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<document id="B90B7F72B6CA81E8725F4B6F4D4E333F" ID-DOI="10.11646/zootaxa.3861.4.5" ID-GBIF-Dataset="d8341a2e-6432-4f3a-a58c-71cae7ec21bf" ID-ISSN="1175-5326" ID-Zenodo-Dep="225326" ID-ZooBank="14A28B09-F9B4-4FF1-A07B-B960463A807B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460911528020" checkinUser="plazi" docAuthor="Tian, Mingyi" docDate="2014" docId="B26C87B5FFB43B1EFF6AFA048C01C121" docLanguage="en" docName="zt03861p362.pdf" docOrigin="Zootaxa 3861 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Guiodytes deharvengi Tian, 2014, n. sp." docType="treatment" docVersion="8" lastPageNumber="361" masterDocId="4E55FFCDFFB63B18FFFDFFD18E60C42D" masterDocTitle="New records and a new species of the cavernicolous genus Guiodytes Tian, 2013 from Guangxi, China (Coleoptera: Carabidae: Scaritinae)" masterLastPageNumber="362" masterPageNumber="355" pageNumber="357" updateTime="1698361309906" updateUser="plazi">
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<mods:title id="0209BC974F0E947AB63840662BBA2691">New records and a new species of the cavernicolous genus Guiodytes Tian, 2013 from Guangxi, China (Coleoptera: Carabidae: Scaritinae)</mods:title>
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<mods:name id="5D5EA9C4EE6AF4280DFE6DEC35A0A490" type="personal">
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<mods:namePart id="E9E8F47CF4FFB645F7FBC76F7EC64DC4">Tian, Mingyi</mods:namePart>
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<mods:title id="D76A66F4D98F5900B9E3CD0DBC2A925C">Zootaxa</mods:title>
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<mods:date id="9B721FA22D3EFA34F2D1B3CB55E057F0">2014</mods:date>
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<treatment id="B26C87B5FFB43B1EFF6AFA048C01C121" ID-DOI="http://doi.org/10.5281/zenodo.6135178" ID-GBIF-Taxon="119588451" ID-Zenodo-Dep="6135178" LSID="urn:lsid:plazi:treatment:B26C87B5FFB43B1EFF6AFA048C01C121" httpUri="http://treatment.plazi.org/id/B26C87B5FFB43B1EFF6AFA048C01C121" lastPageId="6" lastPageNumber="361" pageId="2" pageNumber="357">
<subSubSection id="72DF6528FFB43B1AFF6AFA048F4EC23D" pageId="2" pageNumber="357" type="nomenclature">
<paragraph id="3A7A36A3FFB43B1AFF6AFA048F87C1C2" blockId="2.[151,487,1493,1552]" box="[151,487,1493,1519]" pageId="2" pageNumber="357">
<heading id="613281CFFFB43B1AFF6AFA048F87C1C2" bold="true" box="[151,487,1493,1519]" fontSize="11" level="1" pageId="2" pageNumber="357" reason="1">
<emphasis id="08B1EAB1FFB43B1AFF6AFA048F87C1C2" bold="true" box="[151,487,1493,1519]" pageId="2" pageNumber="357">
<taxonomicName id="FDC54D20FFB43B1AFF6AFA048FFDC1C2" ID-CoL="3HLDW" box="[151,413,1493,1519]" class="Insecta" family="Carabidae" genus="Guiodytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="357" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
<emphasis id="08B1EAB1FFB43B1AFF6AFA048FFDC1C2" bold="true" box="[151,413,1493,1519]" italics="true" pageId="2" pageNumber="357">Guiodytes deharvengi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="138257CAFFB43B1AFE58FA048F87C1C2" box="[421,487,1493,1519]" pageId="2" pageNumber="357" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="3A7A36A3FFB43B1AFF6AFA268F4EC23D" blockId="2.[151,487,1493,1552]" box="[151,302,1527,1552]" pageId="2" pageNumber="357">
(
<figureCitation id="A2FE2A26FFB43B1AFF62FA268E8CC23D" box="[159,236,1527,1552]" captionStart="FIGURE 1" captionStartId="1.[151,250,1414,1436]" captionTargetBox="[151,1435,411,1392]" captionTargetId="figure@1.[151,1435,411,1392]" captionTargetPageId="1" captionText="FIGURE 1. Distributional map of the genus Guiodytes Tian, 2013 (a. showing the location of Guangxi within China; square: G. b e d o s a e Tian; star: G. cavicola Tian; round: G. deharvengi n. sp.)" httpUri="https://zenodo.org/record/225327/files/figure.png" pageId="2" pageNumber="357">Figs. 1</figureCitation>
,
<figureCitation id="A2FE2A26FFB43B1AFF07FA298F45C23D" box="[250,293,1528,1552]" captionStart-0="FIGURES 4 6" captionStart-1="FIGURE 7" captionStartId-0="4.[151,264,1955,1978]" captionStartId-1="5.[151,250,1954,1976]" captionTargetBox-0="[321,1262,199,1925]" captionTargetBox-1="[240,1331,256,1838]" captionTargetId-0="figure@4.[311,1275,193,1934]" captionTargetId-1="figure@5.[165,1421,193,1932]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 4 6. Guiodytes deharvengi n. sp. (4. habitus, female, paratype; 5. median lobe, lateral view; 6. right and left parameres, lateral view)." captionText-1="FIGURE 7. Female genitalia of Guiodytes deharvengi n. sp., ventral view." httpUri-0="https://zenodo.org/record/225329/files/figure.png" httpUri-1="https://zenodo.org/record/225330/files/figure.png" pageId="2" pageNumber="357">47</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="72DF6528FFB43B1EFF6AF9EE8C3BC7C0" lastPageId="6" lastPageNumber="361" pageId="2" pageNumber="357" type="materials_examined">
<paragraph id="3A7A36A3FFB43B1AFF6AF9EE8F44C2E8" blockId="2.[151,1436,1599,2021]" pageId="2" pageNumber="357">
<typeStatus id="E57E8801FFB43B1AFF6AF9EE8F64C275" box="[151,260,1599,1624]" pageId="2" pageNumber="357" type="holotype">Holotype</typeStatus>
: male, Guangxi: Huanjiang Maonan Autonomous County: Shuiyuan Zhen: Sancai Cun: a cave named as Shuiku Dong,
<geoCoordinate id="5FF15064FFB43B1AFEBBF9B58FBBC251" box="[326,475,1636,1660]" direction="north" orientation="latitude" pageId="2" pageNumber="357" precision="1" value="24.823833">24°49.430N</geoCoordinate>
,
<geoCoordinate id="5FF15064FFB43B1AFE11F9B58CEBC250" box="[492,651,1636,1661]" direction="east" orientation="longitude" pageId="2" pageNumber="357" precision="1" value="108.091736">108°05.504E</geoCoordinate>
, altitude
<quantity id="FD3D9B46FFB43B1AFD03F9B58D2DC251" box="[766,845,1636,1660]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="357" unit="m" value="280.0">280 m</quantity>
,
<date id="4E7B1063FFB43B1AFCA2F9B58D81C251" box="[863,993,1636,1660]" pageId="2" pageNumber="357" value="2012-12-04">2012-XII-4</date>
, leg. Louis Deharveng, Anne Bedos, Arnaud Faille and Mingyi Tian, in
<collectionCode id="5CD4AE66FFB43B1AFDCDF9598C1EC28D" box="[560,638,1672,1696]" pageId="2" pageNumber="357">SCAU</collectionCode>
;
<typeStatus id="E57E8801FFB43B1AFD71F9598C9CC28D" box="[652,764,1672,1696]" pageId="2" pageNumber="357" type="paratype">paratypes</typeStatus>
:
<specimenCount id="2CC3FD2AFFB43B1AFCF4F9598D37C28D" box="[777,855,1671,1696]" pageId="2" pageNumber="357" type="male">1 male</specimenCount>
and
<specimenCount id="2CC3FD2AFFB43B1AFC69F9598A66C28D" box="[916,1030,1671,1696]" pageId="2" pageNumber="357" type="female">2 females</specimenCount>
, ibid, in
<collectionCode id="5CD4AE66FFB43B1AFB8DF9598ADBC28D" box="[1136,1211,1672,1696]" pageId="2" pageNumber="357">SCAU</collectionCode>
and
<collectionCode id="5CD4AE66FFB43B1AFB0AF9598B2FC2B2" LSID="urn:lsid:biocol.org:col:34988" box="[1271,1359,1672,1695]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="357">MNHN</collectionCode>
, Paris, respectively.
</paragraph>
<paragraph id="3A7A36A3FFB43B1AFF3AF91E8B16C2C5" blockId="2.[151,1436,1599,2021]" box="[199,1398,1743,1768]" pageId="2" pageNumber="357">
Length (from apex of left mandible to end of elytra):
<quantity id="FD3D9B46FFB43B1AFCE1F91E8DC6C2CA" box="[796,934,1743,1768]" metricMagnitude="-3" metricUnit="m" metricValue="9.35" metricValueMax="9.5" metricValueMin="9.2" pageId="2" pageNumber="357" unit="mm" value="9.35" valueMax="9.5" valueMin="9.2">9.29.5 mm</quantity>
; width:
<quantity id="FD3D9B46FFB43B1AFBFDF91E8AEAC2CA" box="[1024,1162,1743,1768]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="2" pageNumber="357" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.52.8 mm</quantity>
. Habitus as in
<figureCitation id="A2FE2A26FFB43B1AFAD3F91E8B12C2C5" box="[1326,1394,1743,1768]" captionStart="FIGURES 4 6" captionStartId="4.[151,264,1955,1978]" captionTargetBox="[321,1262,199,1925]" captionTargetId="figure@4.[311,1275,193,1934]" captionTargetPageId="4" captionText="FIGURES 4 6. Guiodytes deharvengi n. sp. (4. habitus, female, paratype; 5. median lobe, lateral view; 6. right and left parameres, lateral view)." httpUri="https://zenodo.org/record/225329/files/figure.png" pageId="2" pageNumber="357">Fig. 4</figureCitation>
.
</paragraph>
<paragraph id="3A7A36A3FFB43B1BFF3AF9258F6CC64D" blockId="2.[151,1436,1599,2021]" lastBlockId="3.[151,1437,151,933]" lastPageId="3" lastPageNumber="358" pageId="2" pageNumber="357">Head stout, from apex of mandible longer than wide, HL/HW=1.162.02, mean 1.18, or slightly longer than wide when measured from apex of clypeus, HL/HW= 0.810.84, mean 0.83; distinctly narrower than pronotum, HW/PW=0.780.81, mean 0.8; clypeus (with wings) strongly transverse, convex, bisetose at base, widely and deeply emarginate at front, making clypeal wings very protruding anteriorly; supra-antennal plates well marked, clearly separated from clypeal wings by a small but obvious notch, nearly rounded, strongly convex, smooth and glabrous; frons and vertex convex; two pairs of supraorbital pores present, at behind of supra-antennal plates and at neck constriction, respectively, and both on supraorbital furrows; supraorbital furrows very deep and wide, parallel-sided at middle of frons, then strongly divergent afterwards, ended to posterior supraorbital pores; supraorbital carinae distinct; frontoclypeal sulcus indistinct; eyes completely disappeared; genae well developed, subparallel-sided, frontal portion strongly broadened laterally; neck constriction distinct, beginning at the level of hind supraorbital pores; labrum conspicuously transverse, distinctly narrower than clypeus excluding clypeal wings, 6-setose, and ciliate on both sides, slightly emarginated at anterior margin; mandible short and stout, right mandible with a distinct median tooth; palps ensiform, glabrous, apical segments of both maxillary and labial palps longer than penultimate ones respectively; labial palpomere 2 bisetose on inner margin; ligula unisetose at apex; labial suture well marked, deeply and widely furrowed in median portion; mentum well developed, with two pairs of setae, one pairs situated beneath mental tooth, the other at base near lateral margin; median tooth simple, blunt at apex, lateral lobes wide, gently and obliquely truncated; submentum narrow, quadrisetose. Antennae filiform, extended to scutellum; antennomeres 1 unisetose at subapex, antennomere 2 asetose, pubescent from antennomere 3; antennomeres 2 and 3 subequal in length; antennomeres 510 subcylindrical, somehow flat, evidently longer than wide.</paragraph>
<paragraph id="3A7A36A3FFB53B1BFF3AFDBC8B2CC788" blockId="3.[151,1437,151,933]" pageId="3" pageNumber="358">
Pronotum peltate in form, much wider than head; disc smooth, moderately convex; slightly wider than long, PL/PW=0.960.99, mean 0.97; apex deeply concave, beaded in median portion, fore angles strongly protruding, sharper than in
<taxonomicName id="FDC54D20FFB53B1BFEBBFD658FA4C6E1" box="[326,452,692,716]" class="Insecta" family="Carabidae" genus="Guiodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="358" phylum="Arthropoda" rank="species" species="cavicola">
<emphasis id="08B1EAB1FFB53B1BFEBBFD658FA4C6E1" box="[326,452,692,716]" italics="true" pageId="3" pageNumber="358">G. cavicola</emphasis>
</taxonomicName>
; widest at about middle, gently and gradually narrowed forwards to apex, and strongly contracted towards basal angles; side with three conspicuous teethed projections before hind angle; hind angle obtuse; two lateral setiferous pores present, anterior seta at about apical quarter, posterior one at about basal third, just before anterior lateral projection; marginal channel wide and uneven, narrower before anterior seta, wider between anterior and posterior lateral setae, ended before posterior seta; base unbordered; median line clear and deep; basal transverse impression deep and wide, rugose, and faintly carinated posteriorly; anterior transversal line free from median line, very close to anterior margin. Peduncle short, base of scutellum visible and rounded.
</paragraph>
<caption id="6EBA662BFFB53B1BFF6AF8748F41C3F7" httpUri="https://zenodo.org/record/225328/files/figure.png" pageId="3" pageNumber="358" targetBox="[201,1398,1011,1912]" targetPageId="3">
<paragraph id="3A7A36A3FFB53B1BFF6AF8748F41C3F7" blockId="3.[151,1436,1957,2010]" pageId="3" pageNumber="358">
<emphasis id="08B1EAB1FFB53B1BFF6AF8748F5AC396" bold="true" box="[151,314,1957,1980]" pageId="3" pageNumber="358">FIGURES 23.</emphasis>
Male genitalia of
<taxonomicName id="FDC54D20FFB53B1BFE0EF8778CF8C391" authority="Tian" authorityName="Tian" box="[499,664,1958,1980]" class="Insecta" family="Carabidae" genus="Guiodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="358" phylum="Arthropoda" rank="species" species="cavicola">
<emphasis id="08B1EAB1FFB53B1BFE0EF8778C05C396" box="[499,613,1958,1979]" italics="true" pageId="3" pageNumber="358">G. cavicola</emphasis>
Tian
</taxonomicName>
(2. median lobe, lateral view; 3. right and left parameres, lateral view. Scale bar: 0.5 mm).
</paragraph>
</caption>
<caption id="6EBA662BFFB23B1CFF6AF8728FF2C3F5" httpUri="https://zenodo.org/record/225329/files/figure.png" pageId="4" pageNumber="359" targetBox="[321,1262,199,1925]" targetPageId="4">
<paragraph id="3A7A36A3FFB23B1CFF6AF8728FF2C3F5" blockId="4.[151,1436,1955,2008]" pageId="4" pageNumber="359">
<emphasis id="08B1EAB1FFB23B1CFF6AF8728F5EC394" bold="true" box="[151,318,1955,1978]" pageId="4" pageNumber="359">FIGURES 46.</emphasis>
<taxonomicName id="FDC54D20FFB23B1CFEB5F8758C4BC394" box="[328,555,1956,1977]" class="Insecta" family="Carabidae" genus="Guiodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="359" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
<emphasis id="08B1EAB1FFB23B1CFEB5F8758C4BC394" box="[328,555,1956,1977]" italics="true" pageId="4" pageNumber="359">Guiodytes deharvengi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="138257CAFFB23B1CFDC8F8758C12C394" box="[565,626,1956,1977]" pageId="4" pageNumber="359" rank="species">
<emphasis id="08B1EAB1FFB23B1CFDC8F8758C12C394" bold="true" box="[565,626,1956,1977]" pageId="4" pageNumber="359">n. sp.</emphasis>
</taxonomicNameLabel>
(4. habitus, female, paratype; 5. median lobe, lateral view; 6. right and left parameres, lateral view).
</paragraph>
</caption>
<caption id="6EBA662BFFB33B1DFF6AF8738DF1C395" box="[151,913,1954,1976]" httpUri="https://zenodo.org/record/225330/files/figure.png" pageId="5" pageNumber="360" targetBox="[240,1331,256,1838]" targetPageId="5">
<paragraph id="3A7A36A3FFB33B1DFF6AF8738DF1C395" blockId="5.[151,913,1954,1976]" box="[151,913,1954,1976]" pageId="5" pageNumber="360">
<emphasis id="08B1EAB1FFB33B1DFF6AF8738F72C39A" bold="true" box="[151,274,1954,1976]" pageId="5" pageNumber="360">FIGURE 7.</emphasis>
Female genitalia of
<taxonomicName id="FDC54D20FFB33B1DFE1EF8738CA2C39A" box="[483,706,1954,1975]" class="Insecta" family="Carabidae" genus="Guiodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="360" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
<emphasis id="08B1EAB1FFB33B1DFE1EF8738CA2C39A" box="[483,706,1954,1975]" italics="true" pageId="5" pageNumber="360">Guiodytes deharvengi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="138257CAFFB33B1DFD35F8738D61C39A" box="[712,769,1954,1975]" pageId="5" pageNumber="360" rank="species">
<emphasis id="08B1EAB1FFB33B1DFD35F8738D61C39A" bold="true" box="[712,769,1954,1975]" pageId="5" pageNumber="360">n. sp.</emphasis>
</taxonomicNameLabel>
, ventral view.
</paragraph>
</caption>
<paragraph id="3A7A36A3FFB03B1EFF3AFF468D91C5FD" blockId="6.[151,1437,151,1293]" pageId="6" pageNumber="361">Elytra elongate ovate, wider than pronotum, EW/PW=1.341.35, mean 1.35, much longer than wide, EL/ EW=1.761.79, mean 1.78; strongly convex; widest at about middle, gently contracted anteriad and posteriad, base finely bisinuate; shoulders broadly obtuse, with two large serrated projections; apex pointed; side not parallelsided, distinctly crenulated from base to apical quarter; elytral striae punctate-striate, striae deep, intervals strongly convex; stria 1 with a small blunt tubercle at the base, just before scutellar pore; intervals 14 unbordered at base, other bordered at base; intervals 7 and 8 jointed before base, then jointed to interval 6 at base, all of them carinated near base; interval 3 with five foveolate setiferous pores at 8/9, 2/7, 1/2, 2/3 and 5/6 from base respectively; marginal channel with uninterrupted series of small setiferous pores and several large pores which bearing much longer setae; scutellar striae and scutellar pores present; hind wings reduced.</paragraph>
<paragraph id="3A7A36A3FFB03B1EFF3AFE0D8D90C6A9" blockId="6.[151,1437,151,1293]" pageId="6" pageNumber="361">Underside of head covered with coarsely rugose and short wrinkles, proepisternum with dense isodiametric punctures, without wrinkle; pro-, meso- and metasterna smooth in middle portion, covered with small and sparse punctures at sides, pro-, meso- and meteepisterna coarsely punctured, epipleuron with a few coarser punctures near base; abdominal ventrites sparsely punctured at sides, each of ventrites IVVI bearing a pair of paramedial setae, and ventrite VII of both male and female with two pairs of paramedial setae.</paragraph>
<paragraph id="3A7A36A3FFB03B1EFF3AFD418C8CC738" blockId="6.[151,1437,151,1293]" pageId="6" pageNumber="361">Legs rather stout and short; fore leg stout, profemur simple, moderately dilated; protibia well developed, quadridentate on outer margin, dorsal carina and sulcus distinct; lateral upper spine elongate ensiform, blunt at apex, much longer and stouter than subapical spur; protarsi short, tarsomere 1 much longer than other, tarsomeres 24 flat and transverse; middle and hind legs slender.</paragraph>
<paragraph id="3A7A36A3FFB03B1EFF3AFCCE8B09C752" blockId="6.[151,1437,151,1293]" pageId="6" pageNumber="361">
Male genitalia (
<figureCitation id="A2FE2A26FFB03B1EFE87FCCE8F89C715" box="[378,489,799,824]" captionStart="FIGURES 4 6" captionStartId="4.[151,264,1955,1978]" captionTargetBox="[321,1262,199,1925]" captionTargetId="figure@4.[311,1275,193,1934]" captionTargetPageId="4" captionText="FIGURES 4 6. Guiodytes deharvengi n. sp. (4. habitus, female, paratype; 5. median lobe, lateral view; 6. right and left parameres, lateral view)." httpUri="https://zenodo.org/record/225329/files/figure.png" pageId="6" pageNumber="361">Figs. 56</figureCitation>
). Weakly sclerotized; median lobe of aedeagus stouter than in
<taxonomicName id="FDC54D20FFB03B1EFB4FFCF18B51C715" box="[1202,1329,800,824]" class="Insecta" family="Carabidae" genus="Guiodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="361" phylum="Arthropoda" rank="species" species="cavicola">
<emphasis id="08B1EAB1FFB03B1EFB4FFCF18B51C715" box="[1202,1329,800,824]" italics="true" pageId="6" pageNumber="361">G. cavicola</emphasis>
</taxonomicName>
, strongly arcuated, ventral margin unisinuate, gently enlarged towards apex which is very broad; basal orifice small; inner sac armed with longitudinal copulatory piece, apical one longer and broader; parameres similar to
<taxonomicName id="FDC54D20FFB03B1EFB19FCB98B02C7AD" box="[1252,1378,872,896]" class="Insecta" family="Carabidae" genus="Guiodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="361" phylum="Arthropoda" rank="species" species="cavicola">
<emphasis id="08B1EAB1FFB03B1EFB19FCB98B02C7AD" box="[1252,1378,872,896]" italics="true" pageId="6" pageNumber="361">G. cavicola</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="3A7A36A3FFB03B1EFF3AFC5D8C3BC7C0" blockId="6.[151,1437,151,1293]" pageId="6" pageNumber="361">
Female genitalia (
<figureCitation id="A2FE2A26FFB03B1EFE6CFC5D8FB5C789" box="[401,469,908,933]" captionStart="FIGURE 7" captionStartId="5.[151,250,1954,1976]" captionTargetBox="[240,1331,256,1838]" captionTargetId="figure@5.[165,1421,193,1932]" captionTargetPageId="5" captionText="FIGURE 7. Female genitalia of Guiodytes deharvengi n. sp., ventral view." httpUri="https://zenodo.org/record/225330/files/figure.png" pageId="6" pageNumber="361">Fig. 7</figureCitation>
). Similar to
<taxonomicName id="FDC54D20FFB03B1EFD99FC5D8D02C789" authority="Tian, 2009" authorityName="Tian" authorityYear="2009" box="[612,866,908,933]" class="Insecta" family="Carabidae" genus="Guiodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="361" phylum="Arthropoda" rank="species" species="cavicola">
<emphasis id="08B1EAB1FFB03B1EFD99FC5D8C82C789" box="[612,738,908,932]" italics="true" pageId="6" pageNumber="361">G. cavicola</emphasis>
<bibRefCitation id="5E544B52FFB03B1EFD14FC5D8D02C789" author="Tian" box="[745,866,908,933]" pageId="6" pageNumber="361" refString="Tian, M. Y. (2009) New records and new species of cavedwelling trechine beetles from Mulun Nature Reserve, northern Guangxi, China (Insecta: Coleoptera: Carabidae: Trechinae). Subterranean Biology, 7, 69 - 73." type="journal article" year="2009">Tian, 2009</bibRefCitation>
</taxonomicName>
though gonocoxite and gonosubcoxite are slightly more slender; bursa copulatrix wide and folded, basally narrower; spermathecal gland very long and large, spermathecal gland duct wide and short.
</paragraph>
</subSubSection>
<subSubSection id="72DF6528FFB03B1EFF3AFC268FFBC018" pageId="6" pageNumber="361" type="etymology">
<paragraph id="3A7A36A3FFB03B1EFF3AFC268FFBC018" blockId="6.[151,1437,151,1293]" pageId="6" pageNumber="361">
<emphasis id="08B1EAB1FFB03B1EFF3AFC268F2EC03D" bold="true" box="[199,334,1015,1040]" pageId="6" pageNumber="361">Etymology.</emphasis>
The name of this new species is dedicated to Prof. Dr. Louis Deharveng (MNHN, Paris), a wellknown biospeleologist.
</paragraph>
</subSubSection>
<subSubSection id="72DF6528FFB03B1EFF3AFBEE8DA7C08D" pageId="6" pageNumber="361" type="discussion">
<paragraph id="3A7A36A3FFB03B1EFF3AFBEE8DA7C08D" blockId="6.[151,1437,151,1293]" pageId="6" pageNumber="361">
<emphasis id="08B1EAB1FFB03B1EFF3AFBEE8F5BC075" bold="true" box="[199,315,1087,1112]" pageId="6" pageNumber="361">Remarks.</emphasis>
<taxonomicName id="FDC54D20FFB03B1EFEB9FB918F84C075" box="[324,484,1088,1112]" class="Insecta" family="Carabidae" genus="Guiodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="361" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
<emphasis id="08B1EAB1FFB03B1EFEB9FB918F84C075" box="[324,484,1088,1112]" italics="true" pageId="6" pageNumber="361">G. deharvengi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="138257CAFFB03B1EFE13FBEE8C4FC075" box="[494,559,1087,1112]" pageId="6" pageNumber="361" rank="species">
<emphasis id="08B1EAB1FFB03B1EFE13FBEE8C4FC075" bold="true" box="[494,559,1087,1112]" pageId="6" pageNumber="361">n. sp.</emphasis>
</taxonomicNameLabel>
is similar to
<taxonomicName id="FDC54D20FFB03B1EFD30FB918D2DC075" box="[717,845,1088,1112]" class="Insecta" family="Carabidae" genus="Guiodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="361" phylum="Arthropoda" rank="species" species="cavicola">
<emphasis id="08B1EAB1FFB03B1EFD30FB918D2DC075" box="[717,845,1088,1112]" italics="true" pageId="6" pageNumber="361">G. cavicola</emphasis>
</taxonomicName>
, but it is easily distinguished from the latter by its depigmented and stouter body, as well as its broader pronotum. Furthermore, elytra interval 3 bearing five dorsal setiferous pores in
<taxonomicName id="FDC54D20FFB03B1EFE96FB598C67C08D" box="[363,519,1160,1184]" class="Insecta" family="Carabidae" genus="Guiodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="361" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
<emphasis id="08B1EAB1FFB03B1EFE96FB598C67C08D" box="[363,519,1160,1184]" italics="true" pageId="6" pageNumber="361">G. deharvengi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="138257CAFFB03B1EFDF2FB568C2DC08D" box="[527,589,1159,1184]" pageId="6" pageNumber="361" rank="species">
<emphasis id="08B1EAB1FFB03B1EFDF2FB568C2DC08D" bold="true" box="[527,589,1159,1184]" pageId="6" pageNumber="361">n. sp.</emphasis>
</taxonomicNameLabel>
(while four pores in
<taxonomicName id="FDC54D20FFB03B1EFCC4FB598DD7C08D" box="[825,951,1160,1184]" class="Insecta" family="Carabidae" genus="Guiodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="361" phylum="Arthropoda" rank="species" species="cavicola">
<emphasis id="08B1EAB1FFB03B1EFCC4FB598DD7C08D" box="[825,951,1160,1184]" italics="true" pageId="6" pageNumber="361">G. cavicola</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="72DF6528FFB03B1EFF3AFB7D8C01C121" pageId="6" pageNumber="361" type="distribution">
<paragraph id="3A7A36A3FFB03B1EFF3AFB7D8C01C121" blockId="6.[151,1437,151,1293]" pageId="6" pageNumber="361">
<emphasis id="08B1EAB1FFB03B1EFF3AFB7D8F00C0E8" bold="true" box="[199,352,1196,1221]" pageId="6" pageNumber="361">Distribution.</emphasis>
<collectingCountry id="42D27633FFB03B1EFE9AFB7D8FCAC0E8" box="[359,426,1196,1221]" name="China" pageId="6" pageNumber="361">China</collectingCountry>
(Guangxi: Huangjiang County) (
<figureCitation id="A2FE2A26FFB03B1EFCE0FB7D8D3EC0E9" box="[797,862,1196,1221]" captionStart="FIGURE 1" captionStartId="1.[151,250,1414,1436]" captionTargetBox="[151,1435,411,1392]" captionTargetId="figure@1.[151,1435,411,1392]" captionTargetPageId="1" captionText="FIGURE 1. Distributional map of the genus Guiodytes Tian, 2013 (a. showing the location of Guangxi within China; square: G. b e d o s a e Tian; star: G. cavicola Tian; round: G. deharvengi n. sp.)" httpUri="https://zenodo.org/record/225327/files/figure.png" pageId="6" pageNumber="361">Fig. 1</figureCitation>
). Known only from Shuiku Cave, the
<typeStatus id="E57E8801FFB03B1EFAF1FB7C8B5CC0E8" box="[1292,1340,1197,1221]" pageId="6" pageNumber="361">type</typeStatus>
locality. In this cave are living also two blind trechines:
<taxonomicName id="FDC54D20FFB03B1EFD52FB018AF6C0C5" authority="Deuve &amp; Tian, 1999" authorityName="Deuve &amp; Tian" authorityYear="1999" box="[687,1174,1231,1256]" class="Insecta" family="Carabidae" genus="Uenotrechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="361" phylum="Arthropoda" rank="species" species="liboensis">
<emphasis id="08B1EAB1FFB03B1EFD52FB018DC9C0C5" box="[687,937,1232,1256]" italics="true" pageId="6" pageNumber="361">Uenotrechus liboensis</emphasis>
<bibRefCitation id="5E544B52FFB03B1EFC52FB008AF6C0C5" author="Deuve" box="[943,1174,1231,1256]" pageId="6" pageNumber="361" refString="Deuve, T. Tian, M. Y., Ran, J. C. (1999) Trois Caraboidea remarquables de la reserve nationale de Maolan, dans le Guizhou meridional, Chine (Coleoptera, Carabidae et Trechidae). Revue francaise d'Entomologie, New Series, 21, 131 - 138." type="journal article" year="1999">Deuve &amp; Tian, 1999</bibRefCitation>
</taxonomicName>
(in
<bibRefCitation id="5E544B52FFB03B1EFB39FB008BF4C0C5" author="Deuve" box="[1220,1428,1231,1256]" pageId="6" pageNumber="361" refString="Deuve, T. Tian, M. Y., Ran, J. C. (1999) Trois Caraboidea remarquables de la reserve nationale de Maolan, dans le Guizhou meridional, Chine (Coleoptera, Carabidae et Trechidae). Revue francaise d'Entomologie, New Series, 21, 131 - 138." type="journal article" year="1999">Deuve et al., 1999</bibRefCitation>
) and
<emphasis id="08B1EAB1FFB03B1EFF3BFB258FBAC121" box="[198,474,1268,1292]" italics="true" pageId="6" pageNumber="361">Pilosaphaenops whitteni</emphasis>
<bibRefCitation id="5E544B52FFB03B1EFE1CFB258C3EC121" author="Tian" box="[481,606,1268,1293]" pageId="6" pageNumber="361" refString="Tian, M. Y. (2009) New records and new species of cavedwelling trechine beetles from Mulun Nature Reserve, northern Guangxi, China (Insecta: Coleoptera: Carabidae: Trechinae). Subterranean Biology, 7, 69 - 73." type="journal article" year="2009">Tian, 2009</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>