fig. 9A</figureCitation>
|
||
). Brownish.
|
||
<emphasis id="B95EC2202004FFE21001F9C9FB4AF981" bold="true" box="[1097,1161,1572,1597]" pageId="12" pageNumber="270">Head</emphasis>
|
||
: (
|
||
<figureCitation id="131102B72004FFE210D7F9C9FB3BF980" box="[1183,1272,1572,1597]" captionStart="FIGURE 9" captionStartId="15.[151,250,1921,1943]" captionTargetBox="[151,1436,193,1899]" captionTargetId="figure@15.[151,1436,193,1899]" captionTargetPageId="15" captionText="FIGURE 9. Typhloiulus clavatus sp. nov. (NHMSC). A. Paratype female, habitus, lateral view. B. Paratype female, anterior part of the body, lateral view. C. Paratype female, telson, lateral view. D. Paratype female, midbody rings, lateral view. E. Holotype male, body rings 6 and 7, lateral view. Arrow indicates ventral margin of body ring 7. Scale lines: 1 mm." httpUri="https://zenodo.org/record/1457187/files/figure.png" pageId="12" pageNumber="270">Fig. 9B</figureCitation>
|
||
). Labrum with three labral teeth, with 4 supralabral and 7+7 labral setae. Gnathochilarium with promentum kite-shaped, elongated; lamellae linguales with 4+3 setae in one row, stipites each with 3 setae. Antennae
|
||
<quantity id="4CD2B3D72004FFE21081F981FACFF938" box="[1225,1292,1644,1668]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="270" unit="mm" value="2.0">2 mm</quantity>
|
||
long, length 148% of vertical diameter of the largest body ring. Length of antennomeres I–VIII (in mm): 0.12 (I), 0.42 (II), 0.36 (III), 0.31 (IV), 0.36 (V), 0.26 (VI), 0.12 (VII), and 0.05 (VIII). Length/width ratio of antennomeres I–VII: 0.75 (I), 2.8 (II), 2.4 (III), 2 (IV), 2 (V), 1.6 (VI), and 1 (VII). Antennomeres V and VI each with a terminal corolla of large bacilliform sensilla; antennomere VII with a terminal corolla of microsetae.
|
||
<emphasis id="B95EC2202004FFE21046F911FB51F8A9" bold="true" box="[1038,1170,1788,1813]" pageId="12" pageNumber="270">Body rings</emphasis>
|
||
: (
|
||
<figureCitation id="131102B72004FFE210E5F911FAC9F8A8" box="[1197,1290,1788,1813]" captionStart="FIGURE 9" captionStartId="15.[151,250,1921,1943]" captionTargetBox="[151,1436,193,1899]" captionTargetId="figure@15.[151,1436,193,1899]" captionTargetPageId="15" captionText="FIGURE 9. Typhloiulus clavatus sp. nov. (NHMSC). A. Paratype female, habitus, lateral view. B. Paratype female, anterior part of the body, lateral view. C. Paratype female, telson, lateral view. D. Paratype female, midbody rings, lateral view. E. Holotype male, body rings 6 and 7, lateral view. Arrow indicates ventral margin of body ring 7. Scale lines: 1 mm." httpUri="https://zenodo.org/record/1457187/files/figure.png" pageId="12" pageNumber="270">Fig. 9D</figureCitation>
|
||
). The whole metazonal area with longitudinal striations. Metazonal setae fallen off.
|
||
<emphasis id="B95EC2202004FFE2179AF8F2FBDFF884" bold="true" box="[978,1052,1823,1848]" pageId="12" pageNumber="270">Telson</emphasis>
|
||
: (
|
||
<figureCitation id="131102B72004FFE21071F8F2FB56F884" box="[1081,1173,1823,1848]" captionStart="FIGURE 9" captionStartId="15.[151,250,1921,1943]" captionTargetBox="[151,1436,193,1899]" captionTargetId="figure@15.[151,1436,193,1899]" captionTargetPageId="15" captionText="FIGURE 9. Typhloiulus clavatus sp. nov. (NHMSC). A. Paratype female, habitus, lateral view. B. Paratype female, anterior part of the body, lateral view. C. Paratype female, telson, lateral view. D. Paratype female, midbody rings, lateral view. E. Holotype male, body rings 6 and 7, lateral view. Arrow indicates ventral margin of body ring 7. Scale lines: 1 mm." httpUri="https://zenodo.org/record/1457187/files/figure.png" pageId="12" pageNumber="270">Fig. 9C</figureCitation>
|
||
). Epiproct with a long preanal process, strongly curved downwards and covered by long dorsal and lateral setae. Paraprocts rounded, with numerous long setae. Hypoproct lenticular, with five long setae.
|
||
<emphasis id="B95EC2202004FFE217CBF88AFC78F83C" bold="true" box="[899,955,1895,1920]" pageId="12" pageNumber="270">Legs</emphasis>
|
||
: First pair of legs modified, hook-shaped (
|
||
<figureCitation id="131102B72004FFE214D7F861FEC4F818" box="[159,263,1932,1957]" captionStart="FIGURE 10" captionStartId="16.[151,250,1894,1916]" captionTargetBox="[164,1383,238,1796]" captionTargetId="figure@16.[151,1436,193,1872]" captionTargetPageId="16" captionText="FIGURE 10. Typhloiulus clavatus sp. nov., holotype male (NHMSC). A. First pair of legs, anterior view. B. Penis, posterior view. C. Promere, posterior view. D. Gonopods, mesal view. E. Mesomere and opisthomere, mesal view. F. Gonopods, lateral view. Abbreviations: l: lamella; m: mesomere; o: opisthomere; p: promere; s: solenomere; t: telopodite; v: velum; ml: mesal lobe. Scale line: 0.3 mm." httpUri="https://zenodo.org/record/1457189/files/figure.png" pageId="12" pageNumber="270">Fig. 10A</figureCitation>
|
||
); coxa, postfemur, and tibiotarsus each with one seta, prefemur with four or five setae, femur with two or three setae. Tibiotarsus with a small tarsal remnant in the form of a prominent dorsal lobe. The tip of the first pair of legs tuberculate. Second pair of legs with poorly developed ventral pads on the tibia.
|
||
<emphasis id="B95EC2202004FFE21024F839FB6FF851" bold="true" box="[1132,1196,2004,2029]" pageId="12" pageNumber="270">Penis</emphasis>
|
||
: (
|
||
<figureCitation id="131102B72004FFE2108BF839FAE5F850" box="[1219,1318,2004,2029]" captionStart="FIGURE 10" captionStartId="16.[151,250,1894,1916]" captionTargetBox="[164,1383,238,1796]" captionTargetId="figure@16.[151,1436,193,1872]" captionTargetPageId="16" captionText="FIGURE 10. Typhloiulus clavatus sp. nov., holotype male (NHMSC). A. First pair of legs, anterior view. B. Penis, posterior view. C. Promere, posterior view. D. Gonopods, mesal view. E. Mesomere and opisthomere, mesal view. F. Gonopods, lateral view. Abbreviations: l: lamella; m: mesomere; o: opisthomere; p: promere; s: solenomere; t: telopodite; v: velum; ml: mesal lobe. Scale line: 0.3 mm." httpUri="https://zenodo.org/record/1457189/files/figure.png" pageId="12" pageNumber="270">Fig. 10B</figureCitation>
|
||
). Bilobed. Lobes prominent, elongated.
|
||
<emphasis id="B95EC2202005FFE315A1FF7AFCA2FF0C" bold="true" box="[489,865,151,176]" pageId="13" pageNumber="271">Ventral margin of body ring 7</emphasis>
|
||
: (
|
||
<figureCitation id="131102B72005FFE31733FF7AFC17FF0C" box="[891,980,151,176]" captionStart="FIGURE 9" captionStartId="15.[151,250,1921,1943]" captionTargetBox="[151,1436,193,1899]" captionTargetId="figure@15.[151,1436,193,1899]" captionTargetPageId="15" captionText="FIGURE 9. Typhloiulus clavatus sp. nov. (NHMSC). A. Paratype female, habitus, lateral view. B. Paratype female, anterior part of the body, lateral view. C. Paratype female, telson, lateral view. D. Paratype female, midbody rings, lateral view. E. Holotype male, body rings 6 and 7, lateral view. Arrow indicates ventral margin of body ring 7. Scale lines: 1 mm." httpUri="https://zenodo.org/record/1457187/files/figure.png" pageId="13" pageNumber="271">Fig. 9E</figureCitation>
|
||
). Well-developed, tapering distad, with rounded tip. Posterior margin right-angled to metazona.
|
||
<emphasis id="B95EC2202005FFE31701FF51FC01FF69" bold="true" box="[841,962,188,213]" pageId="13" pageNumber="271">Gonopods</emphasis>
|
||
: (
|
||
<figureCitation id="131102B72005FFE317A9FF51FBACFF68" box="[993,1135,188,213]" captionStart="FIGURE 10" captionStartId="16.[151,250,1894,1916]" captionTargetBox="[164,1383,238,1796]" captionTargetId="figure@16.[151,1436,193,1872]" captionTargetPageId="16" captionText="FIGURE 10. Typhloiulus clavatus sp. nov., holotype male (NHMSC). A. First pair of legs, anterior view. B. Penis, posterior view. C. Promere, posterior view. D. Gonopods, mesal view. E. Mesomere and opisthomere, mesal view. F. Gonopods, lateral view. Abbreviations: l: lamella; m: mesomere; o: opisthomere; p: promere; s: solenomere; t: telopodite; v: velum; ml: mesal lobe. Scale line: 0.3 mm." httpUri="https://zenodo.org/record/1457189/files/figure.png" pageId="13" pageNumber="271">Figs 10C–F</figureCitation>
|
||
,
|
||
<figureCitation id="131102B72005FFE310CBFF51FB77FF69" box="[1155,1204,188,213]" captionStart="FIGURE 15" captionStartId="23.[151,250,1539,1561]" captionTargetBox="[199,1385,193,1520]" captionTargetId="figure@23.[199,1385,193,1520]" captionTargetPageId="23" captionText="FIGURE 15. Gonopods of representatives of the subgenus Inversotyphlus Strasser, 1962, mesal view. A. Typhloiulus clavatus sp. nov., holotype male (NHMSC). B. Typhloiulus gracilis sp. nov., paratype male (IZB). C. Typhloiulus opisthonodus sp. nov., holotype male (NHMSC). D. Typhloiulus lobifer Attems, 1951, male from Bitumen mine Minjera, island of Brač. E. Typhloiulus gellianae Makarov & Rađa, 2006, holotype male (ZDR). Scale lines: 0.1 mm." httpUri="https://zenodo.org/record/1457199/files/figure.png" pageId="13" pageNumber="271">15A</figureCitation>
|
||
). Promere (
|
||
<emphasis id="B95EC2202005FFE31103FF51FA99FF69" bold="true" box="[1355,1370,188,213]" pageId="13" pageNumber="271">p</emphasis>
|
||
) and mesomere (
|
||
<emphasis id="B95EC2202005FFE31554FF32FEF0FF44" bold="true" box="[284,307,223,248]" pageId="13" pageNumber="271">m</emphasis>
|
||
) robust, both shorter than opisthomere (
|
||
<emphasis id="B95EC2202005FFE3174AFF32FCD3FF44" bold="true" box="[770,784,223,248]" pageId="13" pageNumber="271">o</emphasis>
|
||
). Promere (
|
||
<emphasis id="B95EC2202005FFE317D2FF32FC6AFF44" bold="true" box="[922,937,223,248]" pageId="13" pageNumber="271">p</emphasis>
|
||
) shorter than
|
||
<emphasis id="B95EC2202005FFE31003FF32FBA1FF44" bold="true" box="[1099,1122,223,248]" pageId="13" pageNumber="271">m</emphasis>
|
||
; apical half slightly curved anteriad; laterally in the form of a shield; mesal lobe (
|
||
<emphasis id="B95EC2202005FFE316A4FEE9FCC8FEA1" bold="true" box="[748,779,260,285]" pageId="13" pageNumber="271">ml</emphasis>
|
||
) wide, subquadrangular, with three apical setae; telopodite (
|
||
<emphasis id="B95EC2202005FFE314E8FECAFF6AFEFC" bold="true" box="[160,169,295,320]" pageId="13" pageNumber="271">t</emphasis>
|
||
) subcylindrical, tapering distad, over high
|
||
<emphasis id="B95EC2202005FFE316E9FECAFD03FEFC" bold="true" box="[673,704,295,320]" pageId="13" pageNumber="271">ml</emphasis>
|
||
. Mesomere (
|
||
<emphasis id="B95EC2202005FFE31713FECAFCB1FEFC" bold="true" box="[859,882,295,320]" pageId="13" pageNumber="271">m</emphasis>
|
||
) clavate in lateral/mesal view, subapically with anterior tuberculation. Opisthomere (
|
||
<emphasis id="B95EC2202005FFE3160DFEA1FD90FED9" bold="true" box="[581,595,332,357]" pageId="13" pageNumber="271">o</emphasis>
|
||
) slender, straight, and elongated; antero-mesally with poorly developed lamella (
|
||
<emphasis id="B95EC2202005FFE314B4FE82FEC7FE34" bold="true" box="[252,260,367,392]" pageId="13" pageNumber="271">l</emphasis>
|
||
); mesally with one spine; velum (
|
||
<emphasis id="B95EC2202005FFE316DBFE82FD62FE34" bold="true" box="[659,673,367,392]" pageId="13" pageNumber="271">v</emphasis>
|
||
) unipartite, well-developed, tapering anteriad, with fimbriated tip. Solenomere (
|
||
<emphasis id="B95EC2202005FFE31565FE79FEFBFE11" bold="true" box="[301,312,404,429]" pageId="13" pageNumber="271">s</emphasis>
|
||
) narrow, distally with several short spines.
|
||
</paragraph>
|
||
<paragraph id="8B951E322005FFE3148FFE55FE73FE49" blockId="13.[151,1436,151,608]" pageId="13" pageNumber="271">
|
||
<typeStatus id="5491A0902005FFE3148FFE55FEEAFE6C" box="[199,297,440,464]" pageId="13" pageNumber="271" type="paratype">Paratype</typeStatus>
|
||
(female)
|
||
<quantity id="4CD2B3D72005FFE315DDFE5AFE27FE73" box="[405,484,439,464]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="13" pageNumber="271" unit="mm" value="25.0">25 mm</quantity>
|
||
long, vertical diameter of the largest body ring
|
||
<quantity id="4CD2B3D72005FFE317B1FE5AFB93FE73" box="[1017,1104,439,464]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="271" unit="mm" value="1.5">1.5 mm</quantity>
|
||
, body with 47 podous rings + 2 apodous rings + telson.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3304DB92005FFE3148FFE12FA95FD81" pageId="13" pageNumber="271" type="distribution">
|
||
<paragraph id="8B951E322005FFE3148FFE12FA54FDA4" blockId="13.[151,1436,151,608]" box="[199,1431,511,536]" pageId="13" pageNumber="271">
|
||
<emphasis id="B95EC2202005FFE3148FFE12FEA3FDA4" bold="true" box="[199,352,511,536]" pageId="13" pageNumber="271">Distribution.</emphasis>
|
||
This species is known only from its
|
||
<typeStatus id="5491A0902005FFE316B4FDEDFCEFFDA4" box="[764,812,512,536]" pageId="13" pageNumber="271">type</typeStatus>
|
||
locality, Klenovac Pit in Dalmatia (
|
||
<figureCitation id="131102B72005FFE310F7FE12FAD1FDA4" box="[1215,1298,511,536]" captionStart="FIGURE 20" captionStartId="28.[151,250,1947,1969]" captionTargetBox="[151,1436,1121,1925]" captionTargetId="figure@28.[151,1436,1121,1925]" captionTargetPageId="28" captionText="FIGURE 20. Distribution of representatives of Inversotyphlus Strasser, 1962 and Typhloiulus edentulus Attems, 1951 (Attemsotyphlus Strasser, 1962)." httpUri="https://zenodo.org/record/1457209/files/figure.png" pageId="13" pageNumber="271">Fig. 20</figureCitation>
|
||
, red circle).
|
||
</paragraph>
|
||
<paragraph id="8B951E322005FFE3148FFDC9FA95FD81" blockId="13.[151,1436,151,608]" box="[199,1366,548,573]" pageId="13" pageNumber="271">
|
||
<emphasis id="B95EC2202005FFE3148FFDC9FED2FD81" bold="true" box="[199,273,548,573]" pageId="13" pageNumber="271">Notes.</emphasis>
|
||
Klenovac Pit is some
|
||
<quantity id="4CD2B3D72005FFE31644FDC9FD86FD80" box="[524,581,548,573]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="13" pageNumber="271" unit="m" value="45.0">45 m</quantity>
|
||
deep and also inhabited by a stygobiotic
|
||
<taxonomicName id="4C2A65B12005FFE3105CFDC9FB4AFD81" box="[1044,1161,548,573]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95EC2202005FFE3105CFDC9FB4AFD81" box="[1044,1161,548,573]" italics="true" pageId="13" pageNumber="271">Niphargus</emphasis>
|
||
</taxonomicName>
|
||
sp. (
|
||
<taxonomicName id="4C2A65B12005FFE310F7FDC9FA88FD81" authorityName="Latreille" authorityYear="1816" box="[1215,1355,548,573]" class="Malacostraca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="271" phylum="Arthropoda" rank="order">Amphipoda</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3304DB92005FFE3148FFDAAFBD1FDDC" box="[199,1042,583,608]" pageId="13" pageNumber="271" type="etymology">
|
||
<paragraph id="8B951E322005FFE3148FFDAAFBD1FDDC" blockId="13.[151,1436,151,608]" box="[199,1042,583,608]" pageId="13" pageNumber="271">
|
||
<emphasis id="B95EC2202005FFE3148FFDAAFE8DFDDC" bold="true" box="[199,334,583,608]" pageId="13" pageNumber="271">Etymology.</emphasis>
|
||
To denote the presence of a club-shaped mesomere. Adjective.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |