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<document id="B9179A55107DAD0EFA5E8BB20894E9BE" ID-CLB-Dataset="256669" ID-DOI="10.5852/ejt.2023.864.2093" ID-GBIF-Dataset="7f562ba5-17db-46bf-93e8-4d1ba49c2800" ID-ISSN="2118-9773" ID-Zenodo-Dep="7872174" ID-ZooBank="D47B6B04-B2AD-4FDD-B7C4-B71CA6A5BB84" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1682513062582" checkinUser="felipe" docAuthor="Brunke, Adam James" docDate="2023" docId="6C38C875FFA65B44FE1BFB22E033FA5B" docLanguage="en" docName="ejt-864_117-145.pdf" docOrigin="European Journal of Taxonomy 864" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Quedius (Microsaurus) impressithorax , O. Scheerpeltz 1965" docType="treatment" docVersion="5" lastPageNumber="135" masterDocId="9001B00DFFB75B56FFDAFFF6E270FF89" masterDocTitle="Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar" masterLastPageNumber="145" masterPageNumber="117" pageNumber="134" updateTime="1699514916083" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6E805586777056B5AD53AF99DD2D6749">Review of Quedius (Coleoptera, Staphylinidae) described from the 1934 expedition by R. Malaise to Myanmar</mods:title>
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<mods:title id="F75F6427828753B51DFF27C03D28FE26">European Journal of Taxonomy</mods:title>
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<mods:date id="AA517972AAB84425CBA291EF4E51E2B9">2023</mods:date>
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<treatment id="6C38C875FFA65B44FE1BFB22E033FA5B" ID-DOI="http://doi.org/10.5281/zenodo.7872307" ID-GBIF-Taxon="209579375" ID-Zenodo-Dep="7872307" LSID="urn:lsid:plazi:treatment:6C38C875FFA65B44FE1BFB22E033FA5B" httpUri="http://treatment.plazi.org/id/6C38C875FFA65B44FE1BFB22E033FA5B" lastPageId="18" lastPageNumber="135" pageId="17" pageNumber="134">
<subSubSection id="AC8B2AE8FFA65B47FE1BFB22E602FB79" box="[449,1138,1236,1264]" pageId="17" pageNumber="134" type="nomenclature">
<paragraph id="E42E7963FFA65B47FE1BFB22E602FB79" blockId="17.[449,1138,1236,1298]" box="[449,1138,1236,1264]" pageId="17" pageNumber="134">
<heading id="BF66CE0FFFA65B47FE1BFB22E602FB79" box="[449,1138,1236,1264]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="134" reason="2">
<taxonomicName id="239102E0FFA65B47FE1BFB22E602FB79" ID-CoL="4R3P8" authority="Scheerpeltz, 1965" authorityName=", O. Scheerpeltz" authorityYear="1965" box="[449,1138,1236,1264]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="134" phylum="Arthropoda" rank="species" species="impressithorax" subGenus="Microsaurus">
<emphasis id="D6E5A571FFA65B47FE1BFB22E056FB66" bold="true" box="[449,550,1236,1263]" italics="true" pageId="17" pageNumber="134">Quedius</emphasis>
(
<emphasis id="D6E5A571FFA65B47FDE2FB23E0A3FB66" bold="true" box="[568,723,1237,1263]" italics="true" pageId="17" pageNumber="134">Microsaurus</emphasis>
)
<emphasis id="D6E5A571FFA65B47FD3EFB22E1E9FB66" bold="true" box="[740,921,1236,1263]" italics="true" pageId="17" pageNumber="134">impressithorax</emphasis>
<bibRefCitation id="80000492FFA65B47FC7AFB23E602FB79" author="Scheerpeltz O." box="[928,1138,1237,1264]" pageId="17" pageNumber="134" pagination="93 - 371" refId="ref13560" refString="Scheerpeltz O. 1965. Wissenschaftliche Ergebnisse der Schwedischen Expedition 1934 nach Indien und Burma. Arkiv for Zoologi 17: 93 - 371." type="journal article" year="1965">Scheerpeltz, 1965</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="AC8B2AE8FFA65B47FD1DFB0EE11CFA9B" box="[711,876,1272,1298]" pageId="17" pageNumber="134" type="description">
<paragraph id="E42E7963FFA65B47FD1DFB0EE11CFA9B" blockId="17.[449,1138,1236,1298]" box="[711,876,1272,1298]" pageId="17" pageNumber="134">
<figureCitation id="7CAA65E6FFA65B47FD1DFB0EE132FA9B" box="[711,834,1272,1298]" captionStart="Fig" captionStartId="16.[189,228,1872,1896]" captionTargetBox="[190,1396,472,1833]" captionTargetId="figure-78@16.[189,1398,471,1834]" captionTargetPageId="16" captionText="Fig. 2. A–E. Dorsal habitus. A. Malaisdius ruficeps (Scheerpeltz, 1965), ♂, paratype (NHW). B. Malaisdius smetanai gen. et. sp. nov., ♂, holotype (NME). C. Quedius (Microsaurus) rutilipennis Scheerpeltz, 1965, ♂, holotype (NHRS). D. Q. (M.) piceolineatus Scheerpeltz, 1965, ♂ holotype (NHRS). E. Q. (M.) impressithorax Scheerpeltz, 1965, ♀ holotype (NHRS). F. Forebody of Q. (M.) impressithorax Scheerpeltz, 1965. Abbreviation: Pf = posterior frontal puncture. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7872182" httpUri="https://zenodo.org/record/7872182/files/figure.png" pageId="17" pageNumber="134">Figs 2E–F</figureCitation>
,
<figureCitation id="7CAA65E6FFA65B47FC97FB0EE11CFA9B" box="[845,876,1272,1298]" captionStart="Fig" captionStartId="25.[188,227,1936,1960]" captionTargetBox="[192,1395,533,1891]" captionTargetId="figure-75@25.[189,1398,529,1894]" captionTargetPageId="25" captionText="Fig. 7. Female tergite X. A. Indoquedius dispersepunctatus (Scheerpeltz, 1965). B. I. parallelicollis (Scheerpeltz, 1965). C. I. recticollis (Scheerpeltz, 1965). D. Malaisdius ruficeps (Scheerpeltz, 1965). E. Quedius (Microsaurus) rutilipennis Scheerpeltz, 1965. F. Q. (M.) impressithorax Scheerpeltz, 1965. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7872200" httpUri="https://zenodo.org/record/7872200/files/figure.png" pageId="17" pageNumber="134">7F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="AC8B2AE8FFA65B47FF67FACFE1D1FADA" box="[189,929,1337,1364]" pageId="17" pageNumber="134" type="reference_group">
<paragraph id="E42E7963FFA65B47FF67FACFE1D1FADA" blockId="17.[189,929,1337,1364]" box="[189,929,1337,1364]" pageId="17" pageNumber="134">
<treatmentCitationGroup id="C4815E4DFFA65B47FF67FACFE1D1FADA" box="[189,929,1337,1364]" pageId="17" pageNumber="134">
<treatmentCitation id="65305F72FFA65B47FF67FACFE1EDFADA" author="Scheerpeltz O." box="[189,925,1337,1364]" page="281" pageId="17" pageNumber="134" year="1965">
<taxonomicName id="239102E0FFA65B47FF67FACFE1EDFADA" ID-CoL="4R3P8" authority="Scheerpeltz, 1965: 281" authorityName=", O. Scheerpeltz" authorityPageNumber="281" authorityYear="1965" box="[189,925,1337,1364]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="134" phylum="Arthropoda" rank="species" species="impressithorax" subGenus="Microsaurus">
<emphasis id="D6E5A571FFA65B47FF67FACFE36FFADA" box="[189,287,1337,1363]" italics="true" pageId="17" pageNumber="134">Quedius</emphasis>
(
<emphasis id="D6E5A571FFA65B47FEEAFACCE3B5FADA" box="[304,453,1338,1363]" italics="true" pageId="17" pageNumber="134">Microsaurus</emphasis>
)
<emphasis id="D6E5A571FFA65B47FE0CFACFE0F5FADA" box="[470,645,1337,1363]" italics="true" pageId="17" pageNumber="134">impressithorax</emphasis>
<bibRefCitation id="80000492FFA65B47FD56FACFE1EDFADA" author="Scheerpeltz O." box="[652,925,1337,1364]" pageId="17" pageNumber="134" pagination="93 - 371" refId="ref13560" refString="Scheerpeltz O. 1965. Wissenschaftliche Ergebnisse der Schwedischen Expedition 1934 nach Indien und Burma. Arkiv for Zoologi 17: 93 - 371." type="journal article" year="1965">Scheerpeltz, 1965: 281</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="AC8B2AE8FFA65B47FF67FA8CE75FF983" pageId="17" pageNumber="134" type="diagnosis">
<paragraph id="E42E7963FFA65B47FF67FA8CE346FA1D" blockId="17.[189,310,1402,1428]" box="[189,310,1402,1428]" pageId="17" pageNumber="134">
<heading id="BF66CE0FFFA65B47FF67FA8CE346FA1D" bold="true" box="[189,310,1402,1428]" fontSize="11" level="3" pageId="17" pageNumber="134" reason="3">
<emphasis id="D6E5A571FFA65B47FF67FA8CE346FA1D" bold="true" box="[189,310,1402,1428]" pageId="17" pageNumber="134">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="E42E7963FFA65B47FF67FA5FE75FF983" blockId="17.[189,1399,1449,1546]" pageId="17" pageNumber="134">
Within the Kiangsiensis group,
<taxonomicName id="239102E0FFA65B47FD9BFA5FE12DFA4A" authorityName=", O. Scheerpeltz" authorityYear="1965" box="[577,861,1449,1475]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="134" phylum="Arthropoda" rank="species" species="impressithorax">
<emphasis id="D6E5A571FFA65B47FD9BFA5FE12DFA4A" box="[577,861,1449,1475]" italics="true" pageId="17" pageNumber="134">Quedius impressithorax</emphasis>
</taxonomicName>
may be distinguished by a combination of: small eyes, temples about 1.5 × as long as eyes (
<figureCitation id="7CAA65E6FFA65B47FD20FA3AE124FA6E" box="[762,852,1484,1511]" captionStart="Fig" captionStartId="16.[189,228,1872,1896]" captionTargetBox="[190,1396,472,1833]" captionTargetId="figure-78@16.[189,1398,471,1834]" captionTargetPageId="16" captionText="Fig. 2. A–E. Dorsal habitus. A. Malaisdius ruficeps (Scheerpeltz, 1965), ♂, paratype (NHW). B. Malaisdius smetanai gen. et. sp. nov., ♂, holotype (NME). C. Quedius (Microsaurus) rutilipennis Scheerpeltz, 1965, ♂, holotype (NHRS). D. Q. (M.) piceolineatus Scheerpeltz, 1965, ♂ holotype (NHRS). E. Q. (M.) impressithorax Scheerpeltz, 1965, ♀ holotype (NHRS). F. Forebody of Q. (M.) impressithorax Scheerpeltz, 1965. Abbreviation: Pf = posterior frontal puncture. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7872182" httpUri="https://zenodo.org/record/7872182/files/figure.png" pageId="17" pageNumber="134">Fig. 2F</figureCitation>
); head slightly elongate, with posterior frontal puncture about three puncture diameters from eye margin (
<figureCitation id="7CAA65E6FFA65B47FCAAFA06E1BAF983" box="[880,970,1520,1546]" captionStart="Fig" captionStartId="16.[189,228,1872,1896]" captionTargetBox="[190,1396,472,1833]" captionTargetId="figure-78@16.[189,1398,471,1834]" captionTargetPageId="16" captionText="Fig. 2. A–E. Dorsal habitus. A. Malaisdius ruficeps (Scheerpeltz, 1965), ♂, paratype (NHW). B. Malaisdius smetanai gen. et. sp. nov., ♂, holotype (NME). C. Quedius (Microsaurus) rutilipennis Scheerpeltz, 1965, ♂, holotype (NHRS). D. Q. (M.) piceolineatus Scheerpeltz, 1965, ♂ holotype (NHRS). E. Q. (M.) impressithorax Scheerpeltz, 1965, ♀ holotype (NHRS). F. Forebody of Q. (M.) impressithorax Scheerpeltz, 1965. Abbreviation: Pf = posterior frontal puncture. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7872182" httpUri="https://zenodo.org/record/7872182/files/figure.png" pageId="17" pageNumber="134">Fig. 2F</figureCitation>
); body entirely dark (
<figureCitation id="7CAA65E6FFA65B47FB1CFA06E752F983" box="[1222,1314,1520,1546]" captionStart="Fig" captionStartId="16.[189,228,1872,1896]" captionTargetBox="[190,1396,472,1833]" captionTargetId="figure-78@16.[189,1398,471,1834]" captionTargetPageId="16" captionText="Fig. 2. A–E. Dorsal habitus. A. Malaisdius ruficeps (Scheerpeltz, 1965), ♂, paratype (NHW). B. Malaisdius smetanai gen. et. sp. nov., ♂, holotype (NME). C. Quedius (Microsaurus) rutilipennis Scheerpeltz, 1965, ♂, holotype (NHRS). D. Q. (M.) piceolineatus Scheerpeltz, 1965, ♂ holotype (NHRS). E. Q. (M.) impressithorax Scheerpeltz, 1965, ♀ holotype (NHRS). F. Forebody of Q. (M.) impressithorax Scheerpeltz, 1965. Abbreviation: Pf = posterior frontal puncture. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7872182" httpUri="https://zenodo.org/record/7872182/files/figure.png" pageId="17" pageNumber="134">Fig. 2E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="AC8B2AE8FFA65B47FF67F9C7E363F8A3" pageId="17" pageNumber="134" type="materials_examined">
<paragraph id="E42E7963FFA65B47FF67F9C7E31CF9C2" blockId="17.[189,364,1585,1611]" box="[189,364,1585,1611]" pageId="17" pageNumber="134">
<heading id="BF66CE0FFFA65B47FF67F9C7E31CF9C2" bold="true" box="[189,364,1585,1611]" fontSize="11" level="3" pageId="17" pageNumber="134" reason="3">
<emphasis id="D6E5A571FFA65B47FF67F9C7E31CF9C2" bold="true" box="[189,364,1585,1611]" pageId="17" pageNumber="134">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="E42E7963FFA65B47FF30F996E32CF9F3" blockId="17.[189,1399,1632,1834]" box="[234,348,1632,1658]" pageId="17" pageNumber="134">
<emphasis id="D6E5A571FFA65B47FF30F996E32CF9F3" bold="true" box="[234,348,1632,1658]" pageId="17" pageNumber="134">
<typeStatus id="3B2AC7C1FFA65B47FF30F996E32CF9F3" box="[234,348,1632,1658]" pageId="17" pageNumber="134" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="E42E7963FFA65B47FF67F975E363F8A3" blockId="17.[189,1399,1632,1834]" pageId="17" pageNumber="134">
<materialsCitation id="54F9733EFFA65B47FF67F975E363F8A3" ID-GBIF-Occurrence="4097213305" collectingDate="1934-05-01" collectionCode="NHRS" collectorName="R. Malaise" country="Myanmar" determinerName="Scheerpeltz" elevation="2134" location="N. E. Burma" municipality="Kambaiti" pageId="17" pageNumber="134" specimenCode="JLKB 000021054" specimenCount="1" specimenCount-female="1" stateProvince="Kachin State" typeStatus="holotype">
<collectingCountry id="9C8639F3FFA65B47FF67F975E326F914" box="[189,342,1667,1693]" name="Myanmar" pageId="17" pageNumber="134">MYANMAR</collectingCountry>
–
<emphasis id="D6E5A571FFA65B47FEAAF975E061F914" bold="true" box="[368,529,1667,1693]" pageId="17" pageNumber="134">
<collectingRegion id="2655B781FFA65B47FEAAF975E061F914" box="[368,529,1667,1693]" country="Myanmar" name="Kachin" pageId="17" pageNumber="134">Kachin State</collectingRegion>
</emphasis>
•
<specimenCount id="F297B2EAFFA65B47FDF3F975E030F914" box="[553,576,1667,1693]" pageId="17" pageNumber="134" type="female">♀</specimenCount>
; “
<location id="E14E2FB8FFA65B47FD83F975E099F914" LSID="urn:lsid:plazi:treatment:6C38C875FFA65B44FE1BFB22E033FA5B:E14E2FB8FFA65B47FD83F975E099F914" box="[601,745,1667,1693]" country="Myanmar" municipality="Kambaiti" name="N. E. Burma" pageId="17" pageNumber="134" stateProvince="Kachin State">N.E. Burma</location>
,
<collectingMunicipality id="044AE319FFA65B47FD29F975E114F914" box="[755,868,1667,1693]" pageId="17" pageNumber="134">Kambaiti</collectingMunicipality>
;
<quantity id="2369D486FFA65B47FCB5F975E1BBF914" box="[879,971,1667,1693]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="17" pageNumber="134" unit="ft" value="7000.0">
<elevation id="6FBC9E50FFA65B47FCB5F975E1BBF914" box="[879,971,1667,1693]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="17" pageNumber="134" unit="ft" value="7000.0">7000 ft.</elevation>
</quantity>
; 1/5.1934;
<collectorName id="49641CB5FFA65B47FB97F975E6BBF914" box="[1101,1227,1667,1693]" pageId="17" pageNumber="134">R. Malaise</collectorName>
[printed label] /
<typeStatus id="3B2AC7C1FFA65B47FF15F950E30DF949" box="[207,381,1702,1728]" pageId="17" pageNumber="134" type="holotype">HOLOTYPUS</typeStatus>
[red handwritten label] / TYPUS
<taxonomicName id="239102E0FFA65B47FCC1F950E686F949" authority=", O. Scheerpeltz" authorityName=", O. Scheerpeltz" authorityYear="1965" box="[795,1270,1702,1728]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="134" phylum="Arthropoda" rank="species" species="impressithorax">Quedius impressithorax, O. Scheerpeltz</taxonomicName>
[dark red-pink label] /
<originalDetermination id="C78FC19FFFA65B47FE82F93FE1D3F96D" box="[344,931,1737,1764]" pageId="17" pageNumber="134">
<taxonomicName id="239102E0FFA65B47FE82F93FE151F96A" authorityName=", O. Scheerpeltz" authorityYear="1965" box="[344,801,1737,1764]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="134" phylum="Arthropoda" rank="species" species="impressithorax" status="sp. nov." subGenus="Microsaurus">Quedius (Microsaurus) impressithorax</taxonomicName>
<taxonomicNameLabel id="CDD6180AFFA65B47FCF0F93CE1D3F96D" box="[810,931,1738,1764]" pageId="17" pageNumber="134" rank="species">nov. spec.</taxonomicNameLabel>
</originalDetermination>
, det.
<determinerName id="11F295A7FFA65B47FC3CF93FE61FF96A" box="[998,1135,1737,1763]" pageId="17" pageNumber="134">Scheerpeltz</determinerName>
[printed label] / 6590 E91 [blue printed label] /
<taxonomicName id="239102E0FFA65B47FE31F91AE159F88F" authority="Scheerpeltz" authorityName=", O. Scheerpeltz" authorityYear="1965" box="[491,809,1772,1798]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="134" phylum="Arthropoda" rank="species" species="impressithorax">impressithorax Scheerpeltz</taxonomicName>
[large, printed, folded label] /
<specimenCode id="B437D118FFA65B47FB4AF91BE718F88E" box="[1168,1384,1772,1799]" pageId="17" pageNumber="134">JLKB 000021054</specimenCode>
”;
<collectionCode id="8280E1A6FFA65B47FF67F8E6E37FF8A3" box="[189,271,1808,1834]" pageId="17" pageNumber="134">NHRS</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="AC8B2AE8FFA65B44FF67F8A7E669FBA9" lastPageId="18" lastPageNumber="135" pageId="17" pageNumber="134" type="description">
<paragraph id="E42E7963FFA65B47FF67F8A7E31BF8E2" blockId="17.[189,363,1873,1899]" box="[189,363,1873,1899]" pageId="17" pageNumber="134">
<heading id="BF66CE0FFFA65B47FF67F8A7E31BF8E2" bold="true" box="[189,363,1873,1899]" fontSize="11" level="3" pageId="17" pageNumber="134" reason="3">
<emphasis id="D6E5A571FFA65B47FF67F8A7E31BF8E2" bold="true" box="[189,363,1873,1899]" pageId="17" pageNumber="134">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="E42E7963FFA65B44FF67F876E3FCFCBA" blockId="17.[189,1399,1920,2017]" lastBlockId="18.[187,1401,265,819]" lastPageId="18" lastPageNumber="135" pageId="17" pageNumber="134">Head, elytra, abdomen, antennae and legs dark brown, pronotum and apices of abdominal tergites slightly paler, dark reddish brown; apical tarsomeres of all legs paler, yellow-brown; abdomen vaguely iridescent. Head slightly longer than wide, with posterior angles indistinct; eyes small and only slightly protruding from lateral head outline, tempora longer than eyes, about 1.5 ×; without interocular punctures or additional punctures near the eye margin; posterior frontal puncture somewhat close to posterior margin of eye, about three puncture diameters, situated closer to margin of eye than to hind margin of head; temporal puncture slightly closer to posterior margin of head than margin of eye; surface of head with fine and dense microsculpture of transverse waves, becoming meshed at middle of disc, with distinct micropunctures moderately densely distributed. Antennae somewhat bulky, antennomere 4 only slightly transverse, 5–10 distinctly transverse, symmetrical, last segment shorter than previous two. Pronotum distinctly transverse, much wide than long, distinctly explanate laterally; dorsal rows with three punctures, sublateral rows with two punctures, not extended posteriad of large lateral puncture; microsculpture similar to that on head. Scutellum entirely and moderately densely punctate. Elytra at base distinctly narrower than pronotum, at both middle and sides, clearly longer than pronotum; punctures rather fine and dense, many punctures touching laterally; pubescence pale, yellow-brown; disc without microsculpture. Wing fully developed. Abdomen with tergite VII with palisade fringe; punctation sparser than that of elytra, punctures clearly separated by their diameter or more; tergite III with very small sparsely punctate area at middle; tergites with extremely fine microsculpture of transverse waves.</paragraph>
<paragraph id="E42E7963FFA55B44FF30FCADE35AFCFC" blockId="18.[189,313,859,920]" box="[234,298,859,885]" pageId="18" pageNumber="135">
<emphasis id="D6E5A571FFA55B44FF30FCADE35AFCFC" bold="true" box="[234,298,859,885]" pageId="18" pageNumber="135">Male</emphasis>
</paragraph>
<paragraph id="E42E7963FFA55B44FF67FC88E349FC11" blockId="18.[189,313,859,920]" box="[189,313,894,920]" pageId="18" pageNumber="135">Unknown.</paragraph>
<paragraph id="E42E7963FFA55B44FF30FC49E335FC50" blockId="18.[189,1398,959,1056]" box="[234,325,959,985]" pageId="18" pageNumber="135">
<emphasis id="D6E5A571FFA55B44FF30FC49E335FC50" bold="true" box="[234,325,959,985]" pageId="18" pageNumber="135">Female</emphasis>
</paragraph>
<paragraph id="E42E7963FFA55B44FF67FC14E669FBA9" blockId="18.[189,1398,959,1056]" pageId="18" pageNumber="135">
Tergite X elongate triangular, strongly narrowed to acutely projected apex, sides of apex sinuate, disc with median darkening from about posterior two-thirds to apex (
<figureCitation id="7CAA65E6FFA55B44FC68FBF3E67CFBA9" box="[946,1036,1029,1056]" captionStart="Fig" captionStartId="25.[188,227,1936,1960]" captionTargetBox="[192,1395,533,1891]" captionTargetId="figure-75@25.[189,1398,529,1894]" captionTargetPageId="25" captionText="Fig. 7. Female tergite X. A. Indoquedius dispersepunctatus (Scheerpeltz, 1965). B. I. parallelicollis (Scheerpeltz, 1965). C. I. recticollis (Scheerpeltz, 1965). D. Malaisdius ruficeps (Scheerpeltz, 1965). E. Quedius (Microsaurus) rutilipennis Scheerpeltz, 1965. F. Q. (M.) impressithorax Scheerpeltz, 1965. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7872200" httpUri="https://zenodo.org/record/7872200/files/figure.png" pageId="18" pageNumber="135">Fig. 7F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="AC8B2AE8FFA55B44FF67FBB1E112FB19" box="[189,866,1095,1168]" pageId="18" pageNumber="135" type="distribution">
<paragraph id="E42E7963FFA55B44FF67FBB1E326FBE8" blockId="18.[189,342,1095,1121]" box="[189,342,1095,1121]" pageId="18" pageNumber="135">
<heading id="BF66CE0FFFA55B44FF67FBB1E326FBE8" bold="true" box="[189,342,1095,1121]" fontSize="11" level="3" pageId="18" pageNumber="135" reason="3">
<emphasis id="D6E5A571FFA55B44FF67FBB1E326FBE8" bold="true" box="[189,342,1095,1121]" pageId="18" pageNumber="135">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="E42E7963FFA55B44FF67FB80E112FB19" blockId="18.[189,866,1141,1168]" box="[189,866,1141,1168]" pageId="18" pageNumber="135">
Known only from the
<typeStatus id="3B2AC7C1FFA55B44FE18FB80E385FB19" box="[450,501,1142,1168]" pageId="18" pageNumber="135">type</typeStatus>
locality in
<collectingCountry id="9C8639F3FFA55B44FDA3FB80E09BFB19" box="[633,747,1142,1168]" name="Myanmar" pageId="18" pageNumber="135">Myanmar</collectingCountry>
(
<collectingRegion id="2655B781FFA55B44FD21FB83E126FB19" box="[763,854,1141,1168]" country="Myanmar" name="Kachin" pageId="18" pageNumber="135">Kachin</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="AC8B2AE8FFA55B44FF67FB41E033FA5B" pageId="18" pageNumber="135" type="discussion">
<paragraph id="E42E7963FFA55B44FF67FB41E35FFB58" blockId="18.[189,303,1207,1233]" box="[189,303,1207,1233]" pageId="18" pageNumber="135">
<heading id="BF66CE0FFFA55B44FF67FB41E35FFB58" bold="true" box="[189,303,1207,1233]" fontSize="11" level="3" pageId="18" pageNumber="135" reason="3">
<emphasis id="D6E5A571FFA55B44FF67FB41E35FFB58" bold="true" box="[189,303,1207,1233]" pageId="18" pageNumber="135">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="E42E7963FFA55B44FF67FB13E033FA5B" blockId="18.[189,1401,1253,1491]" pageId="18" pageNumber="135">
The posterior frontal puncture is located at a distance from the eye that is similar to
<taxonomicName id="239102E0FFA55B44FB55FB10E288FAAB" authority="Smetana, 1996" authorityName="Smetana" authorityYear="1996" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="135" phylum="Arthropoda" rank="species" species="kubani">
<emphasis id="D6E5A571FFA55B44FB55FB10E773FB76" box="[1167,1283,1253,1280]" italics="true" pageId="18" pageNumber="135">Q. kubani</emphasis>
Smetana, 1996
</taxonomicName>
and
<taxonomicName id="239102E0FFA55B44FEEDFAFFE0D4FAAA" authority="Smetana, 2013" authorityName="Smetana" authorityYear="2013" box="[311,676,1288,1315]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="135" phylum="Arthropoda" rank="species" species="kuatunensis">
<emphasis id="D6E5A571FFA55B44FEEDFAFFE398FAAB" box="[311,488,1288,1315]" italics="true" pageId="18" pageNumber="135">Q. kuatunensis</emphasis>
Smetana, 2013
</taxonomicName>
but in these species the head is clearly transverse, the eyes are larger and the apical antennomeres are asymmetrical.
<taxonomicName id="239102E0FFA55B44FCA9FADAE6FDFACF" authorityName=", O. Scheerpeltz" authorityYear="1965" box="[883,1165,1324,1350]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="135" phylum="Arthropoda" rank="species" species="impressithorax">
<emphasis id="D6E5A571FFA55B44FCA9FADAE6FDFACF" box="[883,1165,1324,1350]" italics="true" pageId="18" pageNumber="135">Quedius impressithorax</emphasis>
</taxonomicName>
keys to
<taxonomicName id="239102E0FFA55B44FB2CFADAE31EFAE0" authority="Smetana, 2012" authorityName="Smetana" authorityYear="2012" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="135" phylum="Arthropoda" rank="species" species="albiorix">
<emphasis id="D6E5A571FFA55B44FB2CFADAE709FACF" box="[1270,1401,1324,1350]" italics="true" pageId="18" pageNumber="135">Q. albiorix</emphasis>
Smetana, 2012
</taxonomicName>
(
<collectingRegion id="2655B781FFA55B44FEA7FAB9E390FAE0" box="[381,480,1359,1385]" country="China" name="Sichuan" pageId="18" pageNumber="135">Sichuan</collectingRegion>
,
<collectingCountry id="9C8639F3FFA55B44FE36FAB9E044FAE3" box="[492,564,1359,1386]" name="China" pageId="18" pageNumber="135">China</collectingCountry>
) in
<bibRefCitation id="80000492FFA55B44FDBBFAB9E159FAE0" author="Smetana A." box="[609,809,1359,1385]" pageId="18" pageNumber="135" refId="ref13920" refString="Smetana A. 2017 a. Quediine Subtribes of Staphylinini (Coleoptera, Staphylinidae, Staphylininae) of Mainland China. J. Farkac, Prague." type="book" year="2017">Smetana (2017a)</bibRefCitation>
but is quite different in habitus, with entirely dark antennae, a longer head and a distinctly less distant posterior frontal puncture (4–5 puncture diameters in
<taxonomicName id="239102E0FFA55B44FF07FA60E32FFA26" authorityName="Smetana" authorityYear="2012" box="[221,351,1429,1456]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="135" phylum="Arthropoda" rank="species" species="albiorix">
<emphasis id="D6E5A571FFA55B44FF07FA60E32FFA26" box="[221,351,1429,1456]" italics="true" pageId="18" pageNumber="135">Q. albiorix</emphasis>
</taxonomicName>
).
<taxonomicName id="239102E0FFA55B44FEA3FA63E0E2FA26" authorityName=", O. Scheerpeltz" authorityYear="1965" box="[377,658,1429,1455]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="135" phylum="Arthropoda" rank="species" species="impressithorax">
<emphasis id="D6E5A571FFA55B44FEA3FA63E0E2FA26" box="[377,658,1429,1455]" italics="true" pageId="18" pageNumber="135">Quedius impressithorax</emphasis>
</taxonomicName>
is here provisionally treated as a valid species until males are found at or near the
<typeStatus id="3B2AC7C1FFA55B44FE71FA4FE3AEFA5A" box="[427,478,1465,1491]" pageId="18" pageNumber="135">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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