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<document id="6A34CC08E49D243D313636F487D3B53D" ID-DOI="10.5281/zenodo.178410" ID-GBIF-Dataset="ba2b7965-13f3-401f-9c5a-1705938112ea" ID-ISSN="1175-5326" ID-Zenodo-Dep="178410" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459893524754" checkinUser="plazi" docAuthor="Asiain, Julieta, Márquez, Juan &amp; Morrone, Juan J." docDate="2007" docId="965DBB53FFC75A782399FF16770B55DA" docLanguage="en" docName="zt01584p053.pdf" docOrigin="Zootaxa 1584" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Plochionocerus ashei Asiain, Marquez &amp; Morrone, sp. nov." docType="treatment" docVersion="8" lastPageNumber="13" masterDocId="6A64C32BFFCB5A74230EFF8175415026" masterDocTitle="Phylogenetic systematics of the genera Plochionocerus Dejean and Agrodes Nordmann (Coleoptera: Staphylinidae: Xantholinini)" masterLastPageNumber="53" masterPageNumber="1" pageNumber="13" updateTime="1698228284375" updateUser="plazi">
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<mods:title id="7E0160907DE08A401753A5B9BA6B185E">Phylogenetic systematics of the genera Plochionocerus Dejean and Agrodes Nordmann (Coleoptera: Staphylinidae: Xantholinini)</mods:title>
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<mods:namePart id="B117C0DA4E7335DC86564061568846A5">Asiain, Julieta</mods:namePart>
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<mods:namePart id="1F131B8BA4C3427C15146B77D6C724F4">Márquez, Juan</mods:namePart>
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<mods:namePart id="3DDA5B831D2CD31B2987D17796F0252E">Morrone, Juan J.</mods:namePart>
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<mods:date id="5C1AC899E05AEDC78BA828D8C276F3C8">2007</mods:date>
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<treatment id="965DBB53FFC75A782399FF16770B55DA" ID-DOI="http://doi.org/10.5281/zenodo.6246763" ID-GBIF-Taxon="119372282" ID-Zenodo-Dep="6246763" LSID="urn:lsid:plazi:treatment:965DBB53FFC75A782399FF16770B55DA" httpUri="http://treatment.plazi.org/id/965DBB53FFC75A782399FF16770B55DA" lastPageNumber="13" pageId="12" pageNumber="13">
<subSubSection id="56EE59CEFFC75A782399FF1676355097" box="[151,884,151,177]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="1E4B0A45FFC75A782399FF1676355097" blockId="12.[151,884,151,177]" box="[151,884,151,177]" pageId="12" pageNumber="13">
<heading id="4503BD29FFC75A782399FF1676355097" bold="true" box="[151,884,151,177]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="2C80D657FFC75A782399FF1676355097" bold="true" box="[151,884,151,177]" pageId="12" pageNumber="13">
<taxonomicName id="D9F471C6FFC75A782399FF1676485097" ID-CoL="6W3NN" authority="Asiain, Marquez &amp; Morrone" authorityName="Asiain, Marquez &amp; Morrone" box="[151,777,151,177]" class="Insecta" family="Staphylinidae" genus="Plochionocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ashei" status="sp. nov.">
<emphasis id="2C80D657FFC75A782399FF1674DA5097" bold="true" box="[151,411,151,177]" italics="true" pageId="12" pageNumber="13">Plochionocerus ashei</emphasis>
Asiain, Márquez &amp; Morrone
</taxonomicName>
,
<taxonomicNameLabel id="37B36B2CFFC75A782019FF1676355097" box="[791,884,151,177]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="56EE59CEFFC75A782399FF63744651BA" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="1E4B0A45FFC75A782399FF63744651BA" blockId="12.[151,1437,226,1532]" pageId="12" pageNumber="13">
<emphasis id="2C80D657FFC75A782399FF63740550DA" bold="true" box="[151,324,226,252]" pageId="12" pageNumber="13">
<typeStatus id="C14FB4E7FFC75A782399FF63759350DA" box="[151,210,226,252]" pageId="12" pageNumber="13">Type</typeStatus>
material
</emphasis>
(
<specimenCount id="08F2C1CCFFC75A782256FF6374AF50DA" box="[344,494,226,252]" pageId="12" pageNumber="13" type="generic">2 specimens</specimenCount>
).
<typeStatus id="C14FB4E7FFC75A78210DFF63773450DA" box="[515,629,226,252]" pageId="12" pageNumber="13" type="holotype">
<emphasis id="2C80D657FFC75A78210DFF63773450DA" bold="true" box="[515,629,226,252]" pageId="12" pageNumber="13">Holotype</emphasis>
</typeStatus>
, male: “
<collectingCountry id="66E34AD5FFC75A7821D6FF6376CC50DA" box="[728,909,226,252]" name="Venezuela" pageId="12" pageNumber="13">VENEZUELA</collectingCountry>
, Aragua, Rancho Grande Biological Station, Pico Periquitos,
<quantity id="D90CA7A0FFC75A782257FE8B74F75102" box="[345,438,266,292]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="13" unit="m" value="1300.0">1300 m</quantity>
,
<geoCoordinate id="7BC06C82FFC75A7822CDFE8B77075102" box="[451,582,266,292]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="10.35">10°210”N</geoCoordinate>
,
<geoCoordinate id="7BC06C82FFC75A78215DFE8B779B5102" box="[595,730,266,292]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-67.683334">67°410”W</geoCoordinate>
,
<date id="6A4A2C85FFC75A7821E9FE8B76235102" box="[743,866,266,292]" pageId="12" pageNumber="13" value="1998-05-13">13.V.1998</date>
, R. Anderson, VEN1A98 0 0 51, ex: cloud forest litter /
<typeStatus id="C14FB4E7FFC75A7823E9FEB37415516A" box="[231,340,306,332]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
<taxonomicName id="D9F471C6FFC75A782252FEB37141516A" authority="Asiain, Marquez &amp; Morrone, 2007" authorityName="Asiain, Marquez &amp; Morrone" authorityYear="2007" box="[348,1024,306,332]" class="Insecta" family="Staphylinidae" genus="Plochionocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ashei">
<emphasis id="2C80D657FFC75A782252FEB3771B516D" box="[348,602,306,331]" italics="true" pageId="12" pageNumber="13">Plochionocerus ashei</emphasis>
Asiain, Márquez &amp; Morrone, 2007
</taxonomicName>
” (
<collectionCode id="78E59280FFC75A782712FEB37133516A" box="[1052,1138,306,332]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="12" pageNumber="13">SEMC</collectionCode>
).
<typeStatus id="C14FB4E7FFC75A782788FEB371B9516A" box="[1158,1272,306,332]" pageId="12" pageNumber="13" type="paratype">
<emphasis id="2C80D657FFC75A782788FEB371B9516A" bold="true" box="[1158,1272,306,332]" pageId="12" pageNumber="13">Paratype</emphasis>
</typeStatus>
, male:
<emphasis id="2C80D657FFC75A78265FFEB37021516A" bold="true" box="[1361,1376,306,332]" pageId="12" pageNumber="13"></emphasis>
Ve n., Caracas, D. F., 1933, G. Vivas-B. /
<typeStatus id="C14FB4E7FFC75A782155FEDB77805152" box="[603,705,346,372]" pageId="12" pageNumber="13" type="paratype">Paratype</typeStatus>
<taxonomicName id="D9F471C6FFC75A7821C0FEDB70D05152" authority="Asiain, Marquez &amp; Morrone, 2007" authorityName="Asiain, Marquez &amp; Morrone" authorityYear="2007" box="[718,1425,346,372]" class="Insecta" family="Staphylinidae" genus="Plochionocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ashei">
<emphasis id="2C80D657FFC75A7821C0FEDB76935155" box="[718,978,346,371]" italics="true" pageId="12" pageNumber="13">Plochionocerus ashei</emphasis>
Asiain, Márquez &amp; Morrone, 2007
</taxonomicName>
” (
<collectionCode id="78E59280FFC75A782391FE0375BA51BA" LSID="urn:lsid:biocol.org:col:34795" box="[159,251,386,412]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="12" pageNumber="13">FMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="56EE59CEFFC75A7823C9FE2B77DA55A2" pageId="12" pageNumber="13" type="description">
<paragraph id="1E4B0A45FFC75A7823C9FE2B710F51E2" blockId="12.[151,1437,226,1532]" box="[199,1102,426,452]" pageId="12" pageNumber="13">
<emphasis id="2C80D657FFC75A7823C9FE2B741E51E2" bold="true" box="[199,351,426,452]" pageId="12" pageNumber="13">Description.</emphasis>
Total length
<quantity id="D90CA7A0FFC75A7822F4FE2B77E951E2" box="[506,680,426,452]" metricMagnitude="-2" metricUnit="m" metricValue="1.76" metricValueMax="1.81" metricValueMin="1.71" pageId="12" pageNumber="13" unit="mm" value="17.6" valueMax="18.1" valueMin="17.1">17.118.1 mm</quantity>
. Body metallic green or blue-violet.
</paragraph>
<paragraph id="1E4B0A45FFC75A7823C9FE5376FD52FA" blockId="12.[151,1437,226,1532]" pageId="12" pageNumber="13">
<emphasis id="2C80D657FFC75A7823C9FE53744751CD" box="[199,262,466,491]" italics="true" pageId="12" pageNumber="13">Head</emphasis>
. Rounded; longer than wide (1.191.30 times), almost as long as pronotum, wider than pronotum (1.32 times); dorsal surface slightly convex; ventral surface flat, with slightly dense, expanded, umbilicate punctures (
<quantity id="D90CA7A0FFC75A782214FDA374C4521A" box="[282,389,546,572]" metricMagnitude="-1" metricUnit="m" metricValue="3.683" metricValueMax="4.826" metricValueMin="2.54" pageId="12" pageNumber="13" unit="in" value="14.5" valueMax="19.0" valueMin="10.0">1019 in</quantity>
each half of the head; similar to
<figureCitation id="86CF16C0FFC75A78201BFDA3762F521A" box="[789,878,546,572]" captionStart="FIGURES 48 53" captionStartId="37.[151,269,1439,1463]" captionTargetBox="[172,1429,211,1390]" captionTargetId="figure@37.[151,1436,194,1415]" captionTargetPageId="37" captionText="FIGURES 48 53. Cephalic ventral view: 48, Renda flagellicornis; 49, Philonthus testaceipennis; 50, Plochionocerus simplicicollis; 51, P. gracilis; 52, P. h e r m a n i; 53, P. puncticeps." httpUri="https://zenodo.org/record/178418/files/figure.png" pageId="12" pageNumber="13">Fig. 52</figureCitation>
), arranged in “v”; first antennomere 1.581.60 times as long as antennomeres 2+3 combined, second antennomere 0.670.73 times as long as third antennomere, antennomeres 410 moderately transverse, gradually larger toward apex, apical antennomere moderately longer than wide (1.26 times), shorter than antennomeres 9+10 combined (0.92 times its length; similar to
<figureCitation id="86CF16C0FFC75A7823BBFD43744A52FA" box="[181,267,706,732]" captionStart="FIGURES 20 31" captionStartId="34.[151,269,1364,1388]" captionTargetBox="[370,1190,207,1330]" captionTargetId="figure@34.[365,1222,194,1340]" captionTargetPageId="34" captionText="FIGURES 20 31. Antennae: 20, Philonthus testaceipennis; 21, Renda flagellicornis; 22, Agrodes conicicollis; 23, A. elegans; 24, Plochionocerus gracilis; 25, P. humeralis; 26, P. i g n e u s; 27, P. m a rq u e z i; 28, P. puncticeps; 29, P. re t i c u - laris; 30, P. s p l e n d e n s; 31, P. transversalis." httpUri="https://zenodo.org/record/178415/files/figure.png" pageId="12" pageNumber="13">Fig. 28</figureCitation>
); mandible with basal external channel slightly developed.
</paragraph>
<paragraph id="1E4B0A45FFC75A7823C9FD6B77515372" blockId="12.[151,1437,226,1532]" pageId="12" pageNumber="13">
<emphasis id="2C80D657FFC75A7823C9FD6B745D5325" box="[199,284,746,771]" italics="true" pageId="12" pageNumber="13">Thorax</emphasis>
. Pronotum 1.481.49 times as long as wide, shorter than elytra (0.850.91 times); with visible microsculpture; with two clearly visible depressed areas in posterior third (
<figureCitation id="86CF16C0FFC75A78270FFC937119530A" box="[1025,1112,786,812]" captionStart="FIGURES 54 62" captionStartId="38.[151,269,1902,1926]" captionTargetBox="[223,1368,197,1867]" captionTargetId="figure@38.[218,1370,194,1878]" captionTargetPageId="38" captionText="FIGURES 54 62. Pronotum: 54, Philonthus testaceipennis; 55, Renda flagellicornis; 56, Plochionocerus simplicicollis; 57, P. reticularis. Elytra: 58, Agrodes spp.; 59, Plochionocerus spp. Midleg: 60, Philonthus testaceipennis; 61, Agrodes spp.; 62, Plochionocerus spp." httpUri="https://zenodo.org/record/178419/files/figure.png" pageId="12" pageNumber="13">Fig. 56</figureCitation>
). Prosternum slightly transverse (length/width: 0.830.84).
</paragraph>
<paragraph id="1E4B0A45FFC75A7823C9FCE3741B53EA" blockId="12.[151,1437,226,1532]" pageId="12" pageNumber="13">
<emphasis id="2C80D657FFC75A7823C9FCE37478535D" box="[199,313,866,891]" italics="true" pageId="12" pageNumber="13">Aedeagus</emphasis>
. Ovate, length
<quantity id="D90CA7A0FFC75A7822E0FCE377DD535A" box="[494,668,866,892]" metricMagnitude="-3" metricUnit="m" metricValue="2.485" metricValueMax="2.6" metricValueMin="2.37" pageId="12" pageNumber="13" unit="mm" value="2.485" valueMax="2.6" valueMin="2.37">2.372.60 mm</quantity>
, with parameres 0.370.38 times as long as the median lobe, apical area of median lobe 0.320.36 times as long as the total length of median lobe, and internal sac moderately visible (
<figureCitation id="86CF16C0FFC75A7823FBFC33740C53EA" box="[245,333,946,972]" captionStart="FIGURES 69 78" captionStartId="41.[151,269,1936,1960]" captionTargetBox="[220,1361,186,1909]" captionTargetId="figure@41.[214,1372,174,1912]" captionTargetPageId="41" captionText="FIGURES 69 78. Ventral (left) and lateral (right) view of aedeagus: 69, Philonthus testaceipennis; 70, Homalolinus flavipennis; 71, Thyreocephalus puncticeps; 72, Renda fimetaria; 73, R. flagellicornis; 74, R. leprieuri; 75, Agrodes conicicollis; 76, A. elegans; 77, Plochionocerus ashei; 78, P. discedens. Scale bar 1 mm." httpUri="https://zenodo.org/record/178422/files/figure.png" pageId="12" pageNumber="13">Fig. 77</figureCitation>
).
</paragraph>
<paragraph id="1E4B0A45FFC75A7823C9FC5B77DB54B2" blockId="12.[151,1437,226,1532]" pageId="12" pageNumber="13">
<emphasis id="2C80D657FFC75A7823C9FC5B740553D2" bold="true" box="[199,324,986,1012]" pageId="12" pageNumber="13">Variation.</emphasis>
The
<typeStatus id="C14FB4E7FFC75A782271FC5B74A453D2" box="[383,485,986,1012]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
is metallic green, the head is short (length/width: 1.19), the expanded, umbilicate punctures on the ventral surface of the head are less dense (1014), and the external mandibular channel is slightly developed. The color of the
<typeStatus id="C14FB4E7FFC75A782131FBAB77E55462" box="[575,676,1066,1092]" pageId="12" pageNumber="13" type="paratype">paratype</typeStatus>
is metallic blue-violet, the head is longer (length/width: 1.3), the expanded, umbilicate punctures on the ventral surface of the head are denser (1519), and the external mandibular channel is more strongly developed.
</paragraph>
<paragraph id="1E4B0A45FFC75A7823C9FB23760F557A" blockId="12.[151,1437,226,1532]" pageId="12" pageNumber="13">
<emphasis id="2C80D657FFC75A7823C9FB2374FF549A" bold="true" box="[199,446,1186,1212]" pageId="12" pageNumber="13">Comparative notes.</emphasis>
The species may be confused with other species with a rounded, dorsally convex head, but only in
<taxonomicName id="D9F471C6FFC75A782268FB4A74B754C2" box="[358,502,1227,1252]" class="Insecta" family="Staphylinidae" genus="Plochionocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="discedens">
<emphasis id="2C80D657FFC75A782268FB4A74B754C2" box="[358,502,1227,1252]" italics="true" pageId="12" pageNumber="13">P. discedens</emphasis>
</taxonomicName>
is the ventral surface of the head flat. The two species can be distinguished by the greater density of expanded, umbilicate punctures, the presence of a basal mandibular channel, and the shorter pronotum in
<emphasis id="2C80D657FFC75A782285FA9A77595512" box="[395,536,1307,1332]" italics="true" pageId="12" pageNumber="13">P. d i s c e d e n s</emphasis>
; whereas in
<taxonomicName id="D9F471C6FFC75A7821BAFA9A76515512" box="[692,784,1307,1332]" class="Insecta" family="Staphylinidae" genus="Plochionocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ashei">
<emphasis id="2C80D657FFC75A7821BAFA9A76515512" box="[692,784,1307,1332]" italics="true" pageId="12" pageNumber="13">P. ashei</emphasis>
</taxonomicName>
the expanded, umbilicate punctures are less dense, the mandibular channel is reduced, and the pronotum is longer.
</paragraph>
<paragraph id="1E4B0A45FFC75A7823C9FAEB77DA55A2" blockId="12.[151,1437,226,1532]" box="[199,667,1386,1412]" pageId="12" pageNumber="13">
<emphasis id="2C80D657FFC75A7823C9FAEB775255A2" bold="true" box="[199,531,1386,1412]" pageId="12" pageNumber="13">Geographical distribution.</emphasis>
<collectingCountry id="66E34AD5FFC75A782114FAEB77D755A2" box="[538,662,1386,1412]" name="Venezuela" pageId="12" pageNumber="13">Venezuela</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="56EE59CEFFC75A7823C9FA13770B55DA" pageId="12" pageNumber="13" type="etymology">
<paragraph id="1E4B0A45FFC75A7823C9FA13770B55DA" blockId="12.[151,1437,226,1532]" pageId="12" pageNumber="13">
<emphasis id="2C80D657FFC75A7823C9FA137415558A" bold="true" box="[199,340,1426,1452]" pageId="12" pageNumber="13">Etymology.</emphasis>
We take pleasure in dedicating this species to the late James Steve Ashe (Snow Museum, Kansas University) for his study of Latin American
<taxonomicName id="D9F471C6FFC75A7821F6FA3B76D855F2" box="[760,921,1466,1492]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
and his fieldwork that resulted in several of the specimens analyzed in this paper.
</paragraph>
</subSubSection>
</treatment>
</document>