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<document id="786CA1B642FBC22B3E48A5872EF0AD0C" ID-CLB-Dataset="256669" ID-DOI="10.5852/ejt.2023.864.2093" ID-GBIF-Dataset="7f562ba5-17db-46bf-93e8-4d1ba49c2800" ID-ISSN="2118-9773" ID-Zenodo-Dep="7872174" ID-ZooBank="D47B6B04-B2AD-4FDD-B7C4-B71CA6A5BB84" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1682513062582" checkinUser="felipe" docAuthor="Brunke, Adam James" docDate="2023" docId="6C38C875FFB15B51FDDBFB31E0D7FD59" docLanguage="en" docName="ejt-864_117-145.pdf" docOrigin="European Journal of Taxonomy 864" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Indoquedius parallelicollis" docType="treatment" docVersion="5" lastPageNumber="124" masterDocId="9001B00DFFB75B56FFDAFFF6E270FF89" masterDocTitle="Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar" masterLastPageNumber="145" masterPageNumber="117" pageNumber="123" updateTime="1699514916083" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="AAB3CF53FA3B2F3DD7A0C6029B35A9C2">Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar</mods:title>
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<treatment id="6C38C875FFB15B51FDDBFB31E0D7FD59" ID-DOI="http://doi.org/10.5281/zenodo.7872280" ID-GBIF-Taxon="209579366" ID-Zenodo-Dep="7872280" LSID="urn:lsid:plazi:treatment:6C38C875FFB15B51FDDBFB31E0D7FD59" httpUri="http://treatment.plazi.org/id/6C38C875FFB15B51FDDBFB31E0D7FD59" lastPageId="7" lastPageNumber="124" pageId="6" pageNumber="123">
<subSubSection id="AC8B2AE8FFB15B50FDDBFB31E642FB6B" box="[513,1074,1223,1250]" pageId="6" pageNumber="123" type="nomenclature">
<paragraph id="E42E7963FFB15B50FDDBFB31E642FB6B" blockId="6.[513,1074,1223,1285]" box="[513,1074,1223,1250]" pageId="6" pageNumber="123">
<heading id="BF66CE0FFFB15B50FDDBFB31E642FB6B" box="[513,1074,1223,1250]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="123" reason="2">
<taxonomicName id="239102E0FFB15B50FDDBFB31E642FB6B" ID-CoL="3PNKJ" authority="(Scheerpeltz, 1965)" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1965" box="[513,1074,1223,1250]" class="Insecta" family="Staphylinidae" genus="Indoquedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="parallelicollis">
<emphasis id="D6E5A571FFB15B50FDDBFB31E135FB6B" bold="true" box="[513,837,1223,1250]" italics="true" pageId="6" pageNumber="123">Indoquedius parallelicollis</emphasis>
(
<bibRefCitation id="80000492FFB15B50FC8EFB31E65AFB6B" author="Scheerpeltz O." box="[852,1066,1223,1250]" pageId="6" pageNumber="123" pagination="93 - 371" refId="ref13560" refString="Scheerpeltz O. 1965. Wissenschaftliche Ergebnisse der Schwedischen Expedition 1934 nach Indien und Burma. Arkiv for Zoologi 17: 93 - 371." type="journal article" year="1965">Scheerpeltz, 1965</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="AC8B2AE8FFB15B50FD18FB1CE102FA8C" box="[706,882,1258,1285]" pageId="6" pageNumber="123" type="description">
<paragraph id="E42E7963FFB15B50FD18FB1CE102FA8C" blockId="6.[513,1074,1223,1285]" box="[706,882,1258,1285]" pageId="6" pageNumber="123">
<figureCitation id="7CAA65E6FFB15B50FD18FB1CE133FA8C" box="[706,835,1258,1285]" captionStart="Fig" captionStartId="4.[189,228,1904,1928]" captionTargetBox="[193,1394,503,1859]" captionTargetId="figure-51@4.[189,1398,500,1863]" captionTargetPageId="4" captionText="Fig. 1. Habitus of Indoquedius described by Scheerpeltz (1965). A. I. malaisei, ♂, holotype (NHRS). B. I. micantiventris, ♂, holotype (NHRS). C. I. dispersepunctatus, ♀, holotype (NHRS). D. I. parallelicollis, ♀, paratype (NHW). E. I. sanguinipennis syn. nov. (= I. parallelicollis), ♀, holotype (NHRS). F. I. recticollis, ♀, holotype (NHRS). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7872180" httpUri="https://zenodo.org/record/7872180/files/figure.png" pageId="6" pageNumber="123">Figs 1D–E</figureCitation>
,
<figureCitation id="7CAA65E6FFB15B50FC95FB1DE102FA8C" box="[847,882,1259,1285]" captionStart="Fig" captionStartId="25.[188,227,1936,1960]" captionTargetBox="[192,1395,533,1891]" captionTargetId="figure-75@25.[189,1398,529,1894]" captionTargetPageId="25" captionText="Fig. 7. Female tergite X. A. Indoquedius dispersepunctatus (Scheerpeltz, 1965). B. I. parallelicollis (Scheerpeltz, 1965). C. I. recticollis (Scheerpeltz, 1965). D. Malaisdius ruficeps (Scheerpeltz, 1965). E. Quedius (Microsaurus) rutilipennis Scheerpeltz, 1965. F. Q. (M.) impressithorax Scheerpeltz, 1965. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7872200" httpUri="https://zenodo.org/record/7872200/files/figure.png" pageId="6" pageNumber="123">7B</figureCitation>
</paragraph>
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<subSubSection id="AC8B2AE8FFB15B50FF67FADAE65BFA38" pageId="6" pageNumber="123" type="reference_group">
<paragraph id="E42E7963FFB15B50FF67FADAE1E3FACE" blockId="6.[189,1067,1324,1457]" box="[189,915,1324,1351]" pageId="6" pageNumber="123">
<treatmentCitationGroup id="C4815E4DFFB15B50FF67FADAE1E3FACE" box="[189,915,1324,1351]" pageId="6" pageNumber="123">
<treatmentCitation id="65305F72FFB15B50FF67FADAE1E0FACE" author="Scheerpeltz O." box="[189,912,1324,1351]" page="295" pageId="6" pageNumber="123" year="1965">
<taxonomicName id="239102E0FFB15B50FF67FADAE1E0FACE" ID-CoL="4R2J6" authority="Scheerpeltz, 1965: 295" authorityName=", O. Scheerpeltz" authorityPageNumber="295" authorityYear="1965" box="[189,912,1324,1351]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="parallelicollis" subGenus="Indoquedius">
<emphasis id="D6E5A571FFB15B50FF67FADAE36FFACF" box="[189,287,1324,1350]" italics="true" pageId="6" pageNumber="123">Quedius</emphasis>
(
<emphasis id="D6E5A571FFB15B50FEEAFADAE3B1FACF" box="[304,449,1324,1350]" italics="true" pageId="6" pageNumber="123">Indoquedius</emphasis>
)
<emphasis id="D6E5A571FFB15B50FE08FADAE007FACF" box="[466,631,1324,1350]" italics="true" pageId="6" pageNumber="123">parallelicollis</emphasis>
<bibRefCitation id="80000492FFB15B50FDA4FADAE1E0FACE" author="Scheerpeltz O." box="[638,912,1324,1351]" pageId="6" pageNumber="123" pagination="93 - 371" refId="ref13560" refString="Scheerpeltz O. 1965. Wissenschaftliche Ergebnisse der Schwedischen Expedition 1934 nach Indien und Burma. Arkiv for Zoologi 17: 93 - 371." type="journal article" year="1965">Scheerpeltz, 1965: 295</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<paragraph id="E42E7963FFB15B50FF67FAB9E67CFAE0" blockId="6.[189,1067,1324,1457]" box="[189,1036,1359,1386]" pageId="6" pageNumber="123">
<treatmentCitationGroup id="C4815E4DFFB15B50FF67FAB9E67CFAE0" box="[189,1036,1359,1386]" pageId="6" pageNumber="123">
<treatmentCitation id="65305F72FFB15B50FF67FAB9E1E9FAE0" author="Scheerpeltz O." box="[189,921,1359,1386]" page="296" pageId="6" pageNumber="123" year="1965">
<taxonomicName id="239102E0FFB15B50FF67FAB9E1E9FAE0" ID-CoL="4R2J8" authority="Scheerpeltz, 1965: 296" authorityName=", O. Scheerpeltz" authorityPageNumber="296" authorityYear="1965" box="[189,921,1359,1386]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="sanguinipennis" subGenus="Indoquedius">
<emphasis id="D6E5A571FFB15B50FF67FAB9E36FFAE0" box="[189,287,1359,1385]" italics="true" pageId="6" pageNumber="123">Quedius</emphasis>
(
<emphasis id="D6E5A571FFB15B50FEEAFAB9E3B1FAE0" box="[304,449,1359,1385]" italics="true" pageId="6" pageNumber="123">Indoquedius</emphasis>
)
<emphasis id="D6E5A571FFB15B50FE08FAA6E0F4FAE0" box="[466,644,1360,1385]" italics="true" pageId="6" pageNumber="123">sanguinipennis</emphasis>
<bibRefCitation id="80000492FFB15B50FD51FAB9E1E9FAE0" author="Scheerpeltz O." box="[651,921,1359,1386]" pageId="6" pageNumber="123" pagination="93 - 371" refId="ref13560" refString="Scheerpeltz O. 1965. Wissenschaftliche Ergebnisse der Schwedischen Expedition 1934 nach Indien und Burma. Arkiv for Zoologi 17: 93 - 371." type="journal article" year="1965">Scheerpeltz, 1965: 296</bibRefCitation>
</taxonomicName>
</treatmentCitation>
<emphasis id="D6E5A571FFB15B50FC7AFAB9E67CFAE0" bold="true" box="[928,1036,1359,1385]" pageId="6" pageNumber="123">
<taxonomicNameLabel id="CDD6180AFFB15B50FC7AFAB9E67CFAE0" box="[928,1036,1359,1385]" pageId="6" pageNumber="123">syn. nov.</taxonomicNameLabel>
</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="E42E7963FFB15B50FF67FA84E152FA04" blockId="6.[189,1067,1324,1457]" box="[189,802,1394,1421]" pageId="6" pageNumber="123">
<treatmentCitationGroup id="C4815E4DFFB15B50FF67FA84E152FA04" box="[189,802,1394,1421]" pageId="6" pageNumber="123">
<treatmentCitation id="65305F72FFB15B50FF67FA84E0DFFA04" author="Smetana A." box="[189,687,1394,1421]" page="184" pageId="6" pageNumber="123" year="2014">
<taxonomicName id="239102E0FFB15B50FF67FA84E0DFFA04" ID-CoL="3PNJR" authority="Smetana, 2014: 184" authorityName="Smetana" authorityPageNumber="184" authorityYear="2014" box="[189,687,1394,1421]" class="Insecta" family="Staphylinidae" genus="Indoquedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="bicoloris">
<emphasis id="D6E5A571FFB15B50FF67FA84E3CDFA05" box="[189,445,1394,1420]" italics="true" pageId="6" pageNumber="123">Indoquedius bicoloris</emphasis>
<bibRefCitation id="80000492FFB15B50FE1FFA85E0DFFA04" author="Smetana A." box="[453,687,1395,1421]" pageId="6" pageNumber="123" pagination="169 - 186" refId="ref13832" refString="Smetana A. 2014. Contributions to the knowledge of the Quediina (Coleoptera: Staphylinidae: Staphylinini) of China. Part 47. Genus Indoquedius Blackwelder, 1952. Section 1. Studies and Reports Taxonomical Series 10: 169 - 186." type="book chapter" year="2014">Smetana, 2014: 184</bibRefCitation>
</taxonomicName>
</treatmentCitation>
<emphasis id="D6E5A571FFB15B50FD6CFA85E152FA04" bold="true" box="[694,802,1395,1421]" pageId="6" pageNumber="123">
<taxonomicNameLabel id="CDD6180AFFB15B50FD6CFA85E152FA04" box="[694,802,1395,1421]" pageId="6" pageNumber="123">syn. nov.</taxonomicNameLabel>
</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="E42E7963FFB15B50FF67FA60E65BFA38" blockId="6.[189,1067,1324,1457]" box="[189,1067,1430,1457]" pageId="6" pageNumber="123">
<treatmentCitationGroup id="C4815E4DFFB15B50FF67FA60E65BFA38" box="[189,1067,1430,1457]" pageId="6" pageNumber="123">
<treatmentCitation id="65305F72FFB15B50FF67FA60E0FAFA39" author="Yan X. &amp; Li Y. &amp; Zheng F." box="[189,650,1430,1456]" page="108" pageId="6" pageNumber="123" year="2017">
<taxonomicName id="239102E0FFB15B50FF67FA60E0FAFA39" ID-CoL="3PNKM" authority="Yan et al. 2017: 108" authorityName="Yan" authorityPageNumber="108" authorityYear="2017" box="[189,650,1430,1456]" class="Insecta" family="Staphylinidae" genus="Indoquedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="qiuae">
<emphasis id="D6E5A571FFB15B50FF67FA60E3E6FA26" box="[189,406,1430,1456]" italics="true" pageId="6" pageNumber="123">Indoquedius qiuae</emphasis>
<bibRefCitation id="80000492FFB15B50FE47FA60E0FAFA39" author="Yan X. &amp; Li Y. &amp; Zheng F." box="[413,650,1430,1456]" pageId="6" pageNumber="123" pagination="107 - 110" refId="ref14058" refString="Yan X., Li Y. &amp; Zheng F. 2017 A new species of the genus Indoquedius (Coleoptera: Staphylinidae: Staphylininae) from Sichuan, China. Entomotaxonomia 39: 107 - 110. https: // doi. org / 10.11680 / entomotax. 2017011" type="journal article" year="2017">
Yan
<emphasis id="D6E5A571FFB15B50FE0BFA60E07CFA39" box="[465,524,1430,1456]" italics="true" pageId="6" pageNumber="123">et al.</emphasis>
2017: 108
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(synonymized by
<bibRefCitation id="80000492FFB15B50FCBEFA60E66EFA38" author="Smetana A." box="[868,1054,1430,1457]" pageId="6" pageNumber="123" pagination="383 - 384" refId="ref13947" refString="Smetana A. 2017 b. A new synonymy in the genus Indoquedius Blackwelder, 1952 and critical comments on the pertinent paper (Coleoptera, Staphylinidae, Staphylininae, Staphylinini, Indoquediina). Elytra 7: 383 - 384." type="journal article" year="2017">Smetana 2017b</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="AC8B2AE8FFB15B50FF67FA21E121F809" pageId="6" pageNumber="123" type="diagnosis">
<paragraph id="E42E7963FFB15B50FF67FA21E346FA78" blockId="6.[189,310,1495,1521]" box="[189,310,1495,1521]" pageId="6" pageNumber="123">
<heading id="BF66CE0FFFB15B50FF67FA21E346FA78" bold="true" box="[189,310,1495,1521]" fontSize="11" level="3" pageId="6" pageNumber="123" reason="3">
<emphasis id="D6E5A571FFB15B50FF67FA21E346FA78" bold="true" box="[189,310,1495,1521]" pageId="6" pageNumber="123">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="E42E7963FFB15B50FF67F9F0E121F809" blockId="6.[189,1399,1541,1920]" pageId="6" pageNumber="123">
Among members of the
<taxonomicName id="239102E0FFB15B50FE31F9F3E007F996" authorityName="Fursch" authorityYear="1987" box="[491,631,1541,1567]" class="Insecta" family="Coccinellidae" genus="Bipunctatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="122" phylum="Arthropoda" rank="genus">Bipunctatus</taxonomicName>
group,
<taxonomicName id="239102E0FFB15B50FD00F9F3E66BF996" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1965" box="[730,1051,1541,1567]" class="Insecta" family="Staphylinidae" genus="Indoquedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="parallelicollis">
<emphasis id="D6E5A571FFB15B50FD00F9F3E66BF996" box="[730,1051,1541,1567]" italics="true" pageId="6" pageNumber="123">Indoquedius parallelicollis</emphasis>
</taxonomicName>
can be distinguished by the darkened femora and tibiae (at least outer face) on all legs, the overall pale antennae without any markedly darkened segments, and the moderately coarse and sparse punctation of the elytra. Sympatric
<taxonomicName id="239102E0FFB15B50FF67F999E0B6F900" authority="Zhao &amp; Zhou, 2010" authorityName="Zhao &amp; Zhou" authorityYear="2010" box="[189,710,1647,1673]" class="Insecta" family="Staphylinidae" genus="Indoquedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="bicornutus">
<emphasis id="D6E5A571FFB15B50FF67F999E3A3F900" box="[189,467,1647,1673]" italics="true" pageId="6" pageNumber="123">Indoquedius bicornutus</emphasis>
<bibRefCitation id="80000492FFB15B50FE01F999E0B6F900" author="Zhao Z. - Y. &amp; Zhou H. - Z." box="[475,710,1647,1673]" pageId="6" pageNumber="123" pagination="27 - 38" refId="ref14108" refString="Zhao Z. - Y. &amp; Zhou H. - Z. 2010. Taxonomy of the genus Indoquedius Blackwelder (Coleoptera: Staphylinidae: Staphylininae) of China with description of four new species. Zootaxa 2619: 27 - 38. https: // doi. org / 10.11646 / zootaxa. 2619.1.3" type="journal article" year="2010">Zhao &amp; Zhou, 2010</bibRefCitation>
</taxonomicName>
is very similar but can be distinguished by the pale tibiae, coarser and denser elytral punctation.
<bibRefCitation id="80000492FFB15B50FDA6F964E133F925" author="Smetana A." box="[636,835,1682,1708]" pageId="6" pageNumber="123" refId="ref13920" refString="Smetana A. 2017 a. Quediine Subtribes of Staphylinini (Coleoptera, Staphylinidae, Staphylininae) of Mainland China. J. Farkac, Prague." type="book" year="2017">Smetana (2017a)</bibRefCitation>
had stated that the elytra of
<taxonomicName id="239102E0FFB15B50FB4AF965E73DF925" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1965" box="[1168,1357,1682,1708]" class="Insecta" family="Staphylinidae" genus="Indoquedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="parallelicollis">
<emphasis id="D6E5A571FFB15B50FB4AF965E73DF925" box="[1168,1357,1682,1708]" italics="true" pageId="6" pageNumber="123">I. parallelicollis</emphasis>
</taxonomicName>
(as
<taxonomicName id="239102E0FFB15B50FF67F940E34EF946" authorityName="Smetana" authorityYear="2014" box="[189,318,1717,1743]" class="Insecta" family="Staphylinidae" genus="Indoquedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="bicoloris">
<emphasis id="D6E5A571FFB15B50FF67F940E34EF946" box="[189,318,1717,1743]" italics="true" pageId="6" pageNumber="123">I. bicoloris</emphasis>
</taxonomicName>
) were either entirely red or entirely dark but among the
<typeStatus id="3B2AC7C1FFB15B50FC05F940E634F959" box="[991,1092,1718,1744]" pageId="6" pageNumber="123" type="paratype">paratype</typeStatus>
specimens examined was
<specimenCount id="F297B2EAFFB15B50FF64F92FE32EF97A" box="[190,350,1753,1779]" pageId="6" pageNumber="123" type="generic">one specimen</specimenCount>
with dark elytra and a faint basolateral red spot, similar to what can occur in
<taxonomicName id="239102E0FFB15B50FB03F92CE71EF97A" authorityName="Zhao &amp; Zhou" authorityYear="2010" box="[1241,1390,1753,1779]" class="Insecta" family="Staphylinidae" genus="Indoquedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="bicornutus">
<emphasis id="D6E5A571FFB15B50FB03F92CE71EF97A" box="[1241,1390,1753,1779]" italics="true" pageId="6" pageNumber="123">I. bicornutus</emphasis>
</taxonomicName>
. The
<typeStatus id="3B2AC7C1FFB15B50FF2FF90BE32AF89E" box="[245,346,1789,1815]" pageId="6" pageNumber="123" type="paratype">paratype</typeStatus>
of
<taxonomicName id="239102E0FFB15B50FE52F90BE039F89F" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1965" box="[392,585,1788,1814]" class="Insecta" family="Staphylinidae" genus="Indoquedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="parallelicollis">
<emphasis id="D6E5A571FFB15B50FE52F90BE039F89F" box="[392,585,1788,1814]" italics="true" pageId="6" pageNumber="123">I. parallelicollis</emphasis>
</taxonomicName>
represents another transitional color pattern with less obvious pale areas of the elytra basally and posteriorly (
<figureCitation id="7CAA65E6FFB15B50FD1DF8E9E157F8B0" box="[711,807,1823,1849]" captionStart="Fig" captionStartId="4.[189,228,1904,1928]" captionTargetBox="[193,1394,503,1859]" captionTargetId="figure-51@4.[189,1398,500,1863]" captionTargetPageId="4" captionText="Fig. 1. Habitus of Indoquedius described by Scheerpeltz (1965). A. I. malaisei, ♂, holotype (NHRS). B. I. micantiventris, ♂, holotype (NHRS). C. I. dispersepunctatus, ♀, holotype (NHRS). D. I. parallelicollis, ♀, paratype (NHW). E. I. sanguinipennis syn. nov. (= I. parallelicollis), ♀, holotype (NHRS). F. I. recticollis, ♀, holotype (NHRS). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7872180" httpUri="https://zenodo.org/record/7872180/files/figure.png" pageId="6" pageNumber="123">Fig. 1D</figureCitation>
). The abdominal coloration is also slightly more variable than previously stated (
<bibRefCitation id="80000492FFB15B50FD9BF8B4E08FF8D5" author="Smetana A." box="[577,767,1858,1884]" pageId="6" pageNumber="123" refId="ref13920" refString="Smetana A. 2017 a. Quediine Subtribes of Staphylinini (Coleoptera, Staphylinidae, Staphylininae) of Mainland China. J. Farkac, Prague." type="book" year="2017">Smetana 2017a</bibRefCitation>
), with the apices of tergites narrowly paler in some specimens, while they are broadly paler in
<taxonomicName id="239102E0FFB15B50FD6EF890E13AF8F6" authorityName="Zhao &amp; Zhou" authorityYear="2010" box="[692,842,1893,1919]" class="Insecta" family="Staphylinidae" genus="Indoquedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="bicornutus">
<emphasis id="D6E5A571FFB15B50FD6EF890E13AF8F6" box="[692,842,1893,1919]" italics="true" pageId="6" pageNumber="123">I. bicornutus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="AC8B2AE8FFB05B51FF67FEFFE0D7FD59" pageId="7" pageNumber="124" type="materials_examined">
<paragraph id="E42E7963FFB05B51FF67FEFFE31CFEAB" blockId="7.[189,364,264,291]" box="[189,364,264,291]" pageId="7" pageNumber="124">
<heading id="BF66CE0FFFB05B51FF67FEFFE31CFEAB" bold="true" box="[189,364,264,291]" fontSize="11" level="3" pageId="7" pageNumber="124" reason="3">
<emphasis id="D6E5A571FFB05B51FF67FEFFE31CFEAB" bold="true" box="[189,364,264,291]" pageId="7" pageNumber="124">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="E42E7963FFB05B51FF30FEC1E32CFED8" blockId="7.[189,1399,311,514]" box="[234,348,311,337]" pageId="7" pageNumber="124">
<emphasis id="D6E5A571FFB05B51FF30FEC1E32CFED8" bold="true" box="[234,348,311,337]" pageId="7" pageNumber="124">
<typeStatus id="3B2AC7C1FFB05B51FF30FEC1E32CFED8" box="[234,348,311,337]" pageId="7" pageNumber="124" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="E42E7963FFB05B51FF67FEACE3C4FD88" blockId="7.[189,1399,311,514]" pageId="7" pageNumber="124">
<materialsCitation id="54F9733EFFB05B51FF67FEACE3C4FD88" ID-GBIF-Occurrence="4097213329" collectingDate="1934-06-04" collectingMethod="Malaise" collectionCode="NHRS" country="Myanmar" determinerName="Scheerpeltz" elevation="2000" location="Kambaiti" municipality="Kambaiti" pageId="7" pageNumber="124" specimenCode="NHRS-JLKB 000021050" specimenCount="1" specimenCount-male="1" stateProvince="Kachin State" typeStatus="holotype">
<collectingCountry id="9C8639F3FFB05B51FF67FEACE326FEFD" box="[189,342,346,372]" name="Myanmar" pageId="7" pageNumber="124">MYANMAR</collectingCountry>
–
<emphasis id="D6E5A571FFB05B51FEA2FEACE06BFEFD" bold="true" box="[376,539,346,372]" pageId="7" pageNumber="124">
<collectingRegion id="2655B781FFB05B51FEA2FEACE06BFEFD" box="[376,539,346,372]" country="Myanmar" name="Kachin" pageId="7" pageNumber="124">Kachin State</collectingRegion>
</emphasis>
•
<specimenCount id="F297B2EAFFB05B51FDEFFEADE03EFEFC" box="[565,590,347,373]" pageId="7" pageNumber="124" type="male">♂</specimenCount>
; “N.E. Burma,
<collectingMunicipality id="044AE319FFB05B51FCDDFEACE108FEFD" box="[775,888,346,372]" pageId="7" pageNumber="124">Kambaiti</collectingMunicipality>
;
<quantity id="2369D486FFB05B51FC5FFEACE193FEFC" box="[901,995,346,373]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="124" unit="m" value="2000.0">
<elevation id="6FBC9E50FFB05B51FC5FFEACE193FEFC" box="[901,995,346,373]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="124" unit="m" value="2000.0">2000 m</elevation>
</quantity>
; 4/6.1934; Malaise [printed label] /
<typeStatus id="3B2AC7C1FFB05B51FF0AFE88E30EFE11" box="[208,382,382,408]" pageId="7" pageNumber="124" type="holotype">HOLOTYPUS</typeStatus>
[red handwritten label] / TYPUS
<taxonomicName id="239102E0FFB05B51FCFEFE8BE683FE1E" authority=", O. Scheerpeltz" authorityName=", O. Scheerpeltz" authorityYear="1965" box="[804,1267,381,408]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="124" phylum="Arthropoda" rank="species" species="parallelicollis">Quedius parallelicollis, O. Scheerpeltz</taxonomicName>
[dark red-pink label] /
<originalDetermination id="C78FC19FFFB05B51FE86FE57E1ECFE32" box="[348,924,417,444]" pageId="7" pageNumber="124">
<taxonomicName id="239102E0FFB05B51FE86FE57E167FE32" authorityName=", O. Scheerpeltz" authorityYear="1965" box="[348,791,417,444]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="124" phylum="Arthropoda" rank="species" species="parallelicollis" status="sp. nov." subGenus="Indoquedius">Quedius (Indoquedius) parallelicollis</taxonomicName>
<taxonomicNameLabel id="CDD6180AFFB05B51FCF8FE57E1ECFE32" box="[802,924,417,443]" pageId="7" pageNumber="124" rank="species">nov. spec.</taxonomicNameLabel>
</originalDetermination>
, det.
<determinerName id="11F295A7FFB05B51FC3BFE57E61AFE32" box="[993,1130,417,443]" pageId="7" pageNumber="124">Scheerpeltz</determinerName>
1944 [printed label] / 6664 E91 [printed blue label] /
<taxonomicName id="239102E0FFB05B51FDECFE32E118FE57" authority="Scheerpeltz" authorityName=", O. Scheerpeltz" authorityYear="1965" box="[566,872,452,478]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="124" phylum="Arthropoda" rank="species" species="parallelicollis">parallelicollis Scheerpeltz</taxonomicName>
[large, printed, folded label] /
<specimenCode id="B437D118FFB05B51FB02FE32E338FD8B" pageId="7" pageNumber="124">NHRS-JLKB 000021050</specimenCode>
”;
<collectionCode id="8280E1A6FFB05B51FE87FE11E3DFFD88" box="[349,431,487,513]" pageId="7" pageNumber="124">NHRS</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="E42E7963FFB05B51FF30FDDFE32CFDCA" blockId="7.[189,1399,553,720]" box="[234,348,553,579]" pageId="7" pageNumber="124">
<emphasis id="D6E5A571FFB05B51FF30FDDFE32CFDCA" bold="true" box="[234,348,553,579]" pageId="7" pageNumber="124">
<typeStatus id="3B2AC7C1FFB05B51FF30FDDFE32CFDCA" box="[234,348,553,579]" pageId="7" pageNumber="124" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="E42E7963FFB05B51FF67FDBAE0D7FD59" blockId="7.[189,1399,553,720]" pageId="7" pageNumber="124">
<materialsCitation id="54F9733EFFB05B51FF67FDBAE0D7FD59" ID-GBIF-Occurrence="4097213315" collectingDate="1934-06-04" collectingMethod="Malaise" collectionCode="NHW" collectorName="N. E. Burma &amp; Scheerpeltz" country="Myanmar" elevation="2000" location="Kambaiti" municipality="Kambaiti" pageId="7" pageNumber="124" specimenCount="1" specimenCount-female="1" stateProvince="Kachin State" typeStatus="allotype">
<collectingCountry id="9C8639F3FFB05B51FF67FDBAE326FDEF" box="[189,342,588,614]" name="Myanmar" pageId="7" pageNumber="124">MYANMAR</collectingCountry>
–
<emphasis id="D6E5A571FFB05B51FEA9FDBDE065FDEC" bold="true" box="[371,533,587,613]" pageId="7" pageNumber="124">
<collectingRegion id="2655B781FFB05B51FEA9FDBDE065FDEC" box="[371,533,587,613]" country="Myanmar" name="Kachin" pageId="7" pageNumber="124">Kachin State</collectingRegion>
</emphasis>
•
<specimenCount id="F297B2EAFFB05B51FDF7FDBAE02AFDEF" box="[557,602,588,614]" pageId="7" pageNumber="124" type="female">1 ♀</specimenCount>
; “
<collectorName id="49641CB5FFB05B51FDAEFDBAE175FDEF" box="[628,773,588,614]" pageId="7" pageNumber="124">N.E. Burma</collectorName>
,
<collectingMunicipality id="044AE319FFB05B51FCCBFDBAE1F2FDEF" box="[785,898,588,614]" pageId="7" pageNumber="124">Kambaiti</collectingMunicipality>
;
<quantity id="2369D486FFB05B51FC57FDBAE199FDEF" box="[909,1001,588,614]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="124" unit="m" value="2000.0">
<elevation id="6FBC9E50FFB05B51FC57FDBAE199FDEF" box="[909,1001,588,614]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="124" unit="m" value="2000.0">2000 m</elevation>
</quantity>
; 4/6.1934; Malaise [printed label] / Schwedisch Indian-Burma Expedition 1934 [printed label] / ex. coll.
<collectorName id="49641CB5FFB05B51FC2DFD99E6F0FD00" box="[1015,1152,623,649]" pageId="7" pageNumber="124">Scheerpeltz</collectorName>
[printed blue label] /
<typeStatus id="3B2AC7C1FFB05B51FF67FD64E318FD25" box="[189,360,658,684]" pageId="7" pageNumber="124" type="allotype">ALLOTYPUS</typeStatus>
[red handwritten label] / TYPUS
<taxonomicName id="239102E0FFB05B51FD27FD64E6B5FD25" authority=", O. Scheerpeltz" authorityName=", O. Scheerpeltz" authorityYear="1965" box="[765,1221,658,684]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="124" phylum="Arthropoda" rank="species" species="parallelicollis">
<originalDetermination id="C78FC19FFFB05B51FD27FD64E67AFD25" box="[765,1034,658,684]" pageId="7" pageNumber="124">Quedius parallelicollis</originalDetermination>
, O. Scheerpeltz
</taxonomicName>
[dark red-pink label] / parallelicollis [pink card]”;
<collectionCode id="8280E1A6FFB05B51FD86FD40E0D3FD59" box="[604,675,694,720]" pageId="7" pageNumber="124">NHW</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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