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<document id="C11507D91278C638A2C68726D200E2B3" ID-CLB-Dataset="7806" ID-DOI="10.5281/zenodo.4503596" ID-GBIF-Dataset="9a3c7406-d4a1-4480-af4b-413daed3c500" ID-ISSN="0374-1036" ID-RefBankID="FF8F4870FFB95708A713FFC9743EFFF5" ID-Zenodo-Dep="4503596" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612461472076" checkinUser="felipe" docAuthor="Hromádka, Lubomír" docDate="2008" docId="E76D87CEC939FFAF677CF1F3FDA1163A" docLanguage="en" docName="ActaEntMusNatPra.48.1.51-65.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 48 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Philonthus lamtoensis subsp. lamtoensis lamtoensis Levasseur 1967" docType="treatment" docVersion="8" lastPageNumber="58" masterDocId="1B54FFB6C93EFFA8663FF129FF501064" masterDocTitle="Revision of Afrotropical species of the Philonthus peripateticus species group (Coleoptera: Staphylinidae: Philonthina)" masterLastPageNumber="65" masterPageNumber="51" pageNumber="58" updateTime="1698927111911" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="26E1A9EBA6177C5FD981297ED2732AEF">Revision of Afrotropical species of the Philonthus peripateticus species group (Coleoptera: Staphylinidae: Philonthina)</mods:title>
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<mods:namePart id="4CB6B303908D555F2392E9CC321F7FB0">Hromádka, Lubomír</mods:namePart>
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<mods:date id="9B9D39DF0A88501917839BCA5CCA7077">2008</mods:date>
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<treatment id="E76D87CEC939FFAF677CF1F3FDA1163A" ID-DOI="http://doi.org/10.5281/zenodo.4594268" ID-GBIF-Taxon="180302218" ID-Zenodo-Dep="4594268" LSID="urn:lsid:plazi:treatment:E76D87CEC939FFAF677CF1F3FDA1163A" httpUri="http://treatment.plazi.org/id/E76D87CEC939FFAF677CF1F3FDA1163A" lastPageNumber="58" pageId="7" pageNumber="58">
<subSubSection id="27DE6553C939FFAF677CF1F3FC3E1097" box="[323,878,218,243]" pageId="7" pageNumber="58" type="nomenclature">
<paragraph id="6F7B36D8C939FFAF677CF1F3FC3E1097" blockId="7.[323,878,218,243]" box="[323,878,218,243]" pageId="7" pageNumber="58">
<heading id="343381B4C939FFAF677CF1F3FC3E1097" bold="true" box="[323,878,218,243]" centered="true" fontSize="8" level="2" pageId="7" pageNumber="58" reason="3">
<taxonomicName id="A8C44D5BC939FFAF677CF1F3FC3E1097" ID-CoL="8YF7W" authority="lamtoensis Levasseur, 1967" authorityName="lamtoensis Levasseur" authorityYear="1967" box="[323,878,218,243]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="subSpecies" species="lamtoensis" subSpecies="lamtoensis">
<emphasis id="5DB0EACAC939FFAF677CF1F3FC3E1097" bold="true" box="[323,878,218,243]" pageId="7" pageNumber="58">
<emphasis id="5DB0EACAC939FFAF677CF1F3FDE41097" bold="true" box="[323,692,218,243]" italics="true" pageId="7" pageNumber="58">Philonthus lamtoensis lamtoensis</emphasis>
Levasseur, 1967
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="27DE6553C939FFAF641EF1D0FDC01168" box="[545,656,249,268]" pageId="7" pageNumber="58" type="description">
<paragraph id="6F7B36D8C939FFAF641EF1D0FDC01168" blockId="7.[113,709,249,303]" box="[545,656,249,268]" pageId="7" pageNumber="58">
(
<figureCitation id="F7FF2A5DC939FFAF6417F1D0FDDA1168" box="[552,650,249,268]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="6.[151,190,1435,1454]" captionStartId-1="9.[113,152,1524,1543]" captionTargetBox-0="[251,1037,219,1372]" captionTargetBox-1="[121,1082,220,1478]" captionTargetId-0="figure-10@6.[231,1047,212,1387]" captionTargetId-1="figure-63@9.[118,1084,219,1479]" captionTargetPageId-0="6" captionTargetPageId-1="9" captionText-0="Figs. 11-19. 11-12 Philonthus dendroaspis sp. nov.; 13-18 P. katangaensis Levasseur, 1967; 19 P. lamtoensis Levasseur, 1967. 11, 18 apical portion of sternite VIII, ventral view; 12 sternite IX, ventral view; 13, 19 head; 14-16 aedeagus: 14 ventral view, 15 dorsal view, 16 lateral view; 17 underside of paramere, with sensory peg setae." captionText-1="Figs. 20-28. 20-24 Philonthus lamtoensis Levasseur, 1967; 25-28 P. madianus Bernhauer, 1936. 20-22, 25- 27 aedeagus: 20, 25 ventral view, 21, 26 dorsal view, 22, 27 lateral view; 23, 28 undersite of paramere, with sensory peg setae; 24 apical portion of sternite VIII, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.4503602" figureDoi-1="http://doi.org/10.5281/zenodo.4503610" httpUri-0="https://zenodo.org/record/4503602/files/figure.png" httpUri-1="https://zenodo.org/record/4503610/files/figure.png" pageId="7" pageNumber="58">Figs. 19-24</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="27DE6553C939FFAF664EF034FD95114B" box="[113,709,284,303]" pageId="7" pageNumber="58" type="reference_group">
<paragraph id="6F7B36D8C939FFAF664EF034FD95114B" blockId="7.[113,709,249,303]" box="[113,709,284,303]" pageId="7" pageNumber="58">
<taxonomicName id="A8C44D5BC939FFAF664EF034FD90114B" ID-CoL="8YF7W" authority="lamtoensis Levasseur, 1967: 1626" authorityName="Levasseur" authorityPageNumber="1626" authorityYear="1967" box="[113,704,284,303]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="subSpecies" species="lamtoensis" subGenus="Raucalius" subSpecies="lamtoensis">
<emphasis id="5DB0EACAC939FFAF664EF034FF9D114B" box="[113,205,285,303]" italics="true" pageId="7" pageNumber="58">Philonthus</emphasis>
(
<emphasis id="5DB0EACAC939FFAF66E5F034FE7F114B" box="[218,303,285,303]" italics="true" pageId="7" pageNumber="58">Raucalius</emphasis>
)
<emphasis id="5DB0EACAC939FFAF6703F034FEA8114B" box="[316,504,285,303]" italics="true" pageId="7" pageNumber="58">lamtoensis lamtoensis</emphasis>
Levasseur, 1967: 1626
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="27DE6553C939FFAF664EF06FFED611EA" pageId="7" pageNumber="58" type="materials_examined">
<paragraph id="6F7B36D8C939FFAF664EF06FFD4C113D" blockId="7.[113,1088,325,398]" box="[113,540,325,345]" pageId="7" pageNumber="58">
<emphasis id="5DB0EACAC939FFAF664EF06FFFBA113D" bold="true" box="[113,234,325,345]" pageId="7" pageNumber="58">
<typeStatus id="B07F887AC939FFAF664EF06FFFCE113D" box="[113,158,326,345]" pageId="7" pageNumber="58">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="17D37648C939FFAF66D0F06FFE06113D" box="[239,342,326,345]" name="Ivory Coast" pageId="7" pageNumber="58">Ivory Coast</collectingCountry>
, Toumodi env., Lamto.
</paragraph>
<paragraph id="6F7B36D8C939FFAF664EF049FED611EA" blockId="7.[113,1088,325,398]" pageId="7" pageNumber="58">
<emphasis id="5DB0EACAC939FFAF664EF049FE021117" bold="true" box="[113,338,351,371]" pageId="7" pageNumber="58">Type material examined.</emphasis>
<materialsCitation id="DFAC3C85C939FFAF6769F048FED611EA" ID-GBIF-Occurrence="3047799305" collectingDate="1962-06-18" collectionCode="MNHN" country="Ivory Coast" location="Lamto" pageId="7" pageNumber="58" specimenCount="1" typeStatus="holotype">
<typeStatus id="B07F887AC939FFAF6769F048FEF91116" box="[342,425,353,372]" pageId="7" pageNumber="58" type="holotype">HOLOTYPE</typeStatus>
: ♁,
<collectingCountry id="17D37648C939FFAF67EEF049FD191117" box="[465,585,352,371]" name="Ivory Coast" pageId="7" pageNumber="58">Côte d´Ivoire</collectingCountry>
,
<location id="6A1B6003C939FFAF646FF048FDD81110" LSID="urn:lsid:plazi:treatment:E76D87CEC939FFAF677CF1F3FDA1163A:6A1B6003C939FFAF646FF048FDD81110" box="[592,648,353,372]" country="Ivory Coast" name="Lamto" pageId="7" pageNumber="58">Lamto</location>
(Toumodi)
<date id="1B7A1018C939FFAF64D3F049FC1A1117" box="[748,842,352,371]" pageId="7" pageNumber="58" value="1962-06-18">
<collectingDate id="0B3EE9F0C939FFAF64D3F049FC1A1117" box="[748,842,352,371]" pageId="7" pageNumber="58" value="1962-06-18">18.vi.1962</collectingDate>
</date>
. [white oblong label with red margin, handwritten] (
<collectionCode id="09D5AE1DC939FFAF6709F052FE2C11EA" box="[310,380,379,398]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="58">MNHN</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="27DE6553C939FFAF664EF08FFEB815FA" pageId="7" pageNumber="58" type="description">
<paragraph id="6F7B36D8C939FFAF664EF08FFE3111BA" blockId="7.[113,1089,422,1631]" pageId="7" pageNumber="58">
<emphasis id="5DB0EACAC939FFAF664EF08FFE4711DA" bold="true" box="[113,279,422,446]" pageId="7" pageNumber="58">Redescription.</emphasis>
Body length
<quantity id="A83C9B3DC939FFAF6798F08FFD5411DA" box="[423,516,422,446]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="7" pageNumber="58" unit="mm" value="11.6">11.6 mm</quantity>
(with stretched abdomen), length of fore body (to the end of elytra)
<quantity id="A83C9B3DC939FFAF6737F0EFFE0D11BA" box="[264,349,454,478]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="7" pageNumber="58" unit="mm" value="5.7">5.7 mm</quantity>
.
</paragraph>
<paragraph id="6F7B36D8C939FFAF66AFF0CFFED1125A" blockId="7.[113,1089,422,1631]" pageId="7" pageNumber="58">Colouration. Head and elytra black, pronotum and abdomen black-brown, clypeus along anterior margin, maxillary and labial palpi, antennae and legs brown-yellow, all tibiae and femora somewhat darker.</paragraph>
<paragraph id="6F7B36D8C939FFAF66AFF36FFCBE12BA" blockId="7.[113,1089,422,1631]" pageId="7" pageNumber="58">
Head (
<figureCitation id="F7FF2A5DC939FFAF66E8F36FFE75123A" box="[215,293,582,606]" captionStart="Figs" captionStartId="6.[151,190,1435,1454]" captionTargetBox="[251,1037,219,1372]" captionTargetId="figure-10@6.[231,1047,212,1387]" captionTargetPageId="6" captionText="Figs. 11-19. 11-12 Philonthus dendroaspis sp. nov.; 13-18 P. katangaensis Levasseur, 1967; 19 P. lamtoensis Levasseur, 1967. 11, 18 apical portion of sternite VIII, ventral view; 12 sternite IX, ventral view; 13, 19 head; 14-16 aedeagus: 14 ventral view, 15 dorsal view, 16 lateral view; 17 underside of paramere, with sensory peg setae." figureDoi="http://doi.org/10.5281/zenodo.4503602" httpUri="https://zenodo.org/record/4503602/files/figure.png" pageId="7" pageNumber="58">Fig. 19</figureCitation>
) markedly transverse (ratio 36: 26), parallel-sided. Eyes slightly convex, hardly longer than temples (ratio 12: 11). Posterior angles of head distinct, with a few bristles. Four coarse punctures between eyes, distance between medial interocular punctures about five times as large as the distance between medial and lateral interocular punctures. Temporal area with several punctures of different size. Surface without microsculpture, shiny.
</paragraph>
<paragraph id="6F7B36D8C939FFAF66AFF3CFFC0B137A" blockId="7.[113,1089,422,1631]" pageId="7" pageNumber="58">Antennae relatively long, reaching third puncture of dorsal rows when reclined. Antennomeres 1-3 longer than wide, remaining antennomeres as long as wide.</paragraph>
<paragraph id="6F7B36D8C939FFAF66AFF20EFCB3131A" blockId="7.[113,1089,422,1631]" pageId="7" pageNumber="58">Pronotum somewhat wider than long (ratio 45: 42), parallel-sided, anterior angles rectangular, posterior angles markedly rounded. Each dorsal row with three almost equidistant punctures; each sublateral row with one puncture. Surface without microsculpture.</paragraph>
<paragraph id="6F7B36D8C939FFAF66AFF2AFFEDD13DA" blockId="7.[113,1089,422,1631]" pageId="7" pageNumber="58">Scutellum very densely and coarsely punctate. Punctures larger than eye-facets, mostly coalescent. Setation black.</paragraph>
<paragraph id="6F7B36D8C939FFAF66AFF2EFFD5C147A" blockId="7.[113,1089,422,1631]" pageId="7" pageNumber="58">Elytra slightly wider than long (ratio 52: 46), parallel-sided, insignificantly widened posteriad. Punctation dense and coarse, punctures much larger than eye-facets. Distance between punctures very short. Setation greyish.</paragraph>
<paragraph id="6F7B36D8C939FFAF66AFF50EFFE4141A" blockId="7.[113,1089,422,1631]" pageId="7" pageNumber="58">Legs. Metatarsus somewhat longer than metatibia (ratio 32: 30). Metatarsomere 1 longer than metatarsomeres 2-4 combined, metatarsomere 5 as long as metatarsomeres 3-4 combined.</paragraph>
<paragraph id="6F7B36D8C939FFAF66AFF5AFFE9B149A" blockId="7.[113,1089,422,1631]" pageId="7" pageNumber="58">Abdomen wide, from third visible tergite slightly narrowed towards both base and apex. Punctation between two basal lines on first three visible tergites coarse and dense. Punctation of all tergites very dense, much finer than that on elytra. Size of punctures slightly larger than eye-facets. Setation brown-grey.</paragraph>
<paragraph id="6F7B36D8C939FFAF66AFF42FFF90151A" blockId="7.[113,1089,422,1631]" pageId="7" pageNumber="58">
Male. Protarsomeres 1-3 moderately dilated, each covered with modified yellow setae ventrally, protarsomere 4 narrower than preceding ones. Sternite VIII (
<figureCitation id="F7FF2A5DC939FFAF6548F40FFC9A155A" box="[887,970,1318,1342]" captionStart="Figs" captionStartId="9.[113,152,1524,1543]" captionTargetBox="[121,1082,220,1478]" captionTargetId="figure-63@9.[118,1084,219,1479]" captionTargetPageId="9" captionText="Figs. 20-28. 20-24 Philonthus lamtoensis Levasseur, 1967; 25-28 P. madianus Bernhauer, 1936. 20-22, 25- 27 aedeagus: 20, 25 ventral view, 21, 26 dorsal view, 22, 27 lateral view; 23, 28 undersite of paramere, with sensory peg setae; 24 apical portion of sternite VIII, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4503610" httpUri="https://zenodo.org/record/4503610/files/figure.png" pageId="7" pageNumber="58">Fig. 24</figureCitation>
), aedeagus with branches of paramere narrowly separated medially, reaching apex of median lobe. (
<figureCitation id="F7FF2A5DC939FFAF6234F46FFFE3151A" captionStart="Figs" captionStartId="9.[113,152,1524,1543]" captionTargetBox="[121,1082,220,1478]" captionTargetId="figure-63@9.[118,1084,219,1479]" captionTargetPageId="9" captionText="Figs. 20-28. 20-24 Philonthus lamtoensis Levasseur, 1967; 25-28 P. madianus Bernhauer, 1936. 20-22, 25- 27 aedeagus: 20, 25 ventral view, 21, 26 dorsal view, 22, 27 lateral view; 23, 28 undersite of paramere, with sensory peg setae; 24 apical portion of sternite VIII, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4503610" httpUri="https://zenodo.org/record/4503610/files/figure.png" pageId="7" pageNumber="58">Figs. 20-23</figureCitation>
).
</paragraph>
<paragraph id="6F7B36D8C939FFAF66AFF4AFFEB815FA" blockId="7.[113,1089,422,1631]" box="[144,488,1414,1438]" pageId="7" pageNumber="58">Female. Unknown to the author.</paragraph>
</subSubSection>
<subSubSection id="27DE6553C939FFAF664EF48FFE36165A" pageId="7" pageNumber="58" type="diagnosis">
<paragraph id="6F7B36D8C939FFAF664EF48FFD65167A" blockId="7.[113,1089,422,1631]" pageId="7" pageNumber="58">
<emphasis id="5DB0EACAC939FFAF664EF48FFE3A15DA" bold="true" box="[113,362,1446,1470]" pageId="7" pageNumber="58">Differential diagnosis.</emphasis>
<taxonomicName id="A8C44D5BC939FFAF6750F48EFD8115DA" authority="lamtoensis" authorityName="lamtoensis Levasseur" authorityYear="1967" box="[367,721,1447,1470]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="subSpecies" species="lamtoensis" subSpecies="lamtoensis">
<emphasis id="5DB0EACAC939FFAF6750F48EFD8115DA" box="[367,721,1447,1470]" italics="true" pageId="7" pageNumber="58">Philonthus lamtoensis lamtoensis</emphasis>
</taxonomicName>
may be distinguished from
<taxonomicName id="A8C44D5BC939FFAF65CBF48EFE7F15BA" authorityName="Hromadka" authorityYear="2005" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="subSpecies" species="lamtoensis" subSpecies="shamnesis">
<emphasis id="5DB0EACAC939FFAF65CBF48EFB5415DA" box="[1012,1028,1447,1470]" italics="true" pageId="7" pageNumber="58">P</emphasis>
.
<emphasis id="5DB0EACAC939FFAF622FF48EFE7F15BA" italics="true" pageId="7" pageNumber="58">lamtoensis shamnesis</emphasis>
</taxonomicName>
by the slender body and paler antennae and legs. It differs from the closely related
<taxonomicName id="A8C44D5BC939FFAF66FFF4CEFE31159A" authorityName="Tottenham" authorityYear="1949" box="[192,353,1511,1534]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="rugulipennis">
<emphasis id="5DB0EACAC939FFAF66FFF4CEFE31159A" box="[192,353,1511,1534]" italics="true" pageId="7" pageNumber="58">P. rugulipennis</emphasis>
</taxonomicName>
and
<taxonomicName id="A8C44D5BC939FFAF67AAF4CEFD69159A" authorityName="Levasseur" authorityYear="1967" box="[405,569,1511,1534]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="katangaensis">
<emphasis id="5DB0EACAC939FFAF67AAF4CEFD69159A" box="[405,569,1511,1534]" italics="true" pageId="7" pageNumber="58">P. katangaensis</emphasis>
</taxonomicName>
by the different shape of the aedeagus, and from
<taxonomicName id="A8C44D5BC939FFAF664EF72EFE43167A" authorityName="Tottenham" authorityYear="1949" box="[113,275,1543,1566]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="rugulipennis">
<emphasis id="5DB0EACAC939FFAF664EF72EFE43167A" box="[113,275,1543,1566]" italics="true" pageId="7" pageNumber="58">P. rugulipennis</emphasis>
</taxonomicName>
also by the paler antennae.
</paragraph>
<paragraph id="6F7B36D8C939FFAF664EF70FFE36165A" blockId="7.[113,1089,422,1631]" box="[113,358,1574,1598]" pageId="7" pageNumber="58">
<emphasis id="5DB0EACAC939FFAF664EF70FFFBF165A" bold="true" box="[113,239,1574,1598]" pageId="7" pageNumber="58">Bionomics.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="27DE6553C939FFAF664EF76FFDA1163A" box="[113,753,1606,1631]" pageId="7" pageNumber="58" type="distribution">
<paragraph id="6F7B36D8C939FFAF664EF76FFDA1163A" blockId="7.[113,1089,422,1631]" box="[113,753,1606,1631]" pageId="7" pageNumber="58">
<emphasis id="5DB0EACAC939FFAF664EF76FFE54163A" bold="true" box="[113,260,1606,1630]" pageId="7" pageNumber="58">Distribution.</emphasis>
<collectingCountry id="17D37648C939FFAF6735F76EFED7163A" box="[266,391,1606,1631]" name="Ivory Coast" pageId="7" pageNumber="58">Ivory Coast</collectingCountry>
(LEVASSEUR 1967,
<bibRefCitation id="0B554B29C939FFAF6471F76EFDB6163A" author="HERMAN L. H." box="[590,742,1606,1630]" pageId="7" pageNumber="58" pagination="2441 - 3020" refId="ref6077" refString="HERMAN L. H. 2001: Catalog of the Staphylinidae (Insecta, Coleoptera); 1758 to the end of the second millennium. V. Staphylinine group (Part 2): Diochini, Maorothiini, Othiini, Platyprosopini, Staphilinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina). Bulletin of the American Museum of Natural History 265: 2441 - 3020." type="journal article" year="2001">HERMAN 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>