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<mods:url id="219E778FF8ADBD4E84FB19B9E9498FB9">http://dx.doi.org/10.3897/phytokeys.240.101716</mods:url>
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<mods:classification id="152169FD9F1B170EBC181E951F419244">journal article</mods:classification>
<mods:identifier id="A8C65E19D73E995A6B192579EB920283" type="DOI">http://dx.doi.org/10.3897/phytokeys.240.101716</mods:identifier>
<mods:identifier id="ADC4BDACC2CD4699861751EBF0AB6111" type="Pensoft-Pub">1314-2003-240-1</mods:identifier>
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<treatment id="B45005DADB6C253444EEAF331F4B53B7" LSID="urn:lsid:plazi:treatment:B45005DADB6C253444EEAF331F4B53B7" httpUri="http://treatment.plazi.org/id/B45005DADB6C253444EEAF331F4B53B7" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection id="7029CF3A0456C03A1ECBAF039D2AB53A" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="17F8268945C3C6770B3F88C7A835DA2F" pageId="0" pageNumber="1">
<taxonomicName id="B4F9A01913215E609AD6E09842A22B8E" ID-CoL="9CJYW" authority="Urb., Repert. Spec. Nov. Regni Veg. 19: 4. 1923." authorityName="Urb., Repert. Spec. Nov. Regni Veg. 19: 4. 1923." class="Magnoliopsida" family="Fabaceae" genus="Arcoa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Arcoa" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Arcoa Urb., Repert. Spec. Nov. Regni Veg. 19: 4. 1923.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="5469BF18E19496B9AF11D01D552F32EB" pageId="0" pageNumber="1" type="description">
<paragraph id="5F99BEC0E7FA767C98FA167ADD5B6FBE" pageId="0" pageNumber="1">
<figureCitation id="B7644B79414AE8015D0A62045F63E0E6" captionStart="Figure 7" captionStartId="F14" captionText="Figure 7. Flower, fruit, and vegetative characters of tribe Ceratonieae A Arcoa gonavensis Urb., foliage and fruits, Dominican Republic B Tetrapterocarpon geayi Humbert, fruits and part of bipinnate leaf, Madagascar (Du Puy M 410) C-E Acrocarpus fraxinifolius Wight &amp; Arn., India, cultivated tree C inflorescence D dehisced fruits and new flush foliage E part of a bipinnate leaf F-J Ceratonia siliqua L. F tree, Jordan, semi-desert near Petra G fruits, Israel, Mt. Scopus Botanical Gardens, Jerusalem H pistillate flowers, Crete I staminate flowers, Crete J staminate inflorescences, Israel, Jerusalem Botanical Gardens. Photo credits A F Jimenez R. B D Du Puy C-E https: // efloraofindia. com / 2011 / 02 / 01 / acrocarpus-fraxinifolius / E D Valke F, G, J O Fragman-Sapir H, I G Lewis." figureDoi="10.3897/phytokeys.240.101716.figure7" httpUri="https://binary.pensoft.net/fig/1014680" pageId="0" pageNumber="1">Figs 7</figureCitation>
<figureCitation id="A6517614426CE575775ED923964B5B91" captionStart="Figure 8" captionStartId="F15" captionText="Figure 8. Distribution of Arcoa based on quality-controlled digitised herbarium records. See Suppl. material 1 for the source of occurrence data." figureDoi="10.3897/phytokeys.240.101716.figure8" httpUri="https://binary.pensoft.net/fig/1014681" pageId="0" pageNumber="1">, 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="5645CE5CA55F1762A3C7F399BF1D8978" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="C3AA01B512022506DBD44BDCD63CE88A" pageId="0" pageNumber="1">
<typeStatus id="A30E2ED8619FC66801355BB6E8AAD754">Type</typeStatus>
.
</paragraph>
<paragraph id="41C79EDAF2E70C04F1F59AF5D6CC41AF" pageId="0" pageNumber="1">
<taxonomicName id="81A1A2E1FF4BB42E8AC58383E7FDC27E" class="Magnoliopsida" family="Leguminosae" genus="Arcoa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Arcoa gonavensis" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="gonavensis">
<emphasis id="C6FA7EEC24803D59CF499FFC399D6B0D" italics="true" pageId="0" pageNumber="1">Arcoa gonavensis</emphasis>
</taxonomicName>
Urb.
</paragraph>
</subSubSection>
<subSubSection id="92795BB99F8FBDE55C888DDFAB86C0C7" pageId="0" pageNumber="1" type="description">
<paragraph id="89BA2FFA95764AC95B3944B07D480936" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="0771E2A033220E51286A87152F4D902A" pageId="0" pageNumber="1">
Shrub or small tree, with well-developed brachylasts. 
<emphasis id="45D3BA28DDE6B9A855C268C312E67E5C" bold="true" pageId="0" pageNumber="1">Stipules</emphasis>
spinescent, caducous. 
<emphasis id="4F0C5189E96CBF0C89E1C6E1742C7BEA" bold="true" pageId="0" pageNumber="1">Leaves</emphasis>
pinnate when juvenile, bipinnate with a terminal pinna when mature, leaflets opposite to subopposite, sessile. 
<emphasis id="5A4945053AA565DC2B91BC9FD7C7F30C" bold="true" pageId="0" pageNumber="1">Inflorescence</emphasis>
a sparsely branching panicle of spikes arising from a thickened woody brachyblast. 
<emphasis id="3EE86C3550B587D1D16A09E61E442C68" bold="true" pageId="0" pageNumber="1">Flowers</emphasis>
unisexual, staminate flowers with a prominent pistillode, pistillate flowers with staminoidia; sepals free to a very short hypanthium; petals 5 (6); a short cupuliform disk present centrally; stamens (or staminodes) number more than twice sepal number (12 or more per flower); pollen markedly irregular and coarsely reticulate, porate with prominent pores; ovary with an appressed rust-coloured indumentum, stigma terminal and capitate. 
<emphasis id="D2374E8BF75C4E4CE6F0895B4E1BD907" bold="true" pageId="0" pageNumber="1">Fruits</emphasis>
oblong-ellipsoid, subterete, thick-walled, indehiscent, 1-few-seeded. 
<emphasis id="675F418F1B9372F34F74792D0EB94E57" bold="true" pageId="0" pageNumber="1">Seeds</emphasis>
ovate, compressed, surrounded by copious pulp, pleurogram lacking (Fig. 
<figureCitation id="34796A93D84AFB0BCE3D887B103230AA" captionStart="Figure 7" captionStartId="F14" captionText="Figure 7. Flower, fruit, and vegetative characters of tribe Ceratonieae A Arcoa gonavensis Urb., foliage and fruits, Dominican Republic B Tetrapterocarpon geayi Humbert, fruits and part of bipinnate leaf, Madagascar (Du Puy M 410) C-E Acrocarpus fraxinifolius Wight &amp; Arn., India, cultivated tree C inflorescence D dehisced fruits and new flush foliage E part of a bipinnate leaf F-J Ceratonia siliqua L. F tree, Jordan, semi-desert near Petra G fruits, Israel, Mt. Scopus Botanical Gardens, Jerusalem H pistillate flowers, Crete I staminate flowers, Crete J staminate inflorescences, Israel, Jerusalem Botanical Gardens. Photo credits A F Jimenez R. B D Du Puy C-E https: // efloraofindia. com / 2011 / 02 / 01 / acrocarpus-fraxinifolius / E D Valke F, G, J O Fragman-Sapir H, I G Lewis." figureDoi="10.3897/phytokeys.240.101716.figure7" httpUri="https://binary.pensoft.net/fig/1014680" pageId="0" pageNumber="1">7A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="12E9AFAFD69C2141A2CE5152B4678927" pageId="0" pageNumber="1" type="chromosome number">
<paragraph id="EC6A7E2B89D725D8174A25804FC64C20" pageId="0" pageNumber="1">Chromosome number.</paragraph>
<paragraph id="297C2EC1A5250968B643DAC740F62535" pageId="0" pageNumber="1">Unknown.</paragraph>
</subSubSection>
<subSubSection id="A117BA4C65C6CAA57C7C9125F010F6CE" pageId="0" pageNumber="1" type="distribution">
<paragraph id="DCD263D0F2199BDEA26D44437F38DABA" pageId="0" pageNumber="1">Included species and geographic distribution.</paragraph>
<paragraph id="FFE6F6FB1AF1A9248890573D97B59A0C" pageId="0" pageNumber="1">
Monospecific (
<taxonomicName id="BFB44FC9049D8055AE236832578C381A" lsidName="A. gonavensis" pageId="0" pageNumber="1" rank="species" species="gonavensis">
<emphasis id="42B23C7841E37450E8BD305147E38284" italics="true" pageId="0" pageNumber="1">A. gonavensis</emphasis>
</taxonomicName>
), endemic to Hispaniola (Dominican Republic, Haiti, and the Island of 
<normalizedToken id="EF72CFE5AAAEDE33DF1333E89295F903" originalValue="Gonâve">Gonave</normalizedToken>
) (Fig. 
<figureCitation id="7880A619F6254D1950475CC9595107AF" captionStart="Figure 8" captionStartId="F15" captionText="Figure 8. Distribution of Arcoa based on quality-controlled digitised herbarium records. See Suppl. material 1 for the source of occurrence data." figureDoi="10.3897/phytokeys.240.101716.figure8" httpUri="https://binary.pensoft.net/fig/1014681" pageId="0" pageNumber="1">8</figureCitation>
).
</paragraph>
<caption id="FBC9C6AE5741CAEE589EFF74DFF7FCB7" doi="10.3897/phytokeys.240.101716.figure8" httpUri="https://binary.pensoft.net/fig/1014681" pageId="0" pageNumber="1" start="Figure 8" startId="F15">
<paragraph id="72D7D27FC1B514D39E6B4DEB98532FE9" pageId="0" pageNumber="1">
<emphasis id="597CED4419D204F130FD37105233C92E" bold="true" pageId="0" pageNumber="1">Figure 8.</emphasis>
Distribution of 
<taxonomicName id="E921989D7B8AF09CFDB56054456EBE1C" class="Magnoliopsida" family="Leguminosae" genus="Arcoa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Arcoa" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="1E7E2FAE8FD85B313FC97A4DFA81B11D" italics="true" pageId="0" pageNumber="1">Arcoa</emphasis>
</taxonomicName>
based on quality-controlled digitised herbarium records. See Suppl. material 1 for the source of occurrence data.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="4C6364114D63F554769B752D2207E35F" pageId="0" pageNumber="1" type="ecology">
<paragraph id="C9498F912C743B6FAB35FBE0ABB642C3" pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph id="0379E2EC6D6D4F9B505A732AF81813F3" pageId="0" pageNumber="1">Arid tropical vegetation on limestone hills and cliffs.</paragraph>
</subSubSection>
<subSubSection id="C94E0A16B9781EB4A9D109B22543134E" pageId="0" pageNumber="1" type="etymology">
<paragraph id="FE82AE7CC19AA89D77A938F31C5EC3FB" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="BE6B510743F8A88F6C11D82468D565B6" pageId="0" pageNumber="1">Named for Count George von Arco (ca. 1903).</paragraph>
</subSubSection>
<subSubSection id="1839B1B5DFBC65E37A7110E7899C31D8" pageId="0" pageNumber="1" type="human uses">
<paragraph id="8976E6B930D666EFFC8753B84F644270" pageId="0" pageNumber="1">Human uses.</paragraph>
<paragraph id="F629DF31DA038403B99B68699BCD4E83" pageId="0" pageNumber="1">Unknown.</paragraph>
</subSubSection>
<subSubSection id="A145AFE5B8E5E49E93043D247EFDF6C0" pageId="0" pageNumber="1" type="notes">
<paragraph id="9350B2A07D1EC55DF89C24C3422EB634" pageId="0" pageNumber="1">Notes.</paragraph>
<paragraph id="F06F331D40A981574EDCCB696711AAE3" pageId="0" pageNumber="1">
Originally placed by 
<bibRefCitation id="FA361E8E8CEB3DCE1FB146B8051C5240" DOI="https://doi.org/10.1002/fedr.19230190102" author="Urban, I" journalOrPublisher="Darwiniana" pageId="0" pageNumber="1" refId="B875" refString="Urban, I, 1923. Arcoa. Repertorium specierum novarum regni vegetabilis 19: 4. https://doi.org/10.1002/fedr.19230190102" title="Arcoa. Repertorium specierum novarum regni vegetabilis 19: 4." url="https://doi.org/10.1002/fedr.19230190102" year="1923">Urban (1923)</bibRefCitation>
in the 
<taxonomicName id="DCFAED316D35C36A465C4D13372005ED" family="Euphorbiaceae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="1" rank="family">Euphorbiaceae</taxonomicName>
, 
<taxonomicName id="ADC3AE01E020148657470BAA15A4C292" class="Magnoliopsida" family="Leguminosae" genus="Arcoa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Arcoa" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="DF05BB6F63188BD9A075D9A20CB0D87D" italics="true" pageId="0" pageNumber="1">Arcoa</emphasis>
</taxonomicName>
was subsequently transferred to the 
<taxonomicName id="A451A6AB516FA5E6EE1DDA7C3FED91E4" family="Leguminosae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="1" rank="family">Leguminosae</taxonomicName>
by 
<bibRefCitation id="48CA44724AA887263117642B66A88049" author="Urban, I" journalOrPublisher="Darwiniana" pageId="0" pageNumber="1" refId="B876" refString="Urban, I, 1928. Plantae Haitiensis et Domingenses novae vel rariores V. cl. EL Ekman 1924-1927. Arkiv foer Botanik 22A(8): 1-98." title="Plantae Haitiensis et Domingenses novae vel rariores V. cl. EL Ekman 1924 - 1927. Arkiv foer Botanik 22 A (8): 1 - 98." year="1928">Urban (1928)</bibRefCitation>
. The genus was placed in the eclectic 
<taxonomicName id="98CB6BF83A31370469FE66DA6EE5B3A8" class="Magnoliopsida" family="Caesalpiniaceae" genus="Dimorphandra" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dimorphandra" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Dimorphandra</taxonomicName>
group of 
<taxonomicName id="0DA65EC64B741A604254194682034D0D" family="Caesalpiniaceae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="1" rank="tribe" tribe="Caesalpinieae">Caesalpinieae</taxonomicName>
by 
<bibRefCitation id="E52EE129970378491C0D9AAF10FA1CC8" author="Polhill, RM" editor="Polhill, RM" journalOrPublisher="Royal Botanic Gardens, Kew, United Kingdom" pageId="0" pageNumber="1" pagination="81 - 95" refId="B669" refString="Polhill, RM, Vidal, JE, 1981. Tribe 1. Caesalpinieae. In: Polhill, RM, Raven, PH, Eds., Advances in Legume Systematics, Part 1. Royal Botanic Gardens, Kew, United Kingdom: 81 - 95" title="Tribe 1. Caesalpinieae." volumeTitle="Advances in Legume Systematics, Part 1." year="1981">Polhill and Vidal (1981)</bibRefCitation>
, along with 
<taxonomicName id="5AF412C4422FD394EE472B4CB372F576" genus="Tetrapetrocarpon" lsidName="Tetrapetrocarpon" pageId="0" pageNumber="1" rank="genus">
<emphasis id="6F562E602EBAECB0019E50411C799468" italics="true" pageId="0" pageNumber="1">Tetrapetrocarpon</emphasis>
</taxonomicName>
, both with unisexual flowers and petals not covered by the sepals in bud, but phylogenetic analyses clearly resolved 
<taxonomicName id="C2EA64C796C6E7CFF9A35F9662B7669A" class="Magnoliopsida" family="Leguminosae" genus="Arcoa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Arcoa" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="21D4947E88793123761E85F2C2276F54" italics="true" pageId="0" pageNumber="1">Arcoa</emphasis>
</taxonomicName>
as part of the &quot; 
<taxonomicName id="C1D088926EF77B40052288084E469AE4" class="Magnoliopsida" family="Fabaceae" genus="Umtiza" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Umtiza" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Umtiza</taxonomicName>
clade&quot;, either sister to the 
<taxonomicName id="995B6F951450872246F8B6686FFB6799" family="Fabaceae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="1" rank="tribe" tribe="Gleditsieae">Gleditsieae</taxonomicName>
genera (
<bibRefCitation id="6390A67C29D7C344600DA091596020E0" DOI="https://doi.org/10.1086/376881" author="Herendeen, PS" journalOrPublisher="Royal Botanic Gardens, Kew, United Kingdom" pageId="0" pageNumber="1" refId="B357" refString="Herendeen, PS, Lewis, GP, Bruneau, A, 2003b. Floral morphology in caesalpinioid legumes: testing the monophyly of the &quot;Umtiza clade&quot;. International Journal of Plant Sciences 164(Supplement): S393-S407. https://doi.org/10.1086/376881" title="Floral morphology in caesalpinioid legumes: testing the monophyly of the &quot; Umtiza clade &quot;. International Journal of Plant Sciences 164 (Supplement): S 393 - S 407." url="https://doi.org/10.1086/376881" year="2003 b">Herendeen et al. 2003b</bibRefCitation>
) or sister to those of 
<taxonomicName id="A6AB1182B3C740528EEF65D70EAD7A28" family="Fabaceae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="1" rank="tribe" tribe="Ceratonieae">Ceratonieae</taxonomicName>
(
<bibRefCitation id="158FE4977726D52136726C9CC3E588BB" DOI="https://doi.org/10.1139/B08-058" author="Bruneau, A" journalOrPublisher="Botany" pageId="0" pageNumber="1" pagination="697 - 718" refId="B131" refString="Bruneau, A, Mercure, M, Lewis, GP, Herendeen, PS, 2008. Phylogenetic patterns and diversification in the caesalpinioid legumes. Botany 86 (7): 697 - 718, DOI: https://doi.org/10.1139/B08-058" title="Phylogenetic patterns and diversification in the caesalpinioid legumes." url="https://doi.org/10.1139/B08-058" volume="86" year="2008">Bruneau et al. 2008</bibRefCitation>
), the latter as in 
<bibRefCitation id="EB413723EBE15DBE86C4D223F9406DD2" DOI="https://doi.org/10.3897/phytokeys.205.85866" author="Ringelberg, JJ" editor="Hughes, CE" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="3 - 58" refId="B712" refString="Ringelberg, JJ, Koenen, EJM, Iganci, JR, Queiroz, LP, Murphy, DJ, Gaudeul, M, Bruneau, A, Luckow, M, Lewis, GP, Hughes, CE, 2022. Phylogenomic analysis of 997 nuclear genes reveals the need for extensive generic re-delimitation in Caesalpinioideae (Leguminosae). In: Hughes, CE, Queiroz, LP, Lewis, GP, Eds., Advances in Legume Systematics 14. Classification of Caesalpinioideae Part 1: New generic delimitations. PhytoKeys 205: 3 - 58, DOI: https://doi.org/10.3897/phytokeys.205.85866" title="Phylogenomic analysis of 997 nuclear genes reveals the need for extensive generic re-delimitation in Caesalpinioideae (Leguminosae)." url="https://doi.org/10.3897/phytokeys.205.85866" volume="205" volumeTitle="Advances in Legume Systematics 14. Classification of Caesalpinioideae Part 1: New generic delimitations." year="2022">Ringelberg et al. (2022)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8BC3FFC6EAACB112684C1206292AA3F7" pageId="0" pageNumber="1" type="taxonomic references">
<paragraph id="28BF32DFD3B7BACC1E5D21E88AEAFC00" pageId="0" pageNumber="1">Taxonomic references.</paragraph>
<paragraph id="FA264B8762ECB974C8F98788600E9A81" pageId="0" pageNumber="1">
<bibRefCitation id="8C24BE33EFAEA07B49650B284D042EC9" author="Lewis, GP" journalOrPublisher="Royal Botanic Gardens, Kew, United Kingdom" pageId="0" pageNumber="1" refId="B471" refString="Lewis, GP, 2005b. Tribe Caesalpinieae. In: Lewis G, Schrire B, Mackinder B, Lock M (Eds) Legumes of the World, Royal Botanic Gardens, Kew, United Kingdom, 127-161." title="Tribe Caesalpinieae. In: Lewis G, Schrire B, Mackinder B, Lock M (Eds) Legumes of the World, Royal Botanic Gardens, Kew, United Kingdom, 127 - 161." year="2005 b">Lewis (2005b)</bibRefCitation>
; 
<bibRefCitation id="81BE359CA54452201EF6E541213CF862" author="Urban, I" journalOrPublisher="Darwiniana" pageId="0" pageNumber="1" refId="B876" refString="Urban, I, 1928. Plantae Haitiensis et Domingenses novae vel rariores V. cl. EL Ekman 1924-1927. Arkiv foer Botanik 22A(8): 1-98." title="Plantae Haitiensis et Domingenses novae vel rariores V. cl. EL Ekman 1924 - 1927. Arkiv foer Botanik 22 A (8): 1 - 98." year="1928">Urban (1928)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>