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<document id="367BAAB5244A6FAC67B7ED83FE1CA548" ID-CLB-Dataset="145591" ID-DOI="10.1649/0010-066X-66.mo4.1" ID-GBIF-Dataset="4476f1cf-3ad0-4538-8407-9c8a6aeb0fe1" ID-ISSN="1938-4394" ID-Zenodo-Dep="4907222" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1623089673881" checkinUser="carolina" docAuthor="Orozco, Jesús" docDate="2012" docId="F449F723D543B20887EB454DECDDFBAE" docLanguage="en" docName="TheColeopBull.66.mo11.1-182.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 11) 66" docStyle="DocumentStyle:4CAA047AD53AEBA766D4E826C2986B94.5:TheColeopBull.2010-2016.monograph" docStyleId="4CAA047AD53AEBA766D4E826C2986B94" docStyleName="TheColeopBull.2010-2016.monograph" docStyleVersion="5" docTitle="Euphoria anneae" docType="treatment" docVersion="11" lastPageNumber="99" masterDocId="08708F5BD522B26A85194065E874FF87" masterDocTitle="Monographic Revision of the American Genus Euphoria Burmeister, 1842 (Coleoptera: Scarabaeidae: Cetoniinae)" masterLastPageNumber="182" masterPageNumber="1" pageNumber="98" updateTime="1699000576926" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="17C429937714662283DB757EBCB01C65">Monographic Revision of the American Genus Euphoria Burmeister, 1842 (Coleoptera: Scarabaeidae: Cetoniinae)</mods:title>
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<mods:name id="D2B0DF590AB37E1B664C722166EEA135" type="personal">
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<mods:namePart id="8FE72347E709D2B687F1BF68ADE6D49B">Orozco, Jesús</mods:namePart>
<mods:affiliation id="236AD7087377E388081AF81AB9CC06C0">Molecular Ecology and Systematics Group Department of Botany and Department of Zoology and Entomology Rhodes University Grahamstown, SOUTH AFRICA 6140</mods:affiliation>
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<mods:typeOfResource id="4DA5B370DB9F9441C4F4A415C18C8F98">text</mods:typeOfResource>
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<mods:title id="4CACDB8DF73593BF57B2EB0F79D61FBC">The Coleopterists Bulletin</mods:title>
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<mods:part id="5F14F292B3F46656A3AD46FE311117D6">
<mods:date id="FDDEAD9991A0272F66880D1C0D57C781">2012</mods:date>
<mods:detail id="75DF51D8DD9D7340BA5C5E61C8782CB4" type="series">
<mods:title id="359C63BFFB65D94E4FD7605A346BA857">mo 11</mods:title>
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<mods:number id="73F67E602A09E3ABFF6C967A95360555">2012-12-20</mods:number>
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<mods:number id="05CCE7FEF9AF92DAA36B71B7A4234F95">66</mods:number>
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<mods:classification id="0F7E1FBA7C47816AC25690DED8C358B8">journal article</mods:classification>
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<mods:identifier id="B09A03193DE6F3FB523E4387CB6BFECB" type="DOI">10.1649/0010-066X-66.mo4.1</mods:identifier>
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<treatment id="F449F723D543B20887EB454DECDDFBAE" ID-DOI="http://doi.org/10.5281/zenodo.7092762" ID-GBIF-Taxon="200629789" ID-Zenodo-Dep="7092762" LSID="urn:lsid:plazi:treatment:F449F723D543B20887EB454DECDDFBAE" httpUri="http://treatment.plazi.org/id/F449F723D543B20887EB454DECDDFBAE" lastPageId="98" lastPageNumber="99" pageId="97" pageNumber="98">
<subSubSection id="34FA15BED543B20B87EB454DEC3BFABA" box="[754,1103,1320,1341]" pageId="97" pageNumber="98" type="nomenclature">
<paragraph id="7C5F4635D543B20B87EB454DEC3BFABA" blockId="97.[754,1103,1320,1368]" box="[754,1103,1320,1341]" pageId="97" pageNumber="98">
<heading id="2717F159D543B20B87EB454DEC3BFABA" box="[754,1103,1320,1341]" centered="true" fontSize="10" level="2" pageId="97" pageNumber="98" reason="2">
<taxonomicName id="BBE03DB6D543B20B87EB454DEC3BFABA" ID-CoL="6HB6L" authority="(Howden, 1955)" baseAuthorityName="Howden" baseAuthorityYear="1955" box="[754,1103,1320,1341]" class="Insecta" family="Scarabaeidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="anneae">
<emphasis id="4E949A27D543B20B87EB454DEC3BFABA" bold="true" box="[754,1103,1320,1341]" pageId="97" pageNumber="98">
<emphasis id="4E949A27D543B20B87EB454DEBEBFABA" bold="true" box="[754,927,1320,1341]" italics="true" pageId="97" pageNumber="98">Euphoria anneae</emphasis>
(
<bibRefCitation id="18713BC4D543B20B86B7454DEC3CFABA" author="Howden H." box="[942,1096,1320,1341]" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="97" pageNumber="98" pagination="257 - 264" part="57" refId="ref123283" refString="Howden H. 1955. Some new species and records of North American Scarabaeidae. Proceedings of the Entomological Society of Washington 57: 257 - 264." title="Some new species and records of North American Scarabaeidae" type="journal article" year="1955">Howden, 1955</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="34FA15BED543B20B862C4526EC78FADF" box="[821,1036,1347,1368]" pageId="97" pageNumber="98" type="description">
<paragraph id="7C5F4635D543B20B862C4526EC78FADF" blockId="97.[754,1103,1320,1368]" box="[821,1036,1347,1368]" pageId="97" pageNumber="98">
(Appendix 4:
<figureCitation id="E4DB5AB0D543B20B86A64526EC71FADF" box="[959,1029,1347,1368]" captionStart="Fig" captionStartId="174.[197,229,1686,1705]" captionTargetBox="[298,1132,196,1662]" captionTargetId="figure-0@174.[174,1192,183,1662]" captionTargetPageId="174" captionText="Fig. 55. Euphoria anneae. a) Habitus, b) Head, c) Parameres, d) Color variation, e) Temporal distribution, f) Geographic distribution in the United States." figureDoi="http://doi.org/10.5281/zenodo.4907368" httpUri="https://zenodo.org/record/4907368/files/figure.png" pageId="97" pageNumber="98">Fig. 55</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="34FA15BED543B20B87B4450BEC08FA1A" pageId="97" pageNumber="98" type="reference_group">
<paragraph id="7C5F4635D543B20B87B4450BEC08FA1A" blockId="97.[685,1172,1390,1437]" pageId="97" pageNumber="98">
<taxonomicName id="BBE03DB6D543B20B87B4450BEC4EFA04" ID-CoL="52CXL" authority="Howden 1955: 263" authorityName="Howden" authorityPageNumber="263" authorityYear="1955" box="[685,1082,1390,1411]" class="Insecta" family="Scarabaeidae" genus="Stephanucha" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="anneae">
<emphasis id="4E949A27D543B20B87B4450BEB01FA04" box="[685,885,1390,1411]" italics="true" pageId="97" pageNumber="98">Stephanucha anneae</emphasis>
<bibRefCitation id="18713BC4D543B20B8664450BEC4EFA04" author="Howden H." box="[893,1082,1390,1411]" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="97" pageNumber="98" pagination="257 - 264" part="57" refId="ref123283" refString="Howden H. 1955. Some new species and records of North American Scarabaeidae. Proceedings of the Entomological Society of Washington 57: 257 - 264." title="Some new species and records of North American Scarabaeidae" type="journal article" year="1955">Howden 1955: 263</bibRefCitation>
</taxonomicName>
. Original combination.
<typeStatus id="A35BF897D543B20B865545EDEBD0FA1A" box="[844,932,1416,1437]" pageId="97" pageNumber="98" type="holotype">Holotype</typeStatus>
at CMNC, examined.
</paragraph>
</subSubSection>
<subSubSection id="34FA15BED543B20887DC45D8E96AFA04" lastPageId="98" lastPageNumber="99" pageId="97" pageNumber="98" type="description">
<paragraph id="7C5F4635D543B20887DC45D8E96AFA04" blockId="97.[685,1172,1469,1729]" lastBlockId="98.[173,660,194,1729]" lastPageId="98" lastPageNumber="99" pageId="97" pageNumber="98">
<emphasis id="4E949A27D543B20B87DC45D8EB16FA55" bold="true" box="[709,866,1469,1490]" pageId="97" pageNumber="98">Description (4).</emphasis>
Length 11.8.
<quantity id="BB18EBD0D543B20B86FC45D8EC20FA55" box="[997,1108,1469,1491]" metricMagnitude="-3" metricUnit="m" metricValue="6.25" metricValueMax="12.5" metricValueMin="0.0" pageId="97" pageNumber="98" unit="mm" value="6.25" valueMax="12.5" valueMin="0.0">012.5 mm</quantity>
; width 7.48.0 mm.
<emphasis id="4E949A27D543B20B863D45BDEB2EFA6A" bold="true" box="[804,858,1496,1517]" pageId="97" pageNumber="98">Color</emphasis>
: Dorsal surface partially tomentous; head, pronotum, scutellum, and pygidium shiny, black. Pronotum mostly shiny, base matte; lateral margin with cretaceous whitish band; irregular, whitish, cretaceous markings frequently on pronotal disc. Elytra yellowish brown, with black markings distributed throughout entire disc.
<emphasis id="4E949A27D543B20B81044612EC25F90B" bold="true" box="[1053,1105,1655,1676]" pageId="97" pageNumber="98">Head</emphasis>
: Frons strongly strigose to strigopunctate, punctures moderate to large, round, deeply impressed, frequently coalescent, densely setose; setae moderate to long, yellowish. Clypeus strongly strigose, sides strongly declivous, weakly to moderately setose; setae short to moderate, yellowish. Apex with 4 short teeth, teeth equidistant or middle ones closer to each other than to lateral teeth. Clypeal margin in front of the eyes strongly raised, developed at times into a sharp denticle. Labium thickened anteriorly. Antennal stem longer in males than in females, club about as long as head in males, half as long or less in females.
<emphasis id="4E949A27D540B208841541A9E902FE66" bold="true" box="[268,374,460,481]" pageId="98" pageNumber="99">Pronotum</emphasis>
: Surface densely punctate, punctures small to moderate in size, lunulate; moderately to densely setose, setae short to long, yellowish. Anterior sides evenly arcuate to moderately tapered anteriorly. Lateral line extending only in anterior half or third. Midline frequently impunctate at base. Base strongly rounded to weakly extended posteriorly. Scutellum longer than wide, impunctate, apex pointed.
<emphasis id="4E949A27D540B208842842C5E904FD32" bold="true" box="[305,368,672,693]" pageId="98" pageNumber="99">Elytra</emphasis>
: Surface weakly to moderately setose, setae short to moderate, yellowish. Punctures moderately dense to dense, lunulate to irregular in shape, randomly distributed thought disc. Costae obsolete to subobsolete. Apex strongly rounded.
<emphasis id="4E949A27D540B20885B44340E965FCBD" bold="true" box="[173,273,805,826]" pageId="98" pageNumber="99">Pygidium</emphasis>
: Surface concentrically striate; striae discontinuous, moderately spaced, moderately impressed; weakly to moderately setose, setae minute to moderate, yellowish.
<emphasis id="4E949A27D540B20884E34310EA5FFC0D" bold="true" box="[506,555,885,906]" pageId="98" pageNumber="99">Legs</emphasis>
: Protibial teeth strongly developed, not sexually dimorphic, first tooth oblique. Mesotibial carina strongly developed into 12 sharp teeth. Metatibiae apically expanded; metatibial spurs frequently thicker in females. Metatarsomeres strongly expanded apically, basal tarsomere strongly expanded distally forming a spine. Claws shorter than last tarsomere.
<emphasis id="4E949A27D540B20885B4442CE884FBD9" bold="true" box="[173,240,1097,1118]" pageId="98" pageNumber="99">Venter</emphasis>
: Mesometasternal process strongly compressed laterally, short, not extending anteriorly beyond the level of mesocoxae, mostly setose. Median sulcus weakly to moderately impressed. Abdominal sternites moderately to densely setose laterally, weakly setose to moderately medially, setae distributed along entire width of segment, moderate to long, yellowish. Abdomen laterally convex in females, flat to concave in males. Dorsal and lateral area of sternites frequently with white, cretaceous markings.
<emphasis id="4E949A27D540B20884634536EA7DFAEF" bold="true" box="[378,521,1363,1384]" pageId="98" pageNumber="99">Male genitalia</emphasis>
: Parameres as in
<figureCitation id="E4DB5AB0D540B20885D1450BE96EFA04" box="[200,282,1390,1411]" captionStart="Fig" captionStartId="174.[197,229,1686,1705]" captionTargetBox="[298,1132,196,1662]" captionTargetId="figure-0@174.[174,1192,183,1662]" captionTargetPageId="174" captionText="Fig. 55. Euphoria anneae. a) Habitus, b) Head, c) Parameres, d) Color variation, e) Temporal distribution, f) Geographic distribution in the United States." figureDoi="http://doi.org/10.5281/zenodo.4907368" httpUri="https://zenodo.org/record/4907368/files/figure.png" pageId="98" pageNumber="99">Fig. 55c</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="34FA15BED540B20885DC45EDEA39F9D0" pageId="98" pageNumber="99" type="diagnosis">
<paragraph id="7C5F4635D540B20885DC45EDEA39F9D0" blockId="98.[173,660,194,1729]" pageId="98" pageNumber="99">
<emphasis id="4E949A27D540B20885DC45EDE959FA1A" bold="true" box="[197,301,1416,1437]" pageId="98" pageNumber="99">Diagnosis.</emphasis>
<taxonomicName id="BBE03DB6D540B208842245EDE996FA1A" baseAuthorityName="Howden" baseAuthorityYear="1955" box="[315,482,1416,1437]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="anneae">
<emphasis id="4E949A27D540B208842245EDE996FA1A" box="[315,482,1416,1437]" italics="true" pageId="98" pageNumber="99">Euphoria anneae</emphasis>
</taxonomicName>
is separated from the other two species in the group based on the black elytral markings not originating from the sutural area as in
<taxonomicName id="BBE03DB6D540B208844445BDE9C2FA6A" box="[349,438,1496,1517]" class="Insecta" family="Scarabaeidae" genus="Stephanucha" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="areata">
<emphasis id="4E949A27D540B208844445BDE91FFA6A" box="[349,363,1496,1517]" italics="true" pageId="98" pageNumber="99">E</emphasis>
.
<emphasis id="4E949A27D540B208846345BDE9C2FA6A" box="[378,438,1496,1517]" italics="true" pageId="98" pageNumber="99">areata</emphasis>
</taxonomicName>
and body not densely setose as in
<taxonomicName id="BBE03DB6D540B20884354596E9D8F98F" baseAuthorityName="Kraatz" baseAuthorityYear="1883" box="[300,428,1523,1544]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="pilipennis">
<emphasis id="4E949A27D540B20884354596E94EF98F" box="[300,314,1523,1544]" italics="true" pageId="98" pageNumber="99">E</emphasis>
.
<emphasis id="4E949A27D540B20884524596E9D8F98F" box="[331,428,1523,1544]" italics="true" pageId="98" pageNumber="99">pilipennis</emphasis>
</taxonomicName>
. The clypeal denticles are not diagnostic in this species as they vary intraspecifically. This is the only species in the group that occurs in southern
<collectingRegion id="BE2488D7D540B20884C84627EA7EF9D0" box="[465,522,1602,1623]" country="United States of America" name="Texas" pageId="98" pageNumber="99">Texas</collectingRegion>
,
<collectingCountry id="04F706A5D540B208870F4627EA3DF9D0" box="[534,585,1602,1623]" name="United States of America" pageId="98" pageNumber="99">USA</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="34FA15BED540B20885DC4638EBD2FD57" pageId="98" pageNumber="99" type="discussion">
<paragraph id="7C5F4635D540B20885DC4638EC49FE41" blockId="98.[173,660,194,1729]" lastBlockId="98.[707,1194,194,1065]" pageId="98" pageNumber="99">
<emphasis id="4E949A27D540B20885DC4638E970F9F5" bold="true" box="[197,260,1629,1650]" pageId="98" pageNumber="99">Notes.</emphasis>
The
<specimenCount id="6AE68DBCD540B20884594638E9A0F9F5" box="[320,468,1629,1650]" pageId="98" pageNumber="99" type="generic">four specimens</specimenCount>
examined differ in the relative positions of the clypeal teeth; all four teeth being equidistant from each other in
<specimenCount id="6AE68DBCD540B20885B446C9E93EF946" box="[173,330,1708,1729]" pageId="98" pageNumber="99" type="generic">three specimens</specimenCount>
and the two middle teeth fused at the base and closer to each other than the outer two teeth in the other. The protibial teeth also show variation in their size, being weakly developed in the
<typeStatus id="A35BF897D540B208862C4177EBF3FEA0" box="[821,903,274,295]" pageId="98" pageNumber="99" type="holotype">holotype</typeStatus>
and strongly developed in the other
<specimenCount id="6AE68DBCD540B20887E14149EBE6FEC6" box="[760,914,300,321]" pageId="98" pageNumber="99" type="generic">three specimens</specimenCount>
. The body of the
<typeStatus id="A35BF897D540B20881594149ECE5FEC6" box="[1088,1169,300,321]" pageId="98" pageNumber="99" type="holotype">holotype</typeStatus>
is moderately tapered posteriorly (as illustrated by
<bibRefCitation id="18713BC4D540B20887DA4107EB28FEF0" author="Howden H." box="[707,860,354,375]" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="98" pageNumber="99" pagination="257 - 264" part="57" refId="ref123283" refString="Howden H. 1955. Some new species and records of North American Scarabaeidae. Proceedings of the Entomological Society of Washington 57: 257 - 264." title="Some new species and records of North American Scarabaeidae" type="journal article" year="1955">Howden [1955]</bibRefCitation>
), a character that is not evident in the other
<specimenCount id="6AE68DBCD540B20886044119EBC0FE16" box="[797,948,380,401]" pageId="98" pageNumber="99" type="generic">three specimens</specimenCount>
examined. The pronotum has small, cretaceous markings in three of the
<specimenCount id="6AE68DBCD540B208819B41F2EB50FE41" pageId="98" pageNumber="99" type="generic">four specimens</specimenCount>
studied (not in the
<typeStatus id="A35BF897D540B20886C441D4EC47FE41" box="[989,1075,433,454]" pageId="98" pageNumber="99" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="7C5F4635D540B20887C241A9EB95FDE1" blockId="98.[707,1194,194,1065]" pageId="98" pageNumber="99">
<emphasis id="4E949A27D540B20887C241A9EBD5FE66" bold="true" box="[731,929,460,481]" pageId="98" pageNumber="99">Taxonomic History.</emphasis>
<bibRefCitation id="18713BC4D540B20886B741A9EC32FE66" author="Howden H." box="[942,1094,460,481]" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="98" pageNumber="99" pagination="257 - 264" part="57" refId="ref123283" refString="Howden H. 1955. Some new species and records of North American Scarabaeidae. Proceedings of the Entomological Society of Washington 57: 257 - 264." title="Some new species and records of North American Scarabaeidae" type="journal article" year="1955">Howden (1955)</bibRefCitation>
described the species in the genus
<taxonomicName id="BBE03DB6D540B20886DD4182EC30FE7B" authorityName="Burmeister" authorityYear="1842" box="[964,1092,487,508]" class="Insecta" family="Scarabaeidae" genus="Stephanucha" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis id="4E949A27D540B20886DD4182EC30FE7B" box="[964,1092,487,508]" italics="true" pageId="98" pageNumber="99">Stephanucha</emphasis>
</taxonomicName>
based on
<specimenCount id="6AE68DBCD540B20887DA4264EB26FD91" box="[707,850,513,534]" pageId="98" pageNumber="99" type="generic">two specimens</specimenCount>
collected in southern
<collectingRegion id="BE2488D7D540B20881374264EC1CFD91" box="[1070,1128,513,534]" country="United States of America" name="Texas" pageId="98" pageNumber="99">Texas</collectingRegion>
,
<collectingCountry id="04F706A5D540B208816A4264ECD2FD91" box="[1139,1190,513,534]" name="United States of America" pageId="98" pageNumber="99">USA</collectingCountry>
.
<bibRefCitation id="18713BC4D540B20887DA4279EBA9FDB6" author="Ratcliffe, B. C. &amp; M. J. Paulsen" box="[707,989,540,561]" journalOrPublisher="Bulletin of the University of Nebraska State Museum" pageId="98" pageNumber="99" pagination="1 - 570" part="22" refId="ref124724" refString="Ratcliffe, B. C., and M. J. Paulsen. 2008. The scarabaeoid beetles of Nebraska (Coleoptera: Scarabaeoidea). Bulletin of the University of Nebraska State Museum 22: 1 - 570." title="The scarabaeoid beetles of Nebraska (Coleoptera: Scarabaeoidea)" type="journal article" year="2008">Ratcliffe and Paulsen (2008)</bibRefCitation>
placed
<taxonomicName id="BBE03DB6D540B20881344279ECDDFDB6" authorityName="Burmeister" authorityYear="1842" box="[1069,1193,540,561]" class="Insecta" family="Scarabaeidae" genus="Stephanucha" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis id="4E949A27D540B20881344279ECDDFDB6" box="[1069,1193,540,561]" italics="true" pageId="98" pageNumber="99">Stephanucha</emphasis>
</taxonomicName>
in synonymy with
<taxonomicName id="BBE03DB6D540B20886954253EB98FDCC" box="[908,1004,566,587]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis id="4E949A27D540B20886954253EB98FDCC" box="[908,1004,566,587]" italics="true" pageId="98" pageNumber="99">Euphoria</emphasis>
</taxonomicName>
, thus transferring
<taxonomicName id="BBE03DB6D540B20887DA4234EB50FDE1" baseAuthorityName="Howden" baseAuthorityYear="1955" box="[707,804,593,614]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="anneae">
<emphasis id="4E949A27D540B20887DA4234EAA5FDE1" box="[707,721,593,614]" italics="true" pageId="98" pageNumber="99">E</emphasis>
.
<emphasis id="4E949A27D540B20887C64234EB50FDE1" box="[735,804,593,614]" italics="true" pageId="98" pageNumber="99">anneae</emphasis>
</taxonomicName>
to the latter genus.
</paragraph>
<paragraph id="7C5F4635D540B20887C2420EEBD2FD57" blockId="98.[707,1194,194,1065]" pageId="98" pageNumber="99">
<emphasis id="4E949A27D540B20887C2420EEBF7FD07" bold="true" box="[731,899,619,640]" pageId="98" pageNumber="99">Natural History.</emphasis>
Adults have been seen flying low over sandy soil (
<bibRefCitation id="18713BC4D540B208868742E3EC45FD1C" author="Howden H." box="[926,1073,646,667]" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="98" pageNumber="99" pagination="257 - 264" part="57" refId="ref123283" refString="Howden H. 1955. Some new species and records of North American Scarabaeidae. Proceedings of the Entomological Society of Washington 57: 257 - 264." title="Some new species and records of North American Scarabaeidae" type="journal article" year="1955">Howden 1955</bibRefCitation>
).
<bibRefCitation id="18713BC4D540B208815F42E3EB33FD32" author="Riley, E. G. &amp; C. S. Wolfe" journalOrPublisher="Southwestern Entomologist, supplement" pageId="98" pageNumber="99" pagination="1 - 37" part="26" refId="ref124803" refString="Riley, E. G., and C. S. Wolfe. 2003. An annotated checklist of the Scarabaeoidea of Texas. Southwestern Entomologist, supplement No. 26: 1 - 37." title="An annotated checklist of the Scarabaeoidea of Texas" type="journal article" year="2003">Riley and Wolfe (2003)</bibRefCitation>
collected both adults and larvae in mounds of
<taxonomicName id="BBE03DB6D540B208862D42DEEBD2FD57" box="[820,934,699,720]" class="Mammalia" family="Geomyidae" genus="Geomys" kingdom="Animalia" order="Rodentia" pageId="98" pageNumber="99" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="4E949A27D540B208862D42DEEBF5FD57" box="[820,897,699,720]" italics="true" pageId="98" pageNumber="99">Geomys</emphasis>
sp.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="34FA15BED540B20887C242B0EC57FCE8" pageId="98" pageNumber="99" type="distribution">
<paragraph id="7C5F4635D540B20887C242B0EB5EFC82" blockId="98.[707,1194,194,1065]" pageId="98" pageNumber="99">
<emphasis id="4E949A27D540B20887C242B0EBA5FD6D" bold="true" box="[731,977,725,746]" pageId="98" pageNumber="99">Temporal Distribution.</emphasis>
May (3), June (1) (
<figureCitation id="E4DB5AB0D540B20887D34295EB6BFC82" box="[714,799,752,773]" captionStart="Fig" captionStartId="174.[197,229,1686,1705]" captionTargetBox="[298,1132,196,1662]" captionTargetId="figure-0@174.[174,1192,183,1662]" captionTargetPageId="174" captionText="Fig. 55. Euphoria anneae. a) Habitus, b) Head, c) Parameres, d) Color variation, e) Temporal distribution, f) Geographic distribution in the United States." figureDoi="http://doi.org/10.5281/zenodo.4907368" httpUri="https://zenodo.org/record/4907368/files/figure.png" pageId="98" pageNumber="99">Fig. 55e</figureCitation>
).
</paragraph>
<paragraph id="7C5F4635D540B20887C2436FEC57FCE8" blockId="98.[707,1194,194,1065]" pageId="98" pageNumber="99">
<emphasis id="4E949A27D540B20887C2436FEBA8FC98" bold="true" box="[731,988,778,799]" pageId="98" pageNumber="99">Geographic Distribution.</emphasis>
Known only from southern
<collectingRegion id="BE2488D7D540B20886044340EB23FCBD" box="[797,855,805,826]" country="United States of America" name="Texas" pageId="98" pageNumber="99">Texas</collectingRegion>
,
<collectingCountry id="04F706A5D540B208867A4340EBE7FCBD" box="[867,915,805,826]" name="United States of America" pageId="98" pageNumber="99">USA</collectingCountry>
(
<figureCitation id="E4DB5AB0D540B20886BA4340EB81FCBD" box="[931,1013,805,826]" captionStart="Fig" captionStartId="174.[197,229,1686,1705]" captionTargetBox="[298,1132,196,1662]" captionTargetId="figure-0@174.[174,1192,183,1662]" captionTargetPageId="174" captionText="Fig. 55. Euphoria anneae. a) Habitus, b) Head, c) Parameres, d) Color variation, e) Temporal distribution, f) Geographic distribution in the United States." figureDoi="http://doi.org/10.5281/zenodo.4907368" httpUri="https://zenodo.org/record/4907368/files/figure.png" pageId="98" pageNumber="99">Fig. 55f</figureCitation>
).
<bibRefCitation id="18713BC4D540B20881114340EB76FCD2" author="Riley, E. G. &amp; C. S. Wolfe" journalOrPublisher="Southwestern Entomologist, supplement" pageId="98" pageNumber="99" pagination="1 - 37" part="26" refId="ref124803" refString="Riley, E. G., and C. S. Wolfe. 2003. An annotated checklist of the Scarabaeoidea of Texas. Southwestern Entomologist, supplement No. 26: 1 - 37." title="An annotated checklist of the Scarabaeoidea of Texas" type="journal article" year="2003">Riley and Wolfe (2003)</bibRefCitation>
recorded the species from Welder Wildlife Refuge,
<collectingCounty id="953E3EB9D540B208860C433FEBADFCE8" box="[789,985,858,879]" pageId="98" pageNumber="99">San Patricio County</collectingCounty>
,
<collectingRegion id="BE2488D7D540B20886FC433FEC6BFCE8" box="[997,1055,858,879]" country="United States of America" name="Texas" pageId="98" pageNumber="99">Texas</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="34FA15BED540B20887C24310ECDDFBAE" pageId="98" pageNumber="99" type="materials_examined">
<paragraph id="7C5F4635D540B20887C24310ECDDFBAE" blockId="98.[707,1194,194,1065]" pageId="98" pageNumber="99">
<emphasis id="4E949A27D540B20887C24310EBA3FC0D" bold="true" box="[731,983,885,906]" pageId="98" pageNumber="99">Specimens Examined (4).</emphasis>
Type material:
<materialsCitation id="CC884C68D540B208816B4310EC16FC72" ID-GBIF-Occurrence="3909183464" collectionCode="CMNC" collectorName="H. F. Howden &amp; W. Cloyd" country="United States of America" county="Cameron Co." location="Kenedy Co." municipality="Padre Island" pageId="98" pageNumber="99" specimenCode="BLNO/ 003345" specimenCount="1" stateProvince="Texas" typeStatus="holotype">
<typeStatus id="A35BF897D540B208816B4310EA99FC23" pageId="98" pageNumber="99" type="holotype">Holotype</typeStatus>
male at
<collectionCode id="1AF1DEF0D540B208865343EAEBE6FC23" box="[842,914,911,932]" pageId="98" pageNumber="99">CMNC</collectionCode>
labeled “Los Olmos Creek (
<quantity id="BB18EBD0D540B20887D343CFEB77FC38" box="[714,771,938,959]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="98" pageNumber="99" unit="mi" value="2.0">2 mi.</quantity>
S.)/ U.S. 77, Tex, May 31,/ 1954
<collectorName id="D11523E3D540B208816C43CFEB6FFC5E" pageId="98" pageNumber="99">H. F. Howden</collectorName>
,
<collectorName id="D11523E3D540B208863243A1EBE5FC5E" box="[811,913,964,985]" pageId="98" pageNumber="99">W. Cloyd</collectorName>
//
<typeStatus id="A35BF897D540B20886B443A1EC7FFC5E" box="[941,1035,964,985]" pageId="98" pageNumber="99" type="holotype">Holotype</typeStatus>
/
<taxonomicName id="BBE03DB6D540B208813943A1ECD5FC5E" authorityName="Burmeister" authorityYear="1842" box="[1056,1185,964,985]" class="Insecta" family="Scarabaeidae" genus="Stephanucha" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">Stephanucha</taxonomicName>
/ anneae/
<collectorName id="D11523E3D540B208860443BAEBE3FC73" box="[797,919,991,1012]" pageId="98" pageNumber="99">H. Howden</collectorName>
//
<specimenCode id="2C46EE4ED540B20886AA43BAEC23FC73" box="[947,1111,991,1012]" collectionCode="BLNO" pageId="98" pageNumber="99">BLNO/ 003345</specimenCode>
</materialsCitation>
Other material:
<emphasis id="4E949A27D540B2088602439FEB3FFB88" bold="true" box="[795,843,1018,1039]" pageId="98" pageNumber="99">
<collectingCountry id="04F706A5D540B2088602439FEB3FFB88" box="[795,843,1018,1039]" name="United States of America" pageId="98" pageNumber="99">USA</collectingCountry>
</emphasis>
(3):
<collectingRegion id="BE2488D7D540B2088667439FEBA4FB88" box="[894,976,1018,1039]" country="United States of America" name="Texas" pageId="98" pageNumber="99">TEXAS</collectingRegion>
:
<collectingCounty id="953E3EB9D540B20886C5439FEC15FB88" box="[988,1121,1018,1039]" pageId="98" pageNumber="99">
<emphasis id="4E949A27D540B20886C5439FEC2FFB88" box="[988,1115,1018,1039]" italics="true" pageId="98" pageNumber="99">Cameron Co</emphasis>
.
</collectingCounty>
:
<collectingMunicipality id="9C3BDC4FD540B208816B439FEA8BFBAE" pageId="98" pageNumber="99">Padre Island</collectingMunicipality>
(1);
<location id="793F10EED540B20886354471EBE9FBAE" LSID="urn:lsid:plazi:treatment:F449F723D543B20887EB454DECDDFBAE:793F10EED540B20886354471EBE9FBAE" box="[812,925,1044,1065]" country="United States of America" county="Cameron Co." municipality="Padre Island" name="Kenedy Co." pageId="98" pageNumber="99" stateProvince="Texas">
<emphasis id="4E949A27D540B20886354471EBE2FBAE" box="[812,918,1044,1065]" italics="true" pageId="98" pageNumber="99">Kenedy Co</emphasis>
.
</location>
:
<location id="793F10EED540B20886B04471EB95FBAE" LSID="urn:lsid:plazi:treatment:F449F723D543B20887EB454DECDDFBAE:793F10EED540B20886B04471EB95FBAE" box="[937,993,1044,1065]" country="United States of America" county="Cameron Co." municipality="Padre Island" name="Sarita" pageId="98" pageNumber="99" stateProvince="Texas">Sarita</location>
(
<quantity id="BB18EBD0D540B20886F74471EC6DFBAE" box="[1006,1049,1044,1065]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="98" pageNumber="99" unit="mi" value="2.0">2 mi</quantity>
N,
<quantity id="BB18EBD0D540B20881254471EC12FBAE" box="[1084,1126,1044,1065]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="98" pageNumber="99" unit="mi" value="7.0">7 mi</quantity>
S) (2).
</paragraph>
</subSubSection>
</treatment>
</document>