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<document id="3BB03D2A820C139B8D1CEA76E472DEB9" ID-CLB-Dataset="7214" ID-DOI="10.11646/zootaxa.4963.1.2" ID-GBIF-Dataset="78e78beb-b1ed-43a4-84c7-8180e61463c5" ID-ISSN="1175-5326" ID-Zenodo-Dep="4711249" ID-ZooBank="E93AB178-3419-498B-9563-A704ABAE5EB9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618560083953" checkinUser="plazi" docAuthor="Bálint, Zsolt, Boyer, Pierre, Cerdeña, José, Larico, Jackie Farfán, Brudecka, Jadwiga Lorenc-, Prieto, Carlos &amp; Pyrcz, Tomasz W." docDate="2021" docId="D5008F50FFE1B306FF76FC2C9379FE60" docLanguage="en" docName="zootaxa.4963.1.2.pdf" docOrigin="Zootaxa 4963 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Johnsonita carpia Balint, Boyer &amp; Pyrcz 2021, sp. n." docType="treatment" docVersion="7" lastPageNumber="21" masterDocId="2939F728FFE9B30CFFE1FFA69629FFCD" masterDocTitle="Contributions to the knowledge of Neotropical Lycaenidae: taxonomy of Johnsonita Salazar &amp; Constantino, 1995 with descriptions of seven new species (Theclinae Eumaeini)" masterLastPageNumber="57" masterPageNumber="11" pageNumber="19" updateTime="1698937134920" updateUser="ExternalLinkService">
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<mods:title id="1148CD4BD34CB8F46C0E2D5C71BCE464">Contributions to the knowledge of Neotropical Lycaenidae: taxonomy of Johnsonita Salazar &amp; Constantino, 1995 with descriptions of seven new species (Theclinae Eumaeini)</mods:title>
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<mods:namePart id="4755B1471910AC7B1A29F7C5CB70B1C9">Bálint, Zsolt</mods:namePart>
<mods:nameIdentifier id="316DAC624C76DD9235CA85202392E6B7" type="ORCID">0000-0001-8174-878X</mods:nameIdentifier>
<mods:affiliation id="63EB8694A9996861CE005B13E7955F95">Hungarian Natural History Museum, Department of Zoology, H- 1088 Budapest, Baross utca 13, Hungary. balint. zsolt @ nhmus. hu; https: // orcid. org / 0000 - 0001 - 8174 - 878 X</mods:affiliation>
<mods:nameIdentifier id="C2568765357E82A374FD481D73639D3F" type="email">balint.zsolt@nhmus.hu</mods:nameIdentifier>
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<mods:namePart id="DE7A6F0E39EF9EBB6E28137D0D847C05">Boyer, Pierre</mods:namePart>
<mods:affiliation id="95608B5298266FBBEEECAB8D5A64EB2E">Lotissement l’Horizon, 13610 Le Puy Sainte Réparade, France.</mods:affiliation>
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<mods:namePart id="C8EBEC8D604FD128F901C15CB6F43EEB">Cerdeña, José</mods:namePart>
<mods:nameIdentifier id="48478B0BC456069A499D59512BFBEAF4" type="ORCID">0000-0003-3930-6480</mods:nameIdentifier>
<mods:affiliation id="CC7799773DA32E35E819DA8146C228FC">Universidad Nacional de San Agustín de Arequipa, Museo de Historia Natural, Av. Alcides Carrión s / n, Arequipa, Peru. cerdenajoseal @ yahoo. es; https: // orcid. org / 0000 - 0003 - 3930 - 6480 jjackie 4 u @ gmail. com; https: // orcid. org / 0000 - 0001 - 5913 - 0756</mods:affiliation>
<mods:nameIdentifier id="3859F0C707B0644576E70076674908BE" type="email">cerdenajoseal@yahoo.es</mods:nameIdentifier>
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<mods:namePart id="709247A8874BCE0362089CB68F0C179C">Larico, Jackie Farfán</mods:namePart>
<mods:nameIdentifier id="0CFDE765F310A54872265D8F83FE41B5" type="ORCID">0000-0003-3930-6480</mods:nameIdentifier>
<mods:affiliation id="6765E157D512B2821533D0564D4BB69B">Universidad Nacional de San Agustín de Arequipa, Museo de Historia Natural, Av. Alcides Carrión s / n, Arequipa, Peru. cerdenajoseal @ yahoo. es; https: // orcid. org / 0000 - 0003 - 3930 - 6480 jjackie 4 u @ gmail. com; https: // orcid. org / 0000 - 0001 - 5913 - 0756</mods:affiliation>
<mods:nameIdentifier id="1321483A9924EEA2E0F9BB891717F1D4" type="email">cerdenajoseal@yahoo.es</mods:nameIdentifier>
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<mods:namePart id="3A1B9868847BFA43E77459159903ED24">Brudecka, Jadwiga Lorenc-</mods:namePart>
<mods:affiliation id="DB8FD23F9AEAD2D84589DEE3BF8CAB8A">Nature Education Centre, Jagiellonian University, Gronostajowa 5, 30 - 387 Kraków, Poland.</mods:affiliation>
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<mods:namePart id="C89079DEF3383040DC57F1ABEAF9F9A2">Prieto, Carlos</mods:namePart>
<mods:affiliation id="2D4AEC312269D48490687C53A128FD40">Corporación Universitaria Autónoma del Cauca, Popayán, Colombia.</mods:affiliation>
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<mods:namePart id="452F44B89F0C4569B092D23F68A273F6">Pyrcz, Tomasz W.</mods:namePart>
<mods:affiliation id="243090D9AD52FF16A12BAF5EF37C9EDA">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland.</mods:affiliation>
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<mods:title id="665848EA66CAFCA9328F6C18CA47CDAF">Zootaxa</mods:title>
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<mods:date id="C8B2A252CAB8170BAFFA95490933EECC">2021</mods:date>
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<mods:number id="8D510EB35DE534F6F055FC5BBE18A510">2021-04-16</mods:number>
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<treatment id="D5008F50FFE1B306FF76FC2C9379FE60" ID-DOI="http://doi.org/10.5281/zenodo.4696412" ID-GBIF-Taxon="181531948" ID-Zenodo-Dep="4696412" LSID="urn:lsid:plazi:treatment:D5008F50FFE1B306FF76FC2C9379FE60" httpUri="http://treatment.plazi.org/id/D5008F50FFE1B306FF76FC2C9379FE60" lastPageId="10" lastPageNumber="21" pageId="8" pageNumber="19">
<subSubSection id="15B36DCDFFE1B304FF76FC2C94F6FC68" box="[151,735,906,933]" pageId="8" pageNumber="19" type="nomenclature">
<paragraph id="5D163E46FFE1B304FF76FC2C94F6FC68" blockId="8.[151,735,906,969]" box="[151,735,906,933]" pageId="8" pageNumber="19">
<heading id="065E892AFFE1B304FF76FC2C94F6FC68" bold="true" box="[151,735,906,933]" fontSize="11" level="1" pageId="8" pageNumber="19" reason="1">
<emphasis id="6FDDE254FFE1B304FF76FC2C94F6FC68" bold="true" box="[151,735,906,933]" pageId="8" pageNumber="19">
<taxonomicName id="9AA945C5FFE1B304FF76FC2C94B9FC68" authority="Balint, Boyer &amp; Pyrcz" authorityName="Balint, Boyer &amp; Pyrcz" authorityYear="2021" box="[151,656,906,933]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="carpia" status="sp. nov.">
<emphasis id="6FDDE254FFE1B304FF76FC2C975AFC68" bold="true" box="[151,371,906,933]" italics="true" pageId="8" pageNumber="19">Johnsonita carpia</emphasis>
Bálint, Boyer &amp; Pyrcz
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,
<taxonomicNameLabel id="74EE5F2FFFE1B304FD7DFC2D94F6FC68" box="[668,735,907,933]" pageId="8" pageNumber="19" rank="species">sp. n.</taxonomicNameLabel>
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<paragraph id="5D163E46FFE1B304FF76FC099439FC04" blockId="8.[151,735,906,969]" box="[151,528,943,969]" pageId="8" pageNumber="19">
(
<figureCitation id="C59222C3FFE1B304FF7EFC0996C1FC04" box="[159,232,943,969]" captionStart="FIGURE 4" captionStartId="9.[151,250,1929,1953]" captionTargetBox="[210,1382,280,1889]" captionTargetId="figure-58@9.[177,1411,273,1906]" captionTargetPageId="9" captionText="FIGURE 4. Johnsonita carpia Bálint, Boyer &amp; Pyrcz, sp. n. imagines (left column: recto, right column: verso). A–B: holotype (Ecuador: Pichincha), C–D: male specimen (Ecuador: Napo), E–F: paratype female (Ecuador: Carchi) (CEP–MZUJ specimens) (scale bar: 10 mm)." figureDoi="http://doi.org/10.5281/zenodo.4711257" httpUri="https://zenodo.org/record/4711257/files/figure.png" pageId="8" pageNumber="19">Figs 4</figureCitation>
,
<figureCitation id="C59222C3FFE1B304FF15FC09976EFC04" box="[244,327,943,969]" captionStart="FIGURE 17" captionStartId="27.[151,250,1943,1967]" captionTargetBox="[272,1345,182,1893]" captionTargetId="figure-18@27.[175,1413,181,1920]" captionTargetPageId="27" captionText="FIGURE 17. Johnsonita genitalia armatures, entire male organs in lateral (left column) and ventral (right column) views.A–B: J. auda, Ecuador, Morona-Santiago, 9 de Octubre-Macas; C–D: J. carpia, Ecuador, Napo, Res. Yanayacu; E–F: J. catadupa, Ecuador, Loja, Yangana; G–H: J. chlamydem, Peru, Pasco, Oxapampa, La Antena (CEP–MZUJ specimens) (scale as indicated)." figureDoi="http://doi.org/10.5281/zenodo.4711291" httpUri="https://zenodo.org/record/4711291/files/figure.png" pageId="8" pageNumber="19">17C–D</figureCitation>
,
<figureCitation id="C59222C3FFE1B304FEB2FC099788FC04" box="[339,417,943,969]" captionStart="FIGURE 20" captionStartId="30.[151,250,1909,1933]" captionTargetBox="[183,1405,181,1885]" captionTargetId="figure-23@30.[183,1405,181,1885]" captionTargetPageId="30" captionText="FIGURE 20. Entire Johnsonita female organs in lateral (left column), ductus in lateral (central column) and ventral (right column) views. A–C: J. auda, Venezuela, Edo. Barinas, Minas de San Isidro; D–F: J. carpia, Ecuador, Prov. Carchi, Res. Forest Golondrinas; G–I: J. catadupa, Peru, Dept. Amazonas, Molinopampa, Tingo Ocol (CEP–MZUJ specimens) (scale as indicated)." figureDoi="http://doi.org/10.5281/zenodo.4711297" httpUri="https://zenodo.org/record/4711297/files/figure.png" pageId="8" pageNumber="19">20D–F</figureCitation>
,
<figureCitation id="C59222C3FFE1B304FE4DFC0997F6FC04" box="[428,479,943,969]" captionStart="FIGURE 23" captionStartId="33.[151,250,1570,1594]" captionTargetBox="[390,1203,198,1539]" captionTargetId="figure-18@33.[378,1210,181,1545]" captionTargetPageId="33" captionText="FIGURE 23. Johnsonita mail trait ventral forewing subcubital scent patch indicated by arrow. A: J. carpia, B: J. catadupa, C: J. chlamydem, D: J. ianusca, E: J. johnsoni, F: J. pardoa, G: J. turquisca, H: J. zubkova." figureDoi="http://doi.org/10.5281/zenodo.4711305" httpUri="https://zenodo.org/record/4711305/files/figure.png" pageId="8" pageNumber="19">23A</figureCitation>
,
<figureCitation id="C59222C3FFE1B304FE0BFC09942EFC04" box="[490,519,943,969]" captionStart="FIGURE 30" captionStartId="39.[151,250,1742,1766]" captionTargetBox="[251,1396,204,1718]" captionTargetId="figure-18@39.[192,1396,181,1718]" captionTargetPageId="39" captionText="FIGURE 30. Johnsonita distributions based on museum material examinded. J. assula = red circle, J. carpia = yellow trinagle, J. catadupa = blue diamond, J. chalamydem = brown triangle, J. ianusca = green square, J. oxalida = purple diamond." figureDoi="http://doi.org/10.5281/zenodo.4711325" httpUri="https://zenodo.org/record/4711325/files/figure.png" pageId="8" pageNumber="19">30</figureCitation>
)
</paragraph>
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<emphasis id="6FDDE254FFE1B304FF76FC519710FBDC" box="[151,313,1015,1041]" italics="true" pageId="8" pageNumber="19">Type material.</emphasis>
<materialsCitation id="EDC1341BFFE1B304FEDEFC519444FBB3" ID-GBIF-Occurrence="3083794328" collectingDate="1996-03-15" collectingDateMax="1998-03-15" collectingDateMin="1996-03-15" collectionCode="CB, CEP, MZUJ" collectorName="P. Boyer &amp; J. Lorenc" country="Ecuador" elevation="2100" location="Nono - Nanegalito" municipality="Prov." pageId="8" pageNumber="19" specimenCount="2" stateProvince="Pichincha" typeStatus="holotype">
<collectionCode id="3BB8A683FFE1B304FEDEFC51974DFBDC" box="[319,356,1015,1041]" country="Australia" name="The CB Rhizobium Collection" pageId="8" pageNumber="19">CB</collectionCode>
(
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–
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)
<typeStatus id="821280E4FFE1B304FDEEFC519458FBDC" box="[527,625,1015,1041]" pageId="8" pageNumber="19" type="holotype">holotype</typeStatus>
male in moderate condition (broken hindwings’ tornal parts), labelled as “Prov. Pichincha, Nono – Nanegalito, km15,
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</quantity>
,
<date id="29171886FFE1B304FD18FBBC9550FBF8" box="[761,889,1050,1077]" pageId="8" pageNumber="19" value="1998-03-15">
<collectingDate id="3953E16EFFE1B304FD18FBBC9550FBF8" box="[761,889,1050,1077]" pageId="8" pageNumber="19" value="1998-03-15">15.III.1998</collectingDate>
</date>
,
<collectorName id="F05C5B90FFE1B304FC67FBBD95C1FBF8" box="[902,1000,1051,1077]" pageId="8" pageNumber="19">P. Boyer</collectorName>
” (white label, printed) (dissected: gen. prep.6639/
<collectorName id="F05C5B90FFE1B304FEF2FB99975FFB97" box="[275,374,1087,1114]" pageId="8" pageNumber="19">J.Lorenc</collectorName>
).
<typeStatus id="821280E4FFE1B304FE6BFB9997DEFB94" box="[394,503,1087,1113]" pageId="8" pageNumber="19" type="paratype">Paratypes</typeStatus>
(n=6; two dissections), all from
<collectingCountry id="25BE7ED6FFE1B304FC8CFB9995D1FB94" box="[877,1016,1087,1113]" name="Ecuador" pageId="8" pageNumber="19">ECUADOR</collectingCountry>
.
<collectingMunicipality id="BD72A43CFFE1B304FBE5FB999216FB94" box="[1028,1087,1087,1113]" pageId="8" pageNumber="19">Prov.</collectingMunicipality>
<collectingRegion id="9F6DF0A4FFE1B304FBA6FB999291FB94" box="[1095,1208,1087,1113]" country="Ecuador" name="Pichincha" pageId="8" pageNumber="19">Pichincha</collectingRegion>
,
<location id="5876689DFFE1B304FB25FB9993B5FB94" LSID="urn:lsid:plazi:treatment:D5008F50FFE1B306FF76FC2C9379FE60:5876689DFFE1B304FB25FB9993B5FB94" box="[1220,1436,1087,1113]" country="Ecuador" municipality="Prov." name="Nono - Nanegalito" pageId="8" pageNumber="19" stateProvince="Pichincha">Nono – Nanegalito</location>
km 15,
<quantity id="9A5193A3FFE1B304FF0CFBC5976DFBB0" box="[237,324,1123,1149]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="19" unit="m" value="2100.0">
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,
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<collectingDate id="3953E16EFFE1B304FEB0FBC497E6FBB0" box="[337,463,1122,1149]" pageId="8" pageNumber="19" value="1996-03-15">15.III.1996</collectingDate>
</date>
(
<collectionCode id="3BB8A683FFE1B304FE3EFBC59420FBB0" box="[479,521,1123,1149]" country="Australia" name="The CB Rhizobium Collection" pageId="8" pageNumber="19">CB</collectionCode>
: female)
</materialsCitation>
;
<materialsCitation id="EDC1341BFFE1B304FD98FBC59218FBB0" ID-GBIF-Occurrence="3083794346" box="[633,1073,1123,1149]" collectorName="Prov." country="Ecuador" location="Nanegalito" municipality="Nono" pageId="8" pageNumber="19" specimenCount="1" stateProvince="Pichincha" typeStatus="holotype">
<collectorName id="F05C5B90FFE1B304FD98FBC5949DFBB0" box="[633,692,1123,1149]" pageId="8" pageNumber="19">Prov.</collectorName>
<collectingRegion id="9F6DF0A4FFE1B304FD5BFBC59502FBB0" box="[698,811,1123,1149]" country="Ecuador" name="Pichincha" pageId="8" pageNumber="19">Pichincha</collectingRegion>
,
<collectingMunicipality id="BD72A43CFFE1B304FCD6FBC5955CFBB0" box="[823,885,1123,1149]" pageId="8" pageNumber="19">Nono</collectingMunicipality>
vers
<location id="5876689DFFE1B304FC52FBC59218FBB0" LSID="urn:lsid:plazi:treatment:D5008F50FFE1B306FF76FC2C9379FE60:5876689DFFE1B304FC52FBC59218FBB0" box="[947,1073,1123,1149]" country="Ecuador" municipality="Nono" name="Nanegalito" pageId="8" pageNumber="19" stateProvince="Pichincha">Nanegalito</location>
</materialsCitation>
,
<date id="29171886FFE1B304FBDDFBC592A6FBB0" box="[1084,1167,1123,1149]" pageId="8" pageNumber="19" value="1999-05">V.1999</date>
,
<materialsCitation id="EDC1341BFFE1B304FB7BFBC59456FB6C" ID-GBIF-Occurrence="3083794325" collectionCode="CB" collectorName="P. Boyer" country="Ecuador" elevation="1750" location="Maldonado" municipality="Prov." pageId="8" pageNumber="19" specimenCount="1" stateProvince="Carchi" typeStatus="holotype">
<collectorName id="F05C5B90FFE1B304FB7BFBC592D0FBB0" box="[1178,1273,1123,1149]" pageId="8" pageNumber="19">P. Boyer</collectorName>
(
<collectionCode id="3BB8A683FFE1B304FAEBFBC5931DFBB0" box="[1290,1332,1123,1149]" country="Australia" name="The CB Rhizobium Collection" pageId="8" pageNumber="19">CB</collectionCode>
: female);
<collectingMunicipality id="BD72A43CFFE1B304FF76FB2196FBFB6C" box="[151,210,1159,1185]" pageId="8" pageNumber="19">Prov.</collectingMunicipality>
<collectingRegion id="9F6DF0A4FFE1B304FF38FB21970EFB6C" box="[217,295,1159,1185]" country="Ecuador" name="Carchi" pageId="8" pageNumber="19">Carchi</collectingRegion>
,
<location id="5876689DFFE1B304FED3FB21979DFB6C" LSID="urn:lsid:plazi:treatment:D5008F50FFE1B306FF76FC2C9379FE60:5876689DFFE1B304FED3FB21979DFB6C" box="[306,436,1159,1185]" country="Ecuador" municipality="Prov." name="Maldonado" pageId="8" pageNumber="19" stateProvince="Carchi">Maldonado</location>
,
<quantity id="9A5193A3FFE1B304FE21FB219474FB6C" box="[448,605,1159,1185]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="8" pageNumber="19" unit="m" value="1750.0" valueMax="1900.0" valueMin="1600.0">
<elevation id="D684D975FFE1B304FE21FB219474FB6C" box="[448,605,1159,1185]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="8" pageNumber="19" unit="m" value="1750.0" valueMax="1900.0" valueMin="1600.0">1600–1900 m</elevation>
</quantity>
,
<date id="29171886FFE1B304FD88FB219456FB6C" box="[617,639,1159,1185]" pageId="8" pageNumber="19" value="1999-05-08">8.</date>
</materialsCitation>
V. 1999 (CB: female);
<materialsCitation id="EDC1341BFFE1B304FC66FB219681FB08" ID-GBIF-Occurrence="3083794311" collectorName="Prov." country="Ecuador" elevation="2500" location="Res. Forest" pageId="8" pageNumber="19" specimenCount="1" stateProvince="Carchi" typeStatus="holotype">
<collectorName id="F05C5B90FFE1B304FC66FB2195E8FB6C" box="[903,961,1159,1185]" pageId="8" pageNumber="19">Prov.</collectorName>
<collectingRegion id="9F6DF0A4FFE1B304FC29FB21923FFB6C" box="[968,1046,1159,1185]" country="Ecuador" name="Carchi" pageId="8" pageNumber="19">Carchi</collectingRegion>
,
<location id="5876689DFFE1B304FBC0FB219289FB6C" LSID="urn:lsid:plazi:treatment:D5008F50FFE1B306FF76FC2C9379FE60:5876689DFFE1B304FBC0FB219289FB6C" box="[1057,1184,1159,1185]" country="Ecuador" name="Res. Forest" pageId="8" pageNumber="19" stateProvince="Carchi">Res. Forest</location>
Golondrinas,
<quantity id="9A5193A3FFE1B304FAA1FB2093BEFB6C" box="[1344,1431,1158,1185]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="19" unit="m" value="2500.0">
<elevation id="D684D975FFE1B304FAA1FB2093BEFB6C" box="[1344,1431,1158,1185]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="19" unit="m" value="2500.0">2500 m</elevation>
</quantity>
,
<date id="29171886FFE1B304FF76FB0D9681FB08" box="[151,168,1195,1221]" pageId="8" pageNumber="19" value="1999-06-03">3.</date>
</materialsCitation>
VI.1999, J. Wojtusiak &amp;
<materialsCitation id="EDC1341BFFE1B304FE29FB0D942BFB24" ID-GBIF-Occurrence="3083794364" collectionCode="T, CEP, MZUJ" collectorName="Pyrcz &amp; J. Lorenc" country="Ecuador" elevation="2500" location="Res. Forest" municipality="Prov." pageId="8" pageNumber="19" specimenCount="1" stateProvince="Carchi" typeStatus="holotype">
<collectionCode id="3BB8A683FFE1B304FE29FB0D97F3FB08" box="[456,474,1195,1221]" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="19">T</collectionCode>
.
<collectorName id="F05C5B90FFE1B304FE07FB0D940FFB08" box="[486,550,1195,1221]" pageId="8" pageNumber="19">Pyrcz</collectorName>
(
<collectionCode id="3BB8A683FFE1B304FDD7FB0D9442FB08" box="[566,619,1195,1221]" pageId="8" pageNumber="19">CEP</collectionCode>
–
<collectionCode id="3BB8A683FFE1B304FD9BFB0D94EEFB08" box="[634,711,1195,1221]" pageId="8" pageNumber="19">MZUJ</collectionCode>
: female; dissected: gen. prep.6396/
<collectorName id="F05C5B90FFE1B304FBB9FB0D9292FB0B" box="[1112,1211,1195,1222]" pageId="8" pageNumber="19">J.Lorenc</collectorName>
);
<collectingMunicipality id="BD72A43CFFE1B304FB2EFB0D9323FB08" box="[1231,1290,1195,1221]" pageId="8" pageNumber="19">Prov.</collectingMunicipality>
<collectingRegion id="9F6DF0A4FFE1B304FAF0FB0D9376FB08" box="[1297,1375,1195,1221]" country="Ecuador" name="Carchi" pageId="8" pageNumber="19">Carchi</collectingRegion>
,
<location id="5876689DFFE1B304FA8AFB0D96F4FB24" LSID="urn:lsid:plazi:treatment:D5008F50FFE1B306FF76FC2C9379FE60:5876689DFFE1B304FA8AFB0D96F4FB24" country="Ecuador" municipality="Prov." name="Res. Forest" pageId="8" pageNumber="19" stateProvince="Carchi">Res. Forest</location>
Golondrinas,
<quantity id="9A5193A3FFE1B304FE9CFB6897FDFB24" box="[381,468,1230,1257]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="19" unit="m" value="2500.0">
<elevation id="D684D975FFE1B304FE9CFB6897FDFB24" box="[381,468,1230,1257]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="19" unit="m" value="2500.0">2500 m</elevation>
</quantity>
,
<date id="29171886FFE1B304FE01FB69942BFB24" box="[480,514,1231,1257]" pageId="8" pageNumber="19" value="1999-06-22">22.</date>
</materialsCitation>
VI.1999, J. Wojtusiak &amp;
<materialsCitation id="EDC1341BFFE1B304FCFBFB69945AFAC0" ID-GBIF-Occurrence="3083794308" collectingDate="2002-02" collectionCode="T, CEP, MZUJ" collectorName="Pyrcz &amp; J. Lorenc" country="Ecuador" elevation="2800" location="Tandapi" municipality="Prov." pageId="8" pageNumber="19" specimenCount="1" stateProvince="Pichincha" typeStatus="holotype">
<collectionCode id="3BB8A683FFE1B304FCFBFB699505FB24" box="[794,812,1231,1257]" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="19">T</collectionCode>
.
<collectorName id="F05C5B90FFE1B304FCD6FB69955EFB24" box="[823,887,1231,1257]" pageId="8" pageNumber="19">Pyrcz</collectorName>
(
<collectionCode id="3BB8A683FFE1B304FC64FB699593FB24" box="[901,954,1231,1257]" pageId="8" pageNumber="19">CEP</collectionCode>
–
<collectionCode id="3BB8A683FFE1B304FC2BFB69923EFB24" box="[970,1047,1231,1257]" pageId="8" pageNumber="19">MZUJ</collectionCode>
: female, gen. prep. 379/
<collectorName id="F05C5B90FFE1B304FAC7FB6993B9FB24" box="[1318,1424,1231,1257]" pageId="8" pageNumber="19">J. Lorenc</collectorName>
);
<collectingMunicipality id="BD72A43CFFE1B304FF76FB5596FBFAC0" box="[151,210,1267,1293]" pageId="8" pageNumber="19">Prov.</collectingMunicipality>
<collectingRegion id="9F6DF0A4FFE1B304FF39FB559760FAC0" box="[216,329,1267,1293]" country="Ecuador" name="Pichincha" pageId="8" pageNumber="19">Pichincha</collectingRegion>
,
<location id="5876689DFFE1B304FEB5FB559798FAC0" LSID="urn:lsid:plazi:treatment:D5008F50FFE1B306FF76FC2C9379FE60:5876689DFFE1B304FEB5FB559798FAC0" box="[340,433,1267,1293]" country="Ecuador" municipality="Prov." name="Tandapi" pageId="8" pageNumber="19" stateProvince="Pichincha">Tandapi</location>
,
<quantity id="9A5193A3FFE1B304FE5DFB55943AFAC0" box="[444,531,1267,1293]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="19" unit="m" value="2800.0">
<elevation id="D684D975FFE1B304FE5DFB55943AFAC0" box="[444,531,1267,1293]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="19" unit="m" value="2800.0">2800 m</elevation>
</quantity>
,
<date id="29171886FFE1B304FDFEFB55945AFAC0" box="[543,627,1267,1293]" pageId="8" pageNumber="19" value="2002-02">
<collectingDate id="3953E16EFFE1B304FDFEFB55945AFAC0" box="[543,627,1267,1293]" pageId="8" pageNumber="19" value="2002-02">II.2002</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="EDC1341BFFE1B304FD9FFB5595DAFAC3" ID-GBIF-Occurrence="3083794333" box="[638,1011,1267,1294]" collectionCode="CEP, MZUJ" collectorName="Wojtusiak" country="Ecuador" pageId="8" pageNumber="19" specimenCount="1" stateProvince="Pichincha" typeStatus="holotype">
J.
<collectorName id="F05C5B90FFE1B304FD76FB559521FAC0" box="[663,776,1267,1293]" pageId="8" pageNumber="19">Wojtusiak</collectorName>
(
<collectionCode id="3BB8A683FFE1B304FCF7FB559562FAC0" box="[790,843,1267,1293]" pageId="8" pageNumber="19">CEP</collectionCode>
–
<collectionCode id="3BB8A683FFE1B304FCBBFB55958EFAC0" box="[858,935,1267,1293]" pageId="8" pageNumber="19">MZUJ</collectionCode>
: male)
</materialsCitation>
.
</paragraph>
<paragraph id="5D163E46FFE1B304FF26FAB195F1FAB4" blockId="8.[151,1437,1015,2013]" pageId="8" pageNumber="19">
<emphasis id="6FDDE254FFE1B304FF26FAB1978AFAFC" box="[199,419,1303,1329]" italics="true" pageId="8" pageNumber="19">Additional material</emphasis>
(n= 2, two dissections),
<materialsCitation id="EDC1341BFFE1B304FD55FAB195FDFAB4" ID-GBIF-Occurrence="3083794321" collectingDate="1998-10-08" collectingDateMax="2011-11-03" collectingDateMin="1998-10-08" collectionCode="NHMUK, T, CEP, MZUJ" collectorName=" J. Wojtusiak &amp; T. Pyrcz " country="Ecuador" elevation="1600" location="Rio Sucio" pageId="8" pageNumber="19" specimenCount="1" stateProvince="Napo">
both from
<collectingCountry id="25BE7ED6FFE1B304FCC8FAB1959DFAFC" box="[809,948,1303,1329]" name="Ecuador" pageId="8" pageNumber="19">ECUADOR</collectingCountry>
. Prov.
<collectingRegion id="9F6DF0A4FFE1B304FC1EFAB19217FAFC" box="[1023,1086,1303,1329]" country="Ecuador" name="Napo" pageId="8" pageNumber="19">Napo</collectingRegion>
,
<location id="5876689DFFE1B304FBA8FAB19292FAFC" LSID="urn:lsid:plazi:treatment:D5008F50FFE1B306FF76FC2C9379FE60:5876689DFFE1B304FBA8FAB19292FAFC" box="[1097,1211,1303,1329]" country="Ecuador" name="Rio Sucio" pageId="8" pageNumber="19" stateProvince="Napo">Río Sucio</location>
,
<quantity id="9A5193A3FFE1B304FB24FAB19332FAFC" box="[1221,1307,1303,1329]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="19" unit="m" value="1600.0">
<elevation id="D684D975FFE1B304FB24FAB19332FAFC" box="[1221,1307,1303,1329]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="19" unit="m" value="1600.0">1600 m</elevation>
</quantity>
,
<date id="29171886FFE1B304FAC6FAB193B1FAFC" box="[1319,1432,1303,1329]" pageId="8" pageNumber="19" value="1998-10-08">
<collectingDate id="3953E16EFFE1B304FAC6FAB193B1FAFC" box="[1319,1432,1303,1329]" pageId="8" pageNumber="19" value="1998-10-08">8.X. 1998</collectingDate>
</date>
,
<collectorName id="F05C5B90FFE1B304FF76FA9D971FFA98" box="[151,310,1339,1365]" pageId="8" pageNumber="19">C.P.J. Samson</collectorName>
(
<collectionCode id="3BB8A683FFE1B304FEA5FA9D979DFA98" box="[324,436,1339,1365]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="19" type="Museum">NHMUK</collectionCode>
: male, dissected, vial no. 5818); Prov.
<collectingRegion id="9F6DF0A4FFE1B304FC82FA9D958BFA98" box="[867,930,1339,1365]" country="Ecuador" name="Napo" pageId="8" pageNumber="19">Napo</collectingRegion>
, Res. Yanayacu,
<quantity id="9A5193A3FFE1B304FB81FA9D929EFA98" box="[1120,1207,1339,1365]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="19" unit="m" value="2000.0">
<elevation id="D684D975FFE1B304FB81FA9D929EFA98" box="[1120,1207,1339,1365]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="19" unit="m" value="2000.0">2000 m</elevation>
</quantity>
,
<date id="29171886FFE1B304FB23FA9C9302FA98" box="[1218,1323,1338,1365]" pageId="8" pageNumber="19" value="2005-02-06">
<collectingDate id="3953E16EFFE1B304FB23FA9C9302FA98" box="[1218,1323,1338,1365]" pageId="8" pageNumber="19" value="2005-02-06">6.II.2005</collectingDate>
</date>
,
<collectorName id="F05C5B90FFE1B304FAD6FA9D96EDFAB4" pageId="8" pageNumber="19">J. Wojtusiak</collectorName>
&amp;
<collectionCode id="3BB8A683FFE1B304FF09FAF996D3FAB4" box="[232,250,1375,1401]" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="19">T</collectionCode>
.
<collectorName id="F05C5B90FFE1B304FEE4FAF9976CFAB4" box="[261,325,1375,1401]" pageId="8" pageNumber="19">Pyrcz</collectorName>
(
<collectionCode id="3BB8A683FFE1B304FEB2FAF997A1FAB4" box="[339,392,1375,1401]" pageId="8" pageNumber="19">CEP</collectionCode>
–
<collectionCode id="3BB8A683FFE1B304FE76FAF997CDFAB4" box="[407,484,1375,1401]" pageId="8" pageNumber="19">MZUJ</collectionCode>
: male; gen. prep. 02/
<date id="29171886FFE1B304FD2CFAF99560FAB4" box="[717,841,1375,1401]" pageId="8" pageNumber="19" value="2011-11-03">
<collectingDate id="3953E16EFFE1B304FD2CFAF99560FAB4" box="[717,841,1375,1401]" pageId="8" pageNumber="19" value="2011-11-03">11.03.2011</collectingDate>
</date>
/
<collectorName id="F05C5B90FFE1B304FCBEFAF995E5FAB4" box="[863,972,1375,1401]" pageId="8" pageNumber="19">J. Lorenc</collectorName>
)
</materialsCitation>
.
</paragraph>
<paragraph id="5D163E46FFE1B304FF26FA2594D5F954" blockId="8.[151,1437,1015,2013]" pageId="8" pageNumber="19">
<emphasis id="6FDDE254FFE1B304FF26FA25977AFA51" box="[199,339,1411,1436]" italics="true" pageId="8" pageNumber="19">Description.</emphasis>
Wings. FW costa length:
<quantity id="9A5193A3FFE1B304FD69FA2494D0FA50" box="[648,761,1410,1437]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="8" pageNumber="19" unit="mm" value="16.5" valueMax="18.0" valueMin="15.0">15–18mm</quantity>
(CEP–MZUJ
<typeStatus id="821280E4FFE1B304FC42FA2595CAFA50" box="[931,995,1411,1437]" pageId="8" pageNumber="19">types</typeStatus>
); FW apex rounded, outer margin not straight; HW vein Cubitus 1 with slightly extended terminus; dorsal wing surface colouration dark violet in both sexes; dorsal FW surface with black border inner margin reaching the middle of vein M3; dorsal HW discal area structurally coloured; ventral wing surface medial pattern composed of light ruptive dashes and inner margin medial area with brown scaling. Genitalia. Male valva with pointed costal margin flip and pointed valval terminus slightly curved (somewhat longer than half valval length). Female ductus with slightly divergent edges and small lower plate, upper plate asymmetric and larger with sclerotized frame with spines or bristles. Androconia present on FW along vein Cubital 2 as wide streak in the median area.
</paragraph>
<paragraph id="5D163E46FFE1B304FF26F9059346F858" blockId="8.[151,1437,1015,2013]" pageId="8" pageNumber="19">
<emphasis id="6FDDE254FFE1B304FF26F9059769F971" box="[199,320,1699,1724]" italics="true" pageId="8" pageNumber="19">Diagnosis.</emphasis>
This is a relatively large species; especially the females. It is similar to
<taxonomicName id="9AA945C5FFE1B304FB91F90592EDF970" authorityName="d'Abrera &amp; Balint" authorityYear="2001" baseAuthorityName="Hewitson" baseAuthorityYear="1867" box="[1136,1220,1699,1725]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="auda">
<emphasis id="6FDDE254FFE1B304FB91F90592EDF970" box="[1136,1220,1699,1725]" italics="true" pageId="8" pageNumber="19">J. auda</emphasis>
</taxonomicName>
and they occur together in
<collectingCountry id="25BE7ED6FFE1B304FEE2F961974BF92C" box="[259,354,1735,1761]" name="Ecuador" pageId="8" pageNumber="19">Ecuador</collectingCountry>
, although they are not syntopic. The two taxa can be identified in the field on the basis of their size (
<taxonomicName id="9AA945C5FFE1B304FF32F94D970CF8C8" authorityName="d'Abrera &amp; Balint" authorityYear="2001" baseAuthorityName="Hewitson" baseAuthorityYear="1867" box="[211,293,1771,1797]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="auda">
<emphasis id="6FDDE254FFE1B304FF32F94D970CF8C8" box="[211,293,1771,1797]" italics="true" pageId="8" pageNumber="19">J. auda</emphasis>
</taxonomicName>
is somewhat smaller), hindwing shape (
<taxonomicName id="9AA945C5FFE1B304FD0EF94D957BF8C9" authorityName="Balint, Boyer &amp; Pyrcz" authorityYear="2021" box="[751,850,1771,1796]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="carpia">
<emphasis id="6FDDE254FFE1B304FD0EF94D957BF8C9" box="[751,850,1771,1796]" italics="true" pageId="8" pageNumber="19">J. carpia</emphasis>
</taxonomicName>
with extended vein Cubitus 1 terminus, what absent in
<taxonomicName id="9AA945C5FFE1B304FF55F8A9972CF8E4" authorityName="d'Abrera &amp; Balint" authorityYear="2001" baseAuthorityName="Hewitson" baseAuthorityYear="1867" box="[180,261,1807,1833]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="auda">
<emphasis id="6FDDE254FFE1B304FF55F8A9972CF8E4" box="[180,261,1807,1833]" italics="true" pageId="8" pageNumber="19">J. auda</emphasis>
</taxonomicName>
) and their dorsal wing suface colouration (
<taxonomicName id="9AA945C5FFE1B304FD04F8A9951EF8E4" authorityName="d'Abrera &amp; Balint" authorityYear="2001" baseAuthorityName="Hewitson" baseAuthorityYear="1867" box="[741,823,1807,1833]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="auda">
<emphasis id="6FDDE254FFE1B304FD04F8A9951EF8E4" box="[741,823,1807,1833]" italics="true" pageId="8" pageNumber="19">J. auda</emphasis>
</taxonomicName>
is lighter). In identifying museum specimens or digital images the following traits provide clue for a positive identification: the colouration of the inner edge of the HW (
<taxonomicName id="9AA945C5FFE1B304FA68F89596E6F8BC" authorityName="d'Abrera &amp; Balint" authorityYear="2001" baseAuthorityName="Hewitson" baseAuthorityYear="1867" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="auda">
<emphasis id="6FDDE254FFE1B304FA68F89596E6F8BC" italics="true" pageId="8" pageNumber="19">J. auda</emphasis>
</taxonomicName>
is orange coloured), male androconia (no androconia in
<taxonomicName id="9AA945C5FFE1B304FCB3F8F1958CF8BC" authorityName="d'Abrera &amp; Balint" authorityYear="2001" baseAuthorityName="Hewitson" baseAuthorityYear="1867" box="[850,933,1879,1905]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="auda">
<emphasis id="6FDDE254FFE1B304FCB3F8F1958CF8BC" box="[850,933,1879,1905]" italics="true" pageId="8" pageNumber="19">J. auda</emphasis>
</taxonomicName>
) and male genitalia (with extremely pointed and straight valval terminus, curved in
<taxonomicName id="9AA945C5FFE1B304FDACF8DD94B6F858" authorityName="d'Abrera &amp; Balint" authorityYear="2001" baseAuthorityName="Hewitson" baseAuthorityYear="1867" box="[589,671,1915,1941]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="auda">
<emphasis id="6FDDE254FFE1B304FDACF8DD94B6F858" box="[589,671,1915,1941]" italics="true" pageId="8" pageNumber="19">J. auda</emphasis>
</taxonomicName>
) and female ductus with lower plate (no lower plate in
<taxonomicName id="9AA945C5FFE1B304FAECF8DD9376F858" authorityName="d'Abrera &amp; Balint" authorityYear="2001" baseAuthorityName="Hewitson" baseAuthorityYear="1867" box="[1293,1375,1915,1941]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="auda">
<emphasis id="6FDDE254FFE1B304FAECF8DD9376F858" box="[1293,1375,1915,1941]" italics="true" pageId="8" pageNumber="19">J. auda</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="5D163E46FFE1B304FF26F839926CF810" blockId="8.[151,1437,1015,2013]" pageId="8" pageNumber="19">
<emphasis id="6FDDE254FFE1B304FF26F8399772F874" box="[199,347,1951,1977]" italics="true" pageId="8" pageNumber="19">Distribution.</emphasis>
Geographical:
<collectingCountry id="25BE7ED6FFE1B304FDF7F83994B6F874" box="[534,671,1951,1977]" name="Ecuador" pageId="8" pageNumber="19">ECUADOR</collectingCountry>
(
<collectingRegion id="9F6DF0A4FFE1B304FD55F839952CF874" box="[692,773,1951,1977]" country="Ecuador" name="Carchi" pageId="8" pageNumber="19">Carchi</collectingRegion>
,
<collectingRegion id="9F6DF0A4FFE1B304FCF7F839957EF874" box="[790,855,1951,1977]" country="Ecuador" name="Napo" pageId="8" pageNumber="19">Napo</collectingRegion>
,
<collectingRegion id="9F6DF0A4FFE1B304FC86F83995F3F877" box="[871,986,1951,1978]" country="Ecuador" name="Pichincha" pageId="8" pageNumber="19">Pichincha</collectingRegion>
) (
<figureCitation id="C59222C3FFE1B304FC16F839927BF874" box="[1015,1106,1951,1977]" captionStart="FIGURE 30" captionStartId="39.[151,250,1742,1766]" captionTargetBox="[251,1396,204,1718]" captionTargetId="figure-18@39.[192,1396,181,1718]" captionTargetPageId="39" captionText="FIGURE 30. Johnsonita distributions based on museum material examinded. J. assula = red circle, J. carpia = yellow trinagle, J. catadupa = blue diamond, J. chalamydem = brown triangle, J. ianusca = green square, J. oxalida = purple diamond." figureDoi="http://doi.org/10.5281/zenodo.4711325" httpUri="https://zenodo.org/record/4711325/files/figure.png" pageId="8" pageNumber="19">Fig. 30</figureCitation>
); spatial: recorded between
<quantity id="9A5193A3FFE1B304FF76F865971CF810" box="[151,309,1987,2013]" metricMagnitude="3" metricUnit="m" metricValue="2.2" metricValueMax="2.8" metricValueMin="1.6" pageId="8" pageNumber="19" unit="m" value="2200.0" valueMax="2800.0" valueMin="1600.0">1600–2800 m</quantity>
a.s.l; temporal: recorded in February–March, May–June and October.
</paragraph>
<paragraph id="5D163E46FFE0B305FF26FF3197DDFF34" blockId="9.[151,1437,151,249]" pageId="9" pageNumber="20">
<emphasis id="6FDDE254FFE0B305FF26FF31976CFF7C" box="[199,325,151,177]" italics="true" pageId="9" pageNumber="20">Etymology.</emphasis>
Named to signify the Ecuadorian provinces, where the species has been collected. The species name „carpia” is freely formed using and combining the first syllable of the departments
<collectingRegion id="9F6DF0A4FFE0B305FBD5FF1D9256FF18" box="[1076,1151,187,213]" country="Ecuador" name="Carchi" pageId="9" pageNumber="20">Carchi</collectingRegion>
and
<collectingRegion id="9F6DF0A4FFE0B305FB54FF1D930FFF18" box="[1205,1318,187,213]" country="Ecuador" name="Pichincha" pageId="9" pageNumber="20">Pichincha</collectingRegion>
, treating it as a female noun in apposition.
</paragraph>
<caption id="09D66ECEFFE0B305FF76F82F9774F824" ID-DOI="http://doi.org/10.5281/zenodo.4711257" ID-Zenodo-Dep="4711257" httpUri="https://zenodo.org/record/4711257/files/figure.png" pageId="9" pageNumber="20" startId="9.[151,250,1929,1953]" targetBox="[210,1382,280,1889]" targetPageId="9">
<paragraph id="5D163E46FFE0B305FF76F82F9774F824" blockId="9.[151,1437,1929,2025]" pageId="9" pageNumber="20">
<emphasis id="6FDDE254FFE0B305FF76F82F973AF86C" bold="true" box="[151,275,1929,1953]" pageId="9" pageNumber="20">FIGURE 4.</emphasis>
<taxonomicName id="9AA945C5FFE0B305FEF8F82F9495F86F" authority="Balint, Boyer &amp; Pyrcz" authorityName="Balint, Boyer &amp; Pyrcz" authorityYear="2021" box="[281,700,1929,1954]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="20" phylum="Arthropoda" rank="species" species="carpia" status="sp. nov.">
<emphasis id="6FDDE254FFE0B305FEF8F82F97F8F86C" box="[281,465,1929,1953]" italics="true" pageId="9" pageNumber="20">Johnsonita carpia</emphasis>
Bálint, Boyer &amp; Pyrcz
</taxonomicName>
,
<taxonomicNameLabel id="74EE5F2FFFE0B305FD27F82C94D5F86C" box="[710,764,1930,1953]" pageId="9" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
imagines (left column: recto, right column: verso). A–B: holotype (Ecuador: Pichincha), C–D: male specimen (Ecuador: Napo), E–F: paratype female (Ecuador: Carchi) (CEP–MZUJ specimens) (scale bar: 10 mm).
</paragraph>
</caption>
<paragraph id="5D163E46FFE3B306FF26FF319379FE60" blockId="10.[151,1436,151,429]" pageId="10" pageNumber="21">
<emphasis id="6FDDE254FFE3B306FF26FF31972DFF7D" box="[199,260,151,176]" italics="true" pageId="10" pageNumber="21">Note.</emphasis>
Our description of
<taxonomicName id="9AA945C5FFE3B306FE06FF319464FF7D" authorityName="Balint, Boyer &amp; Pyrcz" authorityYear="2021" box="[487,589,151,176]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="carpia">
<emphasis id="6FDDE254FFE3B306FE06FF319464FF7D" box="[487,589,151,176]" italics="true" pageId="10" pageNumber="21">J. carpia</emphasis>
</taxonomicName>
is based on material originating exclusively from the Ecuadorian western slope of the Andes (
<collectingRegion id="9F6DF0A4FFE3B306FE94FF1D97E8FF18" box="[373,449,187,213]" country="Ecuador" name="Carchi" pageId="10" pageNumber="21">Carchi</collectingRegion>
and
<collectingRegion id="9F6DF0A4FFE3B306FE19FF1D9445FF1B" box="[504,620,187,214]" country="Ecuador" name="Pichincha" pageId="10" pageNumber="21">Pichincha</collectingRegion>
). Further investigations are necessary for mapping the actual distribution and variability of the species. We have examined
<specimenCount id="4BAFF5CFFFE3B306FD20FF79950EFF34" box="[705,807,223,249]" pageId="10" pageNumber="21" type="male">two male</specimenCount>
specimens from the eastern slopes (
<collectingRegion id="9F6DF0A4FFE3B306FB5DFF7992D6FF34" box="[1212,1279,223,249]" country="Ecuador" name="Napo" pageId="10" pageNumber="21">Napo</collectingRegion>
). One of them seems to be identical with the males collected on the western slopes (
<figureCitation id="C59222C3FFE3B306FC4DFEA595DAFED0" box="[940,1011,259,285]" captionStart="FIGURE 4" captionStartId="9.[151,250,1929,1953]" captionTargetBox="[210,1382,280,1889]" captionTargetId="figure-58@9.[177,1411,273,1906]" captionTargetPageId="9" captionText="FIGURE 4. Johnsonita carpia Bálint, Boyer &amp; Pyrcz, sp. n. imagines (left column: recto, right column: verso). A–B: holotype (Ecuador: Pichincha), C–D: male specimen (Ecuador: Napo), E–F: paratype female (Ecuador: Carchi) (CEP–MZUJ specimens) (scale bar: 10 mm)." figureDoi="http://doi.org/10.5281/zenodo.4711257" httpUri="https://zenodo.org/record/4711257/files/figure.png" pageId="10" pageNumber="21">Fig. 4</figureCitation>
). The NHMUK male from
<collectingRegion id="9F6DF0A4FFE3B306FACBFEA5934EFED0" box="[1322,1383,259,285]" country="Ecuador" name="Napo" pageId="10" pageNumber="21">Napo</collectingRegion>
also has androconia as described for the species, but it is somewhat different in the ventral wing surface pattern: the submedian line erects close to the median pattern. In this respect
<taxonomicName id="9AA945C5FFE3B306FC8FFEED95F8FEA9" authorityName="Balint, Boyer &amp; Pyrcz" authorityYear="2021" box="[878,977,331,356]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="carpia">
<emphasis id="6FDDE254FFE3B306FC8FFEED95F8FEA9" box="[878,977,331,356]" italics="true" pageId="10" pageNumber="21">J. carpia</emphasis>
</taxonomicName>
is variable. The specimen has a red label with black frame, handwritten and printed in black: “
<emphasis id="6FDDE254FFE3B306FD0CFEC99553FE44" box="[749,890,367,393]" italics="true" pageId="10" pageNumber="21">
<typeStatus id="821280E4FFE3B306FD0CFEC99553FE44" box="[749,890,367,393]" pageId="10" pageNumber="21" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
[//] m# [//]
<taxonomicName id="9AA945C5FFE3B306FC1EFEC99250FE44" authorityName="Salazar &amp; Constantino" authorityYear="1995" box="[1023,1145,367,393]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="6FDDE254FFE3B306FC1EFEC99250FE44" box="[1023,1145,367,393]" italics="true" pageId="10" pageNumber="21">Johnsonita</emphasis>
</taxonomicName>
[//]
<emphasis id="6FDDE254FFE3B306FB4BFEC992D7FE45" box="[1194,1278,367,392]" italics="true" pageId="10" pageNumber="21">samson</emphasis>
[//]
<emphasis id="6FDDE254FFE3B306FACEFEC9935AFE44" box="[1327,1395,367,393]" italics="true" pageId="10" pageNumber="21">Bálint</emphasis>
[//]
<emphasis id="6FDDE254FFE3B306FF76FE359762FE60" box="[151,331,403,429]" italics="true" pageId="10" pageNumber="21">London, VIII 23</emphasis>
[//] Zs. Bálint, 2000”, which testifies that the species has been recognized two decades ago.
</paragraph>
</subSubSection>
</treatment>
</document>
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