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<document id="4444B41FE8A0F573B031DA5E38F7A324" ID-DOI="10.11646/zootaxa.3605.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5259280" ID-ZooBank="19CFDC67-4FCB-431D-8BF2-80EEB9EC76A4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629837201830" checkinUser="felipe" docAuthor="Gimmel, Matthew L." docDate="2013" docId="8C75C266106E28032286FA9F7F6CCE27" docLanguage="en" docName="zt03605p147.pdf" docOrigin="Zootaxa 3605 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3605.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Sveculus Gimmel 2013, gen. nov." docType="treatment" docVersion="4" lastPageNumber="74" masterDocId="704CBA1E1029284A2211FFE37E7CC850" masterDocTitle="&lt;strong&gt; Genus-level revision of the family Phalacridae (Coleoptera: Cucujoidea) &lt;/ strong&gt;" masterLastPageNumber="147" masterPageNumber="1" pageNumber="72" updateTime="1699119309817" updateUser="plazi">
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<mods:title id="D44C618E66F60A2C5458D88B11E44F8D">&lt;strong&gt; Genus-level revision of the family Phalacridae (Coleoptera: Cucujoidea) &lt;/ strong&gt;</mods:title>
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<mods:name id="EFE63AE6856E146271822B2B7C24B31B" type="personal">
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<mods:namePart id="C76E1095115EBEA351933CDC627880B0">Gimmel, Matthew L.</mods:namePart>
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<mods:title id="201C220A0430614BFBCF77FD2079FBB7">Zootaxa</mods:title>
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<mods:date id="3528FF36C469A7052FE52162D35C21AB">2013</mods:date>
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<treatment id="8C75C266106E28032286FA9F7F6CCE27" LSID="urn:lsid:plazi:treatment:8C75C266106E28032286FA9F7F6CCE27" httpUri="http://treatment.plazi.org/id/8C75C266106E28032286FA9F7F6CCE27" lastPageId="73" lastPageNumber="74" pageId="71" pageNumber="72">
<subSubSection id="4CC620FB106E280D2286FA9F7C6BCDC6" box="[151,535,1404,1430]" pageId="71" pageNumber="72" type="nomenclature">
<paragraph id="04637370106E280D2286FA9F7C6BCDC6" blockId="71.[151,535,1404,1463]" box="[151,535,1404,1430]" pageId="71" pageNumber="72">
<emphasis id="36A8AF62106E280D2286FA9F7C6BCDC6" bold="true" box="[151,535,1404,1430]" pageId="71" pageNumber="72">
18.
<taxonomicName id="C3DC08F3106E280D22D2FA9F7FE0CDC6" authority="Gimmel" authorityName="Gimmel" authorityYear="2013" box="[195,412,1404,1430]" class="Insecta" family="Phalacridae" genus="Sveculus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="36A8AF62106E280D22D2FA9F7F52CDC6" bold="true" box="[195,302,1404,1430]" italics="true" pageId="71" pageNumber="72">Sveculus</emphasis>
Gimmel
</taxonomicName>
,
<taxonomicNameLabel id="2D9B1219106E280D23B8FA9F7C6BCDC6" box="[425,535,1404,1430]" pageId="71" pageNumber="72" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="4CC620FB106E280D2286FA7C7C0ECDAD" pageId="71" pageNumber="72" type="description">
<paragraph id="04637370106E280D2286FA7C7F28CDE7" blockId="71.[151,535,1404,1463]" box="[151,340,1439,1463]" pageId="71" pageNumber="72">
(
<figureCitation id="9CE76FF5106E280D228EFA7C7E81CDE7" box="[159,253,1439,1463]" captionStart="FIGURE 21" captionStartId="72.[151,250,1900,1923]" captionTargetBox="[202,1385,184,1858]" captionTargetId="figure-98@72.[169,1417,151,1879]" captionTargetPageId="72" captionText="FIGURE 21. Sveculus lewisi, male. (a) Left mandible, dorsal; (b) left antenna (scale bar = 0.5 mm). (c) Left protibia and tarsus, dorsal; (d) right metatibia and tarsus, ventral (scale bar = 0.5 mm). (e) Hind wing (scale bar = 1.0 mm). (f) Meso- and metaventrite, ventral (scale bar = 0.5 mm). (g) Metendosternite (scale bar = 0.5 mm). (h) Tegmen, ventral; (i) penis, ventral (scale bar = 0.5 mm). Female. (j) Spermatheca (scale bar = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.5259322" httpUri="https://zenodo.org/record/5259322/files/figure.png" pageId="71" pageNumber="72">Figs. 21</figureCitation>
;
<figureCitation id="9CE76FF5106E280D2318FA7C7F37CDE7" box="[265,331,1439,1463]" captionStart="FIGURE 40" captionStartId="125.[151,250,1874,1897]" captionTargetBox="[169,1391,221,1841]" captionTargetId="figure-59@125.[169,1417,211,1843]" captionTargetPageId="125" captionText="FIGURE 40. Photographs. Olibroporus punctatus: (a) dorsal. Pycinus sp.: (b) dorsal; (c) ventral. Ochrolitus rubens: (d) dorsal; (e) ventral. Sveculus lewisi, holotype: (f) dorsal; (g) ventral. Tolyphus (Tolyphus) granulatus: (h) dorsal; (i) head and pronotum laterally, showing eye facets." figureDoi="http://doi.org/10.5281/zenodo.5259428" httpUri="https://zenodo.org/record/5259428/files/figure.png" pageId="71" pageNumber="72">40f, g</figureCitation>
)
</paragraph>
<paragraph id="04637370106E280D2286FA047C0ECDAD" blockId="71.[151,626,1511,1533]" box="[151,626,1511,1533]" pageId="71" pageNumber="72">
<typeStatus id="DB67CDD2106E280D2286FA047EB7CDAC" box="[151,203,1511,1532]" pageId="71" pageNumber="72">Type</typeStatus>
species:
<taxonomicName id="C3DC08F3106E280D2337FA047FBCCDAD" authorityName="Gimmel" authorityYear="2013" box="[294,448,1511,1533]" class="Insecta" family="Phalacridae" genus="Sveculus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="lewisi">
<emphasis id="36A8AF62106E280D2337FA047FBCCDAD" box="[294,448,1511,1533]" italics="true" pageId="71" pageNumber="72">Sveculus lewisi</emphasis>
</taxonomicName>
, here designated.
</paragraph>
</subSubSection>
<subSubSection id="4CC620FB106E280D2286F9CA7C8FCE11" box="[151,755,1577,1602]" pageId="71" pageNumber="72" type="materials_examined">
<paragraph id="04637370106E280D2286F9CA7C8FCE11" blockId="71.[151,1437,1577,2033]" box="[151,755,1577,1602]" pageId="71" pageNumber="72">
<emphasis id="36A8AF62106E280D2286F9CA7F39CE12" bold="true" box="[151,325,1577,1602]" pageId="71" pageNumber="72">
<typeStatus id="DB67CDD2106E280D2286F9CA7EADCE11" box="[151,209,1577,1601]" pageId="71" pageNumber="72">Type</typeStatus>
material.
</emphasis>
See account of
<taxonomicName id="C3DC08F3106E280D23E6F9CA7CDDCE11" authorityName="Gimmel" authorityYear="2013" box="[503,673,1577,1601]" class="Insecta" family="Phalacridae" genus="Sveculus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="lewisi">
<emphasis id="36A8AF62106E280D23E6F9CA7CDDCE11" box="[503,673,1577,1601]" italics="true" pageId="71" pageNumber="72">Sveculus lewisi</emphasis>
</taxonomicName>
below.
</paragraph>
</subSubSection>
<subSubSection id="4CC620FB106E280D22D6F9AE7BE1CED9" pageId="71" pageNumber="72" type="diagnosis">
<paragraph id="04637370106E280D22D6F9AE7BE1CED9" blockId="71.[151,1437,1577,2033]" pageId="71" pageNumber="72">
<emphasis id="36A8AF62106E280D22D6F9AE7F42CE36" bold="true" box="[199,318,1613,1638]" pageId="71" pageNumber="72">Diagnosis.</emphasis>
This is the only genus of
<taxonomicName id="C3DC08F3106E280D2045F9AE7CA8CE35" authorityName="Leach" authorityYear="1815" box="[596,724,1613,1637]" class="Insecta" family="Phalacridae" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="family">Phalacridae</taxonomicName>
with the following combination of characters: protibia with long ctenidium, prosternal process with apical transparent laminar process, and metatarsomeres I and II subequal in length.
</paragraph>
</subSubSection>
<subSubSection id="4CC620FB106E280322D6F9767F25CCBA" lastPageId="73" lastPageNumber="74" pageId="71" pageNumber="72" type="description">
<paragraph id="04637370106E280D22D6F9767D9ECE81" blockId="71.[151,1437,1577,2033]" pageId="71" pageNumber="72">
<emphasis id="36A8AF62106E280D22D6F9767F24CEFE" bold="true" box="[199,344,1685,1710]" pageId="71" pageNumber="72">Description.</emphasis>
Very small to small, total length 1.12.0 mm. Dorsal color solid testaceous to rufo-testaceous (
<figureCitation id="9CE76FF5106E280D228EF95A7F5FCE81" box="[159,291,1721,1745]" captionStart="FIGURE 40" captionStartId="125.[151,250,1874,1897]" captionTargetBox="[169,1391,221,1841]" captionTargetId="figure-59@125.[169,1417,211,1843]" captionTargetPageId="125" captionText="FIGURE 40. Photographs. Olibroporus punctatus: (a) dorsal. Pycinus sp.: (b) dorsal; (c) ventral. Ochrolitus rubens: (d) dorsal; (e) ventral. Sveculus lewisi, holotype: (f) dorsal; (g) ventral. Tolyphus (Tolyphus) granulatus: (h) dorsal; (i) head and pronotum laterally, showing eye facets." figureDoi="http://doi.org/10.5281/zenodo.5259428" httpUri="https://zenodo.org/record/5259428/files/figure.png" pageId="71" pageNumber="72">Figs. 40f, g</figureCitation>
). Tibial spur formula 2-2-2, tarsal formula 5-5-
<quantity id="C324DE95106E280D2122F95A7D22CE81" box="[819,862,1721,1745]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="71" pageNumber="72" unit="in" value="5.0">5 in</quantity>
both sexes.
</paragraph>
<paragraph id="04637370106E280D22D6F93D7FF8CFA1" blockId="71.[151,1437,1577,2033]" pageId="71" pageNumber="72">
<emphasis id="36A8AF62106E280D22D6F93D7F77CEA6" box="[199,267,1758,1782]" italics="true" pageId="71" pageNumber="72">Head.</emphasis>
<collectingRegion id="C618BD92106E280D2303F93D7F40CEA5" box="[274,316,1758,1781]" country="Cameroon" name="North" pageId="71" pageNumber="72">Not</collectingRegion>
constricted behind eyes. Eyes small to medium-sized; facets flat; interfacetal setae absent; weakly emarginate medially; without posterior emargination; periocular groove absent; with transverse setose groove ventrally behind eye. Frontoclypeus not or barely emarginate above antennal insertion; clypeal apex arcuatetruncate. Antennal club 3-segmented, club weakly asymmetrical; antennomere XI constricted on anterior edge (
<figureCitation id="9CE76FF5106E280D228EF88E7F7DCFD5" box="[159,257,1901,1925]" captionStart="FIGURE 21" captionStartId="72.[151,250,1900,1923]" captionTargetBox="[202,1385,184,1858]" captionTargetId="figure-98@72.[169,1417,151,1879]" captionTargetPageId="72" captionText="FIGURE 21. Sveculus lewisi, male. (a) Left mandible, dorsal; (b) left antenna (scale bar = 0.5 mm). (c) Left protibia and tarsus, dorsal; (d) right metatibia and tarsus, ventral (scale bar = 0.5 mm). (e) Hind wing (scale bar = 1.0 mm). (f) Meso- and metaventrite, ventral (scale bar = 0.5 mm). (g) Metendosternite (scale bar = 0.5 mm). (h) Tegmen, ventral; (i) penis, ventral (scale bar = 0.5 mm). Female. (j) Spermatheca (scale bar = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.5259322" httpUri="https://zenodo.org/record/5259322/files/figure.png" pageId="71" pageNumber="72">Fig. 21b</figureCitation>
). Mandible (
<figureCitation id="9CE76FF5106E280D239FF88E7F97CFD5" box="[398,491,1901,1925]" captionStart="FIGURE 21" captionStartId="72.[151,250,1900,1923]" captionTargetBox="[202,1385,184,1858]" captionTargetId="figure-98@72.[169,1417,151,1879]" captionTargetPageId="72" captionText="FIGURE 21. Sveculus lewisi, male. (a) Left mandible, dorsal; (b) left antenna (scale bar = 0.5 mm). (c) Left protibia and tarsus, dorsal; (d) right metatibia and tarsus, ventral (scale bar = 0.5 mm). (e) Hind wing (scale bar = 1.0 mm). (f) Meso- and metaventrite, ventral (scale bar = 0.5 mm). (g) Metendosternite (scale bar = 0.5 mm). (h) Tegmen, ventral; (i) penis, ventral (scale bar = 0.5 mm). Female. (j) Spermatheca (scale bar = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.5259322" httpUri="https://zenodo.org/record/5259322/files/figure.png" pageId="71" pageNumber="72">Fig. 21a</figureCitation>
) with apex bidentate; retinaculum absent; mandible without ventral ridge. Maxillary palpomere IV fusiform, short, nearly symmetrical; galea short, rounded; lacinia with two stout spines.
<materialsCitation id="B4B4792D106E280D272CF8717FFCCFA1" collectionCode="III" location="Mentum" pageId="71" pageNumber="72" specimenCount="1" stateProvince="Chin">
<collectingRegion id="C618BD92106E280D272CF8717BE0CFF9" box="[1341,1436,1938,1961]" country="Myanmar" name="Chin" pageId="71" pageNumber="72">Mentum</collectingRegion>
with sides divergent toward apex; labial palpomere
<collectionCode id="62CDEBB5106E280D20CDF8557C84CF9D" box="[732,760,1974,1997]" pageId="71" pageNumber="72">III</collectionCode>
fusiform. Labrum with apical margin arcuate. Gular sutures short, barely evident
</materialsCitation>
.
</paragraph>
<caption id="50A323F8106128022286F88F7D32CFB8" ID-DOI="http://doi.org/10.5281/zenodo.5259322" ID-Zenodo-Dep="5259322" httpUri="https://zenodo.org/record/5259322/files/figure.png" pageId="72" pageNumber="73" startId="72.[151,250,1900,1923]" targetBox="[202,1385,184,1858]" targetPageId="72">
<paragraph id="04637370106128022286F88F7D32CFB8" blockId="72.[151,1436,1900,2024]" pageId="72" pageNumber="73">
<emphasis id="36A8AF62106128022286F88F7F63CFD3" bold="true" box="[151,287,1900,1923]" pageId="72" pageNumber="73">FIGURE 21.</emphasis>
<taxonomicName id="C3DC08F3106128022335F88E7FC2CFD3" authorityName="Gimmel" authorityYear="2013" box="[292,446,1901,1923]" class="Insecta" family="Phalacridae" genus="Sveculus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="lewisi">
<emphasis id="36A8AF62106128022335F88E7FC2CFD3" box="[292,446,1901,1923]" italics="true" pageId="72" pageNumber="73">Sveculus lewisi</emphasis>
</taxonomicName>
, male. (a) Left mandible, dorsal; (b) left antenna (scale bar = 0.5 mm). (c) Left protibia and tarsus, dorsal; (d) right metatibia and tarsus, ventral (scale bar = 0.5 mm). (e) Hind wing (scale bar = 1.0 mm). (f) Meso- and metaventrite, ventral (scale bar = 0.5 mm). (g) Metendosternite (scale bar = 0.5 mm). (h) Tegmen, ventral; (i) penis, ventral (scale bar = 0.5 mm). Female. (j) Spermatheca (scale bar = 0.5 mm).
</paragraph>
</caption>
<paragraph id="046373701060280322D6FF7B7FCACBD2" blockId="73.[151,1437,151,1439]" pageId="73" pageNumber="74">
<emphasis id="36A8AF621060280322D6FF7B7F63C8E0" box="[199,287,152,176]" italics="true" pageId="73" pageNumber="74">Thorax.</emphasis>
Pronotum with microsetae present, distinct; with scutellar lobe weakly to moderately developed.
<taxonomicName id="C3DC08F3106028032286FF5F7F64C883" box="[151,280,188,211]" class="Insecta" family="Elateridae" genus="Prosternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="134" phylum="Arthropoda" rank="genus">Prosternum</taxonomicName>
anteriorly with continuous row of marginal setae, setae flattened at base; procoxal cavity without anterolateral notchlike extension; prosternal process angulate in lateral view, sometimes conspicuously setose preapically, with broad horizontal translucent process at apex, apex without row of spinelike setae. Protrochanter without setae; protibia with ctenidium on kickface, extending about two-thirds length of tibia (
<figureCitation id="9CE76FF51060280326D2FEC47B58C96F" box="[1219,1316,295,319]" captionStart="FIGURE 21" captionStartId="72.[151,250,1900,1923]" captionTargetBox="[202,1385,184,1858]" captionTargetId="figure-98@72.[169,1417,151,1879]" captionTargetPageId="72" captionText="FIGURE 21. Sveculus lewisi, male. (a) Left mandible, dorsal; (b) left antenna (scale bar = 0.5 mm). (c) Left protibia and tarsus, dorsal; (d) right metatibia and tarsus, ventral (scale bar = 0.5 mm). (e) Hind wing (scale bar = 1.0 mm). (f) Meso- and metaventrite, ventral (scale bar = 0.5 mm). (g) Metendosternite (scale bar = 0.5 mm). (h) Tegmen, ventral; (i) penis, ventral (scale bar = 0.5 mm). Female. (j) Spermatheca (scale bar = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.5259322" httpUri="https://zenodo.org/record/5259322/files/figure.png" pageId="73" pageNumber="74">Fig. 21c</figureCitation>
). Scutellar shield small. Elytron with spectral iridescence present or absent; with one weak sutural stria or stria absent; disc without even rudimentary striae or rows of punctures; with weak to strong transverse strigae, or strigae absent; lateral margin with row of tiny, sawtooth-like setae. Mesoventral plate (
<figureCitation id="9CE76FF51060280321E7FE777A2BC9FC" box="[1014,1111,404,428]" captionStart="FIGURE 21" captionStartId="72.[151,250,1900,1923]" captionTargetBox="[202,1385,184,1858]" captionTargetId="figure-98@72.[169,1417,151,1879]" captionTargetPageId="72" captionText="FIGURE 21. Sveculus lewisi, male. (a) Left mandible, dorsal; (b) left antenna (scale bar = 0.5 mm). (c) Left protibia and tarsus, dorsal; (d) right metatibia and tarsus, ventral (scale bar = 0.5 mm). (e) Hind wing (scale bar = 1.0 mm). (f) Meso- and metaventrite, ventral (scale bar = 0.5 mm). (g) Metendosternite (scale bar = 0.5 mm). (h) Tegmen, ventral; (i) penis, ventral (scale bar = 0.5 mm). Female. (j) Spermatheca (scale bar = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.5259322" httpUri="https://zenodo.org/record/5259322/files/figure.png" pageId="73" pageNumber="74">Fig. 21f</figureCitation>
) deeply notched anteriorly, extending posteriorly to metaventrite, not forming procoxal rests; mesanepisternum with complete transverse carina; mesocoxal cavities separated by slightly more than half width of a coxal cavity. Mesotarsomere III not bilobed. Metaventral process (
<figureCitation id="9CE76FF51060280323FEFE1C7C36CA47" box="[495,586,511,535]" captionStart="FIGURE 21" captionStartId="72.[151,250,1900,1923]" captionTargetBox="[202,1385,184,1858]" captionTargetId="figure-98@72.[169,1417,151,1879]" captionTargetPageId="72" captionText="FIGURE 21. Sveculus lewisi, male. (a) Left mandible, dorsal; (b) left antenna (scale bar = 0.5 mm). (c) Left protibia and tarsus, dorsal; (d) right metatibia and tarsus, ventral (scale bar = 0.5 mm). (e) Hind wing (scale bar = 1.0 mm). (f) Meso- and metaventrite, ventral (scale bar = 0.5 mm). (g) Metendosternite (scale bar = 0.5 mm). (h) Tegmen, ventral; (i) penis, ventral (scale bar = 0.5 mm). Female. (j) Spermatheca (scale bar = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.5259322" httpUri="https://zenodo.org/record/5259322/files/figure.png" pageId="73" pageNumber="74">Fig. 21f</figureCitation>
) not extending to anterior level of mesocoxae; metaventral postcoxal lines narrowly separated from mesocoxal cavity margin; discrimen short, extending less than halfway to anterior margin of metaventral process; metendosternite (
<figureCitation id="9CE76FF5106028032064FDA47CABCA0F" box="[629,727,583,607]" captionStart="FIGURE 21" captionStartId="72.[151,250,1900,1923]" captionTargetBox="[202,1385,184,1858]" captionTargetId="figure-98@72.[169,1417,151,1879]" captionTargetPageId="72" captionText="FIGURE 21. Sveculus lewisi, male. (a) Left mandible, dorsal; (b) left antenna (scale bar = 0.5 mm). (c) Left protibia and tarsus, dorsal; (d) right metatibia and tarsus, ventral (scale bar = 0.5 mm). (e) Hind wing (scale bar = 1.0 mm). (f) Meso- and metaventrite, ventral (scale bar = 0.5 mm). (g) Metendosternite (scale bar = 0.5 mm). (h) Tegmen, ventral; (i) penis, ventral (scale bar = 0.5 mm). Female. (j) Spermatheca (scale bar = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.5259322" httpUri="https://zenodo.org/record/5259322/files/figure.png" pageId="73" pageNumber="74">
Fig.
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) with anterior tendons moderately separated, ventral process intersecting ventral longitudinal flange behind anterior margin. Anterior margin of metacoxa with emargination sublaterally; metacoxal plate with transverse line; metatibial foreface with apical ctenidium roughly perpendicular overall to long axis of tibia, or slightly oblique; spurs cylindrical, longest spur subequal to width of tibial apex; metatarsomere I subequal to metatarsomere II or I slightly shorter, joint between I and II rigid (
<figureCitation id="9CE76FF51060280326F8FD347B32CABF" box="[1257,1358,727,751]" captionStart="FIGURE 21" captionStartId="72.[151,250,1900,1923]" captionTargetBox="[202,1385,184,1858]" captionTargetId="figure-98@72.[169,1417,151,1879]" captionTargetPageId="72" captionText="FIGURE 21. Sveculus lewisi, male. (a) Left mandible, dorsal; (b) left antenna (scale bar = 0.5 mm). (c) Left protibia and tarsus, dorsal; (d) right metatibia and tarsus, ventral (scale bar = 0.5 mm). (e) Hind wing (scale bar = 1.0 mm). (f) Meso- and metaventrite, ventral (scale bar = 0.5 mm). (g) Metendosternite (scale bar = 0.5 mm). (h) Tegmen, ventral; (i) penis, ventral (scale bar = 0.5 mm). Female. (j) Spermatheca (scale bar = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.5259322" httpUri="https://zenodo.org/record/5259322/files/figure.png" pageId="73" pageNumber="74">Fig. 21d</figureCitation>
). Hind wing (
<figureCitation id="9CE76FF51060280322CFFD1F7F40CB44" box="[222,316,764,788]" captionStart="FIGURE 21" captionStartId="72.[151,250,1900,1923]" captionTargetBox="[202,1385,184,1858]" captionTargetId="figure-98@72.[169,1417,151,1879]" captionTargetPageId="72" captionText="FIGURE 21. Sveculus lewisi, male. (a) Left mandible, dorsal; (b) left antenna (scale bar = 0.5 mm). (c) Left protibia and tarsus, dorsal; (d) right metatibia and tarsus, ventral (scale bar = 0.5 mm). (e) Hind wing (scale bar = 1.0 mm). (f) Meso- and metaventrite, ventral (scale bar = 0.5 mm). (g) Metendosternite (scale bar = 0.5 mm). (h) Tegmen, ventral; (i) penis, ventral (scale bar = 0.5 mm). Female. (j) Spermatheca (scale bar = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.5259322" httpUri="https://zenodo.org/record/5259322/files/figure.png" pageId="73" pageNumber="74">Fig. 21e</figureCitation>
) with distinct, ovate anal lobe; leading edge with incomplete row of long setae at level of
<collectingRegion id="C618BD92106028032723FD1F7B27CB44" box="[1330,1371,764,788]" country="Malta" name="Luqa" pageId="73" pageNumber="74">RA</collectingRegion>
+ScP; AA
<subScript id="985871351060280322AEFCCC7EABCB6D" attach="left" box="[191,215,815,829]" fontSize="6" pageId="73" pageNumber="74">3+4</subScript>
absent; cubitoanal system unbranched apically; MP
<subScript id="9858713510602803210BFCCC7D4ECB6D" attach="left" box="[794,818,815,829]" fontSize="6" pageId="73" pageNumber="74">3+4</subScript>
with long, unbranched distal remnant; r4 absent; flecks present in apical field distal to rp-mp2; long transverse proximal sclerite and faint triangular sclerite present just distal to end of radial bar.
</paragraph>
<paragraph id="046373701060280322D6FC6C7C19CC67" blockId="73.[151,1437,151,1439]" pageId="73" pageNumber="74">
<emphasis id="36A8AF621060280322D6FC6C7F44CBF7" box="[199,312,911,935]" italics="true" pageId="73" pageNumber="74">Abdomen.</emphasis>
Abdominal ventrite I without paired lines or calli; spiracles apparently absent from segment VII. Male with aedeagus upright in repose; tegmen (
<figureCitation id="9CE76FF51060280320A3FC517D6ECB9A" box="[690,786,946,970]" captionStart="FIGURE 21" captionStartId="72.[151,250,1900,1923]" captionTargetBox="[202,1385,184,1858]" captionTargetId="figure-98@72.[169,1417,151,1879]" captionTargetPageId="72" captionText="FIGURE 21. Sveculus lewisi, male. (a) Left mandible, dorsal; (b) left antenna (scale bar = 0.5 mm). (c) Left protibia and tarsus, dorsal; (d) right metatibia and tarsus, ventral (scale bar = 0.5 mm). (e) Hind wing (scale bar = 1.0 mm). (f) Meso- and metaventrite, ventral (scale bar = 0.5 mm). (g) Metendosternite (scale bar = 0.5 mm). (h) Tegmen, ventral; (i) penis, ventral (scale bar = 0.5 mm). Female. (j) Spermatheca (scale bar = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.5259322" httpUri="https://zenodo.org/record/5259322/files/figure.png" pageId="73" pageNumber="74">Fig. 21h</figureCitation>
) with symmetrical anterior margin and parameres hinged to basal piece, parameres without medial longitudinal division; penis (
<figureCitation id="9CE76FF51060280321D6FC347A58CBBF" box="[967,1060,983,1007]" captionStart="FIGURE 21" captionStartId="72.[151,250,1900,1923]" captionTargetBox="[202,1385,184,1858]" captionTargetId="figure-98@72.[169,1417,151,1879]" captionTargetPageId="72" captionText="FIGURE 21. Sveculus lewisi, male. (a) Left mandible, dorsal; (b) left antenna (scale bar = 0.5 mm). (c) Left protibia and tarsus, dorsal; (d) right metatibia and tarsus, ventral (scale bar = 0.5 mm). (e) Hind wing (scale bar = 1.0 mm). (f) Meso- and metaventrite, ventral (scale bar = 0.5 mm). (g) Metendosternite (scale bar = 0.5 mm). (h) Tegmen, ventral; (i) penis, ventral (scale bar = 0.5 mm). Female. (j) Spermatheca (scale bar = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.5259322" httpUri="https://zenodo.org/record/5259322/files/figure.png" pageId="73" pageNumber="74">Fig. 21i</figureCitation>
) narrowed apically, with pair of endophallic sclerites, apex simple; spiculum gastrale V-shaped, arms free, with oblique anterior extension. Female ovipositor weakly sclerotized, palpiform.
</paragraph>
<paragraph id="046373701060280322D6FBA07C6ECC0A" blockId="73.[151,1437,151,1439]" box="[199,530,1090,1115]" pageId="73" pageNumber="74">
<emphasis id="36A8AF621060280322D6FBA07FE9CC0B" bold="true" box="[199,405,1091,1115]" pageId="73" pageNumber="74">Immature stages.</emphasis>
Unknown.
</paragraph>
<paragraph id="046373701060280322D6FB857F25CCBA" blockId="73.[151,1437,151,1439]" pageId="73" pageNumber="74">
<emphasis id="36A8AF621060280322D6FB857F36CC2F" bold="true" box="[199,330,1126,1151]" pageId="73" pageNumber="74">Bionomics.</emphasis>
Most specimens with ecological data were collected using Malaise or flight intercept traps or by fogging. Two Mindanao specimens were collected “under bark of log,” while one was collected “on decaying fleshy, gilled bracket fungus.” No material has been identified in the adult gut. None are known to have been collected at light.
</paragraph>
</subSubSection>
<subSubSection id="4CC620FB1060280322D6FB157DF2CDB7" pageId="73" pageNumber="74" type="distribution">
<paragraph id="046373701060280322D6FB157B60CD2A" blockId="73.[151,1437,151,1439]" pageId="73" pageNumber="74">
<emphasis id="36A8AF621060280322D6FB157C7DCD5F" bold="true" box="[199,513,1270,1295]" pageId="73" pageNumber="74">Distribution and diversity.</emphasis>
Southeast Asia from
<collectingCountry id="7CCB33E01060280320E0FB147D28CD5F" box="[753,852,1271,1295]" name="Thailand" pageId="73" pageNumber="74">Thailand</collectingCountry>
and the Malay Peninsula through Borneo, Sulawesi, and Mindanao, to
<collectingRegion id="C618BD9210602803237EFAF97F8ACD62" box="[367,502,1306,1330]" country="Australia" name="Queensland" pageId="73" pageNumber="74">Queensland</collectingRegion>
,
<collectingRegion id="C618BD92106028032014FAF87CA5CD62" box="[517,729,1306,1330]" country="Australia" name="New South Wales" pageId="73" pageNumber="74">New South Wales</collectingRegion>
, and
<collectingRegion id="C618BD9210602803210DFAF97A21CD62" box="[796,1117,1306,1330]" country="Australia" name="Australian Capital Territory" pageId="73" pageNumber="74">Australian Capital Territory</collectingRegion>
,
<collectingCountry id="7CCB33E010602803267DFAF97AABCD62" box="[1132,1239,1306,1330]" name="Australia" pageId="73" pageNumber="74">Australia</collectingCountry>
. This genus also occurs in
<collectingCountry id="7CCB33E010602803231BFADC7FEFCD07" box="[266,403,1343,1367]" name="Madagascar" pageId="73" pageNumber="74">Madagascar</collectingCountry>
. Despite having examined a large amount of material from New
<collectingCountry id="7CCB33E0106028032699FADC7AA7CD07" box="[1160,1243,1343,1367]" name="Guinea-Bissau" pageId="73" pageNumber="74">Guinea</collectingCountry>
, I have not seen specimens from that island. There are several morphospecies in collections, all apparently undescribed.
</paragraph>
<paragraph id="046373701060280322D6FA647FC5CDCF" blockId="73.[151,1437,151,1439]" box="[199,441,1414,1439]" pageId="73" pageNumber="74">
<emphasis id="36A8AF621060280322D6FA647FF6CDCF" bold="true" box="[199,394,1414,1439]" pageId="73" pageNumber="74">Included species</emphasis>
(1):
</paragraph>
<paragraph id="046373701060280322D6FA2C7DF2CDB7" blockId="73.[199,910,1487,1511]" box="[199,910,1487,1511]" pageId="73" pageNumber="74">
<taxonomicName id="C3DC08F31060280322D6FA2C7FABCDB7" authority="Gimmel" authorityName="Gimmel" authorityYear="2013" box="[199,471,1487,1511]" class="Insecta" family="Phalacridae" genus="Sveculus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="lewisi" status="sp. nov.">
<emphasis id="36A8AF621060280322D6FA2C7F0DCDB7" box="[199,369,1487,1511]" italics="true" pageId="73" pageNumber="74">Sveculus lewisi</emphasis>
Gimmel
</taxonomicName>
,
<emphasis id="36A8AF621060280323F3FA2C7C47CDB7" bold="true" box="[482,571,1487,1511]" pageId="73" pageNumber="74">
<taxonomicNameLabel id="2D9B12191060280323F3FA2C7C47CDB7" box="[482,571,1487,1511]" pageId="73" pageNumber="74" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Distribution: southeast Asia)
</paragraph>
</subSubSection>
<subSubSection id="4CC620FB1060280322D6F9F47F6CCE27" pageId="73" pageNumber="74" type="etymology">
<paragraph id="046373701060280322D6F9F47F6CCE27" blockId="73.[151,1436,1559,1655]" pageId="73" pageNumber="74">
<emphasis id="36A8AF621060280322D6F9F47F32CE7F" bold="true" box="[199,334,1559,1583]" pageId="73" pageNumber="74">Etymology.</emphasis>
This genus is named in honor of Dr. Zdeněk Švec of
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,
<collectingCountry id="7CCB33E01060280321EDF9F47ACCCE7F" box="[1020,1200,1559,1583]" name="Czech Republic" pageId="73" pageNumber="74">Czech Republic</collectingCountry>
, in recognition of his significant contributions to understanding the phalacrid fauna of the Old World. The gender of the name is masculine.
</paragraph>
</subSubSection>
</treatment>
</document>