307 lines
41 KiB
XML
307 lines
41 KiB
XML
<document ID-CLB-Dataset="57657" ID-DOI="http://dx.doi.org/10.3897/zookeys.1169.98681" ID-GBIF-Dataset="a6237e5a-06a8-4406-b236-ca4fe4183db1" ID-PMC="PMC10848866" ID-Pensoft-Pub="1313-2970-1169-47" ID-Pensoft-UUID="357C8407C0B25174B4953AC11828B559" ID-PubMed="38328028" ID-ZooBank="4E151EFF8B4A4159802C1C17869CDEFC" ModsDocID="1313-2970-1169-47" checkinTime="1688698756328" checkinUser="pensoft" docAuthor="Nugraha, Fitra Arya Dwi, Ahda, Yuni, Tjong, Djong Hon, Kurniawan, Nia, Riyanto, Awal, Fauzi, Muhammad Alif & Lin, Si-Min" docDate="2023" docId="DD6147C025CE519098ED4EE7EA7A0E53" docLanguage="en" docName="ZooKeys 1169: 47-64" docOrigin="ZooKeys 1169" docPubDate="2023-07-06" docSource="http://dx.doi.org/10.3897/zookeys.1169.98681" docTitle="Cyrtodactylus gonjong Nugraha & Ahda & Tjong & Kurniawan & Riyanto & Fauzi & Lin 2023, sp. nov." docType="treatment" docUuid="EFDA2781-5260-43A7-9A59-B5BBE26D5348" docUuidSource="ZooBank" docVersion="4" id="357C8407C0B25174B4953AC11828B559" lastPageNumber="47" masterDocId="357C8407C0B25174B4953AC11828B559" masterDocTitle="Common but ignored: a new species of Cyrtodactylus (Chordata, Reptilia, Squamata, Gekkonidae) from lowland Sumatra Barat, Indonesia" masterLastPageNumber="64" masterPageNumber="47" pageNumber="47" updateTime="1732824454477" updateUser="ExternalLinkService">
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<mods:title id="63F01BFCEBD530808E0950D4698A5E42">Common but ignored: a new species of Cyrtodactylus (Chordata, Reptilia, Squamata, Gekkonidae) from lowland Sumatra Barat, Indonesia</mods:title>
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<mods:name id="CD4CDE3DB72975A57B514D2DE10CA4B5" type="personal">
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<mods:roleTerm id="C3CA0CC2591282A3991BF1F80607C3EE">Author</mods:roleTerm>
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<mods:namePart id="B45CBF3B96CA31C0E2797C8AAC681A40">Nugraha, Fitra Arya Dwi</mods:namePart>
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<mods:nameIdentifier id="F22D56689065949D04D21A07B1D97819" type="ORCID">https://orcid.org/0000-0002-0048-8515</mods:nameIdentifier>
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<mods:affiliation id="3EBA6C1DD4FF708E4E1A4046A1163C2F">Department of Biology, Faculty of Mathematics and Natural Sciences, Universitas Negeri Padang, Jl. Hamka, Kota Padang 25132, Sumatra Barat, Indonesia</mods:affiliation>
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<mods:nameIdentifier id="D1D67AF4ED4E6797E06CE6B0B8E9E25A" type="email">fitraaryadn@fmipa.unp.ac.id</mods:nameIdentifier>
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<mods:namePart id="6561C16B717D51BB850DF56A9876E626">Ahda, Yuni</mods:namePart>
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<mods:nameIdentifier id="B917A21F08B0668387E031A85ED45E02" type="ORCID">https://orcid.org/0000-0003-0545-2965</mods:nameIdentifier>
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<mods:affiliation id="7195426DD9D2C3FEE958BDF0D0587A98">Department of Biology, Faculty of Mathematics and Natural Sciences, Universitas Negeri Padang, Jl. Hamka, Kota Padang 25132, Sumatra Barat, Indonesia</mods:affiliation>
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</mods:name>
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<mods:namePart id="15C30A6C74B508720C07C81D75ADF67C">Tjong, Djong Hon</mods:namePart>
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<mods:nameIdentifier id="A7712236DC8295E89958853479BCBF0B" type="ORCID">https://orcid.org/0000-0002-4743-9964</mods:nameIdentifier>
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<mods:affiliation id="67BA9AD42C15FD85E3713661FD1AE007">Department of Biology, Faculty of Mathematics and Natural Sciences, Universitas Andalas, Limau Manis, Pauh, Kota Padang 25175, Sumatra Barat, Indonesia</mods:affiliation>
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</mods:name>
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<mods:roleTerm id="720CF916A238F651917E8F08695FA2B4">Author</mods:roleTerm>
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<mods:namePart id="B00FCBE9D85AC1576E7FE94F9C8C194B">Kurniawan, Nia</mods:namePart>
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<mods:affiliation id="EB342A823B135A0C15FEED7B5DD46306">Department of Biology, Faculty of Mathematics and Natural Sciences, Universitas Brawijaya, Jl. Veteran 65145, Ketawanggede, Lowokwaru, Kota Malang, Jawa Timur, Indonesia</mods:affiliation>
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</mods:name>
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<mods:namePart id="0CE914CA83658343C62662BB7A132A4D">Riyanto, Awal</mods:namePart>
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<mods:nameIdentifier id="FA272E55A7E66CF63BEF7C25B897AE48" type="ORCID">https://orcid.org/0000-0002-6887-9352</mods:nameIdentifier>
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<mods:affiliation id="610EBC4ED908FBCE7B59289783F10AE6">Herpetology Group Research, Biosystematic and Evolution Research Center - National Research and Innovation Agency (BRIN) - (" Museum Zoologicum Bogoriense "), Jakarta, Indonesia</mods:affiliation>
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</mods:name>
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<mods:namePart id="8251B89AD88DB29539762AF62C55D79E">Fauzi, Muhammad Alif</mods:namePart>
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<mods:nameIdentifier id="2C2FC5D8720D808FEDCE1FC9096DF9CA" type="ORCID">https://orcid.org/0000-0003-0975-9681</mods:nameIdentifier>
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<mods:affiliation id="3E16945FCD8CBE9BDEAD25AD8142A01B">Department of Biology, Faculty of Mathematics and Natural Sciences, Universitas Brawijaya, Jl. Veteran 65145, Ketawanggede, Lowokwaru, Kota Malang, Jawa Timur, Indonesia</mods:affiliation>
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</mods:role>
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<mods:namePart id="C7702D97645C1214C71355824C5BD375">Lin, Si-Min</mods:namePart>
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<mods:nameIdentifier id="B30C27AA20CCA6893280BC575CC434EB" type="ORCID">https://orcid.org/0000-0001-7080-706X</mods:nameIdentifier>
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<mods:affiliation id="1ED1B961095E439A06CA8DBAD999A544">Department of Life Science, National Taiwan Normal University, Taipei, Taiwan</mods:affiliation>
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<mods:title id="1DB909E91B58D746417789C1F00EA26D">ZooKeys</mods:title>
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<mods:part id="7F4C1DFA4E82419AA6E9C392D19E678D">
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<mods:date id="4B2167576754E210A503E6C6FF733B5C">2023</mods:date>
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<mods:number id="A433BBDA87EA43315363198FCA44CF3F">2023-07-06</mods:number>
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<mods:number id="A66B8FFDC9CADE515C06C31009EEC013">1169</mods:number>
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<mods:start id="F20F5F33516FD3F9A04BC85A2CD48A5B">47</mods:start>
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<mods:end id="0BFBEA136BFA13A7120799AFBA52BD09">64</mods:end>
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<mods:url id="14406D543CDA160BAFB542CFD7666360">http://dx.doi.org/10.3897/zookeys.1169.98681</mods:url>
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<mods:classification id="AFFA0BD0E023A6FF893CB67E86BF677B">journal article</mods:classification>
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<mods:identifier id="449FFF5283C3091972BC309FDE8836C7" type="DOI">http://dx.doi.org/10.3897/zookeys.1169.98681</mods:identifier>
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<treatment id="DD6147C025CE519098ED4EE7EA7A0E53" ID-GBIF-Taxon="210885575" LSID="urn:lsid:zoobank.org:act:EFDA2781-5260-43A7-9A59-B5BBE26D5348" httpUri="http://treatment.plazi.org/id/DD6147C025CE519098ED4EE7EA7A0E53" lastPageNumber="47" pageId="0" pageNumber="47" scope_class="Squamata" scope_family="Gekkonidae" scope_phylum="Chordata">
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<subSubSection id="4C0BA08ADFFF5974BD219411611D334F" pageId="0" pageNumber="47" type="nomenclature">
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<paragraph id="760416B270160F5F5BEFC808F749670E" pageId="0" pageNumber="47">
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<taxonomicName id="1F04D609938D8D8CEB1B87E7E4FA9FED" LSID="https://zoobank.org/EFDA2781-5260-43A7-9A59-B5BBE26D5348" authority="Nugraha & Ahda & Tjong & Kurniawan & Riyanto & Fauzi & Lin, 2023" authorityName="Nugraha & Ahda & Tjong & Kurniawan & Riyanto & Fauzi & Lin" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Cyrtodactylus gonjong" order="Squamata" pageId="0" pageNumber="47" rank="species" species="gonjong" status="sp. nov.">Cyrtodactylus gonjong</taxonomicName>
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<taxonomicNameLabel id="693E872C18A4F9ADC963AD25C63672EF" pageId="0" pageNumber="47">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="0E9146F27FFD56F1E976038362D6B870" pageId="0" pageNumber="47" type="type material">
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<paragraph id="7540DCEDF09FEB8E0C65FD4F42FB8CC0" pageId="0" pageNumber="47">Type material.</paragraph>
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<paragraph id="ECD7DFAA50B4182F41C2B1F521560B23" pageId="0" pageNumber="47">
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||
<emphasis id="D725825DEF0F803DA1E9B7C47D0ED97C" bold="true" pageId="0" pageNumber="47">
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||
<emphasis id="61A6A4FC468BB1097053C961BAE9A16A" italics="true" pageId="0" pageNumber="47">Holotype</emphasis>
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.
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</emphasis>
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UNP193 (Fig.
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<figureCitation id="9BEC189FFBE4BFAEEF04B0C1A5DCF421" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype (UNP 193, adult female, SVL 65.1 mm) A rostral view B gular view C precloacal scales D dorsal E ventrolateral F ventral view. A-D, F were taken after 1 month preservation E is a live individual." figureDoi="10.3897/zookeys.1169.98681.figure3" httpUri="https://binary.pensoft.net/fig/873865" pageId="0" pageNumber="47">3</figureCitation>
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) adult female, collected from Langkuik Tamiang waterfall (
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<geoCoordinate id="25F60B3100B2D5E4C8C4AB52C2AFCAC5" degrees="0" direction="south" minutes="26" orientation="latitude" precision="1" seconds="0.7404" value="-0.433539">0°26'0.7404"S</geoCoordinate>
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,
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<geoCoordinate id="AD2DD28EBC829EE330451A9205E7F557" degrees="100" direction="east" minutes="16" orientation="longitude" precision="1" seconds="44.5404" value="100.279045">100°16'44.5404"E</geoCoordinate>
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), Nagari (Village) Malalak Selatan, Kecamatan (District) Malalak, Kabupaten (Regency) Agam, Sumatra Barat Province, Indonesia, on 22 July 2022 by Fitra Arya Dwi Nugraha (FADN), Yunico Amardi (YA) and Mahesa Rafi.
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<emphasis id="AFCEDE75434C50192CB3CFE1F49E63CE" bold="true" italics="true" pageId="0" pageNumber="47">Paratypes</emphasis>
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(Fig.
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<figureCitation id="A563612F9A35420A8C2E4182DAAEA9D2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. The representative paratypes of C. gonjong sp. nov." figureDoi="10.3897/zookeys.1169.98681.figure4" httpUri="https://binary.pensoft.net/fig/873866" pageId="0" pageNumber="47">4</figureCitation>
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). One adult male (UNP045) collected from Anai Dempoland tourism object (
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<geoCoordinate id="088A109B3774B7228BAF1A1E9E48FD84" degrees="0" direction="south" minutes="29" orientation="latitude" precision="1" seconds="46.8312" value="-0.49634197">0°29'46.8312"S</geoCoordinate>
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,
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<geoCoordinate id="CFB12C1BA69B6A6D9D6A8712998D7A8B" degrees="100" direction="east" minutes="20" orientation="longitude" precision="1" seconds="7.638" value="100.33546">100°20'7.638"E</geoCoordinate>
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), village Kayu Tanam, district 2
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<normalizedToken id="137159B4DFFA925A6A32856F46322391" originalValue="×">x</normalizedToken>
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11 Kayu Tanam, regency Padang Pariaman, Sumatra Barat Province on 3 June 2020 by FADN and YA. Two adult males (UNP052, UNP053) collected from village Sungai Barameh (
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<geoCoordinate id="BBAEA0DF165BF0AE4043676841837A71" degrees="1" direction="south" minutes="1" orientation="latitude" precision="1" seconds="54.71" value="-1.0318639">1°1'54.71"S</geoCoordinate>
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,
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<geoCoordinate id="1B408AF5C3D883404A552FE7C9E6927D" degrees="100" direction="east" minutes="24" orientation="longitude" precision="1" seconds="43.69" value="100.41214">100°24'43.69"E</geoCoordinate>
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), district Padang Selatan, Padang city, Sumatra Barat Province on 31 January 2021 by FADN and YA. One adult male (UNP060) collected from Lembah Anai Nature Reserve (
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<geoCoordinate id="201C65060FE4DC79E831D8BA1A0E0228" degrees="0" direction="south" minutes="29" orientation="latitude" precision="15" seconds="24" value="-0.48999998">0°29'24"S</geoCoordinate>
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,
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<geoCoordinate id="47D44B9B5A839140EC24250CAF763C06" degrees="100" direction="east" minutes="20" orientation="longitude" precision="15" seconds="24" value="100.340004">100°20'24"E</geoCoordinate>
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), district X Koto, regency Tanah Datar, Sumatra Barat Province on 10 June 2021 by Mahesa Rafi, Katon Agusdi, Fadhil Raid and Fachrul Rozi Octavian. One adult male (UNP148) collected from Sarasah Uwak Waterfall (
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<geoCoordinate id="0760C65E343E15DB75A9E55D589B5D69" degrees="0" direction="south" minutes="54" orientation="latitude" precision="15" seconds="28" value="-0.9077777">0°54'28"S</geoCoordinate>
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,
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<geoCoordinate id="0C696FA0F8C9883F0551877C51BBBAA6" degrees="100" direction="east" minutes="28" orientation="longitude" precision="15" seconds="54" value="100.48167">100°28'54"E</geoCoordinate>
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), village Limau Manis, district Pauh, Padang city, Sumatra Barat Province on 31 May 2022 by FADN. Two adult females (UNP165, UNP167) collected from village Sungai Sirah (
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<geoCoordinate id="DBDAD1A814BB8B7F89C5FA3D1F65AB46" degrees="0" direction="south" minutes="24" orientation="latitude" precision="1" seconds="8.8128" value="-0.402448">0°24'8.8128"S</geoCoordinate>
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,
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<geoCoordinate id="515E7899EE61993E329EC6755D002E9A" degrees="100" direction="east" minutes="8" orientation="longitude" precision="1" seconds="37.6728" value="100.1438">100°8'37.6728"E</geoCoordinate>
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), district Sungai Geringging, regency Padang Pariaman, Sumatra Barat Province on 7 June 2022 by FADN and YA. One juvenile (UNP194), same data as holotype. Three adult males (UNP196, UNP199, UNP203) collected from village Koto Malintang (
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<geoCoordinate id="87C186FFA7E62F2CDE3641C29795D661" degrees="0" direction="south" minutes="17" orientation="latitude" precision="1" seconds="56.3748" value="-0.298993">0°17'56.3748"S</geoCoordinate>
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,
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<geoCoordinate id="FE5F123BB75627FAC7C0EB29379BA28A" degrees="100" direction="east" minutes="7" orientation="longitude" precision="1" seconds="42.5064" value="100.12848">100°7'42.5064"E</geoCoordinate>
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), district Tanjung Raya, regency Agam, Sumatra Barat Province.
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</paragraph>
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<caption id="BEDDF30C76B8195C496138345CDC53F5" doi="10.3897/zookeys.1169.98681.figure3" httpUri="https://binary.pensoft.net/fig/873865" pageId="0" pageNumber="47" start="Figure 3" startId="F3">
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||
<paragraph id="610105B19B377A3CEAF571015BAEC719" pageId="0" pageNumber="47">
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<emphasis id="12D34B47BD66279B2FD1A60716F526F5" bold="true" pageId="0" pageNumber="47">Figure 3.</emphasis>
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The holotype (UNP193, adult female, SVL 65.1 mm)
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<emphasis id="1B28A237502B74086CD7AF017D5D89B1" bold="true" pageId="0" pageNumber="47">A</emphasis>
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rostral view
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<emphasis id="49289038B66F60A1C672EAE258CF8561" bold="true" pageId="0" pageNumber="47">B</emphasis>
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gular view
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<emphasis id="AC428BE8428A65539C3EEDC21F1DB929" bold="true" pageId="0" pageNumber="47">C</emphasis>
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precloacal scales
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||
<emphasis id="F775400E4B542CB33B1D833FCD1F33B9" bold="true" pageId="0" pageNumber="47">D</emphasis>
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dorsal
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<emphasis id="71858A2F6D8DF2F80800F6D4DFB6A881" bold="true" pageId="0" pageNumber="47">E</emphasis>
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ventrolateral
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<emphasis id="7CA795A89E25891D07950FD7830A65E0" bold="true" pageId="0" pageNumber="47">F</emphasis>
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ventral view.
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<emphasis id="8698765534AAFF3E70F20AB01163E727" bold="true" pageId="0" pageNumber="47">A-D, F</emphasis>
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were taken after 1 month preservation
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<emphasis id="9E503CFDFB4C828BE98FA44C0538BA4B" bold="true" pageId="0" pageNumber="47">E</emphasis>
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is a live individual.
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</paragraph>
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</caption>
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<caption id="EA5B3315DB01D9D003F04A7644772EB1" doi="10.3897/zookeys.1169.98681.figure4" httpUri="https://binary.pensoft.net/fig/873866" pageId="0" pageNumber="47" start="Figure 4" startId="F4">
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||
<paragraph id="A0C2BF28C0576C73C1163FDF945DB692" pageId="0" pageNumber="47">
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<emphasis id="BF394FE593F06243838E42E7EB08DE13" bold="true" pageId="0" pageNumber="47">Figure 4.</emphasis>
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The representative paratypes of
|
||
<taxonomicName id="D77F5CE8A945D1E294E723240A1656BA" class="Amphibia" kingdom="Animalia" lsidName="C. gonjong" order="Anura" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="gonjong">
|
||
<emphasis id="283FD93C67E3B10703C9382AB5C7CE43" italics="true" pageId="0" pageNumber="47">C. gonjong</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="530771A2433DBF251B5D2923FB7ECF87" pageId="0" pageNumber="47" type="etymology">
|
||
<paragraph id="110F268EE53CCBB960BC36C4FB6DB06F" pageId="0" pageNumber="47">Etymology.</paragraph>
|
||
<paragraph id="B6B379347A482F57F5C53E2B80A19532" pageId="0" pageNumber="47">
|
||
The specific ephitet
|
||
<emphasis id="3812DF342A7F4100654504375DFA9C8A" italics="true" pageId="0" pageNumber="47">gonjong</emphasis>
|
||
is taken from the name of the roof style of the typical house in Sumatra Barat created by its ethnic people, called
|
||
<emphasis id="58CDB84370D1769AAF58DAF5CD1C0D7E" italics="true" pageId="0" pageNumber="47">Minang</emphasis>
|
||
. The house itself, called
|
||
<emphasis id="B124BD615D4B16514727A767FCDF803F" italics="true" pageId="0" pageNumber="47">Rumah Gadang</emphasis>
|
||
, and the unique style of the roof shape was inspired by the horn of buffalo, the most respected animal in Minang ethnology. The
|
||
<emphasis id="97BC23308A5193BB6CBFFB89BA14A9BE" italics="true" pageId="0" pageNumber="47">gonjong</emphasis>
|
||
has become a symbol used by the Minang people to show their ethnic identity outside Sumatra Barat and now is used not only for the house but also for government buildings, restaurant buildings, hotels or other public venues. This name is given as an honour to the Minang people because they were helpful during our survey.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="44DF06711025EFDF54FA023CB0053892" pageId="0" pageNumber="47" type="diagnosis">
|
||
<paragraph id="7087D0F7282A7C6B01D94600484C8F4B" pageId="0" pageNumber="47">Diagnosis.</paragraph>
|
||
<paragraph id="E08904941FAF53C0E5D7CEAAC3F3ED4E" pageId="0" pageNumber="47">
|
||
<taxonomicName id="22856760D1B411DC4F6FD239092D9176" authorityName="Nugraha & Ahda & Tjong & Kurniawan & Riyanto & Fauzi & Lin" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" lsidName="Cyrtodactylus gonjong" order="Squamata" pageId="0" pageNumber="47" rank="species" species="gonjong">
|
||
<emphasis id="D8AA40CDB03EB0AB6D18AD77C1E6C4CB" italics="true" pageId="0" pageNumber="47">Cyrtodactylus gonjong</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is assigned to the
|
||
<taxonomicName id="B8690CA90CBF2A135C199BAC75A15C6B" class="Amphibia" kingdom="Animalia" lsidName="C. agamensis" order="Anura" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="agamensis">
|
||
<emphasis id="6FB2417F41927333842A70A319FF2876" italics="true" pageId="0" pageNumber="47">C. agamensis</emphasis>
|
||
</taxonomicName>
|
||
group on the basis of its recovered phylogenetic position (Fig.
|
||
<figureCitation id="99539EB7EE2A4D1346AC392570CC0D0A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The maximum likelihood (ML) tree topology of the new species with other Cyrtodactylus inferred by the ND 2 gene sequences. The light orange box indicates the C. agamensis group members (the sample indicated by an asterisk denotes the holotype). The numbers on the branches are bootstrap values." figureDoi="10.3897/zookeys.1169.98681.figure2" httpUri="https://binary.pensoft.net/fig/873864" pageId="0" pageNumber="47">2</figureCitation>
|
||
). This species can be differentiated from all congeners within the
|
||
<taxonomicName id="3E89DBD77748008E54A5AE062E853007" class="Amphibia" kingdom="Animalia" lsidName="C. agamensis" order="Anura" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="agamensis">
|
||
<emphasis id="CF23493B2DE8B5E39E7725F065D3F4CC" italics="true" pageId="0" pageNumber="47">C. agamensis</emphasis>
|
||
</taxonomicName>
|
||
group by having the following combination of characters: (1) a medium-sized
|
||
<taxonomicName id="4B7E458CCB2DC9525EB25CEF7E3BDB58" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" lsidName="Cyrtodactylus" order="Squamata" pageId="0" pageNumber="47" rank="genus">
|
||
<emphasis id="00AFE997E9664841D7659424ED1D9311" italics="true" pageId="0" pageNumber="47">Cyrtodactylus</emphasis>
|
||
</taxonomicName>
|
||
, 54.1-77.7 mm in adult males, 65.1-76.7 mm in adult females; (2) 10-13 enlarged supralabial and 9-13 enlarged infralabial scales; (3) 3-5 internasal scales; (4) antebrachium tuberculated, brachium not tuberculated; (5) 32-41 paravertebral tubercles; (6) 16-19 longitudinal rows of dorsum tubercles; (7) 38-46 ventral scales; (8) 22-24 subdigital lamellae on the fourth toe; (9) 34-46 enlarged femoroprecloacal scales; (10) femoroprecloacal pores present in both sexes, 13-36 in adult males, 0-18 in adult females; (11) precloacal groove or depression present; (12) enlarged median subcaudals absent; (13) distinct ventrolateral folds; (14) subconical prominent tubercles on body that extend to the base of the tail; (15) 2-3 postcloacal tubercles; (16) two postocular stripes fused to a U-shaped mark on occiput; (17) postocular stripes extended beyond the arms; (18) 7-8 dark bands on trunk; (19) tail ringed by beige and black; and (20) labials yellow or dark with yellow/beige spots.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AE90AFA892E3E6E75D18E09286114863" pageId="0" pageNumber="47" type="description">
|
||
<paragraph id="A99DD21D1C855A19F76B5E1AF95C88CD" pageId="0" pageNumber="47">Description of holotype.</paragraph>
|
||
<paragraph id="E2FA53F6E23A8E4A78CD7910ACBB303D" pageId="0" pageNumber="47">Adult female with 65.1 mm SVL; head moderately in length (HL/SVL 0.29), wide (HW/HL 0.60), slightly flattened (HH/HL 0.34), distinct from neck, and triangular shape from dorsal view; lores concave; canthus rostralis rounded; snout elongated (SL/HL 0.40), rounded in rostral region, eye to snout distance larger than head depth; eyes large (OD/ HL 0.22), obtrusive and appeared beyond labials in dorsal view, eye diameter less than the eye to ear distance, pupil vertical; ear oppening small (EL/HL 0.03), elliptical, oriented obliquely leaning posteriorly; rostral large, subrectangular in shape, medial posterior edge interupted by an subhexagonal internasal scale that embedded within rostral, posteriorly bordered by three internasal scales, laterodorsally by nostril opening and lateroventrally by first supralabial; external nares directed lateroposteriorly, bordered anteriorly by rostral, posteriorly by two postnasals: one subcircular, one crescent shaped, dorsally by large supranasal, and ventrally by first supralabial; supranasal subrectangular, separated by one subhexagonal scale and a smaller subrectangular scale that piled up, the smaller one above the subhexagonal one, supranasal laterally bordered by nostril, right supranasal anteriorly bordered by five significantly smaller scales, left supranasal anteriorly bordered by four significantly smaller scales; one internasal scale, subhexgonal in shape, embedded within rostral, bordered by two supranasal scales, one smaller scale right posteriorly, and rostral anteriorly; 9/10 (right/left) supralabial scales to below center of the eye, 10/11 (right/left) to the posteriormost enlarged scale, 8 infralabial scales to below center of the eye, 10/11 (right/left) to the posteriormost enlarged scale; scales of frontonasal, prefrontal and lores small, juxtaposed, relatively raise; weak tubercles on the supraorbital region; prominent tubercles above the ear opening, larger than those on supraorbital, the tubercles gradually increased in size posteriorly.</paragraph>
|
||
<paragraph id="9BE1DD6E93D40027F3B3632ADC5C6F85" pageId="0" pageNumber="47">Body slender, relatively short (AX/SVL 0.47), with distinct ventrolateral fold; scales on dorsum small, homogenous, interspersed by rounded to trihedral tubercles; tubercles present from the occiput region to the base of the tail but no further than 1/3 of the tail, being more dense gradually posteriorly until the base of the tail; 37 paravertebral tubercles; 17 tubercles transversally in the middle of the trunk; imbricate and smooth ventral scales, scales in middle larger than those on lateral and dorsal, 42 scales across the center of the trunk from one ventrolateral fold to another; femoral scales enlarged, extending until 2/3 portion of femora, contiguous with enlarged precloacal scales, forming a reverse V-shaped mark in the middle of hindlimb; a greatly enlarged scale at the apex, larger than other precloacal and femoral scale, rather long than wide, each right and left bordered by three smaller scales; 46 continuous enlarged precloacal and femoral scales, all similar in size except the scale at apex; precloacal groove or depression absent.</paragraph>
|
||
<paragraph id="9B3293935FCC4DB9EDE503B4B4629F30" pageId="0" pageNumber="47">Limbs moderately slender; forelimbs relatively short (FL/SVL 0.15); scales on forelimbs dorsum larger than on body dorsum, rostrum or frontal, domed to subconical in shape, scales near the junction of limbs with trunk being smaller; round to subconical tubercles present on antebrachium, concentrated in the middle of antebrachium, similar in size with those on nape; brachium not tuberculated; fingers relatively long, well developed, no webbing; claw well developed, relatively short; hind limbs more robust than fore limbs, moderate in length (TBL/SVL 0.19); scales on dorsum domed to subconical, interspersed with trihedral tubercles; tubercles on dorsum approximately similar in size with those on posterior trunk; anterior ventral scales small, gradually increase in size posteriorly, the scales after enlarged femoral scales much smaller than those on anterior part; 22 subdigital lamellae on fourth finger; toes relatively long, claw well developed, relatively short.</paragraph>
|
||
<paragraph id="5B478A50982A4FA24212063A039B01A4" pageId="0" pageNumber="47">Tail 78.9 mm in length, longer than SVL (TL/SVL 1.21), 5.4 mm in width at base, cylindrical, decreasing in size posteriorly; scales on tail dorsum small, approximately similar in size with those on the distal of femoral; tail dorsum tuberculated at its base, no more than 1/3 of tail; postcloacal tubercles at each side, left tubercles slightly searated, right tubercles in contact one another; no enlarged median subcaudal scale; subcaudal scales larger than on tail dorsum.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EA2D04DCBE5BB4FFB47208F987CF090D" pageId="0" pageNumber="47" type="coloration in life">
|
||
<paragraph id="37C1C00F4BC6F188FB261538836F104D" pageId="0" pageNumber="47">Coloration in life.</paragraph>
|
||
<paragraph id="C95CA977C30DE6D1BB0DE6EEA6E0636C" pageId="0" pageNumber="47">Dorsal ground color of head, neck, trunk, limbs and tail beige to weak yellow; yellow mottling along the ventrolateral fold, on the lateral part of neck, on supra and infralabial scales; ventral of head, neck, trunk, limbs whitish to grey pale; palmar, metatarsal, fingers and toes darker; basal subcaudal ground color whitish to pale grey with dense yellow mottling; most of the original tail except basal subcaudal encircled by beige and black; labials yellow or dark with beige or yellow spots; weak black stripe between nostril and eye (sometimes absent); lateral stronger black stripe extends from posterior margin of the eye to approximately second black band on dorsum; wide U-shaped band around occiput connects with lateral stripe; the U-shaped band usually continuous (only one exception in UNP165 which is discontinuous in the middle); the Y-shaped pattern on occiput present in two individuals (UNP045, UNP193), the others lack Y-shaped pattern instead of irregular network or spots; bands on trunk dorsum sometimes clearly lying transversally and sometimes blotched; strong nape black spot only present in one individual (UNP045) where left portion fused with lateral stripe, fainted nape black spot presents in two individuals (UNP148, UNP165), the others lack nape spot; black lateral spots present positioned parallel with dorsum bands or between them; 5-6 irregular black bands on forelimbs and hindlimbs; beige to yellow spots on the base fingers (except finger 1); dorsum first finger lighter than other fingers; the clear beige and black bands on subcaudal appear after the one third portion from basal.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4EFF3527545032BC82350C71B9AFC948" pageId="0" pageNumber="47" type="comparisons">
|
||
<paragraph id="6235FB92A4DD767D192A9204DA5F7D72" pageId="0" pageNumber="47">Comparison.</paragraph>
|
||
<paragraph id="CB1B67FE19303AC39CC1C07C9DA6AE7C" pageId="0" pageNumber="47">
|
||
<taxonomicName id="5A4376518C8FA249CF559E1B20DA9672" authorityName="Nugraha & Ahda & Tjong & Kurniawan & Riyanto & Fauzi & Lin" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" lsidName="Cyrtodactylus gonjong" order="Squamata" pageId="0" pageNumber="47" rank="species" species="gonjong">
|
||
<emphasis id="043D0752561E4EEAB227715BD0442232" italics="true" pageId="0" pageNumber="47">Cyrtodactylus gonjong</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be differentiated from all other congeners in the
|
||
<taxonomicName id="E3F0C481459F768563E582EAB16F5ADB" class="Amphibia" kingdom="Animalia" lsidName="C. agamensis" order="Anura" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="agamensis">
|
||
<emphasis id="64022C5A815B0F0B0D95F87D81192424" italics="true" pageId="0" pageNumber="47">C. agamensis</emphasis>
|
||
</taxonomicName>
|
||
group based on a combination of morphological characters.
|
||
</paragraph>
|
||
<paragraph id="37BD0907E4464CA3AB957EE35218C52B" pageId="0" pageNumber="47">
|
||
<taxonomicName id="B5D751453F5B9AB6780974764631BC63" authorityName="Nugraha & Ahda & Tjong & Kurniawan & Riyanto & Fauzi & Lin" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" lsidName="Cyrtodactylus gonjong" order="Squamata" pageId="0" pageNumber="47" rank="species" species="gonjong">
|
||
<emphasis id="5A71A8BBBBD49A6204A2C6D2EB711837" italics="true" pageId="0" pageNumber="47">Cyrtodactylus gonjong</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differs from its most close relative,
|
||
<taxonomicName id="07A05F2F9D97566955920EB991D3372E" class="Amphibia" kingdom="Animalia" lsidName="C. semenanjungensis" order="Anura" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="semenanjungensis">
|
||
<emphasis id="AAF46A9A3AA499E278E5A0F4E438BA2F" italics="true" pageId="0" pageNumber="47">C. semenanjungensis</emphasis>
|
||
</taxonomicName>
|
||
from Peninsular Malaysia by having a larger body size, SVL maximum 77.7 in adult male and 76.7 in adult female (vs 62.1 mm in adult male and 69 mm in adult female); 8-10 supralabial scales to center of eye (vs 11-15); 16-19 DTR (vs 18-20); maximum PVT reachs 41 (vs 37); 38-46 ventral scales (vs 49-53); enlarged femoral scales present (vs absent); femoral and precloacal pores present in both sexes (vs absent in both sexes); 22-24 subdigital lamellae under fourth toe (vs 20-21); a black spot on the nape absent (vs present); posterior end of lateral stripe far beyond the arm insertion to the body, approximately reaching the second band on dorsum (vs between the arms insertion); posterior end of lateral stripe in contact with dorsum band that is separated (vs lateral stripe in contact with clear, not separated band or non-blotched band, creating a box encircled the nape spot); mostly lack nape spot, if present the spot is faint (vs strong black nape spot present).
|
||
</paragraph>
|
||
<paragraph id="399BA99BDD29657193F017854D11755A" pageId="0" pageNumber="47">
|
||
<taxonomicName id="8DCBD8671E8D7C6D4A19C6D1FE8F08B5" authorityName="Nugraha & Ahda & Tjong & Kurniawan & Riyanto & Fauzi & Lin" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" lsidName="Cyrtodactylus gonjong" order="Squamata" pageId="0" pageNumber="47" rank="species" species="gonjong">
|
||
<emphasis id="1D64FC8A26FBA314EAB3D035E60042B0" italics="true" pageId="0" pageNumber="47">Cyrtodactylus gonjong</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differs from
|
||
<taxonomicName id="F8FB35131E85089CAF58C790995D5379" class="Amphibia" kingdom="Animalia" lsidName="C. semicinctus" order="Anura" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="semicinctus">
|
||
<emphasis id="D433D43CB20A9A89E34830599DD42A47" italics="true" pageId="0" pageNumber="47">C. semicinctus</emphasis>
|
||
</taxonomicName>
|
||
by being smaller in the adult female (76.7 mm vs 89 mm) and larger in the adult male (77.7 mm vs 75 mm); having fewer DTR (16-19 vs 29-35); having more PVT (32-41 vs 24-27); maximum ventral scales 46 (vs 44); 0-36 precloacal and femoral pores (vs 36-38); 22-24 subdigital lamellae under the fourth toe (vs 19-22);10-13 enlarged supralabial scales (vs 8-11); 10-13 enlarged infralabial scales (vs 8-10); lateral stripe extending from behind the eye to behind arm present (vs absent); labials with beige or yellow spot (vs without spot); beige and black band encircled tail and subcaudal (ringed tail) (vs only dorsum tail - not ringed tail).
|
||
</paragraph>
|
||
<paragraph id="1528677C00F6E759F538DC8130B2113E" pageId="0" pageNumber="47">
|
||
<taxonomicName id="B144135C2C28A3A86568616746B21977" authorityName="Nugraha & Ahda & Tjong & Kurniawan & Riyanto & Fauzi & Lin" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" lsidName="Cyrtodactylus gonjong" order="Squamata" pageId="0" pageNumber="47" rank="species" species="gonjong">
|
||
<emphasis id="155B601D4FA868F0B2BFF8E2FCAE8CE8" italics="true" pageId="0" pageNumber="47">Cyrtodactylus gonjong</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differs from
|
||
<taxonomicName id="5E1F7FBC2FEB7372BCBE583522F3CF80" class="Amphibia" kingdom="Animalia" lsidName="C. psarops" order="Anura" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="psarops">
|
||
<emphasis id="B35B68728EBDDBDD8B22E04A8DD773FF" italics="true" pageId="0" pageNumber="47">C. psarops</emphasis>
|
||
</taxonomicName>
|
||
by being smaller in the adult female (76.7 mm vs 82 mm) and larger in the adult male (77.7 mm vs 74 mm); 16-19 dorsal tubercles transversally (vs 28-38); 32-41 longitudinal row tubercles on the middle of the body (vs 23-26); maximum number of ventral scales 46 (vs 49); 0-36 pores on precloacal and femoral region (vs 28-32); 2-3 postcloacal tubercles (vs usually single); 10-13 enlarged supralabials (vs 9-12); 10-13 enlarged infralabial scales (vs 8-11); brachium not tuberculated (vs tuberculated); postocular stripes, left and right, fused to form U-shaped mark on the occiput (vs usually does not fuse to form U-shaped mark); labials usually pale grey or yellow (vs charcoal or dark).
|
||
</paragraph>
|
||
<paragraph id="BBA1C38E35D5CB324F91FEE08D5A31EC" pageId="0" pageNumber="47">
|
||
<taxonomicName id="D8C9EC1F7E9147FA8DF7C35452D39257" authorityName="Nugraha & Ahda & Tjong & Kurniawan & Riyanto & Fauzi & Lin" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" lsidName="Cyrtodactylus gonjong" order="Squamata" pageId="0" pageNumber="47" rank="species" species="gonjong">
|
||
<emphasis id="FFAB67D0E7DBE4879A3EA7AA1BFE6677" italics="true" pageId="0" pageNumber="47">Cyrtodactylus gonjong</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differs from
|
||
<taxonomicName id="3CF2272DB82EDE19D4336CD564E5D477" class="Amphibia" kingdom="Animalia" lsidName="C. agamensis" order="Anura" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="agamensis">
|
||
<emphasis id="FA5D7F988DFA9B35318A22B7B9E0828E" italics="true" pageId="0" pageNumber="47">C. agamensis</emphasis>
|
||
</taxonomicName>
|
||
by being being smaller in the adult female (76.7 mm vs 86.8 mm) and larger in the adult male (77.7 mm vs 74.9 mm); 10-13 infralabial scales (vs 9-12); maximum number of dorsal tubercles transversally (21 vs 19); maximum number of paravertebral tubercles 41 (vs 37); 38-46 ventral scales (vs 50-67); 13-36 precloacal and femoral pores in males (vs 9-10), 0-18 in females (vs 0-7); 7-8 body black bands on trunk (vs 6-7); postocular stripe extends beyond the arms present (vs absent).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B01C3DB512C3BA1DC6C11655AF45B1C6" pageId="0" pageNumber="47" type="variation">
|
||
<paragraph id="E13B8ED8D5BC624BD6C657653E2CDE19" pageId="0" pageNumber="47">Variations.</paragraph>
|
||
<paragraph id="4A3FD3C18612D6A95D439C6A90D94200" pageId="0" pageNumber="47">
|
||
Dorsal ground color of head, neck, trunk, limbs and tail beige to weak yellow. Labials yellow (UNP148, UNP165, UNP167) or dark on the first three supralabials but pale grey for the rest (UNP193, UNP194, UNP199, UNP203) or dark on first eight supralabials (UNP196) (Fig.
|
||
<figureCitation id="DA358A643085C333C59280E760ACE8B0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Variation of labial color, stripe between eye and nostril and body bands in lateral view A UNP 167 B UNP 165 C UNP 196." figureDoi="10.3897/zookeys.1169.98681.figure5" httpUri="https://binary.pensoft.net/fig/873867" pageId="0" pageNumber="47">5</figureCitation>
|
||
). Stripe between eye and nostril absent (UNP045, UNP052, UNP060, UNP148, UNP165, UNP193), faint (UNP167, UNP194, UNP199, UNP203), or stronger (UNP196). Lateral stripe that extends beyond the arm could be slightly discontinuous right in the arm insertion (UNP167), while the others are continuous. The medium part of U-shaped mark is discontinuous in UNP165, while the others are continuous (Fig.
|
||
<figureCitation id="B42B656FDF6DE44AEF922A343DE235A9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. The representative paratypes of C. gonjong sp. nov." figureDoi="10.3897/zookeys.1169.98681.figure4" httpUri="https://binary.pensoft.net/fig/873866" pageId="0" pageNumber="47">4</figureCitation>
|
||
). The bands on the trunk continuous (UNP045, UNP193, UNP167, UNP194, UNP196), while others are irregular (UNP052, UNP199, UNP203) or blotched (UNP060, UNP148, UNP165).
|
||
</paragraph>
|
||
<caption id="A210B38AE450465D23F9D394F8A231CB" doi="10.3897/zookeys.1169.98681.figure5" httpUri="https://binary.pensoft.net/fig/873867" pageId="0" pageNumber="47" start="Figure 5" startId="F5">
|
||
<paragraph id="406A281AB9B938F10FDAA6D01669DE72" pageId="0" pageNumber="47">
|
||
<emphasis id="9A78B74B799A627FE94A8710A0062199" bold="true" pageId="0" pageNumber="47">Figure 5.</emphasis>
|
||
Variation of labial color, stripe between eye and nostril and body bands in lateral view
|
||
<emphasis id="E01F5DBEE12A544EC54CDE5F73D33203" bold="true" pageId="0" pageNumber="47">A</emphasis>
|
||
UNP167
|
||
<emphasis id="32FDDC684FD5D9FE566BCA30C2078572" bold="true" pageId="0" pageNumber="47">B</emphasis>
|
||
UNP165
|
||
<emphasis id="048107883EB4CE7774A865E52B75BA82" bold="true" pageId="0" pageNumber="47">C</emphasis>
|
||
UNP196.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="5DA8E76D35CBB89D41A835F167AE3602" pageId="0" pageNumber="47" type="distribution">
|
||
<paragraph id="C9D1D5585D6C885687F425E7C1FAC09A" pageId="0" pageNumber="47">Distribution, habitat and natural history.</paragraph>
|
||
<paragraph id="7159D4FEE77C2654119001DBF2B914BC" pageId="0" pageNumber="47">
|
||
All individuals were captured in Sumatra Barat Province, from the location between c. 83 meter above sea level (m asl) to c. 700 m asl. In Sungai Barameh, UNP052 was captured when it was sticking on a vertical cement wall approximately 70 cm above the ground; UNP053 was found on a vertical metal roofing sheet which was similar to UNP052 in height; UNP055, a juvenile, was found perching on a fern leave. In Malibo Anai, UNP045, UNP047, UNP048 were found in a similar habitat, sticking on a vertical stem tree approximately 1 m from the ground. In Lembah Anai Nature Reserve, UNP060-062 were found similar to those found in Malibo Anai. However, in Lembah Anai Nature Reserve, we observed an uncollected individual on the forest floor with leaf litter. In Sarasah Uwak Waterfall, we found UNP146 perching on a horizontal branch of a herb approximately 1 m above the ground. In village Sungai Sirah, UNP165 and UNP167 were found perching on a vertical stem of a tree approximately 1 m above the ground. In Langkuik Tamiang waterfall, UNP192 was found on a rock in the wall of a small rocky stream; UNP193 was found on herb branch; and UNP194 was found on a horizontal dead tree approximately 1.2 m above the ground. In village Koto Malintang, we found UNP199 and UNP203 perching on a herb branch approximately 1 m above the ground. There was one female carrying two eggs that can be seen inside the abdomen (UNP165; see Fig.
|
||
<figureCitation id="3B17B6E58837450FCE26320342D216E2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. The representative paratypes of C. gonjong sp. nov." figureDoi="10.3897/zookeys.1169.98681.figure4" httpUri="https://binary.pensoft.net/fig/873866" pageId="0" pageNumber="47">4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |