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<document id="0A548863268C66F83CD8D8BDE0A62F3C" ID-DOI="10.11646/phytotaxa.646.2.3" ID-ISSN="1179-3163" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722605598178" checkinUser="felipe" docAuthor="Akhani, Hossein, Abdullahi, Hussaini, Ürker, Okan &amp; Malekmohammadi, Maryam" docDate="2024" docId="F9050F4FFF96FFD6FF371E41FAA7FCEA" docLanguage="en" docName="phytotaxa.646.2.3.pdf" docOrigin="Phytotaxa 646 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.646.2.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Asparagus azerbaijanensis Hamdi &amp; Assadi X 2017" docType="treatment" docVersion="1" lastPageNumber="136" masterDocId="053C7737FF9DFFDDFFBF1F13FFA3FFAA" masterDocTitle="Molecular phylogenetic studies on Asparagus (Asparagaceae) revealed the position of the highly endangered Irano-Turkish halophytic species A. lycaonicus P. H. Davis related to A. officinalis L." masterLastPageNumber="142" masterPageNumber="125" pageNumber="136" updateTime="1722802309564" updateUser="felipe" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A7229AC45D18FE567165B532A31EEC02">Molecular phylogenetic studies on Asparagus (Asparagaceae) revealed the position of the highly endangered Irano-Turkish halophytic species A. lycaonicus P. H. Davis related to A. officinalis L.</mods:title>
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<mods:namePart id="6EBC11E7A93824BFD23ED5A394A11323">Akhani, Hossein</mods:namePart>
<mods:nameIdentifier id="1381D37A18B307F2A4CDF9EB83637869" type="ORCID">0000-0002-9023-0357</mods:nameIdentifier>
<mods:affiliation id="26CF1CE2A2284C8BDFA6EA396D884624">Halophytes and C &amp; Plants Research Laboratory, Department of Plant Sciences, School of Biology, College of Sciences, University of Tehran, P. O. Box 14155 - 6455, Tehran, Iran</mods:affiliation>
<mods:nameIdentifier id="545A4D6D248F5F8CF9754B8FAC0A4852" type="email">hakhani@ut.ac.ir</mods:nameIdentifier>
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<mods:namePart id="237707955E6FBA5424C6F05B5116DE11">Abdullahi, Hussaini</mods:namePart>
<mods:nameIdentifier id="83FA4D83A0AABF68D20D184C0D24D990" type="ORCID">0009-0006-2115-1213</mods:nameIdentifier>
<mods:affiliation id="82DD822C315B4A069640E03FCD5576DA">Halophytes and C</mods:affiliation>
<mods:nameIdentifier id="2A6565E015BB719F81384E634D73E863" type="email">abdhus1972@gmail.com</mods:nameIdentifier>
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<mods:name id="54223877F29D650900DD664CB52C8B53" type="personal">
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<mods:namePart id="A9A86D28B6085F0DF4DE23B012F50929">Ürker, Okan</mods:namePart>
<mods:nameIdentifier id="53D2DB3AD7C8662764CDBEDAFCF6ABBD" type="ORCID">0000-0002-5103-7757</mods:nameIdentifier>
<mods:affiliation id="ED8EF21A04F54B1682429CD92595DA0C">Environmental Health Program, Şabanözü Vocational School, Çankırı Karatekin University, P. O. Box 18650, Çankırı, Türkiye</mods:affiliation>
<mods:nameIdentifier id="6378EC74DAEA8839BCAC56F2C378AF06" type="email">okan.urker@gmail.com</mods:nameIdentifier>
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<mods:name id="8F6422ACDAF83F8AA5DCD1F022A77B4F" type="personal">
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<mods:namePart id="35889E88C279CA5CE7FFFCD74BC2A0C3">Malekmohammadi, Maryam</mods:namePart>
<mods:nameIdentifier id="431242BBBD940DD4D7B65D6B233E6493" type="ORCID">0000-0001-2345-6785</mods:nameIdentifier>
<mods:affiliation id="A36F1003A66AA6724A4CACC0CE4273C6">Halophytes and C &amp; Halophytes and C &amp; Botanischer Garten und Botanisches Museum Berlin, Freie Universität Berlin, Königin-Luise-Strasse 6 - 8, 14195 Berlin, Germany &amp; Halophytes and C</mods:affiliation>
<mods:nameIdentifier id="F21CC024947ED93F11A4636EA2906F66" type="email">maryam.malekmohammady@gmail.com</mods:nameIdentifier>
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<mods:title id="978B1CF12AF156CB1A88DAC4EDF4DA6D">Phytotaxa</mods:title>
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<mods:date id="D813896DA2E7B768F24347E165B54204">2024</mods:date>
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<mods:classification id="77CE77DDD143298D33C1FAF7E5042BAC">journal article</mods:classification>
<mods:identifier id="73AEABEDFDAD6781FBA60FE73190F2B1" type="DOI">10.11646/phytotaxa.646.2.3</mods:identifier>
<mods:identifier id="4077316145EAB038CD752D8D55BC4414" type="ISSN">1179-3163</mods:identifier>
</mods:mods>
<treatment id="F9050F4FFF96FFD6FF371E41FAA7FCEA" LSID="urn:lsid:plazi:treatment:F9050F4FFF96FFD6FF371E41FAA7FCEA" httpUri="http://treatment.plazi.org/id/F9050F4FFF96FFD6FF371E41FAA7FCEA" lastPageNumber="136" pageId="11" pageNumber="136">
<subSubSection id="39B6EDD2FF96FFD6FF371E41FCA1FEC6" box="[136,770,338,365]" pageId="11" pageNumber="136" type="nomenclature">
<paragraph id="7113BE59FF96FFD6FF371E41FCA1FEC6" blockId="11.[136,770,338,365]" box="[136,770,338,365]" pageId="11" pageNumber="136">
<heading id="2A5B0935FF96FFD6FF371E41FCA1FEC6" box="[136,770,338,365]" fontSize="11" level="2" pageId="11" pageNumber="136" reason="3">
<taxonomicName id="B6ACC5DAFF96FFD6FF371E41FD5DFEC6" ID-CoL="8WRF4" authority="Hamdi &amp; Assadi (2017: 93)" authorityName="Hamdi &amp; Assadi X" authorityPageNumber="93" authorityYear="2017" box="[136,766,338,365]" class="Liliopsida" family="Asparagaceae" genus="Asparagus" kingdom="Plantae" order="Asparagales" pageId="11" pageNumber="136" phylum="Tracheophyta" rank="species" species="azerbaijanensis">
<emphasis id="43D8624BFF96FFD6FF371E41FE1EFEC7" bold="true" box="[136,445,338,365]" italics="true" pageId="11" pageNumber="136">Asparagus azerbaijanensis</emphasis>
Hamdi &amp; Assadi (2017: 93)
</taxonomicName>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="39B6EDD2FF96FFD6FF371E65FAA7FCEA" pageId="11" pageNumber="136" type="materials_examined">
<paragraph id="7113BE59FF96FFD6FF371E65FD2CFE1E" blockId="11.[136,1452,374,833]" pageId="11" pageNumber="136">
<materialsCitation id="C1C4B404FF96FFD6FF371E65FD2CFE1E" collectingDate="1976-06-12" collectionCode="TARI" collectorName="Assadi &amp; Maassoumi" country="Iran" elevation="1400" location="Tabriz" municipality="Tabriz" pageId="11" pageNumber="136" specimenCount="1" stateProvince="Azarbayjan-e Sharqi" typeStatus="holotype">
<typeStatus id="AE1700FBFF96FFD6FF371E65FF50FE3A" box="[136,243,374,400]" pageId="11" pageNumber="136" type="holotype">Holotype</typeStatus>
:—
<collectingCountry id="09BBFEC9FF96FFD6FEAA1E65FEFEFE3A" box="[277,349,374,400]" name="Iran" pageId="11" pageNumber="136">IRAN</collectingCountry>
. [Eastern]
<collectingRegion id="B36870BBFF96FFD6FE6E1E65FD33FE3B" box="[465,656,374,401]" country="Iran" name="Azarbayjan-e Sharqi" pageId="11" pageNumber="136">Azerbaijan prov.</collectingRegion>
:
<locationDeviation id="60A1A77BFF96FFD6FD261E65FCBFFE3A" box="[665,796,374,400]" location="Ahar" pageId="11" pageNumber="136">
<quantity id="B65413BCFF96FFD6FD261E65FD7CFE3A" box="[665,735,374,400]" metricMagnitude="4" metricUnit="m" metricValue="6.2" pageId="11" pageNumber="136" unit="km" value="62.0">62 km</quantity>
Ahar
</locationDeviation>
to
<collectingMunicipality id="91772423FF96FFD6FC811E65FC2BFE3A" box="[830,904,374,400]" pageId="11" pageNumber="136">Tabriz</collectingMunicipality>
, margin of road of
<location id="7473E882FF96FFD6FBE51E65FB07FE3A" LSID="urn:lsid:plazi:treatment:F9050F4FFF96FFD6FF371E41FAA7FCEA:7473E882FF96FFD6FBE51E65FB07FE3A" box="[1114,1188,374,400]" country="Iran" municipality="Tabriz" name="Tabriz" pageId="11" pageNumber="136" stateProvince="Azarbayjan-e Sharqi">Tabriz</location>
,
<quantity id="B65413BCFF96FFD6FB101E65FAA6FE3B" box="[1199,1285,374,401]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="11" pageNumber="136" unit="m" value="1400.0">
<elevation id="FA81596AFF96FFD6FB101E65FAA6FE3B" box="[1199,1285,374,401]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="11" pageNumber="136" unit="m" value="1400.0">1400 m</elevation>
</quantity>
,
<date id="05129899FF96FFD6FAAF1E65FA04FE3A" box="[1296,1447,374,401]" pageId="11" pageNumber="136" value="1976-06-12">
<collectingDate id="15566171FF96FFD6FAAF1E65FA04FE3A" box="[1296,1447,374,401]" pageId="11" pageNumber="136" value="1976-06-12">12 June 1976</collectingDate>
</date>
,
<emphasis id="43D8624BFF96FFD6FF371E89FEDBFE1E" box="[136,376,410,437]" italics="true" pageId="11" pageNumber="136">
<collectorName id="DC59DB8FFF96FFD6FF371E89FF70FE1E" box="[136,211,410,436]" pageId="11" pageNumber="136">Assadi</collectorName>
&amp;
<collectorName id="DC59DB8FFF96FFD6FF481E88FEDBFE1E" box="[247,376,411,436]" pageId="11" pageNumber="136">Maassoumi</collectorName>
</emphasis>
20591 (
<collectionCode id="17BD269CFF96FFD6FE6C1E89FDB4FE1E" box="[467,535,410,436]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="11" pageNumber="136">TARI</collectionCode>
, not seen).
</materialsCitation>
</paragraph>
<paragraph id="7113BE59FF96FFD6FF021EADFCEAFD8A" blockId="11.[136,1452,374,833]" pageId="11" pageNumber="136">
<emphasis id="43D8624BFF96FFD6FF021EADFDF9FE72" bold="true" box="[189,602,446,472]" pageId="11" pageNumber="136">Additional specimens reported:—</emphasis>
<materialsCitation id="C1C4B404FF96FFD6FDE41EADFC30FE56" collectingDate="1991-08-16" collectionCode="TARI" country="Iran" elevation="1800" location="Kelisakandi" municipality="Makou-Bazargan" pageId="11" pageNumber="136" specimenCount="1" stateProvince="Azarbayjan-e Sharqi">
<collectingCountry id="09BBFEC9FF96FFD6FDE41EADFD00FE72" box="[603,675,446,472]" name="Iran" pageId="11" pageNumber="136">IRAN</collectingCountry>
.
<collectingRegion id="B36870BBFF96FFD6FD0F1EADFC3EFE72" box="[688,925,446,472]" country="Iran" name="Azarbayjan-e Sharqi" pageId="11" pageNumber="136">Azerbaijan province</collectingRegion>
:
<quantity id="B65413BCFF96FFD6FC131EADFC5BFE72" box="[940,1016,446,473]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="136" unit="km" value="15.0">15 km</quantity>
from main road of
<collectingMunicipality id="91772423FF96FFD6FB561EADFA08FE72" box="[1257,1451,446,472]" pageId="11" pageNumber="136">Makou-Bazargan</collectingMunicipality>
towards
<location id="7473E882FF96FFD6FF571EF1FED2FE56" LSID="urn:lsid:plazi:treatment:F9050F4FFF96FFD6FF371E41FAA7FCEA:7473E882FF96FFD6FF571EF1FED2FE56" box="[232,369,482,508]" country="Iran" municipality="Makou-Bazargan" name="Kelisakandi" pageId="11" pageNumber="136" stateProvince="Azarbayjan-e Sharqi">Kelisakandi</location>
,
<quantity id="B65413BCFF96FFD6FEC31EF1FE70FE57" box="[380,467,482,509]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="11" pageNumber="136" unit="m" value="1800.0">
<elevation id="FA81596AFF96FFD6FEC31EF1FE70FE57" box="[380,467,482,509]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="11" pageNumber="136" unit="m" value="1800.0">1800 m</elevation>
</quantity>
,
<date id="05129899FF96FFD6FE601EF1FD31FE56" box="[479,658,482,508]" pageId="11" pageNumber="136" value="1991-08-16">
<collectingDate id="15566171FF96FFD6FE601EF1FD31FE56" box="[479,658,482,508]" pageId="11" pageNumber="136" value="1991-08-16">16 August 1991</collectingDate>
</date>
,
<emphasis id="43D8624BFF96FFD6FD201EF1FC95FE56" box="[671,822,482,508]" italics="true" pageId="11" pageNumber="136">Assadi 70831</emphasis>
(
<collectionCode id="17BD269CFF96FFD6FCFA1EF1FC28FE56" box="[837,907,482,508]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="11" pageNumber="136">TARI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C1C4B404FF96FFD6FC211EF1FCE6FD8A" collectingDate="1975-06-28" collectionCode="TARI" country="Iran" elevation="1300" location="Marand" municipality="Makou" pageId="11" pageNumber="136" specimenCount="1">
<quantity id="B65413BCFF96FFD6FC211EF1FC46FE56" box="[926,997,482,508]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="136" unit="km" value="13.0">13 km</quantity>
to
<collectingMunicipality id="91772423FF96FFD6FBB61EF1FBF8FE56" box="[1033,1115,482,508]" pageId="11" pageNumber="136">Makou</collectingMunicipality>
, on road from
<location id="7473E882FF96FFD6FABF1EF1FAF9FE56" LSID="urn:lsid:plazi:treatment:F9050F4FFF96FFD6FF371E41FAA7FCEA:7473E882FF96FFD6FABF1EF1FAF9FE56" box="[1280,1370,482,508]" country="Iran" municipality="Makou" name="Marand" pageId="11" pageNumber="136">Marand</location>
,
<quantity id="B65413BCFF96FFD6FADA1EF1FF7CFD8B" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="11" pageNumber="136" unit="m" value="1300.0" valueMax="1400.0" valueMin="1200.0">
<elevation id="FA81596AFF96FFD6FADA1EF1FF7CFD8B" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="11" pageNumber="136" unit="m" value="1300.0" valueMax="1400.0" valueMin="1200.0">1200 1400 m</elevation>
</quantity>
,
<date id="05129899FF96FFD6FF541D15FE20FD8B" box="[235,387,518,545]" pageId="11" pageNumber="136" value="1975-06-28">
<collectingDate id="15566171FF96FFD6FF541D15FE20FD8B" box="[235,387,518,545]" pageId="11" pageNumber="136" value="1975-06-28">28 June 1975</collectingDate>
</date>
,
<emphasis id="43D8624BFF96FFD6FE311D15FD4BFD8A" box="[398,744,518,544]" italics="true" pageId="11" pageNumber="136">Assadi and Mozaffarian 30105</emphasis>
(
<collectionCode id="17BD269CFF96FFD6FD481D15FC9EFD8A" box="[759,829,518,544]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="11" pageNumber="136">TARI</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="7113BE59FF96FFD6FF021D39FAA7FCEA" blockId="11.[136,1452,374,833]" pageId="11" pageNumber="136">
The new species was described from marl-gypsum hills around the road. The authors compared their species with
<taxonomicName id="B6ACC5DAFF96FFD6FF371D5CFE8AFDC2" box="[136,297,591,617]" class="Liliopsida" family="Asparagaceae" genus="Asparagus" kingdom="Plantae" order="Asparagales" pageId="11" pageNumber="136" phylum="Tracheophyta" rank="species" species="touranensis">
<emphasis id="43D8624BFF96FFD6FF371D5CFE8AFDC2" box="[136,297,591,617]" italics="true" pageId="11" pageNumber="136">A. touranensis</emphasis>
</taxonomicName>
and
<taxonomicName id="B6ACC5DAFF96FFD6FEE31D5CFD10FDC2" authority="Baker (1875: 607)" authorityName="Baker" authorityPageNumber="607" authorityYear="1875" box="[348,691,590,617]" class="Liliopsida" family="Asparagaceae" genus="Asparagus" kingdom="Plantae" order="Asparagales" pageId="11" pageNumber="136" phylum="Tracheophyta" rank="species" species="capitatus">
<emphasis id="43D8624BFF96FFD6FEE31D5CFE43FDC2" box="[348,480,591,617]" italics="true" pageId="11" pageNumber="136">A. capitatus</emphasis>
<bibRefCitation id="153DC3A8FF96FFD6FE5A1D5DFD10FDC2" author="Baker, J. G." box="[485,691,590,617]" pageId="11" pageNumber="136" pagination="594 - 629" refId="ref10369" refString="Baker, J. G. (1875) Revision of the genera and species of the Asparagaceae. Journal of the Linnean Society of London 14: 594 - 629. https: // doi. org / 10.1111 / j. 1095 - 8339.1875. tb 00349. x" type="journal article" year="1875">Baker (1875: 607)</bibRefCitation>
</taxonomicName>
.Apart from the
<typeStatus id="AE1700FBFF96FFD6FCDC1D5CFC30FDC3" box="[867,915,591,617]" pageId="11" pageNumber="136">type</typeStatus>
samples, two additional samples are listed in the protologue (Hamdi &amp; Assadi 2017). None of these specimens are cited in the Flora of
<collectingCountry id="09BBFEC9FF96FFD6FBFC1D60FBD3FD27" box="[1091,1136,627,653]" name="Iran" pageId="11" pageNumber="136">Iran</collectingCountry>
(
<bibRefCitation id="153DC3A8FF96FFD6FBC11D61FAB5FD26" author="Assadi, M." box="[1150,1302,626,652]" pageId="11" pageNumber="136" refId="ref10328" refString="Assadi, M. (2023) Asparagaceae. In: Assadi &amp; Maassoumi, A. A. (Eds.) Flora of Iran, 174. Research Institute of Forests and Rangelands, Iran, 22 pp." type="book" year="2023">Assadi, 2023</bibRefCitation>
). The authors described the species as having only one cladode per fascicle. But the illustration (
<figureCitation id="E997A2DCFF96FFD6FB9E1D85FBC0FD1A" box="[1057,1123,662,688]" captionStart="FIGURE 1" captionStartId="2.[136,229,960,982]" captionTargetBox="[196,1391,187,933]" captionTargetId="figure-23@2.[196,1391,187,933]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of Asparagus lycaonicus P.H. Davis in Iran and Turkey (triangle)." pageId="11" pageNumber="136">Fig. 1</figureCitation>
, p. 94) shows a long and 45 smaller cladodes per fascicle. In the gypsum-marl area between
<collectingRegion id="B36870BBFF96FFD6FCDF1DA9FC0DFD7E" box="[864,942,698,724]" country="Iran" name="Zanjan" pageId="11" pageNumber="136">Zanjan</collectingRegion>
and Marand, two species of
<taxonomicName id="B6ACC5DAFF96FFD6FB481DA8FACCFD7F" box="[1271,1391,699,725]" class="Liliopsida" family="Asparagaceae" genus="Asparagus" kingdom="Plantae" order="Asparagales" pageId="11" pageNumber="136" phylum="Tracheophyta" rank="genus">
<emphasis id="43D8624BFF96FFD6FB481DA8FACCFD7F" box="[1271,1391,699,725]" italics="true" pageId="11" pageNumber="136">Asparagus</emphasis>
</taxonomicName>
have been collected by the first author (
<emphasis id="43D8624BFF96FFD6FDB71DCDFC8FFD52" box="[520,812,734,761]" italics="true" pageId="11" pageNumber="136">Akhani &amp; Salimian 14412</emphasis>
and
<emphasis id="43D8624BFF96FFD6FCDC1DCDFC0AFD52" box="[867,937,734,760]" italics="true" pageId="11" pageNumber="136">14213</emphasis>
, see
<tableCitation id="3C2E8BE2FF96FFD6FC5E1DCDFB97FD52" box="[993,1076,734,760]" captionStart="TABLE 2" captionStartId="2.[136,224,1718,1742]" captionText="TABLE 2. List of Asparagus and outgroup species studied in this paper and their GenBank accession numbers. N: Norup et" pageId="11" pageNumber="136">Table 2</tableCitation>
for details on the collection site). We have identified one of the specimens as
<taxonomicName id="B6ACC5DAFF96FFD6FDD61C10FC9FFCB6" box="[617,828,770,797]" class="Liliopsida" family="Asparagaceae" genus="Asparagus" isUncertain="true" kingdom="Plantae" order="Asparagales" pageId="11" pageNumber="136" phylum="Tracheophyta" rank="species" species="breslerianus">
<emphasis id="43D8624BFF96FFD6FDD61C10FC9FFCB6" box="[617,828,770,797]" italics="true" pageId="11" pageNumber="136">A. aff. breslerianus</emphasis>
</taxonomicName>
(
<emphasis id="43D8624BFF96FFD6FCF41C11FC32FCB6" box="[843,913,770,796]" italics="true" pageId="11" pageNumber="136">14213</emphasis>
) and the other as probably
<taxonomicName id="B6ACC5DAFF96FFD6FB021C10FA0FFCB6" authorityName="Hamdi &amp; Assadi X" authorityYear="2017" box="[1213,1452,770,797]" class="Liliopsida" family="Asparagaceae" genus="Asparagus" isUncertain="true" kingdom="Plantae" order="Asparagales" pageId="11" pageNumber="136" phylum="Tracheophyta" rank="species" species="azerbaijanensis">
<emphasis id="43D8624BFF96FFD6FB021C10FA0FFCB6" box="[1213,1452,770,797]" italics="true" pageId="11" pageNumber="136">A. cf. azerbaijanensis</emphasis>
</taxonomicName>
(
<emphasis id="43D8624BFF96FFD6FF2E1C35FE8DFCEA" box="[145,302,806,832]" italics="true" pageId="11" pageNumber="136">Akhani 14212</emphasis>
).
<collectionCode id="17BD269CFF96FFD6FEFC1C35FEF5FCEA" box="[323,342,806,832]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="11" pageNumber="136" type="Herbarium">A</collectionCode>
final judgment requires further studies including examination of the
<typeStatus id="AE1700FBFF96FFD6FBE41C34FB28FCEB" box="[1115,1163,807,833]" pageId="11" pageNumber="136">type</typeStatus>
specimen.
</paragraph>
</subSubSection>
</treatment>
</document>