496 lines
72 KiB
XML
496 lines
72 KiB
XML
<document id="45E673EAB45D9A991D080738A1C0544B" ID-DOI="10.5852/ejt.2024.975.2769" ID-ISSN="2118-9773" ID-Zenodo-Dep="14508730" ID-ZooBank="23F0D60C-3B5B-4AB1-B608-AD875EB424DF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1734447201131" checkinUser="plazi" docAuthor="Constant, Jérôme & Pham, Hong Thai" docDate="2024" docId="CB3887FB716F621EFB4AFC14FB9DFC79" docLanguage="en" docName="EJT.2024.975.1-64.pdf" docOrigin="European Journal of Taxonomy 975" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2769/12667" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pusulissus bachmaensis Constant & Pham, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="60" masterDocId="3701FF83715A6225F91DFFD3FFF7FFAD" masterDocTitle="Issid planthoppers from Bach Ma and Phong Dien in Central Vietnam. I. Tribe Parahiraciini (Hemiptera: Fulgoromorpha: Issidae)" masterLastPageNumber="64" masterPageNumber="1" pageNumber="54" updateTime="1734447759699" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="A85DA1404E0C3390DE6C32CAB71C0539" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="F8BBA52F930E7A6B67F189D9A13F88E6">Issid planthoppers from Bach Ma and Phong Dien in Central Vietnam. I. Tribe Parahiraciini (Hemiptera: Fulgoromorpha: Issidae)</mods:title>
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<mods:name id="8FE423694499E3703881F961C1BA4BBA" type="personal">
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<mods:role id="E3C528C02D103A4B90FF73426AF03BB5">
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<mods:roleTerm id="6881335B1CD78AE8AE020A13C0673D8F">Author</mods:roleTerm>
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<mods:namePart id="3F76E6A06A9457F76FF79C5C7F8815C3">Constant, Jérôme</mods:namePart>
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<mods:nameIdentifier id="5F3B89FBEC2245472D907FC0BB253187" type="LSID">6E6072A1-9415-4C8D-8E60-2504444DB290</mods:nameIdentifier>
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<mods:affiliation id="0789B1894BF7CC40EAE1DF62FE147314">Royal Belgian Institute of Natural Sciences, O. D. Phylogeny and Taxonomy, Entomology, Vautier Street 29, B- 1000 Brussels, Belgium.</mods:affiliation>
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<mods:nameIdentifier id="EC7F3C2940C03A2165A2E7BEB4A96074" type="email">jerome.constant@naturalsciences.be</mods:nameIdentifier>
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<mods:name id="C0FEF9CC6332D0DF2775354C174E014E" type="personal">
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<mods:roleTerm id="01220A116CC739A452E61E2512F3CB05">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="28DD511F1BBE36013308F1DB8E8720CA">Pham, Hong Thai</mods:namePart>
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<mods:affiliation id="E2D3AF7A898C7719DB9996C0B921AC1B">Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Street, Hanoi, Vietnam. & Graduate School of Science and Technology, Vietnam Academy of Science and Technology, Hanoi, Vietnam.</mods:affiliation>
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<mods:nameIdentifier id="66CA47C48B0BFEB16DDD63523F59C6A6" type="email">jerome.constant@naturalsciences.be</mods:nameIdentifier>
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<mods:typeOfResource id="B565E2134C2907BDE6147D3A6D84C459">text</mods:typeOfResource>
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<mods:title id="D0EE488CFF9249375D2C3FAC6A4E51BB">European Journal of Taxonomy</mods:title>
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<mods:part id="7E9754C1D2F7B3D27183C53BF4C31FA5">
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<mods:date id="346EBBE481F65F07F638ED581738E76E">2024</mods:date>
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<mods:number id="D49DC15DAD36533CAE5BCE1AC300EAE9">2024-12-17</mods:number>
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<mods:number id="46B7D7D5D2AF86EFD02774AFEB0C8C14">975</mods:number>
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<mods:url id="8D5987AF104D7A69705C3D65EDEC35A0">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2769/12667</mods:url>
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<mods:identifier id="9D7DDDDADCBD503C2FD4D89DA8A75A53" type="DOI">10.5852/ejt.2024.975.2769</mods:identifier>
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<treatment id="CB3887FB716F621EFB4AFC14FB9DFC79" LSID="urn:lsid:plazi:treatment:CB3887FB716F621EFB4AFC14FB9DFC79" httpUri="http://treatment.plazi.org/id/CB3887FB716F621EFB4AFC14FB9DFC79" lastPageId="59" lastPageNumber="60" pageId="53" pageNumber="54">
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<subSubSection id="0B8B6566716F6210FB4AFC14FB37FBA8" pageId="53" pageNumber="54" type="nomenclature">
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<paragraph id="432E36ED716F6210FB4AFC14FC2AFC4F" blockId="53.[372,1216,967,1064]" box="[599,989,967,994]" pageId="53" pageNumber="54">
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<heading id="18668181716F6210FB4AFC14FC2AFC4F" box="[599,989,967,994]" centered="true" fontSize="11" level="2" pageId="53" pageNumber="54" reason="2">
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<taxonomicName id="84914D6E716F6210FB4AFC14FC8CFC4F" authorityName="Constant & Pham" authorityYear="2024" box="[599,891,967,994]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="bachmaensis" status="sp. nov.">
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<emphasis id="71E5EAFF716F6210FB4AFC14FC8CFC4F" bold="true" box="[599,891,967,994]" italics="true" pageId="53" pageNumber="54">Pusulissus bachmaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6AD65784716F6210FA9EFC1BFC2AFC4F" box="[899,989,968,994]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="432E36ED716F6210F869FC39FB37FBA8" blockId="53.[372,1216,967,1064]" box="[372,1216,1002,1029]" pageId="53" pageNumber="54">
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urn:lsid:zoobank.org:act:
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<uuid id="37370C38716F6210FB8DFC39FB37FBA8" box="[656,1216,1002,1029]" pageId="53" pageNumber="54">541CA790-5C5F-4407-BA9A-099000E2154E</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="0B8B6566716F6210FBADFBDEFC74FB85" box="[688,899,1037,1064]" pageId="53" pageNumber="54" type="description">
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<paragraph id="432E36ED716F6210FBADFBDEFC74FB85" blockId="53.[372,1216,967,1064]" box="[688,899,1037,1064]" pageId="53" pageNumber="54">
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<figureCitation id="DBAA2A68716F6210FBADFBDEFD0EFB8A" box="[688,761,1037,1063]" captionStart="Fig" captionStartId="6.[189,232,1998,2024]" captionTargetBox="[189,1398,729,1950]" captionTargetId="figure-138@6.[189,1398,729,1950]" captionTargetPageId="6" captionText="Fig. 1. Map of the sampled localities in Thừa Thiên-Huế Province, Vietnam." figureDoi="http://doi.org/10.5281/zenodo.14508732" httpUri="https://zenodo.org/record/14508732/files/figure.png" pageId="53" pageNumber="54">Figs 1</figureCitation>
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,
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<figureCitation id="DBAA2A68716F6210FA1AFBDEFCD9FB8A" box="[775,814,1037,1063]" captionStart="Fig" captionStartId="7.[190,233,1963,1989]" captionTargetBox="[189,1398,1011,1918]" captionTargetId="figure-245@7.[189,1398,1011,1918]" captionTargetPageId="7" captionText="Fig. 2. Typical habitats in Bach Ma National Park, May 2023. A. Pheasant trail. B. Roadside (1200– 1300 m a.s.l.). C. Rhododendron trail. D. Summit (1300–1400 m a.s.l.)." figureDoi="http://doi.org/10.5281/zenodo.14508734" httpUri="https://zenodo.org/record/14508734/files/figure.png" pageId="53" pageNumber="54">2D</figureCitation>
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,
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<figureCitation id="DBAA2A68716F6210FA27FBDEFC74FB85" box="[826,899,1037,1064]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="55.[189,232,1928,1954]" captionStartId-1="56.[189,232,1892,1918]" captionStartId-2="57.[189,232,1892,1918]" captionStartId-3="58.[189,232,1963,1989]" captionTargetBox-0="[221,1374,273,1866]" captionTargetBox-1="[203,1384,453,1847]" captionTargetBox-2="[258,1331,285,1845]" captionTargetBox-3="[189,1398,1110,1917]" captionTargetId-0="figure-11@55.[211,1377,265,1885]" captionTargetId-1="figure-63@56.[203,1384,442,1850]" captionTargetId-2="figure-11@57.[257,1331,265,1845]" captionTargetId-3="figure-331@58.[189,1398,1110,1917]" captionTargetPageId-0="55" captionTargetPageId-1="56" captionTargetPageId-2="57" captionTargetPageId-3="58" captionText-0="Fig. 32. Pusulissus bachmaensis sp. nov., dissected holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Right hind leg, ventral view. E. Metatarsus and apex of metatibia, ventral view." captionText-1="Fig. 33. Pusulissus bachmaensis sp. nov. A–E. Holotype ♂ (RBINS). A. Head and thorax, dorsal view. B. Frons, perpendicular view. C. Head and thorax, lateral view. D. Head and thorax, anterolateral view. E. Right hind wing. F. Paratype ♀ (VNMN), terminalia, ventral view. Abbreviations: see Material and methods." captionText-2="Fig. 34. Pusulissus bachmaensis sp. nov., holotype, ♂ (RBINS). A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E–L. Aedeagus. E. Posteroventral view. F. Left lateral view. G. Left laterodorsal view. H. Anterodorsal view. I. Dorsal view. J. Ventral view. K. Anteroventral view. Abbreviations: see Material and methods." captionText-3="Fig. 35. Pusulissus bachmaensis sp. nov., paratype, ♀ (VNMN) from Bach Ma National Park, summit, 21 May 2023. A–D. Live specimen (photographed in cage)." figureDoi-0="http://doi.org/10.5281/zenodo.14508802" figureDoi-1="http://doi.org/10.5281/zenodo.14508804" figureDoi-2="http://doi.org/10.5281/zenodo.14508806" figureDoi-3="http://doi.org/10.5281/zenodo.14508808" httpUri-0="https://zenodo.org/record/14508802/files/figure.png" httpUri-1="https://zenodo.org/record/14508804/files/figure.png" httpUri-2="https://zenodo.org/record/14508806/files/figure.png" httpUri-3="https://zenodo.org/record/14508808/files/figure.png" pageId="53" pageNumber="54">32–35</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0B8B6566716F6210F9A0FB9DFAE7F8D8" pageId="53" pageNumber="54" type="diagnosis">
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<paragraph id="432E36ED716F6210F9A0FB9DFEC1FBC5" blockId="53.[189,310,1102,1128]" box="[189,310,1102,1128]" pageId="53" pageNumber="54">
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<heading id="18668181716F6210F9A0FB9DFEC1FBC5" bold="true" box="[189,310,1102,1128]" fontSize="11" level="3" pageId="53" pageNumber="54" reason="3">
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<emphasis id="71E5EAFF716F6210F9A0FB9DFEC1FBC5" bold="true" box="[189,310,1102,1128]" pageId="53" pageNumber="54">Diagnosis</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="432E36ED716F6210F9A0FBAFFCB6FB3B" blockId="53.[189,1399,1148,1521]" box="[189,833,1148,1174]" pageId="53" pageNumber="54">
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<taxonomicName id="84914D6E716F6210F9A0FBAFFE2FFB3B" authorityName="Constant & Pham" authorityYear="2024" box="[189,472,1148,1174]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="bachmaensis" status="sp. nov.">
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<emphasis id="71E5EAFF716F6210F9A0FBAFFE2FFB3B" box="[189,472,1148,1174]" italics="true" pageId="53" pageNumber="54">Pusulissus bachmaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6AD65784716F6210F8C2FBAFFDCDFB3B" box="[479,570,1148,1174]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
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can be recognized by:
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</paragraph>
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<paragraph id="432E36ED716F6210F9A0FB4CFC09FB14" blockId="53.[189,1399,1148,1521]" box="[189,1022,1182,1209]" pageId="53" pageNumber="54">
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(1) the frons elongate, distinctly constricted between eyes (
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<figureCitation id="DBAA2A68716F6210FA99FB4DFC07FB14" box="[900,1008,1182,1209]" captionStart="Fig" captionStartId="56.[189,232,1892,1918]" captionTargetBox="[203,1384,453,1847]" captionTargetId="figure-63@56.[203,1384,442,1850]" captionTargetPageId="56" captionText="Fig. 33. Pusulissus bachmaensis sp. nov. A–E. Holotype ♂ (RBINS). A. Head and thorax, dorsal view. B. Frons, perpendicular view. C. Head and thorax, lateral view. D. Head and thorax, anterolateral view. E. Right hind wing. F. Paratype ♀ (VNMN), terminalia, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.14508804" httpUri="https://zenodo.org/record/14508804/files/figure.png" pageId="53" pageNumber="54">Fig. 33B</figureCitation>
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);
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</paragraph>
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<paragraph id="432E36ED716F6210F9A0FB12FE17FA8C" blockId="53.[189,1399,1148,1521]" pageId="53" pageNumber="54">
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(2) the anal tube in dorsal aspect widely spatulate with distal portion nearly circular, nearly 1.3 times as long in midline as wide with apical margin shortly truncate in middle, without distinct lateral flaps (
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<emphasis id="71E5EAFF716F6210F81EFAD4FED3FA8C" box="[259,292,1287,1313]" italics="true" pageId="53" pageNumber="54">An</emphasis>
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–
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<figureCitation id="DBAA2A68716F6210F85CFAD5FE25FA8C" box="[321,466,1286,1313]" captionStart="Fig" captionStartId="57.[189,232,1892,1918]" captionTargetBox="[258,1331,285,1845]" captionTargetId="figure-11@57.[257,1331,265,1845]" captionTargetPageId="57" captionText="Fig. 34. Pusulissus bachmaensis sp. nov., holotype, ♂ (RBINS). A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E–L. Aedeagus. E. Posteroventral view. F. Left lateral view. G. Left laterodorsal view. H. Anterodorsal view. I. Dorsal view. J. Ventral view. K. Anteroventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.14508806" httpUri="https://zenodo.org/record/14508806/files/figure.png" pageId="53" pageNumber="54">Fig. 34A–D</figureCitation>
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);
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</paragraph>
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<paragraph id="432E36ED716F6210F9A0FAFAFE83FACB" blockId="53.[189,1399,1148,1521]" pageId="53" pageNumber="54">
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(3) the neck of the capitulum of the gonostyli rather short, less high than head of capitulum (
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<emphasis id="71E5EAFF716F6210FC5CFAF9FAAAFAEE" box="[1345,1373,1322,1347]" italics="true" pageId="53" pageNumber="54">ca</emphasis>
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–
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<figureCitation id="DBAA2A68716F6210F9E4FA9FFE91FACB" box="[249,358,1356,1382]" captionStart="Fig" captionStartId="57.[189,232,1892,1918]" captionTargetBox="[258,1331,285,1845]" captionTargetId="figure-11@57.[257,1331,265,1845]" captionTargetPageId="57" captionText="Fig. 34. Pusulissus bachmaensis sp. nov., holotype, ♂ (RBINS). A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E–L. Aedeagus. E. Posteroventral view. F. Left lateral view. G. Left laterodorsal view. H. Anterodorsal view. I. Dorsal view. J. Ventral view. K. Anteroventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.14508806" httpUri="https://zenodo.org/record/14508806/files/figure.png" pageId="53" pageNumber="54">Fig. 34A</figureCitation>
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);
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</paragraph>
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<paragraph id="432E36ED716F6210F9A0FABCFDD8FA5C" blockId="53.[189,1399,1148,1521]" pageId="53" pageNumber="54">
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(4) the shape of the lateroventral processes of the aedeagus slightly inflated basally, then constricted, directed cephalad and moderately, evenly upcurved, remaining under the ventral lobe of the periandrium and reaching to basiventral lobes of periandrium; distal portion tapering and curved mesad (
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<emphasis id="71E5EAFF716F6210F848FA05FE8EFA5D" box="[341,377,1494,1520]" italics="true" pageId="53" pageNumber="54">lvp</emphasis>
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–
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<figureCitation id="DBAA2A68716F6210F88AFA05FDD3FA5C" box="[407,548,1494,1521]" captionStart="Fig" captionStartId="57.[189,232,1892,1918]" captionTargetBox="[258,1331,285,1845]" captionTargetId="figure-11@57.[257,1331,265,1845]" captionTargetPageId="57" captionText="Fig. 34. Pusulissus bachmaensis sp. nov., holotype, ♂ (RBINS). A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E–L. Aedeagus. E. Posteroventral view. F. Left lateral view. G. Left laterodorsal view. H. Anterodorsal view. I. Dorsal view. J. Ventral view. K. Anteroventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.14508806" httpUri="https://zenodo.org/record/14508806/files/figure.png" pageId="53" pageNumber="54">Fig. 34F–K</figureCitation>
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).
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</paragraph>
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<paragraph id="432E36ED716F6210F9A0F9C4FE3EF99C" blockId="53.[189,457,1559,1585]" box="[189,457,1559,1585]" pageId="53" pageNumber="54">
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<heading id="18668181716F6210F9A0F9C4FE3EF99C" bold="true" box="[189,457,1559,1585]" fontSize="11" level="3" pageId="53" pageNumber="54" reason="3">
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<emphasis id="71E5EAFF716F6210F9A0F9C4FE3EF99C" bold="true" box="[189,457,1559,1585]" pageId="53" pageNumber="54">Differential diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="432E36ED716F6210F9A0F996FAE7F8D8" blockId="53.[188,1401,1605,1909]" pageId="53" pageNumber="54">
|
||
<taxonomicName id="84914D6E716F6210F9A0F996FE21F9F2" authorityName="Constant & Pham" authorityYear="2024" box="[189,470,1605,1631]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="bachmaensis" status="sp. nov.">
|
||
<emphasis id="71E5EAFF716F6210F9A0F996FE21F9F2" box="[189,470,1605,1631]" italics="true" pageId="53" pageNumber="54">Pusulissus bachmaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6AD65784716F6210F8C6F995FDC3F9CD" box="[475,564,1606,1632]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be separated from
|
||
<taxonomicName id="84914D6E716F6210FA25F995FC02F9F2" authorityName="Bourgoin & Wang" authorityYear="2020" box="[824,1013,1606,1631]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="coronomensis">
|
||
<emphasis id="71E5EAFF716F6210FA25F995FC02F9F2" box="[824,1013,1606,1631]" italics="true" pageId="53" pageNumber="54">P. coronomensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="84914D6E716F6210FD1EF995FB41F9F2" baseAuthorityName="Meng, Qin & Wang" baseAuthorityYear="2020" box="[1027,1206,1605,1631]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="guizhouensis">
|
||
<emphasis id="71E5EAFF716F6210FD1EF995FB41F9F2" box="[1027,1206,1605,1631]" italics="true" pageId="53" pageNumber="54">P. guizhouensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="84914D6E716F6210FDDFF995FA81F9F2" authorityName="Bourgoin & Wang" authorityYear="2020" box="[1218,1398,1605,1631]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="phiaoacensis">
|
||
<emphasis id="71E5EAFF716F6210FDDFF995FA81F9F2" box="[1218,1398,1605,1631]" italics="true" pageId="53" pageNumber="54">P. phiaoacensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="84914D6E716F6210F9EFF9BAFE3CF92F" authorityName="Constant & Pham" authorityYear="2024" box="[242,459,1640,1666]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="quangninhensis">
|
||
<emphasis id="71E5EAFF716F6210F9EFF9BAFE3CF92F" box="[242,459,1640,1666]" italics="true" pageId="53" pageNumber="54">P. quangninhensis</emphasis>
|
||
</taxonomicName>
|
||
by the round shape of the anal tube without lateral flaps (anal tube more or less rhomboidal with lateral flaps projecting ventrad in all three other species). The constricted frons between eyes and the rather strongly curved aedeagus in lateral aspect, also separates
|
||
<taxonomicName id="84914D6E716F6210FDDFF97DFA8EF96A" authorityName="Constant & Pham" authorityYear="2024" box="[1218,1401,1709,1735]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="bachmaensis">
|
||
<emphasis id="71E5EAFF716F6210FDDFF97DFA8EF96A" box="[1218,1401,1709,1735]" italics="true" pageId="53" pageNumber="54">P. bachmaensis</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="84914D6E716F6210F9E0F902FE5DF947" baseAuthorityName="Che, Zhang & Wang" baseAuthorityYear="2020" box="[253,426,1744,1770]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="menglaensis">
|
||
<emphasis id="71E5EAFF716F6210F9E0F902FE5DF947" box="[253,426,1744,1770]" italics="true" pageId="53" pageNumber="54">P. menglaensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="84914D6E716F6210F8FCF902FD9CF947" baseAuthorityName="Che, Zhang & Wang" baseAuthorityYear="2020" box="[481,619,1744,1770]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="ruiliensis">
|
||
<emphasis id="71E5EAFF716F6210F8FCF902FD9CF947" box="[481,619,1744,1770]" italics="true" pageId="53" pageNumber="54">P. ruiliensis</emphasis>
|
||
</taxonomicName>
|
||
, which do not show a distinct constriction of the frons between the eyes, and have a much less curved aedeagus;
|
||
<taxonomicName id="84914D6E716F6210FBC9F927FC7EF8A0" authorityName="Constant & Pham" authorityYear="2024" box="[724,905,1779,1805]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="bachmaensis">
|
||
<emphasis id="71E5EAFF716F6210FBC9F927FC7EF8A0" box="[724,905,1779,1805]" italics="true" pageId="53" pageNumber="54">P. bachmaensis</emphasis>
|
||
</taxonomicName>
|
||
also differs from the two latter species by its subparallel, slender lateroventral processes of the aedeagus reaching before basiventral lobes of the ventral lobe of the periandrium, the processes being longer (reaching to middle of lobe), more robust and curved mesad in distal half in
|
||
<taxonomicName id="84914D6E716F6210FB02F88FFD3BF8D8" baseAuthorityName="Che, Zhang & Wang" baseAuthorityYear="2020" box="[543,716,1883,1909]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="menglaensis">
|
||
<emphasis id="71E5EAFF716F6210FB02F88FFD3BF8D8" box="[543,716,1883,1909]" italics="true" pageId="53" pageNumber="54">P. menglaensis</emphasis>
|
||
</taxonomicName>
|
||
, shorter and overlapping ventrally in
|
||
<taxonomicName id="84914D6E716F6210FD9AF88FFAFFF8D8" box="[1159,1288,1883,1909]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="ruilensis">
|
||
<emphasis id="71E5EAFF716F6210FD9AF88FFAFFF8D8" box="[1159,1288,1883,1909]" italics="true" pageId="53" pageNumber="54">P. ruilensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0B8B6566716F6210F9A0F84FFB35F849" box="[189,1218,1948,2020]" pageId="53" pageNumber="54" type="etymology">
|
||
<paragraph id="432E36ED716F6210F9A0F84FFEB3F81B" blockId="53.[189,324,1948,1974]" box="[189,324,1948,1974]" pageId="53" pageNumber="54">
|
||
<heading id="18668181716F6210F9A0F84FFEB3F81B" bold="true" box="[189,324,1948,1974]" fontSize="11" level="3" pageId="53" pageNumber="54" reason="3">
|
||
<emphasis id="71E5EAFF716F6210F9A0F84FFEB3F81B" bold="true" box="[189,324,1948,1974]" pageId="53" pageNumber="54">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="432E36ED716F6210F9A0F819FB35F849" blockId="53.[189,1218,1994,2020]" box="[189,1218,1994,2020]" pageId="53" pageNumber="54">The species epithet refers to Bach Ma National Park, where the species was discovered.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0B8B6566716C6213F9A0FEDAFAD7FDE9" pageId="54" pageNumber="55" type="materials_examined">
|
||
<paragraph id="432E36ED716C6213F9A0FEDAFE9BFE8F" blockId="54.[189,364,264,291]" box="[189,364,264,291]" pageId="54" pageNumber="55">
|
||
<heading id="18668181716C6213F9A0FEDAFE9BFE8F" bold="true" box="[189,364,264,291]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
|
||
<emphasis id="71E5EAFF716C6213F9A0FEDAFE9BFE8F" bold="true" box="[189,364,264,291]" pageId="54" pageNumber="55">Type material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="432E36ED716C6213F9F7FEE4FEABFEFC" blockId="54.[189,1399,311,443]" box="[234,348,311,337]" pageId="54" pageNumber="55">
|
||
<emphasis id="71E5EAFF716C6213F9F7FEE4FEABFEFC" bold="true" box="[234,348,311,337]" pageId="54" pageNumber="55">
|
||
<typeStatus id="9C2A884F716C6213F9F7FEE4FEABFEFC" box="[234,348,311,337]" pageId="54" pageNumber="55" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="432E36ED716C6213F9A0FE88FE4FFE16" blockId="54.[189,1399,311,443]" pageId="54" pageNumber="55">
|
||
<materialsCitation id="F3F93CB0716C6213F9A0FE88FE4FFE16" collectingDate="2011-07-15" collectingDateMax="2011-07-16" collectingDateMin="2011-07-15" collectionCode="RBINS" collectorName="J. Constant & J. Bresseel" country="Vietnam" elevation="1350" latitude="16.2" location="Bach Ma National Park" longLatPrecision="1282" longitude="107.86667" pageId="54" pageNumber="55" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<collectingCountry id="3B86767D716C6213F9A0FE88FEB3FED8" box="[189,324,347,373]" name="Vietnam" pageId="54" pageNumber="55">VIETNAM</collectingCountry>
|
||
•
|
||
<specimenCount id="5597FD64716C6213F843FE88FE83FED8" box="[350,372,347,373]" count="1" pageId="54" pageNumber="55" type="male">♂</specimenCount>
|
||
(
|
||
<figureCitation id="DBAA2A68716C6213F899FE89FDF9FED8" box="[388,526,346,373]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="55.[189,232,1928,1954]" captionStartId-1="56.[189,232,1892,1918]" captionStartId-2="57.[189,232,1892,1918]" captionTargetBox-0="[221,1374,273,1866]" captionTargetBox-1="[203,1384,453,1847]" captionTargetBox-2="[258,1331,285,1845]" captionTargetId-0="figure-11@55.[211,1377,265,1885]" captionTargetId-1="figure-63@56.[203,1384,442,1850]" captionTargetId-2="figure-11@57.[257,1331,265,1845]" captionTargetPageId-0="55" captionTargetPageId-1="56" captionTargetPageId-2="57" captionText-0="Fig. 32. Pusulissus bachmaensis sp. nov., dissected holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Right hind leg, ventral view. E. Metatarsus and apex of metatibia, ventral view." captionText-1="Fig. 33. Pusulissus bachmaensis sp. nov. A–E. Holotype ♂ (RBINS). A. Head and thorax, dorsal view. B. Frons, perpendicular view. C. Head and thorax, lateral view. D. Head and thorax, anterolateral view. E. Right hind wing. F. Paratype ♀ (VNMN), terminalia, ventral view. Abbreviations: see Material and methods." captionText-2="Fig. 34. Pusulissus bachmaensis sp. nov., holotype, ♂ (RBINS). A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E–L. Aedeagus. E. Posteroventral view. F. Left lateral view. G. Left laterodorsal view. H. Anterodorsal view. I. Dorsal view. J. Ventral view. K. Anteroventral view. Abbreviations: see Material and methods." figureDoi-0="http://doi.org/10.5281/zenodo.14508802" figureDoi-1="http://doi.org/10.5281/zenodo.14508804" figureDoi-2="http://doi.org/10.5281/zenodo.14508806" httpUri-0="https://zenodo.org/record/14508802/files/figure.png" httpUri-1="https://zenodo.org/record/14508804/files/figure.png" httpUri-2="https://zenodo.org/record/14508806/files/figure.png" pageId="54" pageNumber="55">Figs 32–34</figureCitation>
|
||
); [Thừa Thiên-Huế Province],
|
||
<location id="464E6036716C6213FA64FE89FB65FED9" LSID="urn:lsid:plazi:treatment:CB3887FB716F621EFB4AFC14FB9DFC79:464E6036716C6213FA64FE89FB65FED9" box="[889,1170,346,373]" country="Vietnam" latitude="16.2" longLatPrecision="1282" longitude="107.86667" name="Bach Ma National Park" pageId="54" pageNumber="55">Bach Ma National Park</location>
|
||
, summit;
|
||
<geoCoordinate id="26A5502A716C6213FC18FE8AFA85FED8" box="[1285,1394,345,373]" degrees="16" direction="north" minutes="12" orientation="latitude" pageId="54" pageNumber="55" precision="925" value="16.2">16°12′ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="26A5502A716C6213F9A0FEAFFEC3FE35" box="[189,308,380,408]" degrees="107" direction="east" minutes="52" orientation="longitude" pageId="54" pageNumber="55" precision="925" value="107.86667">107°52′ E</geoCoordinate>
|
||
; [
|
||
<elevation id="C8BCD1DE716C6213F85BFEADFDDEFE35" box="[326,553,382,408]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="54" pageNumber="55" unit="m" value="1350.0" valueMax="1400.0" valueMin="1300.0">
|
||
<quantity id="84699B08716C6213F85BFEADFE1DFE35" box="[326,490,382,408]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="54" pageNumber="55" unit="m" value="1350.0" valueMax="1400.0" valueMin="1300.0">1300–1400 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
];
|
||
<date id="372F102D716C6213FB22FEADFD0BFE35" box="[575,764,382,409]" pageId="54" pageNumber="55" value="2011-07-15" valueMax="2011-07-16" valueMin="2011-07-15">
|
||
<collectingDate id="276BE9C5716C6213FB22FEADFD0BFE35" box="[575,764,382,409]" pageId="54" pageNumber="55" value="2011-07-15" valueMax="2011-07-16" valueMin="2011-07-15">15–16 Jul. 2011</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="EE64533B716C6213FA1AFEADFC7FFE35" box="[775,904,382,408]" pageId="54" pageNumber="55">J. Constant</collectorName>
|
||
and
|
||
<collectorName id="EE64533B716C6213FAA2FEADFBCBFE35" box="[959,1084,382,408]" pageId="54" pageNumber="55">J. Bresseel</collectorName>
|
||
leg.; day [time] collecting; I.G.: 31.933;
|
||
<collectionCode id="2580AE28716C6213F847FE72FE43FE16" box="[346,436,417,443]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="54" pageNumber="55" type="Museum">RBINS</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="432E36ED716C6213F9F7FE30FEABFE50" blockId="54.[189,1399,483,580]" box="[234,348,483,509]" pageId="54" pageNumber="55">
|
||
<emphasis id="71E5EAFF716C6213F9F7FE30FEABFE50" bold="true" box="[234,348,483,509]" pageId="54" pageNumber="55">
|
||
<typeStatus id="9C2A884F716C6213F9F7FE30FEABFE50" box="[234,348,483,509]" pageId="54" pageNumber="55" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="432E36ED716C6213F9A0FDD5FAD7FDE9" blockId="54.[189,1399,483,580]" pageId="54" pageNumber="55">
|
||
<materialsCitation id="F3F93CB0716C6213F9A0FDD5FAEAFDE9" collectingDate="2023-05-11" collectingDateMax="2023-05-21" collectingDateMin="2023-05-11" collectionCode="VNMN" collectorName="J. Constant & L. Semeraro" country="Vietnam" county="Bach Ma National Park" elevation="1350" latitude="16.188332" location="Th" longLatPrecision="21" longitude="107.84889" municipality="Province" pageId="54" pageNumber="55" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<collectingCountry id="3B86767D716C6213F9A0FDD5FEB3FD8D" box="[189,324,518,544]" name="Vietnam" pageId="54" pageNumber="55">VIETNAM</collectingCountry>
|
||
•
|
||
<specimenCount id="5597FD64716C6213F842FDD5FE7CFD8C" box="[351,395,518,545]" count="1" pageId="54" pageNumber="55" type="female">1 ♀</specimenCount>
|
||
(
|
||
<figureCitation id="DBAA2A68716C6213F881FDD5FE02FD8C" box="[412,501,518,545]" captionStart="Fig" captionStartId="58.[189,232,1963,1989]" captionTargetBox="[189,1398,1110,1917]" captionTargetId="figure-331@58.[189,1398,1110,1917]" captionTargetPageId="58" captionText="Fig. 35. Pusulissus bachmaensis sp. nov., paratype, ♀ (VNMN) from Bach Ma National Park, summit, 21 May 2023. A–D. Live specimen (photographed in cage)." figureDoi="http://doi.org/10.5281/zenodo.14508808" httpUri="https://zenodo.org/record/14508808/files/figure.png" pageId="54" pageNumber="55">Fig. 35</figureCitation>
|
||
);
|
||
<location id="464E6036716C6213FB16FDD5FDDCFD8D" LSID="urn:lsid:plazi:treatment:CB3887FB716F621EFB4AFC14FB9DFC79:464E6036716C6213FB16FDD5FDDCFD8D" box="[523,555,518,544]" country="Vietnam" county="Bach Ma National Park" latitude="16.188332" longLatPrecision="21" longitude="107.84889" municipality="Province" name="Th" pageId="54" pageNumber="55">Th</location>
|
||
ừa Thiên-Huế
|
||
<collectingMunicipality id="A34AAC97716C6213FBCAFDD5FCB5FD8D" box="[727,834,518,544]" pageId="54" pageNumber="55">Province</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="AA4F4E61716C6213FA53FDD5FB9FFD8D" box="[846,1128,518,545]" pageId="54" pageNumber="55">Bach Ma National Park</collectingCounty>
|
||
, summit;
|
||
<geoCoordinate id="26A5502A716C6213FDC0FDD7FA85FD8C" box="[1245,1394,516,545]" degrees="16" direction="north" minutes="11" orientation="latitude" pageId="54" pageNumber="55" precision="15" seconds="18" value="16.188332">16°11′18″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="26A5502A716C6213F9A0FDFBFEA9FDE9" box="[189,350,552,580]" degrees="107" direction="east" minutes="50" orientation="longitude" pageId="54" pageNumber="55" precision="15" seconds="56" value="107.84889">107°50′56″ E</geoCoordinate>
|
||
;
|
||
<elevation id="C8BCD1DE716C6213F876FDF9FDBAFDEE" box="[363,589,553,580]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="54" pageNumber="55" unit="m" value="1350.0" valueMax="1400.0" valueMin="1300.0">
|
||
<quantity id="84699B08716C6213F876FDF9FDFAFDE9" box="[363,525,554,580]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="54" pageNumber="55" unit="m" value="1350.0" valueMax="1400.0" valueMin="1300.0">1300–1400 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<date id="372F102D716C6213FB47FDF9FCD2FDE9" box="[602,805,554,580]" pageId="54" pageNumber="55" value="2023-05-11" valueMax="2023-05-21" valueMin="2023-05-11">
|
||
<collectingDate id="276BE9C5716C6213FB47FDF9FCD2FDE9" box="[602,805,554,580]" pageId="54" pageNumber="55" value="2023-05-11" valueMax="2023-05-21" valueMin="2023-05-11">11–21 May 2023</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="EE64533B716C6213FA2CFDF9FC44FDE9" box="[817,947,554,580]" pageId="54" pageNumber="55">J. Constant</collectorName>
|
||
and
|
||
<collectorName id="EE64533B716C6213FAF0FDF9FB8AFDE9" box="[1005,1149,554,580]" pageId="54" pageNumber="55">L. Semeraro</collectorName>
|
||
leg.;
|
||
<collectionCode id="2580AE28716C6213FDA2FDF9FAEAFDE9" box="[1215,1309,554,580]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="54" pageNumber="55">VNMN</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0B8B6566716C621EF9A0FDB8FDFAFDCC" lastPageId="59" lastPageNumber="60" pageId="54" pageNumber="55" type="description">
|
||
<paragraph id="432E36ED716C6213F9A0FDB8FEB9FD28" blockId="54.[189,334,619,645]" box="[189,334,619,645]" pageId="54" pageNumber="55">
|
||
<heading id="18668181716C6213F9A0FDB8FEB9FD28" bold="true" box="[189,334,619,645]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
|
||
<emphasis id="71E5EAFF716C6213F9A0FDB8FEB9FD28" bold="true" box="[189,334,619,645]" pageId="54" pageNumber="55">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="432E36ED716C6213F9A0FD49FD17FD7A" blockId="54.[189,1399,666,728]" pageId="54" pageNumber="55">
|
||
<smallCapsWord id="45C8A031716C6213F9A0FD49FE99FD1F" baselines="685,685" box="[189,366,666,692]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="54" pageNumber="55">MEASUREMENTS</smallCapsWord>
|
||
<smallCapsWord id="45C8A031716C6213F86BFD4CFE54FD1F" baselines="685" box="[374,419,671,690]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="54" pageNumber="55">AND</smallCapsWord>
|
||
<smallCapsWord id="45C8A031716C6213F8B7FD4CFE04FD1F" baselines="685" box="[426,499,671,690]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="ratios" pageId="54" pageNumber="55">RATIOS</smallCapsWord>
|
||
. LT:
|
||
<specimenCount id="5597FD64716C6213FB2CFD49FDB0FD19" box="[561,583,666,692]" count="1" pageId="54" pageNumber="55" type="male">♂</specimenCount>
|
||
(n = 1): 7.0 mm,
|
||
<specimenCount id="5597FD64716C6213FA0AFD49FCDAFD19" box="[791,813,666,692]" count="1" pageId="54" pageNumber="55" type="female">♀</specimenCount>
|
||
(n = 1):
|
||
<quantity id="84699B08716C6213FA89FD49FC06FD19" box="[916,1009,666,692]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="54" pageNumber="55" unit="mm" value="7.8">7.8 mm</quantity>
|
||
; LT/BB = 1.48; LTg/BTg = 2.08; LW/BW = 1.43; BV/LV = 2.35; LF/BF = 1.19.
|
||
</paragraph>
|
||
<paragraph id="432E36ED716C6213F9A0FD2CFE01FAAA" blockId="54.[187,1401,766,1287]" pageId="54" pageNumber="55">
|
||
<smallCapsWord id="45C8A031716C6213F9A0FD2CFF0BFCBA" baselines="786,786" box="[189,252,767,793]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="54" pageNumber="55">HEAD</smallCapsWord>
|
||
(
|
||
<figureCitation id="DBAA2A68716C6213F813FD2DFE68FCB4" box="[270,415,766,793]" captionStart="Fig" captionStartId="56.[189,232,1892,1918]" captionTargetBox="[203,1384,453,1847]" captionTargetId="figure-63@56.[203,1384,442,1850]" captionTargetPageId="56" captionText="Fig. 33. Pusulissus bachmaensis sp. nov. A–E. Holotype ♂ (RBINS). A. Head and thorax, dorsal view. B. Frons, perpendicular view. C. Head and thorax, lateral view. D. Head and thorax, anterolateral view. E. Right hind wing. F. Paratype ♀ (VNMN), terminalia, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.14508804" httpUri="https://zenodo.org/record/14508804/files/figure.png" pageId="54" pageNumber="55">Fig. 33A–D</figureCitation>
|
||
). Vertex variegated brown with anterior angles black, distinctly broader than long in midline and with lateral margins longer than midline, with all margins carinate; anterior margin nearly straight with median carina of frons distinct in middle, posterior margin deeply concave, angularly rounded, and lateral ones subparallel; disc shallowly excavate with obsolete median carina. Side of head yellowish brown with black marking at anterodorsal angle extending on a distance along anterior margin, darker area between anteroventral portion of eye and anterior margin, followed by pale yellowish area extending to base of antenna and anterior margin of gena; pale yellowish marking along clypeal suture; anteroventral angle moderately projecting anteriorly in a small round lobe. Frons elongate and generally smooth, variegated brown with dorsal angles blackish, distinct yellowish spot in middle on each side of median carina and more or less disctinct yellowish band along frontoclypeal suture; median carina extending from dorsal margin down nearly to rounded frontoclypeal suture, brown on disc; no distinct row of tubercles along dorsal and lateral margins; dorsal margin distinctly concave, lateral margins distinctly sinuate. Clypeus variegated dark brown, weakly elevated medially. Labium brown with last segment longer than broad, and shorter than penultimate. Scape short, ring-shaped, yellowish; pedicel bulbous, yellowish brown.
|
||
</paragraph>
|
||
<paragraph id="432E36ED716C6213F9A0FAFCFBE7F92A" blockId="54.[188,1400,1327,1671]" pageId="54" pageNumber="55">
|
||
<smallCapsWord id="45C8A031716C6213F9A0FAFCFEEEFAEA" baselines="1346,1346" box="[189,281,1327,1353]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="54" pageNumber="55">THORAX</smallCapsWord>
|
||
(
|
||
<figureCitation id="DBAA2A68716C6213F834FAFCFE2BFAE4" box="[297,476,1327,1353]" captionStart="Fig" captionStartId="56.[189,232,1892,1918]" captionTargetBox="[203,1384,453,1847]" captionTargetId="figure-63@56.[203,1384,442,1850]" captionTargetPageId="56" captionText="Fig. 33. Pusulissus bachmaensis sp. nov. A–E. Holotype ♂ (RBINS). A. Head and thorax, dorsal view. B. Frons, perpendicular view. C. Head and thorax, lateral view. D. Head and thorax, anterolateral view. E. Right hind wing. F. Paratype ♀ (VNMN), terminalia, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.14508804" httpUri="https://zenodo.org/record/14508804/files/figure.png" pageId="54" pageNumber="55">Fig. 33A, C–D</figureCitation>
|
||
). Pronotum variegated yellowish brown; about 0.6 times as long as mesonotum in midline; anterior margin carinate, strongly, angularly protruding anteriorly between eyes, with carinae directed obliquely posteriorly, not reaching hind margin of pronotum; posterior margin almost straight, raised; no median carina but with impressed point on each side of median line; lateral portion behind eye very narrow, laminate; blunt, pale yellowish tubercles scattered on disc with row following anterior margin, and on outer margin of paranotal lobes; paranotal lobes (lateral view) broad, brown in outer portion, pale yellowish in inner portion and with large black marking behind lower portion of eye, and with posteroventral angle straight. Mesonotum variegated yellowish brown with longitudinal carinae obsolete but marked by dark brown lines; smooth, slightly depressed before scutellum; scutellum paler; nearly complete transverse carina along anterior margin. Tegulae brown.
|
||
</paragraph>
|
||
<paragraph id="432E36ED716C621FF9A0F97CFDA7FEC7" blockId="54.[188,1399,1710,2026]" lastBlockId="58.[189,1399,265,362]" lastPageId="58" lastPageNumber="59" pageId="54" pageNumber="55">
|
||
<smallCapsWord id="45C8A031716C6213F9A0F97CFED5F96A" baselines="1730,1730" box="[189,290,1711,1737]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Tegmina" pageId="54" pageNumber="55">TEGMINA</smallCapsWord>
|
||
(
|
||
<figureCitation id="DBAA2A68716C6213F82FF97DFE34F965" box="[306,451,1710,1737]" captionStart="Fig" captionStartId="55.[189,232,1928,1954]" captionTargetBox="[221,1374,273,1866]" captionTargetId="figure-11@55.[211,1377,265,1885]" captionTargetPageId="55" captionText="Fig. 32. Pusulissus bachmaensis sp. nov., dissected holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Right hind leg, ventral view. E. Metatarsus and apex of metatibia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14508802" httpUri="https://zenodo.org/record/14508802/files/figure.png" pageId="54" pageNumber="55">Figs 32A–C</figureCitation>
|
||
,
|
||
<figureCitation id="DBAA2A68716C6213F8D2F97DFDDFF965" box="[463,552,1710,1737]" captionStart="Fig" captionStartId="58.[189,232,1963,1989]" captionTargetBox="[189,1398,1110,1917]" captionTargetId="figure-331@58.[189,1398,1110,1917]" captionTargetPageId="58" captionText="Fig. 35. Pusulissus bachmaensis sp. nov., paratype, ♀ (VNMN) from Bach Ma National Park, summit, 21 May 2023. A–D. Live specimen (photographed in cage)." figureDoi="http://doi.org/10.5281/zenodo.14508808" httpUri="https://zenodo.org/record/14508808/files/figure.png" pageId="54" pageNumber="55">35A–C</figureCitation>
|
||
). Variegated yellowish brown (greener in live specimens); subcoriaceous with longitudinal veins yellow brown, elevated and with a dense reticulum of pale yellow veinlets; shape elongate and convex with sides broadly rounded, widest at basal ⅓, about 2.1 times as long as broad; narrowly rounded apically. Postclaval margin weakly rounded on distal half and weakly notched at apex of clavus. Clavus closed, reaching slightly over midlength of tegmen. Venation: ScP+R forking close to base after rather short common stem, ScP+RA and RP running more or less parallel to costal margin and not forked; MP forked rather close to base after common stem, resulting veins both forked again further; CuA simple, sinuate, more or less parallel to claval joint, then to sutural margin and merging with latter before apex of tegmen; Pcu and A
|
||
<subScript id="DF1534A8716C6213FB59F80FFDBAF847" attach="left" box="[580,589,2012,2026]" fontSize="6" pageId="54" pageNumber="55">1</subScript>
|
||
fused at apical ⅓ of clavus, resulting Pcu+A
|
||
<subScript id="DF1534A8716C6213FD52F80FFBAFF847" attach="left" box="[1103,1112,2012,2026]" fontSize="6" pageId="54" pageNumber="55">1</subScript>
|
||
reaching apex of clavus.
|
||
<smallCapsWord id="45C8A0317162621DF9A0FEDAFF00FE8F" baselines="284,285" box="[189,247,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Hind" pageId="56" pageNumber="57">HIND</smallCapsWord>
|
||
<smallCapsWord id="45C8A0317162621DF81FFEDDFEB1FE8C" baselines="284" box="[258,326,270,289]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="wings" pageId="56" pageNumber="57">WINGS</smallCapsWord>
|
||
(
|
||
<figureCitation id="DBAA2A687162621DF847FEDAFE31FE8E" box="[346,454,265,291]" captionStart="Fig" captionStartId="56.[189,232,1892,1918]" captionTargetBox="[203,1384,453,1847]" captionTargetId="figure-63@56.[203,1384,442,1850]" captionTargetPageId="56" captionText="Fig. 33. Pusulissus bachmaensis sp. nov. A–E. Holotype ♂ (RBINS). A. Head and thorax, dorsal view. B. Frons, perpendicular view. C. Head and thorax, lateral view. D. Head and thorax, anterolateral view. E. Right hind wing. F. Paratype ♀ (VNMN), terminalia, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.14508804" httpUri="https://zenodo.org/record/14508804/files/figure.png" pageId="56" pageNumber="57">Fig. 33E</figureCitation>
|
||
). Brown with large basicostal portion of lobe Sc-R-MP-CuA and base of lobe CuP-Pcu-A
|
||
<subScript id="DF1534A87162621DF859FEECFEBAFEE0" attach="left" box="[324,333,319,333]" fontSize="6" pageId="56" pageNumber="57">1</subScript>
|
||
yellow-brown; venation darker than corresponding background; wing broader than tegmen and deeply bilobed at CuP; costal margin moderately sinuate; CuP-Pcu-A1 lobe broadly rounded along postclaval margin and about 1.5 times as wide as Sc-R-MP-CuA lobe; both lobes approximately the same length, angularly rounded at apex; A2 lobe reduced and narrow. Venation: longitudinal veins ScP-R-MP-Cu well distinct; Pcu and A
|
||
</paragraph>
|
||
<caption id="17EE6665716D6212F9A0F85BFDC6F845" ID-DOI="http://doi.org/10.5281/zenodo.14508802" ID-Zenodo-Dep="14508802" httpUri="https://zenodo.org/record/14508802/files/figure.png" pageId="55" pageNumber="56" startId="55.[189,232,1928,1954]" targetBox="[221,1374,273,1866]" targetPageId="55">
|
||
<paragraph id="432E36ED716D6212F9A0F85BFDC6F845" blockId="55.[189,1399,1928,2025]" pageId="55" pageNumber="56">
|
||
<emphasis id="71E5EAFF716D6212F9A0F85BFEE9F80F" bold="true" box="[189,286,1928,1954]" pageId="55" pageNumber="56">Fig. 32.</emphasis>
|
||
<taxonomicName id="84914D6E716D6212F834F85BFDBFF80F" authorityName="Constant & Pham" authorityYear="2024" box="[297,584,1928,1954]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="bachmaensis" status="sp. nov.">
|
||
<emphasis id="71E5EAFF716D6212F834F85BFDBFF80F" box="[297,584,1928,1954]" italics="true" pageId="55" pageNumber="56">Pusulissus bachmaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6AD65784716D6212FB4EF85BFD44F80F" box="[595,691,1928,1954]" pageId="55" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, dissected holotype, ♂ (RBINS).
|
||
<emphasis id="71E5EAFF716D6212FD4DF85BFB92F80F" bold="true" box="[1104,1125,1928,1954]" pageId="55" pageNumber="56">A</emphasis>
|
||
. Habitus, dorsal view.
|
||
<emphasis id="71E5EAFF716D6212F9A0F878FF26F868" bold="true" box="[189,209,1963,1989]" pageId="55" pageNumber="56">B</emphasis>
|
||
. Habitus, ventral view.
|
||
<emphasis id="71E5EAFF716D6212F8ECF878FDF3F868" bold="true" box="[497,516,1963,1989]" pageId="55" pageNumber="56">C</emphasis>
|
||
. Habitus, lateral view.
|
||
<emphasis id="71E5EAFF716D6212FA01F878FCD8F868" bold="true" box="[796,815,1963,1989]" pageId="55" pageNumber="56">D</emphasis>
|
||
. Right hind leg, ventral view.
|
||
<emphasis id="71E5EAFF716D6212FDBDF878FB45F868" bold="true" box="[1184,1202,1963,1989]" pageId="55" pageNumber="56">E</emphasis>
|
||
. Metatarsus and apex of metatibia, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="17EE66657162621DF9A0F8B7FEDFF845" ID-DOI="http://doi.org/10.5281/zenodo.14508804" ID-Zenodo-Dep="14508804" httpUri="https://zenodo.org/record/14508804/files/figure.png" pageId="56" pageNumber="57" startId="56.[189,232,1892,1918]" targetBox="[203,1384,453,1847]" targetPageId="56">
|
||
<paragraph id="432E36ED7162621DF9A0F8B7FEDFF845" blockId="56.[189,1399,1892,2024]" pageId="56" pageNumber="57">
|
||
<emphasis id="71E5EAFF7162621DF9A0F8B7FEEDF8D2" bold="true" box="[189,282,1892,1919]" pageId="56" pageNumber="57">Fig. 33.</emphasis>
|
||
<taxonomicName id="84914D6E7162621DF83CF8B6FDCAF8D2" authorityName="Constant & Pham" authorityYear="2024" box="[289,573,1893,1919]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="bachmaensis" status="sp. nov.">
|
||
<emphasis id="71E5EAFF7162621DF83CF8B6FDCAF8D2" box="[289,573,1893,1919]" italics="true" pageId="56" pageNumber="57">Pusulissus bachmaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6AD657847162621DFB5EF8B6FD69F8D2" box="[579,670,1893,1919]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="71E5EAFF7162621DFBB8F8B7FD2BF8D3" bold="true" box="[677,732,1892,1918]" pageId="56" pageNumber="57">A–E</emphasis>
|
||
. Holotype ♂ (RBINS).
|
||
<emphasis id="71E5EAFF7162621DFAE8F8B7FBFDF8D3" bold="true" box="[1013,1034,1892,1918]" pageId="56" pageNumber="57">A</emphasis>
|
||
. Head and thorax, dorsal view.
|
||
<emphasis id="71E5EAFF7162621DF9A0F85BFF26F80F" bold="true" box="[189,209,1928,1954]" pageId="56" pageNumber="57">B</emphasis>
|
||
. Frons, perpendicular view.
|
||
<emphasis id="71E5EAFF7162621DFB02F85BFDC3F80F" bold="true" box="[543,564,1928,1954]" pageId="56" pageNumber="57">C</emphasis>
|
||
. Head and thorax, lateral view.
|
||
<emphasis id="71E5EAFF7162621DFAB7F85BFC48F80F" bold="true" box="[938,959,1928,1954]" pageId="56" pageNumber="57">D</emphasis>
|
||
. Head and thorax, anterolateral view.
|
||
<emphasis id="71E5EAFF7162621DF9A0F878FF26F868" bold="true" box="[189,209,1963,1989]" pageId="56" pageNumber="57">E</emphasis>
|
||
. Right hind wing.
|
||
<emphasis id="71E5EAFF7162621DF8B2F878FE37F868" bold="true" box="[431,448,1963,1989]" pageId="56" pageNumber="57">F</emphasis>
|
||
. Paratype ♀ (VNMN), terminalia, ventral view. Abbreviations: see Material and methods.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="17EE66657163621CF9A0F8B7FB49F845" ID-DOI="http://doi.org/10.5281/zenodo.14508806" ID-Zenodo-Dep="14508806" httpUri="https://zenodo.org/record/14508806/files/figure.png" pageId="57" pageNumber="58" startId="57.[189,232,1892,1918]" targetBox="[258,1331,285,1845]" targetPageId="57">
|
||
<paragraph id="432E36ED7163621CF9A0F8B7FB49F845" blockId="57.[187,1399,1892,2025]" pageId="57" pageNumber="58">
|
||
<emphasis id="71E5EAFF7163621CF9A0F8B7FED4F8D2" bold="true" box="[189,291,1892,1919]" pageId="57" pageNumber="58">Fig. 34.</emphasis>
|
||
<taxonomicName id="84914D6E7163621CF82EF8B6FDA0F8D2" authorityName="Constant & Pham" authorityYear="2024" box="[307,599,1893,1919]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="bachmaensis" status="sp. nov.">
|
||
<emphasis id="71E5EAFF7163621CF82EF8B6FDA0F8D2" box="[307,599,1893,1919]" italics="true" pageId="57" pageNumber="58">Pusulissus bachmaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6AD657847163621CFB7BF8B6FD3BF8D2" box="[614,716,1893,1919]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂ (RBINS).
|
||
<emphasis id="71E5EAFF7163621CFD18F8B7FBC8F8D3" bold="true" box="[1029,1087,1892,1918]" pageId="57" pageNumber="58">A–D</emphasis>
|
||
. Pygofer, anal tube and gonostyli.
|
||
<emphasis id="71E5EAFF7163621CF825F85BFEBAF80F" bold="true" box="[312,333,1928,1954]" pageId="57" pageNumber="58">A</emphasis>
|
||
. Left lateral view.
|
||
<emphasis id="71E5EAFF7163621CFB34F85BFDCAF80F" bold="true" box="[553,573,1928,1954]" pageId="57" pageNumber="58">B</emphasis>
|
||
. Posterolateral view.
|
||
<emphasis id="71E5EAFF7163621CFA25F85BFCBAF80F" bold="true" box="[824,845,1928,1954]" pageId="57" pageNumber="58">C</emphasis>
|
||
. Caudal view.
|
||
<emphasis id="71E5EAFF7163621CFAE6F85BFBE7F80F" bold="true" box="[1019,1040,1928,1954]" pageId="57" pageNumber="58">D</emphasis>
|
||
. Dorsal view.
|
||
<emphasis id="71E5EAFF7163621CFDA4F85BFB19F80F" bold="true" box="[1209,1262,1928,1954]" pageId="57" pageNumber="58">E–L</emphasis>
|
||
. Aedeagus.
|
||
<emphasis id="71E5EAFF7163621CF9A6F878FF38F868" bold="true" box="[187,207,1963,1989]" pageId="57" pageNumber="58">E</emphasis>
|
||
. Posteroventral view.
|
||
<emphasis id="71E5EAFF7163621CF8C5F878FE1EF868" bold="true" box="[472,489,1963,1989]" pageId="57" pageNumber="58">F</emphasis>
|
||
. Left lateral view.
|
||
<emphasis id="71E5EAFF7163621CFBD7F878FD17F868" bold="true" box="[714,736,1963,1989]" pageId="57" pageNumber="58">G</emphasis>
|
||
. Left laterodorsal view.
|
||
<emphasis id="71E5EAFF7163621CFD1FF878FBEFF868" bold="true" box="[1026,1048,1963,1989]" pageId="57" pageNumber="58">H</emphasis>
|
||
. Anterodorsal view.
|
||
<emphasis id="71E5EAFF7163621CFC10F878FAEFF868" bold="true" box="[1293,1304,1963,1989]" pageId="57" pageNumber="58">I</emphasis>
|
||
. Dorsal view.
|
||
<emphasis id="71E5EAFF7163621CF81EF81DFEE5F845" bold="true" box="[259,274,1998,2024]" pageId="57" pageNumber="58">J</emphasis>
|
||
. Ventral view.
|
||
<emphasis id="71E5EAFF7163621CF8DCF81DFE2FF845" bold="true" box="[449,472,1998,2024]" pageId="57" pageNumber="58">K</emphasis>
|
||
. Anteroventral view. Abbreviations: see Material and methods.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="432E36ED7160621FFB4DFEB0FDAEFEDC" box="[592,601,355,369]" pageId="58" pageNumber="59">1</paragraph>
|
||
<paragraph id="432E36ED7160621FFB7DFE83FB7BFEC7" blockId="58.[189,1399,265,362]" box="[608,1164,336,362]" pageId="58" pageNumber="59">separated; numerous cross-veins; A2 indistinct.</paragraph>
|
||
<paragraph id="432E36ED7160621FF9A0FE41FCDFFDB8" blockId="58.[187,1399,401,533]" pageId="58" pageNumber="59">
|
||
<smallCapsWord id="45C8A0317160621FF9A0FE41FF01FE07" baselines="421,421" box="[189,246,402,428]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="58" pageNumber="59">LEGS</smallCapsWord>
|
||
(
|
||
<figureCitation id="DBAA2A687160621FF818FE42FE65FE06" box="[261,402,401,428]" captionStart="Fig" captionStartId="55.[189,232,1928,1954]" captionTargetBox="[221,1374,273,1866]" captionTargetId="figure-11@55.[211,1377,265,1885]" captionTargetPageId="55" captionText="Fig. 32. Pusulissus bachmaensis sp. nov., dissected holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Right hind leg, ventral view. E. Metatarsus and apex of metatibia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14508802" httpUri="https://zenodo.org/record/14508802/files/figure.png" pageId="58" pageNumber="59">Fig. 32A–E</figureCitation>
|
||
). Moderately elongate and slender, yellowish brown femora darker than corresponding tibiae; apex of femora and base and pro- and mesotibiae yellowish; posterior margin of pro- and mesofemora with row of acute irregular minute teeth. Metatibiae with 2 lateral spines in distal half and 8 apical spines. Metatibiotarsal formula: (2) 8 / 8 / 2.
|
||
</paragraph>
|
||
<paragraph id="432E36ED7160621FF9A0FDEEFBFAFDFA" blockId="58.[189,1037,573,599]" box="[189,1037,573,599]" pageId="58" pageNumber="59">
|
||
<smallCapsWord id="45C8A0317160621FF9A0FDEEFEDBFDF8" baselines="592,592" box="[189,300,573,599]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="58" pageNumber="59">ABDOMEN</smallCapsWord>
|
||
(
|
||
<figureCitation id="DBAA2A687160621FF823FDEEFE5EFDFA" box="[318,425,573,599]" captionStart="Fig" captionStartId="55.[189,232,1928,1954]" captionTargetBox="[221,1374,273,1866]" captionTargetId="figure-11@55.[211,1377,265,1885]" captionTargetPageId="55" captionText="Fig. 32. Pusulissus bachmaensis sp. nov., dissected holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Right hind leg, ventral view. E. Metatarsus and apex of metatibia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14508802" httpUri="https://zenodo.org/record/14508802/files/figure.png" pageId="58" pageNumber="59">Fig. 32B</figureCitation>
|
||
). Yellowish with wide, black brown band in middle.
|
||
</paragraph>
|
||
<paragraph id="432E36ED7160621EF9A0FD53FC20FE75" blockId="58.[187,1400,639,1061]" lastBlockId="59.[189,1399,265,472]" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
|
||
<smallCapsWord id="45C8A0317160621FF9A0FD53FF08FD35" baselines="659,659" box="[189,255,640,666]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="58" pageNumber="59">MALE</smallCapsWord>
|
||
<smallCapsWord id="45C8A0317160621FF818FD56FE71FD35" baselines="659" box="[261,390,645,664]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="terminalia" pageId="58" pageNumber="59">TERMINALIA</smallCapsWord>
|
||
. Pygofer (
|
||
<emphasis id="71E5EAFF7160621FF8E2FD53FDE9FD34" box="[511,542,640,665]" italics="true" pageId="58" pageNumber="59">Py</emphasis>
|
||
–
|
||
<figureCitation id="DBAA2A687160621FFB2AFDACFD34FD34" box="[567,707,639,665]" captionStart="Fig" captionStartId="57.[189,232,1892,1918]" captionTargetBox="[258,1331,285,1845]" captionTargetId="figure-11@57.[257,1331,265,1845]" captionTargetPageId="57" captionText="Fig. 34. Pusulissus bachmaensis sp. nov., holotype, ♂ (RBINS). A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E–L. Aedeagus. E. Posteroventral view. F. Left lateral view. G. Left laterodorsal view. H. Anterodorsal view. I. Dorsal view. J. Ventral view. K. Anteroventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.14508806" httpUri="https://zenodo.org/record/14508806/files/figure.png" pageId="58" pageNumber="59">Fig. 34A–D</figureCitation>
|
||
) about 2.2 times as high as long in lateral view, with anterior margin sinuate and posterior margin more or less straight in middle portion, rounded to posterodorsal angle, and obliquely excavate in ventral ⅓; posterior margin dorsally deeply excavate (U-shaped excavation). Gonostyli (
|
||
<emphasis id="71E5EAFF7160621FF8C5FD38FE1AFCA8" box="[472,493,747,773]" italics="true" pageId="58" pageNumber="59">G</emphasis>
|
||
–
|
||
<figureCitation id="DBAA2A687160621FFB17FD38FD62FCA8" box="[522,661,747,773]" captionStart="Fig" captionStartId="57.[189,232,1892,1918]" captionTargetBox="[258,1331,285,1845]" captionTargetId="figure-11@57.[257,1331,265,1845]" captionTargetPageId="57" captionText="Fig. 34. Pusulissus bachmaensis sp. nov., holotype, ♂ (RBINS). A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E–L. Aedeagus. E. Posteroventral view. F. Left lateral view. G. Left laterodorsal view. H. Anterodorsal view. I. Dorsal view. J. Ventral view. K. Anteroventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.14508806" httpUri="https://zenodo.org/record/14508806/files/figure.png" pageId="58" pageNumber="59">Fig. 34A–C</figureCitation>
|
||
) (in lateral view) strongly elongate (without dorsal capitulum), projecting posteriorly in a distinct elongate lobe rounded apically, and concave; capitulum (
|
||
<emphasis id="71E5EAFF7160621FFC0EFCC3FAD8FC84" box="[1299,1327,784,809]" italics="true" pageId="58" pageNumber="59">ca</emphasis>
|
||
) with neck rather short and wide, less high than head of capitulum, with head massive, falcate in lateral view with apical hook directed cephalad, with strong tooth on inner margin directed mesad, and with lateral laminate projection in dorsal half with outer margin under it strongly concave in caudal view. Aedeagus (
|
||
<figureCitation id="DBAA2A687160621FF9D8FC4CFEA6FC14" box="[197,337,927,953]" captionStart="Fig" captionStartId="57.[189,232,1892,1918]" captionTargetBox="[258,1331,285,1845]" captionTargetId="figure-11@57.[257,1331,265,1845]" captionTargetPageId="57" captionText="Fig. 34. Pusulissus bachmaensis sp. nov., holotype, ♂ (RBINS). A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E–L. Aedeagus. E. Posteroventral view. F. Left lateral view. G. Left laterodorsal view. H. Anterodorsal view. I. Dorsal view. J. Ventral view. K. Anteroventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.14508806" httpUri="https://zenodo.org/record/14508806/files/figure.png" pageId="58" pageNumber="59">Fig. 34E–K</figureCitation>
|
||
) symmetrical, rather strongly curved dorsad (in lateral view), but with distal ¼ more or less straight; pair of elongate lateroventral processes (
|
||
<emphasis id="71E5EAFF7160621FFBE3FC10FCD5FC70" box="[766,802,963,989]" italics="true" pageId="58" pageNumber="59">lvp</emphasis>
|
||
) slightly inflated basally, then constricted, directed cephalad and moderately, evenly upcurved, remaining under ventral lobe of periandrium and reaching to basiventral lobes of periandrium; distal portion tapering and curved mesad; ventral lobe of periandrium (
|
||
<emphasis id="71E5EAFF7161621EF9DAFEDAFF2BFE8E" box="[199,220,265,291]" italics="true" pageId="59" pageNumber="60">vl</emphasis>
|
||
) with pair of longitudinal basiventral lobe-shaped processes, ventral lobe evenly widening posterior to lateroventral processes, then gently tapering to generally rounded apical margin with small acutely rounded projection in middle. Connective (
|
||
<emphasis id="71E5EAFF7161621EFBDCFE81FD2AFEC6" box="[705,733,338,363]" italics="true" pageId="59" pageNumber="60">co</emphasis>
|
||
) elongate, curved, with tectiductus (
|
||
<emphasis id="71E5EAFF7161621EFD8EFE81FB5FFEC6" box="[1171,1192,338,363]" italics="true" pageId="59" pageNumber="60">te</emphasis>
|
||
) well developed. Anal tube (
|
||
<emphasis id="71E5EAFF7161621EF823FEA6FEA8FE22" box="[318,351,373,399]" italics="true" pageId="59" pageNumber="60">An</emphasis>
|
||
–
|
||
<figureCitation id="DBAA2A687161621EF864FEA6FDF2FE22" box="[377,517,373,399]" captionStart="Fig" captionStartId="57.[189,232,1892,1918]" captionTargetBox="[258,1331,285,1845]" captionTargetId="figure-11@57.[257,1331,265,1845]" captionTargetPageId="57" captionText="Fig. 34. Pusulissus bachmaensis sp. nov., holotype, ♂ (RBINS). A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E–L. Aedeagus. E. Posteroventral view. F. Left lateral view. G. Left laterodorsal view. H. Anterodorsal view. I. Dorsal view. J. Ventral view. K. Anteroventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.14508806" httpUri="https://zenodo.org/record/14508806/files/figure.png" pageId="59" pageNumber="60">Fig. 34A–D</figureCitation>
|
||
) dorsoventrally flattened, in dorsal view widely spatulate with distal portion nearly circular, nearly 1.3 times as long in midline as wide, with apical margin shortly truncate in middle, without distinct lateral flaps; anal opening at about basal ⅓.
|
||
</paragraph>
|
||
<caption id="17EE66657160621FF9A0F878FC8CF845" ID-DOI="http://doi.org/10.5281/zenodo.14508808" ID-Zenodo-Dep="14508808" httpUri="https://zenodo.org/record/14508808/files/figure.png" pageId="58" pageNumber="59" startId="58.[189,232,1963,1989]" targetBox="[189,1398,1110,1917]" targetPageId="58">
|
||
<paragraph id="432E36ED7160621FF9A0F878FC8CF845" blockId="58.[189,1398,1963,2025]" pageId="58" pageNumber="59">
|
||
<emphasis id="71E5EAFF7160621FF9A0F878FEECF868" bold="true" box="[189,283,1963,1989]" pageId="58" pageNumber="59">Fig. 35.</emphasis>
|
||
<taxonomicName id="84914D6E7160621FF83FF878FDCAF868" authorityName="Constant & Pham" authorityYear="2024" box="[290,573,1963,1989]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="bachmaensis" status="sp. nov.">
|
||
<emphasis id="71E5EAFF7160621FF83FF878FDCAF868" box="[290,573,1963,1989]" italics="true" pageId="58" pageNumber="59">Pusulissus bachmaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6AD657847160621FFB59F87FFD56F86B" box="[580,673,1964,1990]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype, ♀ (VNMN) from Bach Ma National Park, summit, 21 May 2023.
|
||
<emphasis id="71E5EAFF7160621FF875F81DFE56F845" bold="true" box="[360,417,1998,2024]" pageId="58" pageNumber="59">A–D</emphasis>
|
||
. Live specimen (photographed in cage).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="432E36ED7161621EF9A0FDD3FDFAFDCC" blockId="59.[188,1398,511,609]" pageId="59" pageNumber="60">
|
||
<smallCapsWord id="45C8A0317161621EF9A0FDD3FEE3FDB5" baselines="531,531" box="[189,276,512,538]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Female" pageId="59" pageNumber="60">FEMALE</smallCapsWord>
|
||
<smallCapsWord id="45C8A0317161621EF806FDD6FE6BFDB5" baselines="531" box="[283,412,517,536]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="terminalia" pageId="59" pageNumber="60">TERMINALIA</smallCapsWord>
|
||
. Hind margin of sternum VII (
|
||
<emphasis id="71E5EAFF7161621EFA1CFDD3FCB4FDB4" box="[769,835,512,538]" italics="true" pageId="59" pageNumber="60">St VII</emphasis>
|
||
–
|
||
<figureCitation id="DBAA2A687161621EFA42FE2CFC34FDB4" box="[863,963,511,537]" captionStart="Fig" captionStartId="56.[189,232,1892,1918]" captionTargetBox="[203,1384,453,1847]" captionTargetId="figure-63@56.[203,1384,442,1850]" captionTargetPageId="56" captionText="Fig. 33. Pusulissus bachmaensis sp. nov. A–E. Holotype ♂ (RBINS). A. Head and thorax, dorsal view. B. Frons, perpendicular view. C. Head and thorax, lateral view. D. Head and thorax, anterolateral view. E. Right hind wing. F. Paratype ♀ (VNMN), terminalia, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.14508804" httpUri="https://zenodo.org/record/14508804/files/figure.png" pageId="59" pageNumber="60">Fig. 33F</figureCitation>
|
||
) concave, with middle portion nearly straight; small digitiform process directed posterad, derived from middle of sternum VII and slightly surpassing posterior margin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0B8B65667161621EF9A0FD5AFCB8FCCE" pageId="59" pageNumber="60" type="biology_ecology">
|
||
<paragraph id="432E36ED7161621EF9A0FD5AFEEBFD0E" blockId="59.[189,284,649,675]" box="[189,284,649,675]" pageId="59" pageNumber="60">
|
||
<heading id="186681817161621EF9A0FD5AFEEBFD0E" bold="true" box="[189,284,649,675]" fontSize="11" level="3" pageId="59" pageNumber="60" reason="3">
|
||
<emphasis id="71E5EAFF7161621EF9A0FD5AFEEBFD0E" bold="true" box="[189,284,649,675]" pageId="59" pageNumber="60">Biology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="432E36ED7161621EF9A0FD6AFCB8FCCE" blockId="59.[188,1401,697,867]" pageId="59" pageNumber="60">
|
||
<taxonomicName id="84914D6E7161621EF9A0FD6AFE20FD7E" authorityName="Constant & Pham" authorityYear="2024" box="[189,471,697,723]" class="Insecta" family="Issidae" genus="Pusulissus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="bachmaensis" status="sp. nov.">
|
||
<emphasis id="71E5EAFF7161621EF9A0FD6AFE20FD7E" box="[189,471,697,723]" italics="true" pageId="59" pageNumber="60">Pusulissus bachmaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6AD657847161621EF8C0FD6AFDC1FD7E" box="[477,566,697,723]" pageId="59" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
|
||
was collected only twice:
|
||
<specimenCount id="5597FD647161621EFA74FD6AFC25FD7E" box="[873,978,697,723]" count="1" pageId="59" pageNumber="60" type="male">one male</specimenCount>
|
||
in
|
||
<date id="372F102D7161621EFAE8FD6AFB90FD7E" box="[1013,1127,697,723]" pageId="59" pageNumber="60" value="2011-07">July 2011</date>
|
||
and
|
||
<specimenCount id="5597FD647161621EFD80FD6AFAE9FD7E" box="[1181,1310,697,723]" count="1" pageId="59" pageNumber="60" type="female">one female</specimenCount>
|
||
in
|
||
<date id="372F102D7161621EFC5CFD6AFF0EFD5A" pageId="59" pageNumber="60" value="2023-05">May 2023</date>
|
||
, both in montane evergreen tropical forest, at about
|
||
<quantity id="84699B087161621EFA49FD0EFC01FD5A" box="[852,1014,733,759]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="59" pageNumber="60" unit="m" value="1350.0" valueMax="1400.0" valueMin="1300.0">1300–1400 m</quantity>
|
||
a.s.l., near the summit (
|
||
<figureCitation id="DBAA2A687161621EFC16FD0EFA9FFD5A" box="[1291,1384,733,759]" captionStart="Fig" captionStartId="6.[189,232,1998,2024]" captionTargetBox="[189,1398,729,1950]" captionTargetId="figure-138@6.[189,1398,729,1950]" captionTargetPageId="6" captionText="Fig. 1. Map of the sampled localities in Thừa Thiên-Huế Province, Vietnam." figureDoi="http://doi.org/10.5281/zenodo.14508732" httpUri="https://zenodo.org/record/14508732/files/figure.png" pageId="59" pageNumber="60">Fig. 1D</figureCitation>
|
||
). The specimens were collected on tree trunks densely covered in lichens and mosses (the female was photographed in a cage –
|
||
<figureCitation id="DBAA2A687161621EF8F5FCF6FDC9FCED" box="[488,574,805,832]" captionStart="Fig" captionStartId="58.[189,232,1963,1989]" captionTargetBox="[189,1398,1110,1917]" captionTargetId="figure-331@58.[189,1398,1110,1917]" captionTargetPageId="58" captionText="Fig. 35. Pusulissus bachmaensis sp. nov., paratype, ♀ (VNMN) from Bach Ma National Park, summit, 21 May 2023. A–D. Live specimen (photographed in cage)." figureDoi="http://doi.org/10.5281/zenodo.14508808" httpUri="https://zenodo.org/record/14508808/files/figure.png" pageId="59" pageNumber="60">Fig. 35</figureCitation>
|
||
). When disturbed, they tend to walk away and hide on the opposite side of the trunk or branch, and jump away as a last solution.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0B8B65667161621EF9A0FC58FB9DFC79" box="[189,1130,907,983]" pageId="59" pageNumber="60" type="distribution">
|
||
<paragraph id="432E36ED7161621EF9A0FC58FEA1FC08" blockId="59.[189,342,907,933]" box="[189,342,907,933]" pageId="59" pageNumber="60">
|
||
<heading id="186681817161621EF9A0FC58FEA1FC08" bold="true" box="[189,342,907,933]" fontSize="11" level="3" pageId="59" pageNumber="60" reason="3">
|
||
<emphasis id="71E5EAFF7161621EF9A0FC58FEA1FC08" bold="true" box="[189,342,907,933]" pageId="59" pageNumber="60">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="432E36ED7161621EF9A0FC69FB9DFC79" blockId="59.[189,1130,954,983]" box="[189,1130,954,983]" pageId="59" pageNumber="60">
|
||
<collectingCountry id="3B86767D7161621EF9A0FC69FED2FC79" box="[189,293,954,980]" name="Vietnam" pageId="59" pageNumber="60">Vietnam</collectingCountry>
|
||
: Thừa Thiên-Huế Province, summit of Bach Ma National Park (
|
||
<figureCitation id="DBAA2A687161621EFD09FC69FBAEFC79" box="[1044,1113,954,980]" captionStart="Fig" captionStartId="6.[189,232,1998,2024]" captionTargetBox="[189,1398,729,1950]" captionTargetId="figure-138@6.[189,1398,729,1950]" captionTargetPageId="6" captionText="Fig. 1. Map of the sampled localities in Thừa Thiên-Huế Province, Vietnam." figureDoi="http://doi.org/10.5281/zenodo.14508732" httpUri="https://zenodo.org/record/14508732/files/figure.png" pageId="59" pageNumber="60">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |