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<mods:title id="A4D9738707C145544A51642D816DE8BF">Annotated catalogue of the types of Triphoridae (Mollusca, Gastropoda) in the Natural History Museum of the United Kingdom, London</mods:title>
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<mods:namePart id="545160B320630FE46D919A5D7362B698">Albano, Paolo G.</mods:namePart>
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<mods:namePart id="E976805F9EF9EE1CEF20B78BEDD081F7">Bakker, Piet A. J.</mods:namePart>
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<treatment id="C7527A3CF74BA32FDB288133D2983253" ID-GBIF-Taxon="189923250" LSID="urn:lsid:plazi:treatment:C7527A3CF74BA32FDB288133D2983253" httpUri="http://treatment.plazi.org/id/C7527A3CF74BA32FDB288133D2983253" lastPageNumber="161" pageId="0" pageNumber="161" scope_class="Gastropoda" scope_family="Triphoridae" scope_phylum="Mollusca">
<subSubSection id="7788B4B4F497BD4E6107112A7B9C8FF2" pageId="0" pageNumber="161" type="nomenclature">
<paragraph id="86F3D293F54F0AC65FC7AA221972A000" pageId="0" pageNumber="161">
<taxonomicName id="451480627CACD152562716AC80A26665" ID-CoL="5RWQK" LSID="FD7C6302-4638-5BD4-B97B-2AB21DAF65C6" authority="Verco, 1909" authorityName="Verco" authorityYear="1909" class="Gastropoda" family="Triphoridae" genus="Triphora" higherTaxonomySource="treatment-meta" infraspecific-rank="var." kingdom="Animalia" lsidName="Triphora albovittata subsp. var. var. mamillata" order="Ptenoglossa" pageId="0" pageNumber="161" phylum="Mollusca" rank="variety" species="albovittata" subSpecies="var." variety="mamillata">Triphora albovittata var. mamillata Verco, 1909</taxonomicName>
</paragraph>
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<subSubSection id="95E04D3D8039920B1141CE82B749845C" pageId="0" pageNumber="161" type="description">
<paragraph id="9474B1B66B20D701063A014CECF117B8" pageId="0" pageNumber="161">
<figureCitation id="ED796C8CF04DF3DAB763CD10735FD21D" captionStart="Figure 109" captionStartId="F109" captionText="Figure 109. Triphora albovittata var. mamillata Verco, 1909, Gulf St. Vincent, South Australia. A, B NHMUK 1910.3.29.49: front (A), back (B). C, D NHMUK 1910.3.29.50 - 51: front views. Scale bar: 0.5 mm (photo courtesy: Kevin Webb, NHMUK Photographic Unit)." figureDoi="10.3897/zse.95.32803.figure109" httpUri="https://binary.pensoft.net/fig/295738" pageId="0" pageNumber="161">Figure 109</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="81EA91D68EA1E5A6D602038FF5B75544" pageId="0" pageNumber="161">
<taxonomicName id="F47C2A3B59284450F5CD30E333C7BE51" authorityName="Verco" authorityYear="1909" class="Gastropoda" family="Triphoridae" genus="Triphora" higherTaxonomySource="treatment-meta" infraspecific-rank="var." kingdom="Animalia" lsidName="Triphora albovittata subsp. var. var. mamillata" order="Ptenoglossa" pageId="0" pageNumber="161" phylum="Mollusca" rank="variety" species="albovittata" subSpecies="var." variety="mamillata">Triphora albovittata var. mamillata</taxonomicName>
<bibRefCitation id="0B11678E3461AF26641828C25C36DB5B" author="Verco, JC" journalOrPublisher="Transactions of the Royal Society of South Australia" pageId="0" pageNumber="161" pagination="277 - 292" refId="B90" refString="Verco, JC, 1909. Notes on South Australian marine Mollusca, with description of new species. Part XI. Transactions of the Royal Society of South Australia 33: 277 - 292" title="Notes on South Australian marine Mollusca, with description of new species. Part XI." volume="33" year="1909">Verco 1909</bibRefCitation>
: 285-286, not illustrated.
</paragraph>
</subSubSection>
<subSubSection id="B7F0B10576CDEEF53AFC045425C532C3" pageId="0" pageNumber="161" type="type locality">
<paragraph id="2F3E786FDAD22E7BA10CF99374E153E5" pageId="0" pageNumber="161">
<typeStatus id="B53EE7EFF1C21253C7CBD828F2EBC567">Type</typeStatus>
locality.
</paragraph>
<paragraph id="975521D9E46C53BED7C098E96A6D0A0C" pageId="0" pageNumber="161">
Gulf St Vincent, South Australia (fide
<bibRefCitation id="573C83D6419A25E82FCE274405EF6F05" DOI="https://doi.org/10.3853/j.0812-7387.2.1983.102" author="Marshall, BA" journalOrPublisher="Records of the Australian Museum Supplement" pageId="0" pageNumber="161" pagination="1 - 119" refId="B45" refString="Marshall, BA, 1983. A revision of the recent Triphoridae of Southern Australia. Records of the Australian Museum Supplement 2: 1 - 119, DOI: https://doi.org/10.3853/j.0812-7387.2.1983.102" title="A revision of the recent Triphoridae of Southern Australia." url="https://doi.org/10.3853/j.0812-7387.2.1983.102" volume="2" year="1983">Marshall 1983</bibRefCitation>
, by
<typeStatus id="D96EA3CF31F197C47047DC84AD487433">lectotype</typeStatus>
designation).
</paragraph>
</subSubSection>
<subSubSection id="7CB6404CFA9FD16DCE3602A862F8263D" pageId="0" pageNumber="161" type="type material">
<paragraph id="FE911D1B7ED2065B12159F2F5F93D278" pageId="0" pageNumber="161">
<typeStatus id="A8F81F2466BED77753B8BF0E82A86785">Type</typeStatus>
material.
</paragraph>
<paragraph id="743D1DACB6E781C5543FFD6CF4ADB73A" pageId="0" pageNumber="161">
<materialsCitation id="48E6801F044F9C187A51C0374099F420" collectionCode="SAM" country="Marshall Islands" typeStatus="Lectotype">
<typeStatus id="0AE69B3CBFCEAB87681124D69F3E61E0">Lectotype</typeStatus>
:
<collectionCode id="AE162FFE3004D6139315691E34DB2636" collectionName="Australia, South Australia, Adelaide, South Australian Museum" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" type="">SAM</collectionCode>
D.13446 (fide
<bibRefCitation id="1BA15E6785BF135BF993BB2BC98FB6C8" DOI="https://doi.org/10.3853/j.0812-7387.2.1983.102" author="Marshall, BA" journalOrPublisher="Records of the Australian Museum Supplement" pageId="0" pageNumber="161" pagination="1 - 119" refId="B45" refString="Marshall, BA, 1983. A revision of the recent Triphoridae of Southern Australia. Records of the Australian Museum Supplement 2: 1 - 119, DOI: https://doi.org/10.3853/j.0812-7387.2.1983.102" title="A revision of the recent Triphoridae of Southern Australia." url="https://doi.org/10.3853/j.0812-7387.2.1983.102" volume="2" year="1983">Marshall 1983</bibRefCitation>
; not seen, see Remarks)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="463AF30760C5701E11211FA1CB551DD5" pageId="0" pageNumber="161" type="materials_examined">
<paragraph id="867D48D4E8BF6341F705C3F1D165EDCF" pageId="0" pageNumber="161">Additional material.</paragraph>
<paragraph id="3676CDC9AC312E26B514B92597F596D0" pageId="0" pageNumber="161">
<materialsCitation id="45C580FBD13A5F7BD5E4218D17A63079" collectingDate="2019-01-01" collectingDateMax="2019-12-31" collectingDateMin="2019-01-01" collectionCode="NHMUK" country="Australia" location="Gulf St Vincent" specimenCount="3" stateProvince="South Australia">
<collectionCode id="A66106943128ADEE7F357853176FE5B7" collectionName="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
1910.3.29.49-51:
<specimenCount id="06FD6595144EDD027466DF84D295EC1E" type="generic">3 specimens</specimenCount>
,
<location id="FB19A07B35D78574FFEF4E3434A1B9EE" LSID="urn:lsid:plazi:treatment:C7527A3CF74BA32FDB288133D2983253:FB19A07B35D78574FFEF4E3434A1B9EE" name="Gulf St Vincent" stateProvince="South Australia">Gulf St Vincent</location>
,
<collectingRegion id="9CE4143272BD57AAC5F800B4D017B630" country="Australia" name="South Australia">
South
<collectingCountry id="852B6DC948B33F7A020322F3B740721D" name="Australia" value="Australia">Australia</collectingCountry>
</collectingRegion>
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.
</paragraph>
</subSubSection>
<subSubSection id="30B8DD8E7C3D28821A444C7D99885237" pageId="0" pageNumber="161" type="description">
<paragraph id="D4843565FD09F1A8C1D9CB2B907B8EDF" pageId="0" pageNumber="161">Original description.</paragraph>
<paragraph id="AE3A6FF45AA6C3EE835BE66FA98E1FFF" pageId="0" pageNumber="161">
<emphasis id="36618492F4B779B7B8E8E635C88D4D91" italics="true" pageId="0" pageNumber="161">
Instead of having the elongate four-whorled protoconch of the
<typeStatus id="6DA4DD4CB6030F371FE67C5533ED8E66">type</typeStatus>
, it has a mamillate two-whorled apex. The first whorl is round and smooth, the second has a central carina and subdistant axial bars. Generally the second is swollen and lies somewhat out of the axis of the shell, causing the mamillate form. Rarely the first whorl may be as large as the second. This protoconch seems complete, and not the base of a spiculate protoconch, whose terminal whorls have fallen. The shell varies in shape, being short, broad, and pupaeform, or long, narrow, and elongate-pyramidal.
</emphasis>
</paragraph>
<paragraph id="5D1249F34B81BB480449991AB0731FAB" pageId="0" pageNumber="161">
<emphasis id="727CA9EF5853741EC4B21924CFA406E4" italics="true" pageId="0" pageNumber="161">Dredged in Gulf St. Vincent, 7 perfect and 7 poor; in 90 fathoms off Cape Jaffa, 2 good; in 150 fathoms off Beachport, 2 poor. Taken on the beach MacDonnell Bay, 1; Gulf St. Vincent, 23, in varying condition; Venus Bay, 2, good.</emphasis>
</paragraph>
<paragraph id="8F0EB0BFC9551FF16495AF70BF64B5FA" pageId="0" pageNumber="161">
<emphasis id="3D8A98CBCE1685FA46A2A6CCC7D2198B" italics="true" pageId="0" pageNumber="161">The very different protoconch makes me diffident about calling this a variety, inasmuch as the characters of the protoconch are generally regarded as very certain specific diagnostics; but the shells are otherwise indistinguishable.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="2C3CDBD1299A064B3EE552743AB735D2" pageId="0" pageNumber="161" type="remarks">
<paragraph id="0A35F21D7E89D4D575913791217E4053" pageId="0" pageNumber="161">Remarks.</paragraph>
<paragraph id="C0E0FC4A69908153856D55B2FC22DFEC" pageId="0" pageNumber="161">
<bibRefCitation id="797B24D9C8599607575340AFF20ACC0F" DOI="https://doi.org/10.3853/j.0812-7387.2.1983.102" author="Marshall, BA" journalOrPublisher="Records of the Australian Museum Supplement" pageId="0" pageNumber="161" pagination="1 - 119" refId="B45" refString="Marshall, BA, 1983. A revision of the recent Triphoridae of Southern Australia. Records of the Australian Museum Supplement 2: 1 - 119, DOI: https://doi.org/10.3853/j.0812-7387.2.1983.102" title="A revision of the recent Triphoridae of Southern Australia." url="https://doi.org/10.3853/j.0812-7387.2.1983.102" volume="2" year="1983">Marshall (1983)</bibRefCitation>
considered this variety a valid taxon:
<taxonomicName id="D0EB8730E54E5025C9708C2BCF42468C" baseAuthorityName="Verco" baseAuthorityYear="1909" class="Gastropoda" family="Triphoridae" genus="Obesula" kingdom="Animalia" lsidName="Obesula mamillata" pageId="0" pageNumber="161" phylum="Mollusca" rank="species" species="mamillata">
<emphasis id="E951E24B70D8626353FB37881A9D0F60" italics="true" pageId="0" pageNumber="161">Obesula mamillata</emphasis>
</taxonomicName>
, distinguished by
<taxonomicName id="E76C386859F521338FA44194AC00D40C" lsidName="O. albovittata" pageId="0" pageNumber="161" rank="species" species="albovittata">
<emphasis id="CDDA3B9D36735399201B617FD9381D88" italics="true" pageId="0" pageNumber="161">O. albovittata</emphasis>
</taxonomicName>
by its paucispiral protoconch. Verco referred to a
<typeStatus id="8FFDCFF23574025A7B520088E87D81A6">type</typeStatus>
series of multiple specimens in his original description.
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<normalizedToken id="68E4A8128B42C3223DA71EC5C0895317" originalValue="Marshalls">Marshall's</normalizedToken>
(1983)
</bibRefCitation>
report of the
<normalizedToken id="E28160E0D4C5516DBCAD2C2734BDF926" originalValue="“holotype”">
&quot;
<typeStatus id="51B1A4F177B1ED2D704665188B5778F2">holotype</typeStatus>
&quot;
</normalizedToken>
in the
<collectionCode id="6811C4C58290582B7FAEBDAC449409C4" collectionName="Australia, South Australia, Adelaide, South Australian Museum" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" type="">SAM</collectionCode>
should be considered a
<typeStatus id="A86568A9947AAD07823E89A70529CE42">lectotype</typeStatus>
designation according to Article 74.6 of the Code (
<bibRefCitation id="2F1AEFDEF264AA327B45746DC1A4835B" author="Hinds, RB" journalOrPublisher="Annals and Magazine of Natural History" pageId="0" pageNumber="161" refId="B29" refString="ICZN (1999) International Code of Zoological Nomenclature, 4th edition.The International Trust for Zoological Nomenclature, London, 306 pp." title="International Code of Zoological Nomenclature, 4 th edition." year="1844">ICZN 1999</bibRefCitation>
).
</paragraph>
<paragraph id="A7318835B21DF6CDC684EAB1E244AB4D" pageId="0" pageNumber="161">
The specimens in
<collectionCode id="933AEA32B25F306B4D2FD49C50F4E9E7" collectionName="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
represent two strikingly different colour forms: a white form (Fig.
<figureCitation id="A02C4BE3A3C28D15A769ED3D8A9FCC8C" captionStart="Figure 109" captionStartId="F109" captionText="Figure 109. Triphora albovittata var. mamillata Verco, 1909, Gulf St. Vincent, South Australia. A, B NHMUK 1910.3.29.49: front (A), back (B). C, D NHMUK 1910.3.29.50 - 51: front views. Scale bar: 0.5 mm (photo courtesy: Kevin Webb, NHMUK Photographic Unit)." figureDoi="10.3897/zse.95.32803.figure109" httpUri="https://binary.pensoft.net/fig/295738" pageId="0" pageNumber="161">109A, B</figureCitation>
) and a form with light brownish shell, white first spiral cord and brown base (Fig.
<figureCitation id="E9D6476F0242406CE68BE6A03F56B2E3" captionStart="Figure 109" captionStartId="F109" captionText="Figure 109. Triphora albovittata var. mamillata Verco, 1909, Gulf St. Vincent, South Australia. A, B NHMUK 1910.3.29.49: front (A), back (B). C, D NHMUK 1910.3.29.50 - 51: front views. Scale bar: 0.5 mm (photo courtesy: Kevin Webb, NHMUK Photographic Unit)." figureDoi="10.3897/zse.95.32803.figure109" httpUri="https://binary.pensoft.net/fig/295738" pageId="0" pageNumber="161">109C, D</figureCitation>
).
<bibRefCitation id="20CEE9BEDCCAAFC4C3F64FF403329A8B" DOI="https://doi.org/10.3853/j.0812-7387.2.1983.102" author="Marshall, BA" journalOrPublisher="Records of the Australian Museum Supplement" pageId="0" pageNumber="161" pagination="1 - 119" refId="B45" refString="Marshall, BA, 1983. A revision of the recent Triphoridae of Southern Australia. Records of the Australian Museum Supplement 2: 1 - 119, DOI: https://doi.org/10.3853/j.0812-7387.2.1983.102" title="A revision of the recent Triphoridae of Southern Australia." url="https://doi.org/10.3853/j.0812-7387.2.1983.102" volume="2" year="1983">Marshall (1983)</bibRefCitation>
reported that also
<taxonomicName id="3D21EA18F179EDCE0663374DBC56ACD9" lsidName="O. albovittata" pageId="0" pageNumber="161" rank="species" species="albovittata">
<emphasis id="049FDBF24D348A2D1E4D0C57CD1BBB8B" italics="true" pageId="0" pageNumber="161">O. albovittata</emphasis>
</taxonomicName>
occurs in two colour forms, similar to the ones illustrated here for
<taxonomicName id="08699AEF9E90005A1889AFE5F025AD61" lsidName="O. mamillata" pageId="0" pageNumber="161" rank="species" species="mamillata">
<emphasis id="C3DF891B7E2C42BB4E25FB0F785CFEA5" italics="true" pageId="0" pageNumber="161">O. mamillata</emphasis>
</taxonomicName>
and suggested that the only differentiating character between the two species is the protoconch
<typeStatus id="DE01FBECB14247A8AD8B89855CDC800D">type</typeStatus>
.
</paragraph>
<paragraph id="C2A0DAA47469156D0D23ABE9233C9991" pageId="0" pageNumber="161">
The
<collectionCode id="4B533686217EFC0EF8F5D70B30CFF47E" collectionName="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
collection register does not indicate that these are
<normalizedToken id="BCA64C19D3DC6247563382A2A162B6B1" originalValue="“Co-types”">&quot;Co-types&quot;</normalizedToken>
, although the specimens were part of
<normalizedToken id="05F4708349DD7B044F3D01A2CB0A4FD0" originalValue="Vercos">Verco's</normalizedToken>
collection and come from the
<typeStatus id="0DE2941BA22A80225E9F1C60E6A67DA5">type</typeStatus>
locality. Until additional evidence is available, we do not consider these specimens as belonging to the original
<typeStatus id="6E5334B2C1512D810A1783AB8F3E21FD">type</typeStatus>
series.
</paragraph>
<caption id="16E9736F0E6FBD54AEA5A295105F61C3" ID-Zenodo-Dep="2654039" doi="10.3897/zse.95.32803.figure109" httpUri="https://binary.pensoft.net/fig/295738" pageId="0" pageNumber="161" start="Figure 109" startId="F109">
<paragraph id="E46DC301581903F74BCD0C97C1072BC2" pageId="0" pageNumber="161">
<emphasis id="71EB9E06E1B782537D3FDCB7369B139D" bold="true" pageId="0" pageNumber="161">Figure 109.</emphasis>
<taxonomicName id="045B5C313FFC5F54939C2F300B962034" authorityName="Verco" authorityYear="1909" class="Gastropoda" family="Triphoridae" genus="Triphora" infraspecific-rank="var." kingdom="Animalia" lsidName="Triphora albovittata subsp. var. var. mamillata" order="Ptenoglossa" pageId="0" pageNumber="161" phylum="Mollusca" rank="variety" species="albovittata" subSpecies="var." variety="mamillata">Triphora albovittata var. mamillata</taxonomicName>
Verco, 1909, Gulf St. Vincent, South Australia.
<emphasis id="FE9B052CCF12BE7D7D69A08A09BEBE12" bold="true" pageId="0" pageNumber="161">A, B</emphasis>
<collectionCode id="18671DA7CFA594BAEC9822ECE0F34C65" collectionName="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
1910.3.29.49: front (
<emphasis id="0A9B637ED7727D5B0BA920AD5C51305A" bold="true" pageId="0" pageNumber="161">A</emphasis>
), back (
<emphasis id="5B43DD63CDCCB54DCDFF59A03FF7FAF8" bold="true" pageId="0" pageNumber="161">B</emphasis>
).
<emphasis id="B0EADE17D9576BCD4C886F530CF68060" bold="true" pageId="0" pageNumber="161">C, D</emphasis>
<collectionCode id="2950D7B68D275ACAB2CB1C5FDACCE4A9" collectionName="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
1910.3.29.50-51: front views. Scale bar:
<quantity id="7E72D0FA392D61D685F86D34CE68F8DA" metricMagnitude="-4" metricUnit="m" metricValue="5.0" unit="mm" value="0.5">0.5 mm</quantity>
(photo courtesy: Kevin Webb,
<collectionCode id="C3D4EE46C8B5082410F93A7E89CE804A" collectionName="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
Photographic Unit).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>