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<document id="083890C65E7CFC2846E8C5A23BFFEABA" ID="10.11646/zootaxa.4273.4.3SLASH11267" ID-CLB-Dataset="32891" ID-DOI="10.11646/zootaxa.4273.4.3" ID-GBIF-Dataset="0935deaf-8f18-434e-b4a0-0468cefc9577" ID-ISSN="1175-5326" ID-Zenodo-Dep="803704" ID-ZooBank="4CD96097-1016-47CB-8DC3-AD6E9EDA330C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1496820946147" checkinUser="plazi" docAuthor="Enghoff, Henrik" docDate="2017" docId="8C70BE3B2447196AFF676A5AAFD8FC8F" docLanguage="en" docName="zootaxa.4273.4.3SLASH11267.pdf" docOrigin="Zootaxa 4273 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tropostreptus microcephalus Enghoff, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="520" masterDocId="7049C64324551979FFF06F72AF21FF9B" masterDocTitle="A new East African genus of spirostreptid millipedes (Diplopoda, Spirostreptida, Spirostreptidae), with notes on their fungal ectoparasite Rickia gigas" masterLastPageNumber="530" masterPageNumber="501" pageNumber="519" updateTime="1698440246528" updateUser="ExternalLinkService">
<mods:mods id="BACF3BC605E3C82B2263E853456ECF7B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7F05C75A02235AB6D86220285B433818">
<mods:title id="61DBD8789C2B70CA8BB63D1775FD50A4">A new East African genus of spirostreptid millipedes (Diplopoda, Spirostreptida, Spirostreptidae), with notes on their fungal ectoparasite Rickia gigas</mods:title>
</mods:titleInfo>
<mods:name id="7A6214EDCD8B2EA33DF39E25818F342C" type="personal">
<mods:role id="5EE8D4489916B67ADBC7DC09773A7F46">
<mods:roleTerm id="397CEE306DC21105867E89466648CD18">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2388F8EA4FB4408A4B887D0BA5DF6B66">Enghoff, Henrik</mods:namePart>
</mods:name>
<mods:typeOfResource id="F4691A3141D2D8B80313A1A0CC4AE75E">text</mods:typeOfResource>
<mods:relatedItem id="7B3A0384B03DDE7E5F08EA307247D210" type="host">
<mods:titleInfo id="304BD2996A9D04954C01E7243B5A40DD">
<mods:title id="C6ED55A5131A9290A6066B86162E1B43">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="AC81A292C55D657FEB2AD8FA2A713CD8">
<mods:date id="6E1307691F168290064AF35273D2CF59">2017</mods:date>
<mods:detail id="31D1196EC1C1A542849C8FD99BF10613" type="volume">
<mods:number id="EE310FD5DBB40C0188BD0FBD3F7A9119">4273</mods:number>
</mods:detail>
<mods:detail id="CC9FA5F544B7874A966F58516CB9B407" type="issue">
<mods:number id="31F3786440A478A6E247E6F9D45AE94A">4</mods:number>
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<mods:start id="18C9D0CCD36310493174C2D555E670E9">501</mods:start>
<mods:end id="98A3416B2D6A810107C2D8E4EAE55830">530</mods:end>
</mods:extent>
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<mods:classification id="580057EF23F584EAA9DAF0CDFE7F26EB">journal article</mods:classification>
<mods:identifier id="8EBBDF17E52813985E143D43FACDDAB0" type="CLB-Dataset">32891</mods:identifier>
<mods:identifier id="1E668F0A39CFF6362641FB89BC89FE94" type="DOI">10.11646/zootaxa.4273.4.3</mods:identifier>
<mods:identifier id="9594537F7581247CCCFE9001416F0A6E" type="GBIF-Dataset">0935deaf-8f18-434e-b4a0-0468cefc9577</mods:identifier>
<mods:identifier id="23732C1241DD0954DA9682CA7EE5F363" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0C349353F7A217C856FA478B3DAC060A" type="Zenodo-Dep">803704</mods:identifier>
<mods:identifier id="0AEAA0470EACD20F9E9A36DAD1CE2F92" type="ZooBank">4CD96097-1016-47CB-8DC3-AD6E9EDA330C</mods:identifier>
</mods:mods>
<treatment id="8C70BE3B2447196AFF676A5AAFD8FC8F" ID-DOI="http://doi.org/10.5281/zenodo.6015537" ID-GBIF-Taxon="131812498" ID-Zenodo-Dep="6015537" LSID="urn:lsid:plazi:treatment:8C70BE3B2447196AFF676A5AAFD8FC8F" httpUri="http://treatment.plazi.org/id/8C70BE3B2447196AFF676A5AAFD8FC8F" lastPageId="19" lastPageNumber="520" pageId="18" pageNumber="519">
<subSubSection id="4CC35CA62447196BFF676A5AAE5BFAF8" pageId="18" pageNumber="519" type="nomenclature">
<paragraph id="04660F2D2447196BFF676A5AAD7AFAD9" blockId="18.[151,603,1320,1380]" box="[151,603,1320,1346]" pageId="18" pageNumber="519">
<heading id="5F2EB8412447196BFF676A5AAD7AFAD9" bold="true" box="[151,603,1320,1346]" fontSize="11" level="1" pageId="18" pageNumber="519" reason="1">
<emphasis id="36ADD33F2447196BFF676A5AAD7AFAD9" bold="true" box="[151,603,1320,1346]" pageId="18" pageNumber="519">
<taxonomicName id="C3D974AE2447196BFF676A5AAED7FAD9" ID-CoL="59D9Y" box="[151,502,1320,1346]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="18" pageNumber="519" phylum="Arthropoda" rank="species" species="microcephalus" status="sp. nov.">
<emphasis id="36ADD33F2447196BFF676A5AAED7FAD9" bold="true" box="[151,502,1320,1346]" italics="true" pageId="18" pageNumber="519">Tropostreptus microcephalus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2D9E6E442447196BFE0D6A5AAD7AFAD9" box="[509,603,1320,1346]" pageId="18" pageNumber="519" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="04660F2D2447196BFF676A39AE5BFAF8" blockId="18.[151,603,1320,1380]" box="[151,378,1355,1380]" pageId="18" pageNumber="519">
<figureCitation id="9CE213A82447196BFF676A39AFFFFAF8" box="[151,222,1355,1380]" captionStart="FIGURE 2" captionStartId="5.[151,250,1182,1204]" captionTargetBox="[151,1436,193,1096]" captionTargetId="figure@5.[151,1436,193,1161]" captionTargetPageId="5" captionText="FIGURE 2. Size of Tropostreptus males, expressed as number of podous rings and midbody vertical diameter. Females reach higher diameters than males. See Figs. 11 and 13 for details on T. hamatus and T. sigmatospinus sp. nov." httpUri="https://zenodo.org/record/803710/files/figure.png" pageId="18" pageNumber="519">Figs 2</figureCitation>
,
<figureCitation id="9CE213A82447196BFF1A6A3EAFD9FAF8" box="[234,248,1356,1379]" captionStart="FIGURE 7" captionStartId="13.[151,250,1869,1891]" captionTargetBox="[207,1387,199,1819]" captionTargetId="figure@13.[151,1436,155,1848]" captionTargetPageId="13" captionText="FIGURE 7. Tropostretus spp., gonopods, anterior view. A: T. hamatus, East Usambara Mts, B: T. hamatus, Udzungwa Mts., left telopodite missing, C: T. sigmatospinus sp. nov., D: T. microcephalus sp. nov., E: T. droides sp. nov., F: T. kipunji sp. nov. Scales 0.5 mm. atp = antetorsal process, bh = birdhead-shaped metaplical process, lpl = apicolateral proplical lobe, map = mesapical metaplical process, mp = metaplica, mpl = mesapical proplical lobe, pp = proplica, px = paracoxite, st = sternum, tlp = telopodite." httpUri="https://zenodo.org/record/803720/files/figure.png" pageId="18" pageNumber="519">7</figureCitation>
D, 8D, 10B
</paragraph>
</subSubSection>
<subSubSection id="4CC35CA62447196BFF676AE1ADB2FA6F" pageId="18" pageNumber="519" type="diagnosis">
<paragraph id="04660F2D2447196BFF676AE1ADB2FA6F" blockId="18.[151,1437,1427,2028]" pageId="18" pageNumber="519">
<emphasis id="36ADD33F2447196BFF676AE1AE30FA37" bold="true" box="[151,273,1427,1452]" pageId="18" pageNumber="519">Diagnosis.</emphasis>
A small species of
<taxonomicName id="C3D974AE2447196BFE096AE6ADB2FA30" box="[505,659,1428,1451]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="18" pageNumber="528" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="36ADD33F2447196BFE096AE6ADB2FA30" box="[505,659,1428,1451]" italics="true" pageId="18" pageNumber="519">Tropostreptus</emphasis>
</taxonomicName>
(
<figureCitation id="9CE213A82447196BFD546AE1ADCBFA30" box="[676,746,1427,1452]" captionStart="FIGURE 2" captionStartId="5.[151,250,1182,1204]" captionTargetBox="[151,1436,193,1096]" captionTargetId="figure@5.[151,1436,193,1161]" captionTargetPageId="5" captionText="FIGURE 2. Size of Tropostreptus males, expressed as number of podous rings and midbody vertical diameter. Females reach higher diameters than males. See Figs. 11 and 13 for details on T. hamatus and T. sigmatospinus sp. nov." httpUri="https://zenodo.org/record/803710/files/figure.png" pageId="18" pageNumber="519">Fig. 2</figureCitation>
). Differs from other species by the combination of size, the presence of two (not three) apical processes (
<emphasis id="36ADD33F2447196BFD7C6ACAAD9CFA54" box="[652,701,1464,1487]" italics="true" pageId="18" pageNumber="519">map</emphasis>
and
<emphasis id="36ADD33F2447196BFD026AC4AC2FFA54" box="[754,782,1462,1487]" italics="true" pageId="18" pageNumber="519">bh</emphasis>
) on the gonopodal metaplica, and the laterad direction and small size of the birdhead-shaped process
<emphasis id="36ADD33F2447196BFD9F6AA9ADB2FA6F" box="[623,659,1499,1524]" italics="true" pageId="18" pageNumber="519">bh.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="4CC35CA62447196BFF376A8CAEB9F9A7" pageId="18" pageNumber="519" type="etymology">
<paragraph id="04660F2D2447196BFF376A8CAEB9F9A7" blockId="18.[151,1437,1427,2028]" pageId="18" pageNumber="519">
<emphasis id="36ADD33F2447196BFF376A8CAE6FF98C" bold="true" box="[199,334,1534,1559]" pageId="18" pageNumber="519">Etymology.</emphasis>
The name is a noun in apposition and refers to the small size of the metaplical birdhead-shaped process of this species.
</paragraph>
</subSubSection>
<subSubSection id="4CC35CA62447196BFF376934AA09F877" pageId="18" pageNumber="519" type="materials_examined">
<paragraph id="04660F2D2447196BFF376934ADFDF9C4" blockId="18.[151,1437,1427,2028]" box="[199,732,1606,1631]" pageId="18" pageNumber="519">
<emphasis id="36ADD33F2447196BFF376934AEACF9C4" bold="true" box="[199,397,1606,1631]" pageId="18" pageNumber="519">Material studied</emphasis>
(total: 7 ♂♂, 12 ♀♀, 71 juv.)
</paragraph>
<paragraph id="04660F2D2447196BFF376919AC30F950" blockId="18.[151,1437,1427,2028]" pageId="18" pageNumber="519">
<materialsCitation id="B4B105702447196BFF376919AC30F950" ID-GBIF-Occurrence="1563407781" collectingDate="2007-01-01" collectingDateMax="2007-01-12" collectingDateMin="2007-01-01" collectionCode="ZMUC" collectorName="L. A. Hansen" country="Tanzania" county="Mahenge District" elevation="1550" latitude="-7.7666664" location="West Kilombero Scarp Forest Reserve" longLatPrecision="1303" longitude="36.483334" municipality="Udzungwa Mts" pageId="18" pageNumber="519" specimenCount="1" specimenCount-male="1" stateProvince="Iringa Region" typeStatus="holotype">
<emphasis id="36ADD33F2447196BFF376919AE1CF91F" bold="true" box="[199,317,1643,1668]" pageId="18" pageNumber="519">
<typeStatus id="DB62B18F2447196BFF376919AE19F91F" box="[199,312,1643,1668]" pageId="18" pageNumber="519" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="12DFC4A42447196BFEB7691EAE7DF918" box="[327,348,1644,1667]" pageId="18" pageNumber="519" type="male">♂</specimenCount>
<collectingCountry id="7CCE4FBD2447196BFE956919AED6F918" box="[357,503,1643,1667]" name="Tanzania" pageId="18" pageNumber="519">TANZANIA</collectingCountry>
,
<collectingRegion id="C61DC1CF2447196BFDF56919AD84F91F" box="[517,677,1643,1668]" country="Tanzania" name="Iringa" pageId="18" pageNumber="519">Iringa Region</collectingRegion>
,
<collectingCounty id="ED0777A12447196BFD436919AC5DF91F" box="[691,892,1643,1668]" pageId="18" pageNumber="519">Mahenge District</collectingCounty>
,
<collectingMunicipality id="E40295572447196BFC796919AB1BF918" box="[905,1082,1643,1668]" pageId="18" pageNumber="519">Udzungwa Mts</collectingMunicipality>
,
<location id="010659F62447196BFBB9691EAFD5F93C" LSID="urn:lsid:plazi:treatment:8C70BE3B2447196AFF676A5AAFD8FC8F:010659F62447196BFBB9691EAFD5F93C" country="Tanzania" county="Mahenge District" latitude="-7.7666664" longLatPrecision="1303" longitude="36.483334" municipality="Udzungwa Mts" name="West Kilombero Scarp Forest Reserve" pageId="18" pageNumber="519" stateProvince="Iringa Region">West Kilombero Scarp Forest Reserve</location>
,
<location id="010659F62447196BFEF569FCAEE7F93C" LSID="urn:lsid:plazi:treatment:8C70BE3B2447196AFF676A5AAFD8FC8F:010659F62447196BFEF569FCAEE7F93C" box="[261,454,1678,1703]" country="Tanzania" county="Mahenge District" latitude="-7.7666664" longLatPrecision="1303" longitude="36.483334" municipality="Udzungwa Mts" name="Ndundulu forest" pageId="18" pageNumber="519" stateProvince="Iringa Region">Ndundulu forest</location>
,
<geoCoordinate id="61ED69EA2447196BFE2669FCAD15F93C" box="[470,564,1678,1703]" direction="south" orientation="latitude" pageId="18" pageNumber="519" precision="925" value="-7.7666664">07°46’S</geoCoordinate>
,
<geoCoordinate id="61ED69EA2447196BFDB769FCAD86F93C" box="[583,679,1678,1703]" direction="east" orientation="longitude" pageId="18" pageNumber="519" precision="925" value="36.483334">36°29’E</geoCoordinate>
, Waller’s camp, tropical semi-evergreen forest,
<elevation id="8FF4E81E2447196BFB1469FCAA4EF93C" box="[1252,1391,1678,1703]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="18" pageNumber="519" unit="m" value="1550.0">
<quantity id="C321A2C82447196BFB1469FCAA61F93C" box="[1252,1344,1678,1703]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="18" pageNumber="519" unit="m" value="1550.0">1550 m</quantity>
asl
</elevation>
,
<collectingDate id="6023D0052447196BFA7069FDAE22F957" pageId="18" pageNumber="519" value="2007-01-01" valueMax="2007-01-12" valueMin="2007-01-01">
<date id="706729ED2447196BFA7069FDAE22F957" pageId="18" pageNumber="519" value="2007-01-01" valueMax="2007-01-12" valueMin="2007-01-01">1– 12.i.2007</date>
</collectingDate>
,
<collectorName id="A92C6AFB2447196BFEFE69C6AE85F950" box="[270,420,1715,1739]" pageId="18" pageNumber="519">L. A. Hansen</collectorName>
&amp; local assistants leg. (
<collectionCode id="62C897E82447196BFD5E69C1AC24F950" box="[686,773,1715,1739]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="18" pageNumber="519">ZMUC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="04660F2D2447196BFF3769A4AB67F838" blockId="18.[151,1437,1427,2028]" pageId="18" pageNumber="519">
<materialsCitation id="B4B105702447196BFF3769A4AD89F888" ID-GBIF-Occurrence="1563407770" collectionCode="ZMUC" country="Tanzania" county="All" location="Udzungwa Mts" municipality="Mahenge District" pageId="18" pageNumber="519" specimenCount="1" stateProvince="Iringa Region" typeStatus="paratype">
<emphasis id="36ADD33F2447196BFF3769A4AE69F974" bold="true" box="[199,328,1750,1775]" pageId="18" pageNumber="519">
<typeStatus id="DB62B18F2447196BFF3769A4AE62F974" box="[199,323,1750,1775]" pageId="18" pageNumber="519" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectingCounty id="ED0777A12447196BFEA169A4AE54F974" box="[337,373,1750,1775]" pageId="18" pageNumber="519">All</collectingCounty>
from
<collectingCountry id="7CCE4FBD2447196BFE4D69A5AD6FF974" box="[445,590,1751,1775]" name="Tanzania" pageId="18" pageNumber="519">TANZANIA</collectingCountry>
,
<collectingRegion id="C61DC1CF2447196BFDAB69A4ADDAF974" box="[603,763,1750,1775]" country="Tanzania" name="Iringa" pageId="18" pageNumber="519">Iringa Region</collectingRegion>
,
<collectingMunicipality id="E40295572447196BFCF769A4ACEEF974" box="[775,975,1750,1775]" pageId="18" pageNumber="519">Mahenge District</collectingMunicipality>
,
<location id="010659F62447196BFC2C69A4ABACF974" LSID="urn:lsid:plazi:treatment:8C70BE3B2447196AFF676A5AAFD8FC8F:010659F62447196BFC2C69A4ABACF974" box="[988,1165,1750,1775]" country="Tanzania" county="All" municipality="Mahenge District" name="Udzungwa Mts" pageId="18" pageNumber="519" stateProvince="Iringa Region">Udzungwa Mts</location>
,
<location id="010659F62447196BFB6A69AAAFFCF888" LSID="urn:lsid:plazi:treatment:8C70BE3B2447196AFF676A5AAFD8FC8F:010659F62447196BFB6A69AAAFFCF888" country="Tanzania" county="All" municipality="Mahenge District" name="West Kilombero Scarp Forest" pageId="18" pageNumber="519" stateProvince="Iringa Region">West Kilombero Scarp Forest</location>
reserve,
<location id="010659F62447196BFEB66989AD20F88F" LSID="urn:lsid:plazi:treatment:8C70BE3B2447196AFF676A5AAFD8FC8F:010659F62447196BFEB66989AD20F88F" box="[326,513,1787,1812]" country="Tanzania" county="All" municipality="Mahenge District" name="Ndundulu forest" pageId="18" pageNumber="519" stateProvince="Iringa Region">Ndundulu forest</location>
, all in
<collectionCode id="62C897E82447196BFDA06989AD85F888" box="[592,676,1787,1811]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="18" pageNumber="519">ZMUC</collectionCode>
.
</materialsCitation>
—2 ♂♂ 5 ♀♀ 45 juv., data as holotype.—
<materialsCitation id="B4B105702447196BFB656989AE7AF8AC" ID-GBIF-Occurrence="1563407778" collectingDate="2006-12" collectionCode="ZMUC" country="Tanzania" county="All" location="Udzungwa Mts" municipality="Mahenge District" pageId="18" pageNumber="519" specimenCount="32" specimenCount-female="5" specimenCount-male="1" stateProvince="Iringa Region" typeStatus="paratype">
<specimenCount id="12DFC4A42447196BFB656989AB9AF888" box="[1173,1211,1787,1812]" pageId="18" pageNumber="519" type="male">1 ♂</specimenCount>
<specimenCount id="12DFC4A42447196BFB336989AA25F888" box="[1219,1284,1787,1812]" pageId="18" pageNumber="519" type="female">5 ♀♀</specimenCount>
<specimenCount id="12DFC4A42447196BFAFD6989AA7AF88F" box="[1293,1371,1787,1812]" pageId="18" pageNumber="519" type="generic">26 juv.</specimenCount>
same data but
<collectingDate id="6023D0052447196BFF07686CAE76F8AC" box="[247,343,1822,1847]" pageId="18" pageNumber="519" value="2006-12">
<date id="706729ED2447196BFF07686CAE76F8AC" box="[247,343,1822,1847]" pageId="18" pageNumber="519" value="2006-12">xii.2006</date>
</collectingDate>
.
</materialsCitation>
-
<materialsCitation id="B4B105702447196BFE82686DADABF8C7" ID-GBIF-Occurrence="1563407774" collectingDate="2007-02-01" collectingDateMax="2007-02-22" collectingDateMin="2007-02-01" collectionCode="ZMUC" collectorName="J. Madanga &amp; L. A. Hansen" country="Tanzania" county="All" elevation="1880" location="Luwala" municipality="Mahenge District" pageId="18" pageNumber="519" specimenCount="1" specimenCount-male="1" stateProvince="Iringa Region" typeStatus="paratype">
<specimenCount id="12DFC4A42447196BFE82686DAEBDF8AC" box="[370,412,1823,1847]" pageId="18" pageNumber="519" type="male">1 ♂</specimenCount>
same data but
<location id="010659F62447196BFDB4686CADB9F8AC" LSID="urn:lsid:plazi:treatment:8C70BE3B2447196AFF676A5AAFD8FC8F:010659F62447196BFDB4686CADB9F8AC" box="[580,664,1822,1847]" name="Luwala" pageId="18" pageNumber="519">Luwala</location>
camp, semi-evergreen tropical, montane virgin forest,
<elevation id="8FF4E81E2447196BFB0E686DAA5CF8AC" box="[1278,1405,1822,1847]" metricMagnitude="3" metricUnit="m" metricValue="1.88" pageId="18" pageNumber="519" unit="m" value="1880.0">
<quantity id="C321A2C82447196BFB0E686DAA72F8AC" box="[1278,1363,1823,1847]" metricMagnitude="3" metricUnit="m" metricValue="1.88" pageId="18" pageNumber="519" unit="m" value="1880.0">1880 m</quantity>
asl
</elevation>
,
<collectingDate id="6023D0052447196BFA72686CAE2AF8C7" pageId="18" pageNumber="519" value="2007-02-01" valueMax="2007-02-22" valueMin="2007-02-01">
<date id="706729ED2447196BFA72686CAE2AF8C7" pageId="18" pageNumber="519" value="2007-02-01" valueMax="2007-02-22" valueMin="2007-02-01">1– 22.ii.2007</date>
</collectingDate>
,
<collectorName id="A92C6AFB2447196BFEE86836AEBBF8C7" box="[280,410,1859,1884]" pageId="18" pageNumber="519">J. Madanga</collectorName>
&amp;
<collectorName id="A92C6AFB2447196BFE306836AD79F8C0" box="[448,600,1859,1883]" pageId="18" pageNumber="519">L. A. Hansen</collectorName>
leg.
</materialsCitation>
—
<materialsCitation id="B4B105702447196BFD5A6831AD0DF8E4" ID-GBIF-Occurrence="1563407772" collectingDate="1993-08" collectorName="L. L. Sorensen" country="Tanzania" elevation="1750" latitude="-7.7833333" location="Udekwa Village" longLatPrecision="1302" longitude="36.483334" pageId="18" pageNumber="519" specimenCount="1" specimenCount-male="1" stateProvince="Iringa Region" typeStatus="paratype">
<specimenCount id="12DFC4A42447196BFD5A6831ADF1F8C0" box="[682,720,1859,1884]" pageId="18" pageNumber="519" type="male">1 ♂</specimenCount>
ESE
<location id="010659F62447196BFCE36831ACEDF8C7" LSID="urn:lsid:plazi:treatment:8C70BE3B2447196AFF676A5AAFD8FC8F:010659F62447196BFCE36831ACEDF8C7" box="[787,972,1859,1884]" country="Tanzania" latitude="-7.7833333" longLatPrecision="1302" longitude="36.483334" name="Udekwa Village" pageId="18" pageNumber="519" stateProvince="Iringa Region">Udekwa Village</location>
,
<geoCoordinate id="61ED69EA2447196BFC286831AB17F8C0" box="[984,1078,1859,1883]" direction="south" orientation="latitude" pageId="18" pageNumber="519" precision="925" value="-7.7833333">07°47’S</geoCoordinate>
,
<geoCoordinate id="61ED69EA2447196BFBB36831AB85F8C7" box="[1091,1188,1859,1884]" direction="east" orientation="longitude" pageId="18" pageNumber="519" precision="925" value="36.483334">36°29’E</geoCoordinate>
,
<elevation id="8FF4E81E2447196BFB406831AA10F8C7" box="[1200,1329,1859,1884]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="18" pageNumber="519" unit="m" value="1750.0">
<quantity id="C321A2C82447196BFB406831AA26F8C0" box="[1200,1287,1859,1884]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="18" pageNumber="519" unit="m" value="1750.0">1750 m</quantity>
asl
</elevation>
, in rotten wood,
<collectingDate id="6023D0052447196BFF136814AE6AF8E4" box="[227,331,1894,1919]" pageId="18" pageNumber="519" value="1993-08">
<date id="706729ED2447196BFF136814AE6AF8E4" box="[227,331,1894,1919]" pageId="18" pageNumber="519" value="1993-08">viii.1993</date>
</collectingDate>
,
<collectorName id="A92C6AFB2447196BFEA6681AAEDAF8E4" box="[342,507,1895,1919]" pageId="18" pageNumber="519">L. L. Sørensen</collectorName>
leg.
</materialsCitation>
-
<materialsCitation id="B4B105702447196BFDB46815AB7FF8E4" ID-GBIF-Occurrence="1563407769" box="[580,1118,1894,1919]" collectingDate="1993-08" collectorName="L. L. Sorensen" country="Tanzania" elevation="1750" latitude="-7.7833333" location="Udekwa Village" longLatPrecision="1302" longitude="36.483334" pageId="18" pageNumber="519" specimenCount="1" specimenCount-male="1" stateProvince="Iringa Region" typeStatus="paratype">
<specimenCount id="12DFC4A42447196BFDB46815AD4FF8E4" box="[580,622,1895,1919]" pageId="18" pageNumber="519" type="male">1 ♂</specimenCount>
same data but
<collectingDate id="6023D0052447196BFCE76814AC5FF8E4" box="[791,894,1894,1919]" pageId="18" pageNumber="519" value="1993-08">
<date id="706729ED2447196BFCE76814AC5FF8E4" box="[791,894,1894,1919]" pageId="18" pageNumber="519" value="1993-08">viii.1993</date>
</collectingDate>
,
<collectorName id="A92C6AFB2447196BFC7A681AAB11F8E4" box="[906,1072,1895,1919]" pageId="18" pageNumber="519">L. L. Sørensen</collectorName>
leg.
</materialsCitation>
—
<materialsCitation id="B4B105702447196BFB8C6814AB67F838" ID-GBIF-Occurrence="1563407771" collectingDate="1993-08" collectorName="L. L. Sorensen" country="Tanzania" elevation="1700" latitude="-7.7833333" location="Udekwa Village" longLatPrecision="1302" longitude="36.483334" pageId="18" pageNumber="519" specimenCount="2" specimenCount-female="2" stateProvince="Iringa Region" typeStatus="paratype">
<specimenCount id="12DFC4A42447196BFB8C6814AB9AF8E4" box="[1148,1211,1894,1919]" pageId="18" pageNumber="519" type="female">2 ♀♀</specimenCount>
same data but
<elevation id="8FF4E81E2447196BFA946815AFF7F83F" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="18" pageNumber="519" unit="m" value="1700.0">
<quantity id="C321A2C82447196BFA946815AF8CF838" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="18" pageNumber="519" unit="m" value="1700.0">1700 m</quantity>
asl
</elevation>
, in rotten wood,
<collectingDate id="6023D0052447196BFE6268F9AEDBF83F" box="[402,506,1931,1956]" pageId="18" pageNumber="519" value="1993-08">
<date id="706729ED2447196BFE6268F9AEDBF83F" box="[402,506,1931,1956]" pageId="18" pageNumber="519" value="1993-08">viii.1993</date>
</collectingDate>
,
<collectorName id="A92C6AFB2447196BFDF468FEAD8BF838" box="[516,682,1931,1955]" pageId="18" pageNumber="519">L. L. Sørensen</collectorName>
leg. (in same sample as
<taxonomicName id="C3D974AE2447196BFC4C68FEAB17F83F" box="[956,1078,1931,1956]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="18" pageNumber="519" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="36ADD33F2447196BFC4C68FEAB17F83F" box="[956,1078,1931,1956]" italics="true" pageId="18" pageNumber="519">T. hamatus</emphasis>
</taxonomicName>
).
</materialsCitation>
</paragraph>
<paragraph id="04660F2D2447196BFF3768DCAA09F877" blockId="18.[151,1437,1427,2028]" pageId="18" pageNumber="519">
<materialsCitation id="B4B105702447196BFF3768DCAA09F877" ID-GBIF-Occurrence="1563407784" country="Tanzania" county="Mahenge District" elevation="1550" latitude="-7.7666664" location="West Kilombero Scarp Forest Reserve" longLatPrecision="1303" longitude="36.483334" municipality="Udzungwa Mts" pageId="18" pageNumber="519" specimenCount="1" stateProvince="Iringa Region" typeStatus="holotype">
<emphasis id="36ADD33F2447196BFF3768DCAE46F85C" bold="true" box="[199,359,1966,1991]" pageId="18" pageNumber="519">
<typeStatus id="DB62B18F2447196BFF3768DCAE23F85C" box="[199,258,1966,1991]" pageId="18" pageNumber="519">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="7CCE4FBD2447196BFE9F68DDAD20F85C" box="[367,513,1967,1991]" name="Tanzania" pageId="18" pageNumber="519">TANZANIA</collectingCountry>
,
<collectingRegion id="C61DC1CF2447196BFDFD68DCAD8CF85C" box="[525,685,1966,1991]" country="Tanzania" name="Iringa" pageId="18" pageNumber="519">Iringa Region</collectingRegion>
,
<collectingCounty id="ED0777A12447196BFD4A68DCACA3F85C" box="[698,898,1966,1991]" pageId="18" pageNumber="519">Mahenge District</collectingCounty>
,
<collectingMunicipality id="E40295572447196BFC7E68DCAB1FF85C" box="[910,1086,1966,1991]" pageId="18" pageNumber="519">Udzungwa Mts</collectingMunicipality>
,
<location id="010659F62447196BFBBC68C2AFD5F870" LSID="urn:lsid:plazi:treatment:8C70BE3B2447196AFF676A5AAFD8FC8F:010659F62447196BFBBC68C2AFD5F870" country="Tanzania" county="Mahenge District" latitude="-7.7666664" longLatPrecision="1303" longitude="36.483334" municipality="Udzungwa Mts" name="West Kilombero Scarp Forest Reserve" pageId="18" pageNumber="519" stateProvince="Iringa Region">West Kilombero Scarp Forest Reserve</location>
,
<location id="010659F62447196BFF0F68A1AE98F877" LSID="urn:lsid:plazi:treatment:8C70BE3B2447196AFF676A5AAFD8FC8F:010659F62447196BFF0F68A1AE98F877" box="[255,441,2003,2028]" country="Tanzania" county="Mahenge District" latitude="-7.7666664" longLatPrecision="1303" longitude="36.483334" municipality="Udzungwa Mts" name="Ndundulu forest" pageId="18" pageNumber="519" stateProvince="Iringa Region">Ndundulu forest</location>
,
<geoCoordinate id="61ED69EA2447196BFE3468A1AD03F877" box="[452,546,2003,2028]" direction="south" orientation="latitude" pageId="18" pageNumber="519" precision="925" value="-7.7666664">07°46’S</geoCoordinate>
,
<geoCoordinate id="61ED69EA2447196BFDDE68A1ADAFF877" box="[558,654,2003,2028]" direction="east" orientation="longitude" pageId="18" pageNumber="519" precision="925" value="36.483334">36°29’E</geoCoordinate>
,
<location id="010659F62447196BFD6968A1ADD6F877" LSID="urn:lsid:plazi:treatment:8C70BE3B2447196AFF676A5AAFD8FC8F:010659F62447196BFD6968A1ADD6F877" box="[665,759,2003,2028]" country="Tanzania" county="Mahenge District" latitude="-7.7666664" longLatPrecision="1303" longitude="36.483334" municipality="Udzungwa Mts" name="Waller's" pageId="18" pageNumber="519" stateProvince="Iringa Region">Waller’s</location>
camp, tropical semi-evergreen forest,
<elevation id="8FF4E81E2447196BFB5668A1AA05F877" box="[1190,1316,2003,2028]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="18" pageNumber="519" unit="m" value="1550.0">
<quantity id="C321A2C82447196BFB5668A1ABDAF870" box="[1190,1275,2003,2028]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="18" pageNumber="519" unit="m" value="1550.0">1550 m</quantity>
asl
</elevation>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="4CC35CA62446196AFF376FE5AE3CFD1E" pageId="19" pageNumber="520" type="description">
<paragraph id="04660F2D2446196AFF376FE5AED5FF4E" blockId="19.[151,1437,151,788]" pageId="19" pageNumber="520">
<emphasis id="36ADD33F2446196AFF376FE5AE78FF2B" bold="true" box="[199,345,151,176]" pageId="19" pageNumber="520">Description.</emphasis>
SIZE (
<figureCitation id="9CE213A82446196AFE5B6FE5AED4FF2B" box="[427,501,151,176]" captionStart="FIGURE 2" captionStartId="5.[151,250,1182,1204]" captionTargetBox="[151,1436,193,1096]" captionTargetId="figure@5.[151,1436,193,1161]" captionTargetPageId="5" captionText="FIGURE 2. Size of Tropostreptus males, expressed as number of podous rings and midbody vertical diameter. Females reach higher diameters than males. See Figs. 11 and 13 for details on T. hamatus and T. sigmatospinus sp. nov." httpUri="https://zenodo.org/record/803710/files/figure.png" pageId="19" pageNumber="520">Fig. 2</figureCitation>
). Length ca.
<quantity id="C321A2C82446196AFD666FEAADF7FF34" box="[662,726,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="19" pageNumber="520" unit="cm" value="8.0">8 cm</quantity>
; vertical body diameter 6.0–
<quantity id="C321A2C82446196AFBDA6FE5ABA4FF34" box="[1066,1157,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="19" pageNumber="520" unit="mm" value="6.6">6.6 mm</quantity>
(males), up to 8.0 mm (females); 47–55 podous rings.
</paragraph>
<paragraph id="04660F2D2446196AFF376F92ADA9FEDB" blockId="19.[151,1437,151,788]" pageId="19" pageNumber="520">COLOUR. After 9 years in alcohol: Head yellowish brown, collum yellowish brown with darker margins. Body rings yellowish from front back to ozopore, behind ozopore first blackish, then amber back to hind margin. Telson yellow. Antennae and legs yellowish brown.</paragraph>
<paragraph id="04660F2D2446196AFF376E3EAB08FEFE" blockId="19.[151,1437,151,788]" box="[199,1065,332,357]" pageId="19" pageNumber="520">SIGILLA. Arranged in one or two rows, not in a broader band as in congeners.</paragraph>
<paragraph id="04660F2D2446196AFF376E02AE3CFD1E" blockId="19.[151,1437,151,788]" pageId="19" pageNumber="520">
GONOPODS (
<figureCitation id="9CE213A82446196AFEA36E1DAEB8FE1C" box="[339,409,367,392]" captionStart="FIGURE 7" captionStartId="13.[151,250,1869,1891]" captionTargetBox="[207,1387,199,1819]" captionTargetId="figure@13.[151,1436,155,1848]" captionTargetPageId="13" captionText="FIGURE 7. Tropostretus spp., gonopods, anterior view. A: T. hamatus, East Usambara Mts, B: T. hamatus, Udzungwa Mts., left telopodite missing, C: T. sigmatospinus sp. nov., D: T. microcephalus sp. nov., E: T. droides sp. nov., F: T. kipunji sp. nov. Scales 0.5 mm. atp = antetorsal process, bh = birdhead-shaped metaplical process, lpl = apicolateral proplical lobe, map = mesapical metaplical process, mp = metaplica, mpl = mesapical proplical lobe, pp = proplica, px = paracoxite, st = sternum, tlp = telopodite." httpUri="https://zenodo.org/record/803720/files/figure.png" pageId="19" pageNumber="520">Figs 7</figureCitation>
D, 8 D, 10 B). Sternum slender tounge-shaped, reaching level of tip of paracoxites. Proplica ending in blunt-triangular process (
<emphasis id="36ADD33F2446196AFDDB6EE6AD74FE36" box="[555,597,404,429]" italics="true" pageId="19" pageNumber="520">mpl</emphasis>
) separated by very shallow sinus from subapico-lateral, sub-semicircular lobe (
<emphasis id="36ADD33F2446196AFF2B6EC5AFD9FE4B" box="[219,248,439,464]" italics="true" pageId="19" pageNumber="520">lpl</emphasis>
). Metaplica in anterior view basally with gently curved ridge flanking sternum; apically with slender, straight, mesal process (
<emphasis id="36ADD33F2446196AFE576EAFAEF6FE6F" box="[423,471,477,500]" italics="true" pageId="19" pageNumber="520">map</emphasis>
) separated by broad sinus from small birdhead-shaped process (
<emphasis id="36ADD33F2446196AFB5D6EAEABE8FE6E" box="[1197,1225,476,501]" italics="true" pageId="19" pageNumber="520">bh</emphasis>
); lateral “beak” of the latter short, pointed, directed laterad. Mesal process reaching further apicad than birdhead-shaped process. Telopodite shortly after emergence from coxa with long, stout, pointed, slightly sigmoid antetorsal process (
<emphasis id="36ADD33F2446196AFA976D57AAAAFDA7" box="[1383,1419,549,572]" italics="true" pageId="19" pageNumber="520">atp</emphasis>
); direction of process basad and slightly laterad. Telopodite distal to antetorsal process slender, simple, without outgrowths.
</paragraph>
</subSubSection>
<subSubSection id="4CC35CA62446196AFF376DFDAFD8FC8F" pageId="19" pageNumber="520" type="distribution">
<paragraph id="04660F2D2446196AFF376DFDAE38FD56" blockId="19.[151,1437,151,788]" pageId="19" pageNumber="520">
<emphasis id="36ADD33F2446196AFF376DFDAE41FD33" bold="true" box="[199,352,655,680]" pageId="19" pageNumber="520">Distribution.</emphasis>
Known only from the Udzungwa Mts,
<collectingRegion id="C61DC1CF2446196AFCD86DE2AC7EFD3C" box="[808,863,656,679]" country="Tanzania" name="Mjini Magharibi" pageId="19" pageNumber="520">West</collectingRegion>
Kilombero Scarp Forest Reserve, altitudes
<quantity id="C321A2C82446196AFAA66DFDAFCDFD57" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.75" metricValueMin="1.55" pageId="19" pageNumber="520" unit="m" value="1650.0" valueMax="1750.0" valueMin="1550.0">1550– 1750 m</quantity>
asl.
</paragraph>
<paragraph id="04660F2D2446196AFF376DA5AFD8FC8F" blockId="19.[151,1437,151,788]" pageId="19" pageNumber="520">
<emphasis id="36ADD33F2446196AFF376DA5AEE5FD6B" bold="true" box="[199,452,727,752]" pageId="19" pageNumber="520">Coexisting congener.</emphasis>
Coexisting with
<taxonomicName id="C3D974AE2446196AFD666DABAC36FD6B" box="[662,791,727,752]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="19" pageNumber="520" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="36ADD33F2446196AFD666DABAC36FD6B" box="[662,791,727,752]" italics="true" pageId="19" pageNumber="520">T. hamatus</emphasis>
</taxonomicName>
in the Udzungwa Mts,
<collectingRegion id="C61DC1CF2446196AFBCE6DAAAB54FD74" box="[1086,1141,728,751]" country="Tanzania" name="Mjini Magharibi" pageId="19" pageNumber="520">West</collectingRegion>
Kilombero Scarp Forest Reserve.
</paragraph>
</subSubSection>
</treatment>
</document>
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