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<document id="6D25B8DCE8DFFAD7F15A78FE57AB466C" ID-DOI="10.26107/RBZ-2020-0052" ID-ISSN="2345-7600" ID-Zenodo-Dep="4576861" ID-ZooBank="C7D4631B-9B02-4E6E-9665-76CCFECA86A0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614780803714" checkinUser="carolina" docAuthor="Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Buczkó, Krisztina &amp; Vermeulen, Jaap J." docDate="2020" docId="9565D05FFFC3FFC3FFE15A94FED49D39" docLanguage="en" docName="RafflesBZool.68.350-368.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Clostophis obtusus Pall-Gergely &amp; Grego 2020, new species" docType="treatment" docVersion="4" lastPageNumber="364" masterDocId="695CA827FFCDFFCDFF9A5A29FFDE9B6C" masterDocTitle="Clostophis Benson, 1860, is not a monotypic diplommatinid but a speciose hypselostomatid (Gastropoda: Eupulmonata), with descriptions of six new species" masterLastPageNumber="368" masterPageNumber="350" pageNumber="364" updateTime="1698878356270" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="DF353641866823E1FB517E0F3874E2EA">Clostophis Benson, 1860, is not a monotypic diplommatinid but a speciose hypselostomatid (Gastropoda: Eupulmonata), with descriptions of six new species</mods:title>
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<mods:namePart id="BD31B1F1507A87D294A4CBAA52D0806D">Páll-Gergely, Barna</mods:namePart>
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<mods:namePart id="B8574DF66F7D8B97B89780928FFB6914">Hunyadi, András</mods:namePart>
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<mods:namePart id="47FA8D28F911E0212A9EFB2681B2BFE5">Grego, Jozef</mods:namePart>
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<mods:namePart id="9F5EFCE22BD72D66FFFB6E20EA1AEFED">Reischütz, Alexander</mods:namePart>
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<mods:namePart id="AF49D8C6750BD0EAEFEBBE7E410281EE">Buczkó, Krisztina</mods:namePart>
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<mods:namePart id="B487B7804B1D4D5A00CFFA5E5E19B312">Vermeulen, Jaap J.</mods:namePart>
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<mods:date id="EE0B4A7C276B5E405E55E78FC81A6E95">2020</mods:date>
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<treatment id="9565D05FFFC3FFC3FFE15A94FED49D39" ID-DOI="http://doi.org/10.5281/zenodo.4576696" ID-Zenodo-Dep="4576696" LSID="urn:lsid:plazi:treatment:9565D05FFFC3FFC3FFE15A94FED49D39" httpUri="http://treatment.plazi.org/id/9565D05FFFC3FFC3FFE15A94FED49D39" lastPageNumber="364" pageId="14" pageNumber="364">
<subSubSection id="55D632C2FFC3FFC3FFE15A94FD399BB9" box="[123,743,188,213]" pageId="14" pageNumber="364" type="nomenclature">
<paragraph id="1D736149FFC3FFC3FFE15A94FD399BB9" blockId="14.[123,743,188,245]" box="[123,743,188,213]" pageId="14" pageNumber="364">
<heading id="463BD625FFC3FFC3FFE15A94FD399BB9" bold="true" box="[123,743,188,213]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="364" reason="2">
<emphasis id="2FB8BD5BFFC3FFC3FFE15A94FD399BB9" bold="true" box="[123,743,188,213]" pageId="14" pageNumber="364">
<taxonomicName id="DACC1ACAFFC3FFC3FFE15A94FD8A9BB9" ID-CoL="7QQQ5" authority="Pall-Gergely &amp; Grego" authorityName="Pall-Gergely &amp; Grego" authorityYear="2020" box="[123,596,188,213]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="364" phylum="Mollusca" rank="species" species="obtusus" status="sp. nov.">
<emphasis id="2FB8BD5BFFC3FFC3FFE15A94FE959BB9" bold="true" box="[123,331,189,213]" italics="true" pageId="14" pageNumber="364">Clostophis obtusus</emphasis>
Páll-Gergely &amp; Grego
</taxonomicName>
,
<taxonomicNameLabel id="348B0020FFC3FFC3FDF85A94FD399BB9" box="[610,743,189,213]" pageId="14" pageNumber="364" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="55D632C2FFC3FFC3FE1B5AF4FE3E9B99" box="[385,480,221,245]" pageId="14" pageNumber="364" type="description">
<paragraph id="1D736149FFC3FFC3FE1B5AF4FE3E9B99" blockId="14.[123,743,188,245]" box="[385,480,221,245]" pageId="14" pageNumber="364">
(
<figureCitation id="85F77DCCFFC3FFC3FE135AF4FE069B99" box="[393,472,221,245]" captionStart="Fig" captionStartId="15.[135,170,2064,2086]" captionTargetBox="[161,1447,189,2048]" captionTargetId="figure-7@15.[161,1447,189,2048]" captionTargetPageId="15" captionText="Fig. 10. Clostophis obtusus Páll-Gergely &amp; Grego, new species (HNHM 104435, holotype). A, ventral view; B, dorsal view; C, apertural view; D, lateral view; E, protoconch sculpture; F, sculpture of the ventral side; G, sculpture of body whorl from standard apertural view." figureDoi="http://doi.org/10.5281/zenodo.4576884" httpUri="https://zenodo.org/record/4576884/files/figure.png" pageId="14" pageNumber="364">Fig. 10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="55D632C2FFC3FFC3FFEB5B34FD0E9959" pageId="14" pageNumber="364" type="materials_examined">
<paragraph id="1D736149FFC3FFC3FFEB5B34FD0E9959" blockId="14.[112,754,285,565]" pageId="14" pageNumber="364">
<emphasis id="2FB8BD5BFFC3FFC3FFEB5B34FE929A59" bold="true" box="[113,332,285,309]" pageId="14" pageNumber="364">Material examined.</emphasis>
<materialsCitation id="ADA46B14FFC3FFC3FED55B34FDBD9AD9" collectingDate="2017-02-11" collectionCode="HNHM" collectorName="J. Grego" country="Laos" elevation="160" latitude="17.563583" location="Earthquake Dome" longLatPrecision="1" longitude="104.871635" municipality="Tham Nam Don Cave" pageId="14" pageNumber="364" specimenCode="HNHM 104435" specimenCount="1" stateProvince="Khammouan" typeStatus="holotype">
<typeStatus id="C277DFEBFFC3FFC3FED55B34FE6F9A59" box="[335,433,285,309]" pageId="14" pageNumber="364" type="holotype">Holotype</typeStatus>
(SH:
<quantity id="DA34CCACFFC3FFC3FE775B34FD929A59" box="[493,588,285,309]" metricMagnitude="-3" metricUnit="m" metricValue="1.39" pageId="14" pageNumber="364" unit="mm" value="1.39">1.39 mm</quantity>
, SW:
<quantity id="DA34CCACFFC3FFC3FD135B34FD399A59" box="[649,743,285,309]" metricMagnitude="-3" metricUnit="m" metricValue="1.22" pageId="14" pageNumber="364" unit="mm" value="1.22">1.22 mm</quantity>
) (
<specimenCode id="4D6AC932FFC3FFC3FFED5B14FEFC9A39" box="[119,290,317,341]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="14" pageNumber="364">HNHM 104435</specimenCode>
),
<collectingCountry id="65DB21D9FFC3FFC3FEAE5B17FEB49A39" box="[308,362,318,341]" name="Laos" pageId="14" pageNumber="364">Laos</collectingCountry>
,
<collectingRegion id="DF08AFABFFC3FFC3FEE95B14FDB19A39" box="[371,623,317,341]" country="Laos" name="Khammouan" pageId="14" pageNumber="364">Khammouane Province</collectingRegion>
,
<collectingMunicipality id="FD17FB33FFC3FFC3FDE25B14FF3D9A19" pageId="14" pageNumber="364">Tham Nam Dôn Cave</collectingMunicipality>
,
<location id="18133792FFC3FFC3FF6B5B74FE629A19" LSID="urn:lsid:plazi:treatment:9565D05FFFC3FFC3FFE15A94FED49D39:18133792FFC3FFC3FF6B5B74FE629A19" box="[241,444,349,373]" country="Laos" latitude="17.563583" longLatPrecision="1" longitude="104.871635" municipality="Tham Nam Don Cave" name="Earthquake Dome" pageId="14" pageNumber="364" stateProvince="Khammouan">Earthquake Dome</location>
, sand sediments at bank of cave river,
<elevation id="96E1867AFFC3FFC3FF7C5B54FEBE9AF9" box="[230,352,381,405]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="14" pageNumber="364" unit="m" value="160.0">
<quantity id="DA34CCACFFC3FFC3FF7C5B54FEF79AF9" box="[230,297,381,405]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="14" pageNumber="364" unit="m" value="160.0">160 m</quantity>
a.s.l.
</elevation>
,
<geoCoordinate id="78F8078EFFC3FFC3FEF65B55FE289AF9" box="[364,502,380,405]" degrees="17" direction="north" minutes="33.815" orientation="latitude" pageId="14" pageNumber="364" precision="1" value="17.563583">17°33.815N</geoCoordinate>
,
<geoCoordinate id="78F8078EFFC3FFC3FD9A5B55FD4C9AF9" box="[512,658,380,405]" degrees="104" direction="east" minutes="52.298" orientation="longitude" pageId="14" pageNumber="364" precision="1" value="104.871635">104°52.298E</geoCoordinate>
(locality code: JG2B), coll.
<collectorName id="B039049FFFC3FFC3FEA05BB7FE499AD9" box="[314,407,413,437]" pageId="14" pageNumber="364">J. Grego</collectorName>
,
<date id="69724789FFC3FFC3FE385BB4FDBD9AD9" box="[418,611,413,437]" pageId="14" pageNumber="364" value="2017-02-11">
<collectingDate id="7936BE61FFC3FFC3FE385BB4FDBD9AD9" box="[418,611,413,437]" pageId="14" pageNumber="364" value="2017-02-11">11 February 2017</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="ADA46B14FFC3FFC3FDF75BB7FD139959" collectingDate="2017-02-09" collectionCode="HNHM" collectorName="J. Grego" country="Laos" elevation="160" latitude="17.5509" location="Pha Soung Cave" longLatPrecision="1" longitude="104.873566" municipality="Tham Nam Don Cave" pageId="14" pageNumber="364" specimenCount="1" stateProvince="Khammouan" typeStatus="paratype">
<typeStatus id="C277DFEBFFC3FFC3FDF75BB7FD079ADA" box="[621,729,414,438]" pageId="14" pageNumber="364" type="paratype">Paratypes</typeStatus>
: 2 shells (coll. JG), same data as preceding; 1 shell (coll. JG),
<collectingCountry id="65DB21D9FFC3FFC3FFE95BF7FF779A99" box="[115,169,478,501]" name="Laos" pageId="14" pageNumber="364">Laos</collectingCountry>
,
<collectingRegion id="DF08AFABFFC3FFC3FF2F5BF4FE519A99" box="[181,399,477,501]" country="Laos" name="Khammouan" pageId="14" pageNumber="364">Khammouane Prov.</collectingRegion>
,
<locationDeviation id="0CC1786BFFC3FFC3FE015BF4FD339A99" box="[411,749,477,501]" location="Ban Na village" pageId="14" pageNumber="364">
<quantity id="DA34CCACFFC3FFC3FE015BF4FE0D9A99" box="[411,467,477,501]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="364" unit="km" value="2.0">2 km</quantity>
WNW of Ban Na village
</locationDeviation>
,
<location id="18133792FFC3FFC3FFE95BD4FEF59979" LSID="urn:lsid:plazi:treatment:9565D05FFFC3FFC3FFE15A94FED49D39:18133792FFC3FFC3FFE95BD4FEF59979" box="[115,299,509,533]" country="Laos" latitude="17.5509" longLatPrecision="1" longitude="104.873566" name="Pha Soung Cave" pageId="14" pageNumber="364" stateProvince="Khammouan">Pha Soung Cave</location>
Frog lake,
<geoCoordinate id="78F8078EFFC3FFC3FE5C5BD5FD8C9979" box="[454,594,508,533]" degrees="17" direction="north" minutes="33.054" orientation="latitude" pageId="14" pageNumber="364" precision="1" value="17.5509">17°33.054N</geoCoordinate>
,
<geoCoordinate id="78F8078EFFC3FFC3FDC55BD5FD2C9979" box="[607,754,508,533]" degrees="104" direction="east" minutes="52.414" orientation="longitude" pageId="14" pageNumber="364" precision="1" value="104.873566">104°52.414E</geoCoordinate>
(locality code: JG3A), leg.
<collectorName id="B039049FFFC3FFC3FE065837FE249959" box="[412,506,541,565]" pageId="14" pageNumber="364">J. Grego</collectorName>
,
<date id="69724789FFC3FFC3FD9D5834FD139959" box="[519,717,541,565]" pageId="14" pageNumber="364" value="2017-02-09">
<collectingDate id="7936BE61FFC3FFC3FD9D5834FD139959" box="[519,717,541,565]" pageId="14" pageNumber="364" value="2017-02-09">09 February 2017</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="55D632C2FFC3FFC3FFEB5874FE2B99D9" pageId="14" pageNumber="364" type="diagnosis">
<paragraph id="1D736149FFC3FFC3FFEB5874FE2B99D9" blockId="14.[112,753,605,693]" pageId="14" pageNumber="364">
<emphasis id="2FB8BD5BFFC3FFC3FFEB5874FF389919" bold="true" box="[113,230,605,629]" pageId="14" pageNumber="364">Diagnosis.</emphasis>
A relatively large, triangular
<taxonomicName id="DACC1ACAFFC3FFC3FDB05874FD429919" authorityName="Benson" authorityYear="1860" box="[554,668,605,629]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="364" phylum="Mollusca" rank="genus">
<emphasis id="2FB8BD5BFFC3FFC3FDB05874FD429919" box="[554,668,605,629]" italics="true" pageId="14" pageNumber="364">Clostophis</emphasis>
</taxonomicName>
species with blunt apex, a relatively narrow umbilicus, and a parietal tooth that is only slightly indicated.
</paragraph>
</subSubSection>
<subSubSection id="55D632C2FFC3FFC3FFEB58F4FF1B9F19" pageId="14" pageNumber="364" type="description">
<paragraph id="1D736149FFC3FFC3FFEB58F4FD3F9F79" blockId="14.[112,753,733,1045]" pageId="14" pageNumber="364">
<emphasis id="2FB8BD5BFFC3FFC3FFEB58F4FED99999" bold="true" box="[113,263,733,757]" pageId="14" pageNumber="364">Description.</emphasis>
Shell conical, higher than wide. Whorls rounded and very slightly shouldered. Tuba not detached from penultimate whorl. Protoconch nearly reaches 1.75 whorls, pitted and spirally striated. Entire shell with 4.5 whorls. Umbilicus narrow. Entire teleoconch with rather equidistant spiral striae, and some weak, irregular radial growth lines. Aperture subcircular with straight parietal part. Parietal tooth very weak, only slightly indicted, but as long as in
<taxonomicName id="DACC1ACAFFC3FFC3FF2B59F7FE8E9899" authority="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen, 2020" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" box="[177,336,989,1013]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="364" phylum="Mollusca" rank="species" species="multiformis" status="sp. nov.">
<emphasis id="2FB8BD5BFFC3FFC3FF2B59F7FE8E9899" box="[177,336,989,1013]" italics="true" pageId="14" pageNumber="364">C. multiformis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="348B0020FFC3FFC3FEFA59F7FE369899" box="[352,488,989,1013]" pageId="14" pageNumber="364" rank="species">new species</taxonomicNameLabel>
. Aperture not adnate to penultimate whorl. Peristome expanded but not reflected.
</paragraph>
<paragraph id="1D736149FFC3FFC3FFEB5E14FF1B9F19" blockId="14.[113,752,1085,1141]" pageId="14" pageNumber="364">
<emphasis id="2FB8BD5BFFC3FFC3FFEB5E14FE5C9F39" bold="true" box="[113,386,1085,1109]" pageId="14" pageNumber="364">Measurements (in mm).</emphasis>
SH = 1.171.39, SW = 1.071.24 (n = 4).
</paragraph>
</subSubSection>
<subSubSection id="55D632C2FFC3FFC3FFEB5EB4FD669FB9" pageId="14" pageNumber="364" type="etymology">
<paragraph id="1D736149FFC3FFC3FFEB5EB4FD669FB9" blockId="14.[113,752,1181,1237]" pageId="14" pageNumber="364">
<emphasis id="2FB8BD5BFFC3FFC3FFEB5EB4FF2D9FD9" bold="true" box="[113,243,1181,1205]" pageId="14" pageNumber="364">Etymology.</emphasis>
The specific epithet “
<taxonomicName id="DACC1ACAFFC3FFC3FE7B5EB4FDEC9FD9" authorityName="Pall-Gergely &amp; Grego" authorityYear="2020" box="[481,562,1181,1205]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="364" phylum="Mollusca" rank="species" species="obtusus">
<emphasis id="2FB8BD5BFFC3FFC3FE7B5EB4FDEC9FD9" box="[481,562,1181,1205]" italics="true" pageId="14" pageNumber="364">obtusus</emphasis>
</taxonomicName>
” (= obtuse, blunt in Latin) refers to the blunt apex of this new species.
</paragraph>
</subSubSection>
<subSubSection id="55D632C2FFC3FFC3FFEB5ED4FDEC9ED9" pageId="14" pageNumber="364" type="diagnosis">
<paragraph id="1D736149FFC3FFC3FFEB5ED4FDEC9ED9" blockId="14.[113,752,1277,1461]" pageId="14" pageNumber="364">
<emphasis id="2FB8BD5BFFC3FFC3FFEB5ED4FEAB9E79" bold="true" box="[113,373,1277,1301]" pageId="14" pageNumber="364">Differential diagnosis.</emphasis>
<taxonomicName id="DACC1ACAFFC3FFC3FEE55ED4FD8C9E79" authority="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen, 2020" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" box="[383,594,1277,1301]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="364" phylum="Mollusca" rank="species" species="obtusus" status="sp. nov.">
<emphasis id="2FB8BD5BFFC3FFC3FEE55ED4FD8C9E79" box="[383,594,1277,1301]" italics="true" pageId="14" pageNumber="364">Clostophis obtusus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="348B0020FFC3FFC3FDF95ED7FD329E79" box="[611,748,1277,1301]" pageId="14" pageNumber="364" rank="species">new species</taxonomicNameLabel>
, differs from
<taxonomicName id="DACC1ACAFFC3FFC3FE915F37FE6A9E59" authority="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen, 2020" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" box="[267,436,1309,1333]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="364" phylum="Mollusca" rank="species" species="multiformis" status="sp. nov.">
<emphasis id="2FB8BD5BFFC3FFC3FE915F37FE6A9E59" box="[267,436,1309,1333]" italics="true" pageId="14" pageNumber="364">C. multiformis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="348B0020FFC3FFC3FE525F37FD8B9E59" box="[456,597,1309,1333]" pageId="14" pageNumber="364" rank="species">new species</taxonomicNameLabel>
(populations northeast of Thakhek), in the following traits: the shell of
<taxonomicName id="DACC1ACAFFC3FFC3FF0B5F77FEDB9E19" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" box="[145,261,1373,1397]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="364" phylum="Mollusca" rank="species" species="obtusus">
<emphasis id="2FB8BD5BFFC3FFC3FF0B5F77FEDB9E19" box="[145,261,1373,1397]" italics="true" pageId="14" pageNumber="364">C. obtusus</emphasis>
</taxonomicName>
is larger, always triangular, not ovoid, apex blunter, umbilicus narrower, aperture never detached from penultimate whorl, parietal tooth weaker.
</paragraph>
</subSubSection>
<subSubSection id="55D632C2FFC3FFC3FFEB5FF4FED49D39" pageId="14" pageNumber="364" type="discussion">
<paragraph id="1D736149FFC3FFC3FFEB5FF4FED49D39" blockId="14.[113,752,1501,1621]" pageId="14" pageNumber="364">
<emphasis id="2FB8BD5BFFC3FFC3FFEB5FF4FF3E9E99" bold="true" box="[113,224,1501,1525]" pageId="14" pageNumber="364">Remarks.</emphasis>
We first considered this species to be fitting into the wide variability of
<taxonomicName id="DACC1ACAFFC3FFC3FEE75FD7FDFF9D79" authority="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen, 2020" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" box="[381,545,1533,1557]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="364" phylum="Mollusca" rank="species" species="multiformis" status="sp. nov.">
<emphasis id="2FB8BD5BFFC3FFC3FEE75FD7FDFF9D79" box="[381,545,1533,1557]" italics="true" pageId="14" pageNumber="364">C. multiformis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="348B0020FFC3FFC3FDA95FD7FD609D79" box="[563,702,1533,1557]" pageId="14" pageNumber="364" rank="species">new species</taxonomicNameLabel>
, but in fact, at one site (locality: JG2B) the two species occur sympatrically.
</paragraph>
</subSubSection>
</treatment>
</document>