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<document id="F99E9CA201B04D399A7B4B6115F576A6" ID-CLB-Dataset="34970" ID-DOI="http://dx.doi.org/10.3897/zookeys.494.8748" ID-GBIF-Dataset="e8b2a5fb-bd85-48c3-b33e-121ddc15bec9" ID-PMC="PMC4400378" ID-Pensoft-Pub="1313-2970-494-69" ID-PubMed="25901115" ID-ZooBank="F73F468E4DF7479E85C4717466635BF9" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-494-69" ModsDocOrigin="ZooKeys 494" ModsDocTitle="Revision of the genus Philonome Chambers and its proposed reassignment to the family Tineidae (Lepidoptera, Tineoidea)" checkinTime="1451244473555" checkinUser="pensoft" docAuthor="Sohn, Jae-Cheon, Davis, Donald R. &amp; Lopez-Vaamonde, Carlos" docDate="2015" docId="CB3CB9331B30EC6AD6B11669333ECC14" docLanguage="en" docName="ZooKeys 494: 69-106" docOrigin="ZooKeys 494" docSource="http://dx.doi.org/10.3897/zookeys.494.8748" docTitle="Philonome clemensella Chambers 1874" docType="treatment" docVersion="5" lastPageNumber="80" masterDocId="3D1D9C71FF96FFF09C07FFAC9A60FF9D" masterDocTitle="Revision of the genus Philonome Chambers and its proposed reassignment to the family Tineidae (Lepidoptera, Tineoidea)" masterLastPageNumber="106" masterPageNumber="69" pageNumber="77" updateTime="1701384965743" updateUser="ExternalLinkService">
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<mods:title id="4852FEBCC4F6936D31D5FC32362BA854">Revision of the genus Philonome Chambers and its proposed reassignment to the family Tineidae (Lepidoptera, Tineoidea)</mods:title>
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<mods:namePart id="16CDEF028F99AA91C5C7FB1B3BF11403">Sohn, Jae-Cheon</mods:namePart>
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<mods:namePart id="930324708ED7B0BB30744A688DD4C873">Davis, Donald R.</mods:namePart>
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<mods:namePart id="0159242C25DE8B8F3FE5DCDAFDDB9D4D">Lopez-Vaamonde, Carlos</mods:namePart>
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<mods:identifier id="6FA5F19397DA50A3023FF4CC94C918CE" type="DOI">http://dx.doi.org/10.3897/zookeys.494.8748</mods:identifier>
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<treatment id="CB3CB9331B30EC6AD6B11669333ECC14" ID-GBIF-Taxon="152059278" LSID="urn:lsid:plazi:treatment:CB3CB9331B30EC6AD6B11669333ECC14" httpUri="http://treatment.plazi.org/id/CB3CB9331B30EC6AD6B11669333ECC14" lastPageId="11" lastPageNumber="80" pageId="8" pageNumber="77">
<subSubSection id="33B2E5BDEB4DBD51475EDD837D713273" pageId="8" pageNumber="77" type="multiple">
<paragraph id="2764A911D53C7B2F64BB13FBEC45C081" pageId="8" pageNumber="77">Taxon classification Animalia Lepidoptera Lyonetiidae</paragraph>
</subSubSection>
<subSubSection id="189913A3D666A941498082FEE6329EC0" pageId="8" pageNumber="77" type="nomenclature">
<paragraph id="ED5F45A6353A5E45A5145093E710A14B" pageId="8" pageNumber="77">
<taxonomicName id="756FE083F236BCEFDF3F005BFD9DCA82" ID-CoL="76XV7" authority="Chambers, 1874" authorityName="Chambers" authorityYear="1874" class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome clemensella" order="Lepidoptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="clemensella">Philonome clemensella Chambers, 1874</taxonomicName>
Figs 5, 34-36, 59-60
</paragraph>
</subSubSection>
<subSubSection id="147F3790295620400AAB8A1900AC3356" pageId="8" pageNumber="77" type="reference_group">
<paragraph id="54122431A468E9579A5F478EC679432A" pageId="8" pageNumber="77">
<taxonomicName id="1093901D4132B246A8497AD1B2C8B3B1" class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome clemensella" order="Lepidoptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="clemensella">Philonome clemensella</taxonomicName>
:
<bibRefCitation id="5AEAFCF7DAE59ECD1C4C51EA06F7DA18" author="Chambers, VT" journalOrPublisher="Canadian Entomologist" pageId="27" pageNumber="96" pagination="96 - 97" title="Micro-Lepidoptera." url="10.4039/Ent696-5" volume="6" year="1874">Chambers 1874</bibRefCitation>
: 97;
<bibRefCitation id="6F43E7FF089E822D4670F0B9A8E9F322" author="Dyar, HG" journalOrPublisher="Bulletin of the United States National Museum" pageId="28" pageNumber="97" pagination="1 - 723" title="A list of North American Lepidoptera and key to the literature of this order of insects." volume="52" year="1903">Dyar 1903</bibRefCitation>
: 563;
<bibRefCitation id="732F801D011191223B1CD0948BBC06D1" author="McDunnough, J" journalOrPublisher="Memoirs of the Southern California Academy of Sciences" pageId="28" pageNumber="97" pagination="1 - 171" title="Check list of the Lepidoptera of Canada and the United States of America. Part II Microlepidoptera." volume="2" year="1939">McDunnough 1939</bibRefCitation>
: 100;
<bibRefCitation id="366F097EBE08BE27C7FD8F507BE5EE34" author="Davis, DR" editor="Hodges, RW" journalOrPublisher="E. W. Classey Limited and the Wedge Entomological Research Foundation, London" pageId="27" pageNumber="96" pagination="8 - 9" title="Lyonetiidae." volumeTitle="Check List of the Lepidoptera of America North of Mexico" year="1983">Davis 1983</bibRefCitation>
: 8.
</paragraph>
<paragraph id="0EEF2555FE5CAD98697252712DA8F699" pageId="8" pageNumber="77">
<taxonomicName id="7EEA8B8004566F184F769F1E4BD013E1" class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome staintonella" order="Lepidoptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="staintonella">Philonome staintonella</taxonomicName>
:
<bibRefCitation id="7FE8ECD9FEC170E902EA2C55085D9EF2" author="Chambers, VT" journalOrPublisher="The Canadian Entomologist" pageId="27" pageNumber="96" pagination="135 - 138" title="Tineina." url="10.4039/Ent8135-7" volume="8" year="1876">Chambers 1876</bibRefCitation>
: 136. Nomen nudum
</paragraph>
</subSubSection>
<subSubSection id="B57514FEF6DCD9BCBD44FFB98F2B2F10" lastPageId="9" lastPageNumber="78" pageId="8" pageNumber="77" type="adult">
<paragraph id="433208FBBAF180463139D1F2AD1EAD45" pageId="8" pageNumber="77">Adult</paragraph>
<paragraph id="EBF247F6CAE317870C4D4D34A09DEBBF" pageId="8" pageNumber="77">
(Fig. 5). Head: Vertex convex medially; scales on vertex orange, as long as antennal scape, slanted forward; semicircular, dome-like scale cap on anterior vertex between antennal scapes with compact appressed, lustrous pale yellow scales; frons smooth, lustrous pale yellow. Antenna 3/5 as long as forewing; scape reddish brown dorsally, lustrous pale yellow ventrally, with fan-shaped scale cap anterioventrally; flagellomeres dark reddish brown dorsally, lustrous pale yellow ventrally. Labial palpus straight, slender, very small, 1/2 as long as antennal scape, pale orange, apex acuminate. Maxillary palpus 3
<normalizedToken id="673F6BC2D05C40E15BC42C1505AF1ED1" originalValue="×">x</normalizedToken>
longer than labial palpus, pale orange.
</paragraph>
<paragraph id="2DA8DE8DED4035BE9B7CFC26BA49BF9D" pageId="9" pageNumber="78">
<pageBreakToken id="092A2B21893F226D85BFE077DB563BD8" pageId="9" pageNumber="78" start="start">Thorax</pageBreakToken>
: Patagium reddish brown; tegula pale orange-white, suffused with reddish brown basally; mesonotum pale orange-white. Foreleg with coxa lustrous orange-white, tinged with gray dorsobasally; femur and tibia dark brownish gray dorsally, lustrous orange-white ventrally; tarsi orange-white, lustrous ventrally. Midleg with coxa lustrous orange-white; femur to tarsi pale orange dorsally, lustrous orange-white. Hindleg with coxa to tarsi lustrous orange-white; femur narrowly tinged with pale orange dorsally; tibia spinose dorsally, with hair-tufts ventrally. Forewing length 2.8-4.4 mm (n = 70), reddish brown; longitudinal fascia, white from base to the middle of forewing, often connected with white dorsal bar; costal fascia at distal 1/3 of costa, white, terminal 1/3 curved to apex, accompanying a row of dark brown scales caudad; subapical spot orange-white; tornal patch white, semicircular, borders blurred; elongate scales on apex, terminal 1/3 of costa, termen, with dark brown tips; fringe orange-white. Hindwing and fringe lustrous pale gray.
</paragraph>
<paragraph id="8CAC85F4BAB171776BC213C37857C7D2" pageId="9" pageNumber="78">Abdomen: Terga lustrous, yellowish gray; sterna lustrous orange-white. Male tergum VIII trapezoidal; male sternum VIII subrectangular, broadly emarginated posteriorly, with oblique furrow laterally and short, stiff coremata posterolaterally.</paragraph>
<paragraph id="35F76A1784E5D84A21CCAFCF257B1E61" pageId="9" pageNumber="78">Male genitalia (Figs 34-36): Tegumen trapezoidal, with subrectangular protrusion apically, strongly sclerotized, digitate process laterally, and short, lanceolate sclerite at center; long setae above teguminal process; tuba analis arising from dorsal area of apical protrusion. Valva deeply divided into two portions; costal portion triangular on basal half, elongate on distal half, sparsely setose, apex protruding; saccular portion elliptical, more densely setose to apex. Anellus extending to middle of phallus. Vinculum broadly sclerotized, capsulate, with a pair of small protrusions on distal margin. Phallus of even width except slightly-swollen at basal 1/6, slightly bifid apically.</paragraph>
<paragraph id="8146FC32C984EA2078DF7B18F1438287" pageId="9" pageNumber="78">
Female genitalia (Figs 59-60): Apophyses posteriores 2
<normalizedToken id="A73E2F83FB49D005FF0664F0B4AF0321" originalValue="×">x</normalizedToken>
longer than apophyses anteriores. Elongate scales on posteroventral margin of A8. Lamella antevaginalis short, cylindrical, with ridge posterolaterally. Ductus bursae as long as corpus bursae, narrow on anterior 1/2; inception of ductus seminalis at middle of ductus bursae, bulged, with a sclerotized ring. Corpus bursae obovate.
</paragraph>
</subSubSection>
<subSubSection id="39E30C1DE85AA85ACC0C16B512A4B6A9" pageId="9" pageNumber="78" type="types">
<paragraph id="6170EE472D1C13E4C6F5AA8FAFAB8366" pageId="9" pageNumber="78">Types.</paragraph>
<paragraph id="C2ECC9FF701C2CB71A8B4CE8C185DB00" pageId="9" pageNumber="78">
Lectotype (designated here): male,
<normalizedToken id="B415509A2253D807AFE7C86D7B1CA28D" originalValue="“48”">&quot;48&quot;</normalizedToken>
[hand-written], &quot;Type No. 522 U.S.N.M.&quot; [red label], &quot;
<taxonomicName id="866139838EEF0C6914FBA2405B05E657" class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome clemmensella" order="Lepidoptera" pageId="9" pageNumber="78" phylum="Arthropoda" rank="species" species="clemmensella">Philonome clemmensella</taxonomicName>
[sic] K[entuck]y. 5961Lis1 Cham[bers]&quot; [hand-written], USNM. Paralectotypes: USA: Kentucky: 4♀, 3 ex., June 14 [no year] (Chambers), Type no. 1311, MCZ.
</paragraph>
</subSubSection>
<subSubSection id="86662A4701E98CE435E7B11E13D1BB8B" lastPageId="11" lastPageNumber="80" pageId="9" pageNumber="78" type="material examined">
<paragraph id="E45F377381E2DA7DFF248DD35A6D95A4" pageId="9" pageNumber="78">Material examined.</paragraph>
<paragraph id="99452090C2929E7B286DF1E10D302DF5" lastPageId="11" lastPageNumber="80" pageId="9" pageNumber="78">
Canada: Ontario: Ottawa-Careton, Dunrobin: 1♀, 9 July 2007, CNCLEP00035968; 28 July, 2007, CNCLEP00038457, (L Scott). Quebec: 2♂, 21 July 2004, CNCLEP00006545, (JF Landry), CNC. USA: Alabama: Monroe Co.: Haines Island Park (
<geoCoordinate id="56D262224BE1198AFB6F48CA52E1C2D8" direction="north" orientation="latitude" precision="15" value="31.723057">31°43'23&quot;N</geoCoordinate>
,
<geoCoordinate id="A6B842DE75843A17BF25028A137C5C54" direction="west" orientation="longitude" precision="15" value="-87.469444">87°28'10&quot;W</geoCoordinate>
): 2♂, 2♀, 26-27 May 1995 (R Brown, J MacGown &amp; D Pollock), MSU. District of Columbia: Unspecified locality: 1♂, no date (Fernald); 1♂, 28 June 1885 (Fernald), &quot;on oak&quot;; 1♀, 21 June 1886 (Fernald); 1♂, 11 July 1896,
<normalizedToken id="D0533014F24955DBFCE3E126A8380D4A" originalValue="“Hickory”">&quot;Hickory&quot;</normalizedToken>
; 3♀, 2-4 June 1897, &quot;from Linden&quot;; 1♀, September 1953, USNM. Florida: Pinellas Co.: Dunedin: Hammock Park: 1♂, 22 April 1987 (LC Dow), [GSN] USNM 96414, USNM. Kentucky: No specified locality: 1♂, no date &amp; collector, USNM. Illinois: Macon Co.: Decatur: 1♂, 8-15 June [no year], USNM. Putnam Co.: 1♀, 30 June 1976 (MO Glenn), USNM. Maryland: Montgomery Co.: Takoma Park: 1♂, 1♀, 8 July 1986 (WE Steiner); 1♂, 7 July 1986, USNM. Wicomico Co.: 1km SW Sharptown at Plum Creek: 1♀, 12 July 1986 (JM Hill et al.), USNM. Massachusetts: Dukes Co.:
<normalizedToken id="3CA57307E5A403B9800FF0E311E0DD8E" originalValue="Marthas">Martha's</normalizedToken>
Vineyard: 1♂, 13 July [no
<pageBreakToken id="ACB3B930749A65E2139AA8A27A877953" pageId="10" pageNumber="79" start="start">year</pageBreakToken>
] (FM Jones); 1 ex., 29 July [no year], all USNM. New Jersey: Burlington Co.: Medford: Lake Pine: 1♂, 13 July 1974 (DC Rentz), USNM. Essex Co.: Caldwell: 3♂, 2♀, 8 July 1900 (WD Kearfott), USNM. Essex County Park: 1♀, 20 May [no year] (WD Kearfott); 1♂, 7 July [no year], GSN: USNM 29977; 1♂, 12 July 1901; 1♂, 15 July [no year], USNM. Montclair: 1♀, 10 July [no year] (WD Kearfott), USNM; 1♂, 18 July [no year] (WD Kearfott), USNM. New York: Tompkins Co.: Ithaca: Six Mile Creek: 1♀, 23 July 1960 (RW Hodges), USNM. Unspecified locality: 1♀, &quot;4971/ WLSM. 1906&quot; (Beutenmueller), USNM. North Carolina: Craven Co.: Cherrypoint:
<pageBreakToken id="17AF3F71BCF7C10EFD60A068FCBBFE3D" pageId="11" pageNumber="80" start="start">1</pageBreakToken>
♀, 3 July 1961 (SS Nicolay); 1♀, 12 July 1961; 1♀, 21 July 1961, all USNM. Harnett Co.: Spout Springs: 1♂, 25 August 1984 (WE Steiner et al.), USNM. Ohio: Hamilton Co.: Cincinnati, 1 ex., 23 June 1906 (A Braun); 1♂, 1♀, 27-28 June 1906, [GSN] USNM 16405 (♂); 1♀, 24 July 1907; 1♂, 3 August 1907; 1♂, 16 June 1908, all USNM. PENNSYLVANIA: Adams Co.: Arendtsville: 5♂, 6 July 1921 (SW Frost), USNM, GSN: USNM 29575. Allegheny Co.: Oak Station: 1♂, 6 July 1907 (F Marloff); 1♂, 11 July 1907, all USNM. Beaver Co.: New Brighton: 2♂, 11 July 1907 (Merrick Museum); 1♂, 23 July 1907; 1♂, 2♀, 26 July 1907, [GSN] USNM 34213 (♀), USNM. South Carolina: Charleston Co.: McClellanville: Wedge Plantation: 1♀, 11 May 1981 (RW Hodges), USNM, GSN: USNM 34212. Tennessee: Cocke Co.: Great Smoky Mt. National Park: Foothills Parkway (
<geoCoordinate id="4046052AD9D01CD8856A1E7BE9C00A85" direction="north" orientation="latitude" precision="15" value="35.816387">35°48'59&quot;N</geoCoordinate>
,
<geoCoordinate id="E7FB12B0640FA39B81E111CE23BDD5F5" direction="west" orientation="longitude" precision="15" value="-83.21972">83°13'11&quot;W</geoCoordinate>
): 3♂, 1♀, 9 June 2002 (RL Brown &amp; SM Lee), MSU. Texas: Harris Co.: 1♂, 20 May 1984 (Bellaire), [GSN] USNM 96415; 1♂, 2 April 1986, all USNM. Virginia: Fairfax Co.: 1km E Fairfax City: 1♂, 9 July 2005 (J Brown), USNM. Unspecified locality: 1♀, 27 June 1886, USNM. West Virginia: Morgan Co.: Sleepy Creek Forest: 2♂, 1 July 2010 (J Glaser); 1♂, 16 July 2011; 1♂, 19 July 2011; 1♂, 21 July 2011, USNM.
</paragraph>
</subSubSection>
<subSubSection id="4E3E0A3CBDE6A7DB20B9F63E82BE3AA1" pageId="11" pageNumber="80" type="distribution">
<paragraph id="4419FBE4C4A978FE44FF52757826F452" pageId="11" pageNumber="80">Distribution.</paragraph>
<paragraph id="EB057EA1FE450ADE6DA3E6372ED3A867" pageId="11" pageNumber="80">Eastern Canada and the United States west to Texas.</paragraph>
</subSubSection>
<subSubSection id="31CC86204FB4426A84012C7F9FDD754E" pageId="11" pageNumber="80" type="host plants">
<paragraph id="B1F93A13BF8E1D617E0E77250AC3A38C" pageId="11" pageNumber="80">Host plants.</paragraph>
<paragraph id="2615F9B6EDFA7B051E5073AB58BD7A66" pageId="11" pageNumber="80">
Hickory (
<taxonomicName id="309D8125B49CCFF7E711B85A327CCE1B" family="Juglandaceae" lsidName="Yponomeutoidea" pageId="11" pageNumber="80" rank="family">Juglandaceae</taxonomicName>
:
<taxonomicName id="E14A9CEECF926978BB75B046B5370E66" class="Magnoliopsida" family="Juglandaceae" genus="Carya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Carya" order="Fagales" pageId="11" pageNumber="80" phylum="Tracheophyta" rank="genus">Carya</taxonomicName>
) and linden (
<taxonomicName id="BB7AED564CC924F61B3CA00C53A13246" family="Tiliaceae" lsidName="Yponomeutoidea" pageId="11" pageNumber="80" rank="family">Tiliaceae</taxonomicName>
: possibly
<taxonomicName id="B111307B7AB554BA50301C229B0CB74F" class="Magnoliopsida" family="Malvaceae" genus="Tilia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tilia americana" order="Malvales" pageId="11" pageNumber="80" phylum="Tracheophyta" rank="species" species="americana">Tilia americana</taxonomicName>
L.) (
<bibRefCitation id="3F1B670583C1F73102B2878F136F17B1" author="Forbes, WTM" journalOrPublisher="Cornell University Press, Ithaca, New York" pageId="28" pageNumber="97" title="Lepidoptera of New York and Neighboring States Part 1. Cornell University Agricultural Experiment Station Memoir 68." year="1923">Forbes 1923</bibRefCitation>
). These are from the label data in the USNM collection. The collection also includes a specimen whose label data states that it came from oak (
<taxonomicName id="04D57D825793432EB7D1123F873AFE3D" family="Fagaceae" lsidName="Yponomeutoidea" pageId="11" pageNumber="80" rank="family">Fagaceae</taxonomicName>
:
<taxonomicName id="887742BE5B3F9B4C05C2D92B50383A08" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus" order="Fagales" pageId="11" pageNumber="80" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
). The label data give no details other than plant common names. Therefore, it is not clear if these records refer to larval host plants or where the adults were collected.
</paragraph>
</subSubSection>
</treatment>
</document>