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<document id="0844CD973355F3F599A414653A77CED4" ID-CLB-Dataset="307263" ID-DOI="10.11646/zootaxa.5566.1.9" ID-GBIF-Dataset="a07c2970-8904-4e48-9903-a610cf98cc4c" ID-ISSN="1175-5326" ID-Zenodo-Dep="14613093" ID-ZooBank="E17139EB-1205-421D-BBF0-C82E83321B7C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1736290453737" checkinUser="plazi" docAuthor="Kong, Xiangfei, Lin, Aili, Bozdoğan, Hakan, Yang, Ding &amp; Li, Weihai" docDate="2025" docId="524E87DE402AFFF38DE02038FCDAF9AB" docLanguage="en" docName="zootaxa.5566.1.9.pdf" docOrigin="Zootaxa 5566 (1)" docSource="https://doi.org/10.11646/zootaxa.5566.1.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amphinemura pentaspina Kong, Yang &amp; Li 2025, sp. nov." docType="treatment" docVersion="3" lastPageNumber="191" masterDocId="AE77FFA64028FFF58D772566FFE0FFA4" masterDocTitle="Two new species of the Amphinemura luteipes species group (Plecoptera: Nemouridae) from Xizang, China" masterLastPageNumber="192" masterPageNumber="185" pageNumber="187" updateTime="1736291903170" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F98F3DBDA0C4011F848DF408C9CA4EDC">Two new species of the Amphinemura luteipes species group (Plecoptera: Nemouridae) from Xizang, China</mods:title>
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<mods:namePart id="7AB5077E0AF2A69F2E2C1D6CE56AC3BF">Kong, Xiangfei</mods:namePart>
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<mods:affiliation id="DE9E062688FE5F543502ED24272D055E">Henan International Joint Laboratory of Taxonomy and Systematic Evolution of Insecta, Henan Institute of Science and Technology, Xinxiang, Henan 453003, China</mods:affiliation>
<mods:nameIdentifier id="43E7858388F240B5D2EF67E0C6C45094" type="email">962828136@qq.com</mods:nameIdentifier>
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<mods:namePart id="F910724CAA239F26F97252CF304B735D">Lin, Aili</mods:namePart>
<mods:nameIdentifier id="FA044F9E0E5F6C14E8A09A45C55A7924" type="ORCID">0000-0001-9480-092X</mods:nameIdentifier>
<mods:affiliation id="A0B059520C1A24995AE1CDCFFB46B248">Henan International Joint Laboratory of Taxonomy and Systematic Evolution of Insecta, Henan Institute of Science and Technology, Xinxiang, Henan 453003, China</mods:affiliation>
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<mods:namePart id="81C82D893EEA12B5957EA6C2270E03F5">Bozdoğan, Hakan</mods:namePart>
<mods:nameIdentifier id="3A27D4DCD84F3E605A6EE6899C852096" type="ORCID">0000-0002-6836-4383</mods:nameIdentifier>
<mods:affiliation id="C84B0603837145CB9B0029335E6BB8EF">Kırşehir Ahi Evran University, Vocational School of Technical Sciences, Department of Plant and Animal Production, 40100, Kırşehir, Turkey</mods:affiliation>
<mods:nameIdentifier id="7A3E3E0D695FD7ACC3CBB859D59EB634" type="email">hakan.bozdogan@ahievran.edu.tr</mods:nameIdentifier>
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<mods:namePart id="0FFF21A1F47912BE9B202C8E2CC5FF9D">Yang, Ding</mods:namePart>
<mods:nameIdentifier id="E296CE6F4E48EEBC8158CBAF312E1528" type="ORCID">0000-0002-7685-3478</mods:nameIdentifier>
<mods:affiliation id="852F7B015DC113FBD9964B783776C617">Department of Entomology, China Agricultural University, 2 Yuanmingyuan West Road, Beijing 100193, China</mods:affiliation>
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<mods:namePart id="ED67FF96D72905B504D5C3EBD55A52C7">Li, Weihai</mods:namePart>
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<mods:affiliation id="123134B5C494816186A19328B3BF86BE">Henan International Joint Laboratory of Taxonomy and Systematic Evolution of Insecta, Henan Institute of Science and Technology, Xinxiang, Henan 453003, China</mods:affiliation>
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<mods:title id="0BD78DD0F4B38D051978CE7DD133B068">Zootaxa</mods:title>
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<mods:date id="54668B151CE2B6C106E03F66C11720D3">2025</mods:date>
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<treatment id="524E87DE402AFFF38DE02038FCDAF9AB" ID-DOI="http://doi.org/10.5281/zenodo.14613135" ID-Zenodo-Dep="14613135" LSID="urn:lsid:plazi:treatment:524E87DE402AFFF38DE02038FCDAF9AB" httpUri="http://treatment.plazi.org/id/524E87DE402AFFF38DE02038FCDAF9AB" lastPageId="6" lastPageNumber="191" pageId="2" pageNumber="187" scope_family="Nemouridae" scope_order="Plecoptera">
<subSubSection id="92FD6543402AFFF78DE02038FCFBFADD" box="[151,795,1374,1401]" pageId="2" pageNumber="187" type="nomenclature">
<paragraph id="DA5836C8402AFFF78DE02038FCFBFADD" blockId="2.[151,795,1374,1437]" box="[151,795,1374,1401]" pageId="2" pageNumber="187">
<heading id="811081A4402AFFF78DE02038FCFBFADD" bold="true" box="[151,795,1374,1401]" fontSize="11" level="1" pageId="2" pageNumber="187" reason="1">
<emphasis id="E893EADA402AFFF78DE02038FCFBFADD" bold="true" box="[151,795,1374,1401]" pageId="2" pageNumber="187">
<taxonomicName id="1DE74D4B402AFFF78DE02038FD50FADC" authority="Kong, Yang &amp; Li" authorityName="Kong, Yang &amp; Li" authorityYear="2025" box="[151,688,1374,1401]" class="Insecta" family="Nemouridae" genus="Amphinemura" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="pentaspina" status="sp. nov.">
<emphasis id="E893EADA402AFFF78DE02038FE33FADC" bold="true" box="[151,467,1374,1401]" italics="true" pageId="2" pageNumber="187">Amphinemura pentaspina</emphasis>
Kong, Yang &amp; Li
</taxonomicName>
,
<taxonomicNameLabel id="F3A057A1402AFFF78FCA2039FCFBFADD" box="[701,795,1375,1401]" pageId="2" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="92FD6543402AFFF68DE020E4FDD2F8DF" lastPageId="3" lastPageNumber="188" pageId="2" pageNumber="187" type="description">
<paragraph id="DA5836C8402AFFF78DE020E4FE90FA38" blockId="2.[151,795,1374,1437]" box="[151,368,1410,1437]" pageId="2" pageNumber="187">
(
<figureCitation id="42DC2A4D402AFFF78DE820E4FF1CFA38" box="[159,252,1410,1436]" captionStart="FIGURE 1" captionStartId="2.[151,250,1232,1256]" captionTargetBox="[298,1289,192,1193]" captionTargetId="figure-30@2.[286,1302,179,1206]" captionTargetPageId="2" captionText="FIGURE 1. Male habitus. a: Amphinemura beibeng sp. nov., male, dorsal view; b: Amphinemura pentaspina sp. nov., male, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14613095" httpUri="https://zenodo.org/record/14613095/files/figure.png" pageId="2" pageNumber="187">Figs. 1b</figureCitation>
,
<figureCitation id="42DC2A4D402AFFF78C7E20E5FEF8FA39" box="[265,280,1411,1437]" captionStart="FIGURE 3" captionStartId="4.[151,250,1553,1577]" captionTargetBox="[198,1389,179,1503]" captionTargetId="figure-30@4.[198,1389,179,1528]" captionTargetPageId="4" captionText="FIGURE 3. Amphinemura pentaspina sp. nov., male. a: terminalia, dorsal view; b: terminalia, ventral view; c: epiproct, dorsal view; d: epiproct, lateral view; e: right paraproct, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14613101" httpUri="https://zenodo.org/record/14613101/files/figure.png" pageId="2" pageNumber="187">3</figureCitation>
,
<figureCitation id="42DC2A4D402AFFF78C5220E4FE88FA38" box="[293,360,1410,1436]" captionStart="FIGURE 4" captionStartId="5.[151,250,1824,1848]" captionTargetBox="[164,1423,192,1787]" captionTargetId="figure-18@5.[151,1436,179,1798]" captionTargetPageId="5" captionText="FIGURE 4. Male terminalia, cleared, Amphinemura beibeng sp. nov. (a–c), Amphinemura pentaspina sp. nov. (d–f). a, d: terminalia, dorsal view; b, e: terminalia, ventral view; c, f: terminalia, lateral view. ds: dorsal sclerite, il: inner lobe of paraproct, la: lateral arm, ml: median lobe of paraproct, ol: outer lobe of paraproct, vs: ventral sclerite," figureDoi="http://doi.org/10.5281/zenodo.14613105" httpUri="https://zenodo.org/record/14613105/files/figure.png" pageId="2" pageNumber="187">4d–4f</figureCitation>
)
</paragraph>
<paragraph id="DA5836C8402AFFF78DE020ADFB35F9F4" blockId="2.[151,1437,1482,2013]" pageId="2" pageNumber="187">
<emphasis id="E893EADA402AFFF78DE020ADFED6FA40" bold="true" box="[151,310,1482,1509]" pageId="2" pageNumber="187">Male habitus</emphasis>
. Body color generally brownish (
<figureCitation id="42DC2A4D402AFFF78FC020ACFCEBFA40" box="[695,779,1482,1508]" captionStart="FIGURE 1" captionStartId="2.[151,250,1232,1256]" captionTargetBox="[298,1289,192,1193]" captionTargetId="figure-30@2.[286,1302,179,1206]" captionTargetPageId="2" captionText="FIGURE 1. Male habitus. a: Amphinemura beibeng sp. nov., male, dorsal view; b: Amphinemura pentaspina sp. nov., male, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14613095" httpUri="https://zenodo.org/record/14613095/files/figure.png" pageId="2" pageNumber="187">Fig. 1b</figureCitation>
). Head dark brown; antennae and palpi yellowish brown; compound eyes black; head slightly wider than pronotum; pronotum dark brown, corners obtuse and lateral margins bulging, with brown dotted and scattered larger rugosities. Legs yellow, tibia of hindlegs with dark distal band. Wing membranes brownish, veins brown. Abdominal segments including terminalia dark brown.
</paragraph>
<paragraph id="DA5836C8402AFFF68DB0233DFD38FF5D" blockId="2.[151,1437,1482,2013]" lastBlockId="3.[151,1436,151,249]" lastPageId="3" lastPageNumber="188" pageId="2" pageNumber="187">
<emphasis id="E893EADA402AFFF78DB0233DFEE4F9D1" bold="true" box="[199,260,1627,1653]" pageId="2" pageNumber="187">Male</emphasis>
. Forewing length 5.9–6.0 mm, hind wing length
<quantity id="1D1F9B2D402AFFF78E4D233CFC23F9D1" box="[826,963,1626,1653]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="4.7" metricValueMin="4.6" pageId="2" pageNumber="187" unit="mm" value="4.65" valueMax="4.7" valueMin="4.6">4.6–4.7 mm</quantity>
(n=2). Tergum 9 lightly sclerotized, with irregular rows of short spines and bristles at posterior margin (
<figureCitation id="42DC2A4D402AFFF78E1D2318FC21F93C" box="[874,961,1662,1688]" captionStart="FIGURE 4" captionStartId="5.[151,250,1824,1848]" captionTargetBox="[164,1423,192,1787]" captionTargetId="figure-18@5.[151,1436,179,1798]" captionTargetPageId="5" captionText="FIGURE 4. Male terminalia, cleared, Amphinemura beibeng sp. nov. (a–c), Amphinemura pentaspina sp. nov. (d–f). a, d: terminalia, dorsal view; b, e: terminalia, ventral view; c, f: terminalia, lateral view. ds: dorsal sclerite, il: inner lobe of paraproct, la: lateral arm, ml: median lobe of paraproct, ol: outer lobe of paraproct, vs: ventral sclerite," figureDoi="http://doi.org/10.5281/zenodo.14613105" httpUri="https://zenodo.org/record/14613105/files/figure.png" pageId="2" pageNumber="187">Fig. 4d</figureCitation>
). Slender vesicle of sternum 9 claviform, length barely 6.0X maximum width, slightly constricted medially (
<figureCitation id="42DC2A4D402AFFF78EE323C4FC13F918" box="[916,1011,1698,1724]" captionStart="FIGURE 3" captionStartId="4.[151,250,1553,1577]" captionTargetBox="[198,1389,179,1503]" captionTargetId="figure-30@4.[198,1389,179,1528]" captionTargetPageId="4" captionText="FIGURE 3. Amphinemura pentaspina sp. nov., male. a: terminalia, dorsal view; b: terminalia, ventral view; c: epiproct, dorsal view; d: epiproct, lateral view; e: right paraproct, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14613101" httpUri="https://zenodo.org/record/14613101/files/figure.png" pageId="2" pageNumber="187">Figs. 3b</figureCitation>
,
<figureCitation id="42DC2A4D402AFFF7897623C5FBFFF919" box="[1025,1055,1699,1725]" captionStart="FIGURE 4" captionStartId="5.[151,250,1824,1848]" captionTargetBox="[164,1423,192,1787]" captionTargetId="figure-18@5.[151,1436,179,1798]" captionTargetPageId="5" captionText="FIGURE 4. Male terminalia, cleared, Amphinemura beibeng sp. nov. (a–c), Amphinemura pentaspina sp. nov. (d–f). a, d: terminalia, dorsal view; b, e: terminalia, ventral view; c, f: terminalia, lateral view. ds: dorsal sclerite, il: inner lobe of paraproct, la: lateral arm, ml: median lobe of paraproct, ol: outer lobe of paraproct, vs: ventral sclerite," figureDoi="http://doi.org/10.5281/zenodo.14613105" httpUri="https://zenodo.org/record/14613105/files/figure.png" pageId="2" pageNumber="187">4e</figureCitation>
). Hypoproct subquadrate at basal half, medial portion gradually tapering toward nipple-like tip (
<figureCitation id="42DC2A4D402AFFF78E2323A0FC51F944" box="[852,945,1734,1760]" captionStart="FIGURE 3" captionStartId="4.[151,250,1553,1577]" captionTargetBox="[198,1389,179,1503]" captionTargetId="figure-30@4.[198,1389,179,1528]" captionTargetPageId="4" captionText="FIGURE 3. Amphinemura pentaspina sp. nov., male. a: terminalia, dorsal view; b: terminalia, ventral view; c: epiproct, dorsal view; d: epiproct, lateral view; e: right paraproct, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14613101" httpUri="https://zenodo.org/record/14613101/files/figure.png" pageId="2" pageNumber="187">Figs. 3b</figureCitation>
,
<figureCitation id="42DC2A4D402AFFF78EC923A1FC3CF945" box="[958,988,1735,1761]" captionStart="FIGURE 4" captionStartId="5.[151,250,1824,1848]" captionTargetBox="[164,1423,192,1787]" captionTargetId="figure-18@5.[151,1436,179,1798]" captionTargetPageId="5" captionText="FIGURE 4. Male terminalia, cleared, Amphinemura beibeng sp. nov. (a–c), Amphinemura pentaspina sp. nov. (d–f). a, d: terminalia, dorsal view; b, e: terminalia, ventral view; c, f: terminalia, lateral view. ds: dorsal sclerite, il: inner lobe of paraproct, la: lateral arm, ml: median lobe of paraproct, ol: outer lobe of paraproct, vs: ventral sclerite," figureDoi="http://doi.org/10.5281/zenodo.14613105" httpUri="https://zenodo.org/record/14613105/files/figure.png" pageId="2" pageNumber="187">4e</figureCitation>
). Tergum 10 sclerotized, a membranous concavity present below the epiproct, bearing many black spines along either side of the concavity (
<figureCitation id="42DC2A4D402AFFF78871238CFA83F8A1" box="[1286,1379,1770,1797]" captionStart="FIGURE 3" captionStartId="4.[151,250,1553,1577]" captionTargetBox="[198,1389,179,1503]" captionTargetId="figure-30@4.[198,1389,179,1528]" captionTargetPageId="4" captionText="FIGURE 3. Amphinemura pentaspina sp. nov., male. a: terminalia, dorsal view; b: terminalia, ventral view; c: epiproct, dorsal view; d: epiproct, lateral view; e: right paraproct, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14613101" httpUri="https://zenodo.org/record/14613101/files/figure.png" pageId="2" pageNumber="187">Figs. 3a</figureCitation>
,
<figureCitation id="42DC2A4D402AFFF78807238CFA70F8A0" box="[1392,1424,1770,1796]" captionStart="FIGURE 4" captionStartId="5.[151,250,1824,1848]" captionTargetBox="[164,1423,192,1787]" captionTargetId="figure-18@5.[151,1436,179,1798]" captionTargetPageId="5" captionText="FIGURE 4. Male terminalia, cleared, Amphinemura beibeng sp. nov. (a–c), Amphinemura pentaspina sp. nov. (d–f). a, d: terminalia, dorsal view; b, e: terminalia, ventral view; c, f: terminalia, lateral view. ds: dorsal sclerite, il: inner lobe of paraproct, la: lateral arm, ml: median lobe of paraproct, ol: outer lobe of paraproct, vs: ventral sclerite," figureDoi="http://doi.org/10.5281/zenodo.14613105" httpUri="https://zenodo.org/record/14613105/files/figure.png" pageId="2" pageNumber="187">4d</figureCitation>
). Cercus membranous, length approximately 2.0X width (
<figureCitation id="42DC2A4D402AFFF78E6D2268FC99F88D" box="[794,889,1806,1833]" captionStart="FIGURE 3" captionStartId="4.[151,250,1553,1577]" captionTargetBox="[198,1389,179,1503]" captionTargetId="figure-30@4.[198,1389,179,1528]" captionTargetPageId="4" captionText="FIGURE 3. Amphinemura pentaspina sp. nov., male. a: terminalia, dorsal view; b: terminalia, ventral view; c: epiproct, dorsal view; d: epiproct, lateral view; e: right paraproct, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14613101" httpUri="https://zenodo.org/record/14613101/files/figure.png" pageId="2" pageNumber="187">Figs. 3a</figureCitation>
,
<figureCitation id="42DC2A4D402AFFF78EF12268FC46F88C" box="[902,934,1806,1832]" captionStart="FIGURE 4" captionStartId="5.[151,250,1824,1848]" captionTargetBox="[164,1423,192,1787]" captionTargetId="figure-18@5.[151,1436,179,1798]" captionTargetPageId="5" captionText="FIGURE 4. Male terminalia, cleared, Amphinemura beibeng sp. nov. (a–c), Amphinemura pentaspina sp. nov. (d–f). a, d: terminalia, dorsal view; b, e: terminalia, ventral view; c, f: terminalia, lateral view. ds: dorsal sclerite, il: inner lobe of paraproct, la: lateral arm, ml: median lobe of paraproct, ol: outer lobe of paraproct, vs: ventral sclerite," figureDoi="http://doi.org/10.5281/zenodo.14613105" httpUri="https://zenodo.org/record/14613105/files/figure.png" pageId="2" pageNumber="187">4d</figureCitation>
). Epiproct nearly rectangular (
<figureCitation id="42DC2A4D402AFFF788732268FA82F88D" box="[1284,1378,1806,1833]" captionStart="FIGURE 3" captionStartId="4.[151,250,1553,1577]" captionTargetBox="[198,1389,179,1503]" captionTargetId="figure-30@4.[198,1389,179,1528]" captionTargetPageId="4" captionText="FIGURE 3. Amphinemura pentaspina sp. nov., male. a: terminalia, dorsal view; b: terminalia, ventral view; c: epiproct, dorsal view; d: epiproct, lateral view; e: right paraproct, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14613101" httpUri="https://zenodo.org/record/14613101/files/figure.png" pageId="2" pageNumber="187">Figs. 3c</figureCitation>
,
<figureCitation id="42DC2A4D402AFFF788072268FA6FF88C" box="[1392,1423,1806,1832]" captionStart="FIGURE 4" captionStartId="5.[151,250,1824,1848]" captionTargetBox="[164,1423,192,1787]" captionTargetId="figure-18@5.[151,1436,179,1798]" captionTargetPageId="5" captionText="FIGURE 4. Male terminalia, cleared, Amphinemura beibeng sp. nov. (a–c), Amphinemura pentaspina sp. nov. (d–f). a, d: terminalia, dorsal view; b, e: terminalia, ventral view; c, f: terminalia, lateral view. ds: dorsal sclerite, il: inner lobe of paraproct, la: lateral arm, ml: median lobe of paraproct, ol: outer lobe of paraproct, vs: ventral sclerite," figureDoi="http://doi.org/10.5281/zenodo.14613105" httpUri="https://zenodo.org/record/14613105/files/figure.png" pageId="2" pageNumber="187">4d</figureCitation>
), apex triangularly notched (
<figureCitation id="42DC2A4D402AFFF78CBA2254FDC4F8E8" box="[461,548,1842,1868]" captionStart="FIGURE 4" captionStartId="5.[151,250,1824,1848]" captionTargetBox="[164,1423,192,1787]" captionTargetId="figure-18@5.[151,1436,179,1798]" captionTargetPageId="5" captionText="FIGURE 4. Male terminalia, cleared, Amphinemura beibeng sp. nov. (a–c), Amphinemura pentaspina sp. nov. (d–f). a, d: terminalia, dorsal view; b, e: terminalia, ventral view; c, f: terminalia, lateral view. ds: dorsal sclerite, il: inner lobe of paraproct, la: lateral arm, ml: median lobe of paraproct, ol: outer lobe of paraproct, vs: ventral sclerite," figureDoi="http://doi.org/10.5281/zenodo.14613105" httpUri="https://zenodo.org/record/14613105/files/figure.png" pageId="2" pageNumber="187">Fig. 4d</figureCitation>
). Dorsal sclerite mostly membranous, lateral arms darkly sclerotized, basally band-like distinctly tapering medially, apical half stripe-like (
<figureCitation id="42DC2A4D402AFFF78E222230FC53F8D5" box="[853,947,1878,1905]" captionStart="FIGURE 3" captionStartId="4.[151,250,1553,1577]" captionTargetBox="[198,1389,179,1503]" captionTargetId="figure-30@4.[198,1389,179,1528]" captionTargetPageId="4" captionText="FIGURE 3. Amphinemura pentaspina sp. nov., male. a: terminalia, dorsal view; b: terminalia, ventral view; c: epiproct, dorsal view; d: epiproct, lateral view; e: right paraproct, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14613101" httpUri="https://zenodo.org/record/14613101/files/figure.png" pageId="2" pageNumber="187">Figs. 3c</figureCitation>
,
<figureCitation id="42DC2A4D402AFFF78EB62230FC01F8D4" box="[961,993,1878,1904]" captionStart="FIGURE 4" captionStartId="5.[151,250,1824,1848]" captionTargetBox="[164,1423,192,1787]" captionTargetId="figure-18@5.[151,1436,179,1798]" captionTargetPageId="5" captionText="FIGURE 4. Male terminalia, cleared, Amphinemura beibeng sp. nov. (a–c), Amphinemura pentaspina sp. nov. (d–f). a, d: terminalia, dorsal view; b, e: terminalia, ventral view; c, f: terminalia, lateral view. ds: dorsal sclerite, il: inner lobe of paraproct, la: lateral arm, ml: median lobe of paraproct, ol: outer lobe of paraproct, vs: ventral sclerite," figureDoi="http://doi.org/10.5281/zenodo.14613105" httpUri="https://zenodo.org/record/14613105/files/figure.png" pageId="2" pageNumber="187">4d</figureCitation>
). Ventral sclerite distinctly sclerotized, wide at base, tapering toward apex, apical portion extended over dorsal sclerite forming a posteriorly directed tube (
<figureCitation id="42DC2A4D402AFFF78DE822F8FF1EF81C" box="[159,254,1950,1976]" captionStart="FIGURE 3" captionStartId="4.[151,250,1553,1577]" captionTargetBox="[198,1389,179,1503]" captionTargetId="figure-30@4.[198,1389,179,1528]" captionTargetPageId="4" captionText="FIGURE 3. Amphinemura pentaspina sp. nov., male. a: terminalia, dorsal view; b: terminalia, ventral view; c: epiproct, dorsal view; d: epiproct, lateral view; e: right paraproct, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14613101" httpUri="https://zenodo.org/record/14613101/files/figure.png" pageId="2" pageNumber="187">Figs. 3d</figureCitation>
,
<figureCitation id="42DC2A4D402AFFF78C7B22F8FEC7F81C" box="[268,295,1950,1976]" captionStart="FIGURE 4" captionStartId="5.[151,250,1824,1848]" captionTargetBox="[164,1423,192,1787]" captionTargetId="figure-18@5.[151,1436,179,1798]" captionTargetPageId="5" captionText="FIGURE 4. Male terminalia, cleared, Amphinemura beibeng sp. nov. (a–c), Amphinemura pentaspina sp. nov. (d–f). a, d: terminalia, dorsal view; b, e: terminalia, ventral view; c, f: terminalia, lateral view. ds: dorsal sclerite, il: inner lobe of paraproct, la: lateral arm, ml: median lobe of paraproct, ol: outer lobe of paraproct, vs: ventral sclerite," figureDoi="http://doi.org/10.5281/zenodo.14613105" httpUri="https://zenodo.org/record/14613105/files/figure.png" pageId="2" pageNumber="187">4f</figureCitation>
), ventral ridge located at apex, arcuate with rows of tiny black spines (
<figureCitation id="42DC2A4D402AFFF7892722F8FB41F81C" box="[1104,1185,1950,1976]" captionStart="FIGURE 4" captionStartId="5.[151,250,1824,1848]" captionTargetBox="[164,1423,192,1787]" captionTargetId="figure-18@5.[151,1436,179,1798]" captionTargetPageId="5" captionText="FIGURE 4. Male terminalia, cleared, Amphinemura beibeng sp. nov. (a–c), Amphinemura pentaspina sp. nov. (d–f). a, d: terminalia, dorsal view; b, e: terminalia, ventral view; c, f: terminalia, lateral view. ds: dorsal sclerite, il: inner lobe of paraproct, la: lateral arm, ml: median lobe of paraproct, ol: outer lobe of paraproct, vs: ventral sclerite," figureDoi="http://doi.org/10.5281/zenodo.14613105" httpUri="https://zenodo.org/record/14613105/files/figure.png" pageId="2" pageNumber="187">Fig. 4f</figureCitation>
). Paraproct inner lobe slightly sclerotized, nearly triangular, most hidden by hypoproct (
<figureCitation id="42DC2A4D402AFFF78EFF22A4FC05F879" box="[904,997,1986,2013]" captionStart="FIGURE 3" captionStartId="4.[151,250,1553,1577]" captionTargetBox="[198,1389,179,1503]" captionTargetId="figure-30@4.[198,1389,179,1528]" captionTargetPageId="4" captionText="FIGURE 3. Amphinemura pentaspina sp. nov., male. a: terminalia, dorsal view; b: terminalia, ventral view; c: epiproct, dorsal view; d: epiproct, lateral view; e: right paraproct, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14613101" httpUri="https://zenodo.org/record/14613101/files/figure.png" pageId="2" pageNumber="187">Figs. 3e</figureCitation>
,
<figureCitation id="42DC2A4D402AFFF78E8322A5FBF2F879" box="[1012,1042,1987,2013]" captionStart="FIGURE 4" captionStartId="5.[151,250,1824,1848]" captionTargetBox="[164,1423,192,1787]" captionTargetId="figure-18@5.[151,1436,179,1798]" captionTargetPageId="5" captionText="FIGURE 4. Male terminalia, cleared, Amphinemura beibeng sp. nov. (a–c), Amphinemura pentaspina sp. nov. (d–f). a, d: terminalia, dorsal view; b, e: terminalia, ventral view; c, f: terminalia, lateral view. ds: dorsal sclerite, il: inner lobe of paraproct, la: lateral arm, ml: median lobe of paraproct, ol: outer lobe of paraproct, vs: ventral sclerite," figureDoi="http://doi.org/10.5281/zenodo.14613105" httpUri="https://zenodo.org/record/14613105/files/figure.png" pageId="2" pageNumber="187">4e</figureCitation>
); median lobe slightly sclerotized, curved upward medially, with three ventrally directed spines apically (
<figureCitation id="42DC2A4D402BFFF68EDB25F1FBE7FF15" box="[940,1031,151,177]" captionStart="FIGURE 3" captionStartId="4.[151,250,1553,1577]" captionTargetBox="[198,1389,179,1503]" captionTargetId="figure-30@4.[198,1389,179,1528]" captionTargetPageId="4" captionText="FIGURE 3. Amphinemura pentaspina sp. nov., male. a: terminalia, dorsal view; b: terminalia, ventral view; c: epiproct, dorsal view; d: epiproct, lateral view; e: right paraproct, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14613101" httpUri="https://zenodo.org/record/14613101/files/figure.png" pageId="3" pageNumber="188">Figs. 3e</figureCitation>
,
<figureCitation id="42DC2A4D402BFFF6896325F1FBD2FF15" box="[1044,1074,151,177]" captionStart="FIGURE 4" captionStartId="5.[151,250,1824,1848]" captionTargetBox="[164,1423,192,1787]" captionTargetId="figure-18@5.[151,1436,179,1798]" captionTargetPageId="5" captionText="FIGURE 4. Male terminalia, cleared, Amphinemura beibeng sp. nov. (a–c), Amphinemura pentaspina sp. nov. (d–f). a, d: terminalia, dorsal view; b, e: terminalia, ventral view; c, f: terminalia, lateral view. ds: dorsal sclerite, il: inner lobe of paraproct, la: lateral arm, ml: median lobe of paraproct, ol: outer lobe of paraproct, vs: ventral sclerite," figureDoi="http://doi.org/10.5281/zenodo.14613105" httpUri="https://zenodo.org/record/14613105/files/figure.png" pageId="3" pageNumber="188">4e</figureCitation>
); outer lobe shorter than median lobe, basal portion stripe-like and distinctly sclerotized, apical half membranous, curved upward entirely fused with median lobe, with two larger apical spines (
<figureCitation id="42DC2A4D402BFFF68FF725B9FD2BFF5D" box="[640,715,223,249]" captionStart="FIGURE 3" captionStartId="4.[151,250,1553,1577]" captionTargetBox="[198,1389,179,1503]" captionTargetId="figure-30@4.[198,1389,179,1528]" captionTargetPageId="4" captionText="FIGURE 3. Amphinemura pentaspina sp. nov., male. a: terminalia, dorsal view; b: terminalia, ventral view; c: epiproct, dorsal view; d: epiproct, lateral view; e: right paraproct, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14613101" httpUri="https://zenodo.org/record/14613101/files/figure.png" pageId="3" pageNumber="188">Fig.3e</figureCitation>
).
</paragraph>
<caption id="8E986640402BFFF68DE023B5FCC6F8AB" ID-DOI="http://doi.org/10.5281/zenodo.14613099" ID-Zenodo-Dep="14613099" httpUri="https://zenodo.org/record/14613099/files/figure.png" pageId="3" pageNumber="188" startId="3.[151,250,1747,1771]" targetBox="[227,1361,270,1700]" targetPageId="3" targetType="figure">
<paragraph id="DA5836C8402BFFF68DE023B5FCC6F8AB" blockId="3.[151,1436,1747,1807]" pageId="3" pageNumber="188">
<emphasis id="E893EADA402BFFF68DE023B5FEF5F94F" bold="true" box="[151,277,1747,1771]" pageId="3" pageNumber="188">FIGURE 2.</emphasis>
<taxonomicName id="1DE74D4B402BFFF68C6B23B5FDE4F94F" authorityName="Kong &amp; Lin &amp; Bozdoğan &amp; Yang &amp; Li" authorityYear="2025" box="[284,516,1747,1771]" class="Insecta" family="Nemouridae" genus="Amphinemura" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="species" species="beibeng" status="sp. nov.">
<emphasis id="E893EADA402BFFF68C6B23B5FDE4F94F" box="[284,516,1747,1771]" italics="true" pageId="3" pageNumber="188">Amphinemura beibeng</emphasis>
</taxonomicName>
<emphasis id="E893EADA402BFFF68F7B23B5FDBFF94F" bold="true" box="[524,607,1747,1771]" pageId="3" pageNumber="188">
<taxonomicNameLabel id="F3A057A1402BFFF68F7B23B5FDBFF94F" box="[524,607,1747,1771]" pageId="3" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. a: terminalia, dorsal view; b: terminalia, ventral view; c: epiproct, dorsal view; d: epiproct, lateral view; e: right paraproct, posterior view.
</paragraph>
</caption>
<paragraph id="DA5836C8402BFFF68DB02204FDD2F8DF" blockId="3.[151,1436,1889,1988]" box="[199,562,1889,1916]" pageId="3" pageNumber="188">
<emphasis id="E893EADA402BFFF68DB02204FE54F8D8" bold="true" box="[199,436,1890,1916]" pageId="3" pageNumber="188">Female and nymph.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="92FD6543402BFFF68DB022E0FAF3F867" pageId="3" pageNumber="188" type="materials_examined">
<paragraph id="DA5836C8402BFFF68DB022E0FAF3F867" blockId="3.[151,1436,1889,1988]" pageId="3" pageNumber="188">
<emphasis id="E893EADA402BFFF68DB022E0FE8CF83B" bold="true" box="[199,364,1925,1952]" pageId="3" pageNumber="188">Type material</emphasis>
.
<materialsCitation id="6A8F3C95402BFFF68C0F22E3FD7DF867" collectingDate="2014-07-28" collectionCode="CAU" collectorName="Tingting Zhang." country="China" county="Motuo County" elevation="885" latitude="29.3288" location="Autonomous region" longLatPrecision="6" longitude="95.3227" pageId="3" pageNumber="188" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
<typeStatus id="055C886A402BFFF68C0F22E3FE00F83B" box="[376,480,1925,1951]" pageId="3" pageNumber="188" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="BCF6AE0D402BFFF68F5E22E0FD8BF804" box="[553,619,1926,1952]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="3" pageNumber="188">CAU</collectionCode>
):
<collectingCountry id="A2F07658402BFFF68F0C22E0FD37F804" box="[635,727,1926,1952]" name="China" pageId="3" pageNumber="188">CHINA</collectingCountry>
,
<collectingRegion id="1823F82A402BFFF68F9722E3FCD1F83B" box="[736,817,1925,1951]" country="China" name="Xizang" pageId="3" pageNumber="188">Xizang</collectingRegion>
<location id="DF386013402BFFF68E4322E0FBF4F83B" LSID="urn:lsid:plazi:treatment:524E87DE402AFFF38DE02038FCDAF9AB:DF386013402BFFF68E4322E0FBF4F83B" box="[820,1044,1925,1952]" country="China" county="Motuo County" latitude="29.3288" longLatPrecision="6" longitude="95.3227" name="Autonomous region" pageId="3" pageNumber="188" stateProvince="Xizang">Autonomous region</location>
,
<collectingCounty id="33394E44402BFFF6896922E0FB21F804" box="[1054,1217,1926,1952]" pageId="3" pageNumber="188">Motuo County</collectingCounty>
,
<quantity id="1D1F9B2D402BFFF689BD22E3FAEDF804" box="[1226,1293,1925,1952]" metricMagnitude="2" metricUnit="m" metricValue="8.85" pageId="3" pageNumber="188" unit="m" value="885.0">
<elevation id="51CAD1FB402BFFF689BD22E3FAEDF804" box="[1226,1293,1925,1952]" metricMagnitude="2" metricUnit="m" metricValue="8.85" pageId="3" pageNumber="188" unit="m" value="885.0">885m</elevation>
</quantity>
,
<geoCoordinate id="BFD3500F402BFFF6886122E0FA78F804" box="[1302,1432,1926,1952]" degrees="29.3288" direction="north" orientation="latitude" pageId="3" pageNumber="188" precision="5" value="29.3288">N 29.3288°</geoCoordinate>
,
<geoCoordinate id="BFD3500F402BFFF68DE022CFFEF2F860" box="[151,274,1961,1988]" degrees="95.3227" direction="east" orientation="longitude" pageId="3" pageNumber="188" precision="5" value="95.3227">E95.3227°</geoCoordinate>
,
<date id="AE591008402BFFF68C6B22CCFE4DF860" box="[284,429,1962,1988]" pageId="3" pageNumber="188" value="2014-07-28">
<collectingDate id="BE1DE9E0402BFFF68C6B22CCFE4DF860" box="[284,429,1962,1988]" pageId="3" pageNumber="188" value="2014-07-28">28-VII-2014</collectingDate>
</date>
, leg.
<collectorName id="7712531E402BFFF68C9F22CFFD7DF867" box="[488,669,1961,1987]" pageId="3" pageNumber="188">Tingting Zhang.</collectorName>
</materialsCitation>
<materialsCitation id="6A8F3C95402BFFF68FD322CCFAF0F867" box="[676,1296,1961,1988]" collectingDate="2014-07-28" collectionCode="CAU" collectorName="Tingting Zhang." country="China" county="Motuo County" elevation="885" latitude="29.3288" location="Autonomous region" longLatPrecision="6" longitude="95.3227" pageId="3" pageNumber="188" specimenCount="1" specimenCount-male="1" stateProvince="Xizang" typeStatus="paratype">
<typeStatus id="055C886A402BFFF68FD322CCFCEAF860" box="[676,778,1962,1988]" pageId="3" pageNumber="188" type="paratype">Paratype</typeStatus>
:
<specimenCount id="CCE1FD41402BFFF68E6322CCFCBFF867" box="[788,863,1961,1988]" count="1" pageId="3" pageNumber="188" type="male">1 male</specimenCount>
(
<collectionCode id="BCF6AE0D402BFFF68E1822CCFC4FF860" box="[879,943,1962,1988]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="3" pageNumber="188">CAU</collectionCode>
), the same data as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="8E986640402CFFF18DE02377FCC6F9EA" ID-DOI="http://doi.org/10.5281/zenodo.14613101" ID-Zenodo-Dep="14613101" httpUri="https://zenodo.org/record/14613101/files/figure.png" pageId="4" pageNumber="189" startId="4.[151,250,1553,1577]" targetBox="[198,1389,179,1503]" targetPageId="4" targetType="figure">
<paragraph id="DA5836C8402CFFF18DE02377FCC6F9EA" blockId="4.[151,1436,1553,1614]" pageId="4" pageNumber="189">
<emphasis id="E893EADA402CFFF18DE02377FEEDF98E" bold="true" box="[151,269,1553,1578]" pageId="4" pageNumber="189">FIGURE 3</emphasis>
.
<taxonomicName id="1DE74D4B402CFFF18C6E2374FDFEF98D" authorityName="Kong &amp; Lin &amp; Bozdoğan &amp; Yang &amp; Li" authorityYear="2025" box="[281,542,1554,1578]" class="Insecta" family="Nemouridae" genus="Amphinemura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="species" species="pentaspina" status="sp. nov.">
<emphasis id="E893EADA402CFFF18C6E2374FDFEF98D" box="[281,542,1554,1578]" italics="true" pageId="4" pageNumber="189">Amphinemura pentaspina</emphasis>
</taxonomicName>
<emphasis id="E893EADA402CFFF18F532374FD95F98E" bold="true" box="[548,629,1554,1578]" pageId="4" pageNumber="189">
<taxonomicNameLabel id="F3A057A1402CFFF18F532374FD95F98E" box="[548,629,1554,1578]" pageId="4" pageNumber="189" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. a: terminalia, dorsal view; b: terminalia, ventral view; c: epiproct, dorsal view; d: epiproct, lateral view; e: right paraproct, posterior view.
</paragraph>
</caption>
<subSubSection id="92FD6543402CFFF18DB023C6FBD8F91E" box="[199,1080,1696,1722]" pageId="4" pageNumber="189" type="etymology">
<paragraph id="DA5836C8402CFFF18DB023C6FBD8F91E" blockId="4.[151,1437,1696,1866]" box="[199,1080,1696,1722]" pageId="4" pageNumber="189">
<emphasis id="E893EADA402CFFF18DB023C6FEA8F91E" bold="true" box="[199,328,1696,1722]" pageId="4" pageNumber="189">Etymology</emphasis>
. The specific name refers to the number of spines of the paraproct.
</paragraph>
</subSubSection>
<subSubSection id="92FD6543402CFFF18DB023A2FCEAF97A" box="[199,778,1732,1758]" pageId="4" pageNumber="189" type="distribution">
<paragraph id="DA5836C8402CFFF18DB023A2FCEAF97A" blockId="4.[151,1437,1696,1866]" box="[199,778,1732,1758]" pageId="4" pageNumber="189">
<emphasis id="E893EADA402CFFF18DB023A2FEB9F97A" bold="true" box="[199,345,1732,1758]" pageId="4" pageNumber="189">Distribution</emphasis>
.
<collectingCountry id="A2F07658402CFFF18C1023A2FE4AF97A" box="[359,426,1732,1758]" name="China" pageId="4" pageNumber="189">China</collectingCountry>
(
<collectingRegion id="1823F82A402CFFF18CCE23A2FDECF97A" box="[441,524,1732,1758]" country="China" name="Xizang" pageId="4" pageNumber="189">Xizang</collectingRegion>
Autonomous Region).
</paragraph>
</subSubSection>
<subSubSection id="92FD6543402CFFF38DB0238EFCDAF9AB" lastPageId="6" lastPageNumber="191" pageId="4" pageNumber="189" type="diagnosis">
<paragraph id="DA5836C8402CFFF18DB0238EFBCDF8EE" blockId="4.[151,1437,1696,1866]" pageId="4" pageNumber="189">
<emphasis id="E893EADA402CFFF18DB0238EFEDAF8A6" bold="true" box="[199,314,1768,1794]" pageId="4" pageNumber="189">Diagnosis</emphasis>
.
<taxonomicName id="1DE74D4B402CFFF18C32238EFDA2F8A6" authorityName="Kong &amp; Lin &amp; Bozdoğan &amp; Yang &amp; Li" authorityYear="2025" box="[325,578,1768,1794]" class="Insecta" family="Nemouridae" genus="Amphinemura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="species" species="beibeng" status="sp. nov.">
<emphasis id="E893EADA402CFFF18C32238EFDA2F8A6" box="[325,578,1768,1794]" italics="true" pageId="4" pageNumber="189">Amphinemura beibeng</emphasis>
</taxonomicName>
<emphasis id="E893EADA402CFFF18F31238EFD7DF8A6" bold="true" box="[582,669,1768,1794]" pageId="4" pageNumber="189">
<taxonomicNameLabel id="F3A057A1402CFFF18F31238EFD7DF8A6" box="[582,669,1768,1794]" pageId="4" pageNumber="189" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
appears to be closely related to
<taxonomicName id="1DE74D4B402CFFF18E82238EFB6CF8A6" authorityName="Kong &amp; Lin &amp; Bozdoğan &amp; Yang &amp; Li" authorityYear="2025" box="[1013,1164,1768,1794]" class="Insecta" family="Nemouridae" genus="Amphinemura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="species" species="pentaspina" status="sp. nov.">
<emphasis id="E893EADA402CFFF18E82238EFB6CF8A6" box="[1013,1164,1768,1794]" italics="true" pageId="4" pageNumber="189">A. pentaspina</emphasis>
</taxonomicName>
<emphasis id="E893EADA402CFFF189E6238EFB08F8A6" bold="true" box="[1169,1256,1768,1794]" pageId="4" pageNumber="189">
<taxonomicNameLabel id="F3A057A1402CFFF189E6238EFB08F8A6" box="[1169,1256,1768,1794]" pageId="4" pageNumber="189" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
after comparing members of the
<taxonomicName id="1DE74D4B402CFFF18C3C226AFE20F882" authorityName="Kimmins" authorityYear="1947" box="[331,448,1804,1830]" class="Insecta" family="Nemouridae" genus="Amphinemura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="species" species="luteipes">
<emphasis id="E893EADA402CFFF18C3C226AFEBCF882" box="[331,348,1804,1830]" italics="true" pageId="4" pageNumber="189">A</emphasis>
.
<emphasis id="E893EADA402CFFF18C1E226AFE20F882" box="[361,448,1804,1830]" italics="true" pageId="4" pageNumber="189">luteipes</emphasis>
</taxonomicName>
species group (
<bibRefCitation id="BE764B39402CFFF18F1A226AFD1FF882" author="Aubert, J." box="[621,767,1804,1830]" pageId="4" pageNumber="189" pagination="209 - 253" refId="ref2659" refString="Aubert, J. (1967) Les Nemouridae de l'Assam (Plecopteres). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 39 (3 - 4), 209 - 253." type="journal article" year="1967">Aubert 1967</bibRefCitation>
). Both species share similarities with regards to the epiproct apical tube and ventral ridge, paraproct characteristics, and pronotal color pattern.
</paragraph>
<caption id="8E986640402DFFF08DE02246FBD9F824" ID-DOI="http://doi.org/10.5281/zenodo.14613105" ID-Zenodo-Dep="14613105" httpUri="https://zenodo.org/record/14613105/files/figure.png" pageId="5" pageNumber="190" startId="5.[151,250,1824,1848]" targetBox="[164,1423,192,1787]" targetPageId="5" targetType="figure">
<paragraph id="DA5836C8402DFFF08DE02246FBD9F824" blockId="5.[151,1437,1824,1920]" pageId="5" pageNumber="190">
<emphasis id="E893EADA402DFFF08DE02246FEF6F89C" bold="true" box="[151,278,1824,1848]" pageId="5" pageNumber="190">FIGURE 4.</emphasis>
Male terminalia, cleared,
<taxonomicName id="1DE74D4B402DFFF08F5C2246FCF4F89C" authorityName="Kong &amp; Lin &amp; Bozdoğan &amp; Yang &amp; Li" authorityYear="2025" box="[555,788,1824,1848]" class="Insecta" family="Nemouridae" genus="Amphinemura" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="beibeng" status="sp. nov.">
<emphasis id="E893EADA402DFFF08F5C2246FCF4F89C" box="[555,788,1824,1848]" italics="true" pageId="5" pageNumber="190">Amphinemura beibeng</emphasis>
</taxonomicName>
<emphasis id="E893EADA402DFFF08E6A2246FC90F89C" bold="true" box="[797,880,1824,1848]" pageId="5" pageNumber="190">
<taxonomicNameLabel id="F3A057A1402DFFF08E6A2246FC90F89C" box="[797,880,1824,1848]" pageId="5" pageNumber="190" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(a–c),
<taxonomicName id="1DE74D4B402DFFF08EC92246FB25F89C" authorityName="Kong &amp; Lin &amp; Bozdoğan &amp; Yang &amp; Li" authorityYear="2025" box="[958,1221,1824,1848]" class="Insecta" family="Nemouridae" genus="Amphinemura" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="pentaspina" status="sp. nov.">
<emphasis id="E893EADA402DFFF08EC92246FB25F89C" box="[958,1221,1824,1848]" italics="true" pageId="5" pageNumber="190">Amphinemura pentaspina</emphasis>
</taxonomicName>
<emphasis id="E893EADA402DFFF089B82246FAC2F89C" bold="true" box="[1231,1314,1824,1848]" pageId="5" pageNumber="190">
<taxonomicNameLabel id="F3A057A1402DFFF089B82246FAC2F89C" box="[1231,1314,1824,1848]" pageId="5" pageNumber="190" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(d–f). a, d: terminalia, dorsal view; b, e: terminalia, ventral view; c, f: terminalia, lateral view. ds: dorsal sclerite, il: inner lobe of paraproct, la: lateral arm, ml: median lobe of paraproct, ol: outer lobe of paraproct, vs: ventral sclerite,
</paragraph>
</caption>
<caption id="8E986640402EFFF38DE021B8FC14FB52" ID-DOI="http://doi.org/10.5281/zenodo.14613109" ID-Zenodo-Dep="14613109" box="[151,1012,1246,1271]" httpUri="https://zenodo.org/record/14613109/files/figure.png" pageId="6" pageNumber="191" startId="6.[151,250,1246,1270]" targetBox="[162,1422,192,1211]" targetPageId="6" targetType="figure">
<paragraph id="DA5836C8402EFFF38DE021B8FC14FB52" blockId="6.[151,1012,1246,1271]" box="[151,1012,1246,1271]" pageId="6" pageNumber="191">
<emphasis id="E893EADA402EFFF38DE021B8FEF3FB52" bold="true" box="[151,275,1246,1270]" pageId="6" pageNumber="191">FIGURE 5.</emphasis>
Distribution of the
<taxonomicName id="1DE74D4B402EFFF38CAC21B8FD20FB52" authorityName="Kimmins" authorityYear="1947" box="[475,704,1246,1270]" class="Insecta" family="Nemouridae" genus="Amphinemura" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="luteipes">
<emphasis id="E893EADA402EFFF38CAC21B8FD20FB52" box="[475,704,1246,1270]" italics="true" pageId="6" pageNumber="191">Amphinemura luteipes</emphasis>
</taxonomicName>
species group (
<bibRefCitation id="BE764B39402EFFF38E2921B8FC09FB52" author="Aubert, J." box="[862,1001,1246,1270]" pageId="6" pageNumber="191" pagination="209 - 253" refId="ref2659" refString="Aubert, J. (1967) Les Nemouridae de l'Assam (Plecopteres). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 39 (3 - 4), 209 - 253." type="journal article" year="1967">Aubert, 1967</bibRefCitation>
).
</paragraph>
</caption>
<paragraph id="DA5836C8402EFFF38DB02027FCDAF9AB" blockId="6.[151,1437,1344,1551]" pageId="6" pageNumber="191">
Paraproct characteristics distinguishes the two species from each other. The apex of the outer paraproct lobe of
<taxonomicName id="1DE74D4B402EFFF38DE02003FEF1FADA" authorityName="Kong &amp; Lin &amp; Bozdoğan &amp; Yang &amp; Li" authorityYear="2025" box="[151,273,1380,1407]" class="Insecta" family="Nemouridae" genus="Amphinemura" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="beibeng" status="sp. nov.">
<emphasis id="E893EADA402EFFF38DE02003FEF1FADA" box="[151,273,1380,1407]" italics="true" pageId="6" pageNumber="191">A. beibeng</emphasis>
</taxonomicName>
<emphasis id="E893EADA402EFFF38C6E2003FE95FADB" bold="true" box="[281,373,1381,1407]" pageId="6" pageNumber="191">
<taxonomicNameLabel id="F3A057A1402EFFF38C6E2003FE95FADB" box="[281,373,1381,1407]" pageId="6" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
terminates into an outcurved projection and forming prominent fork with the fingerlike apex of the median lobe (
<figureCitation id="42DC2A4D402EFFF38C0E20EEFE36FA07" box="[377,470,1416,1443]" captionStart="FIGURE 2" captionStartId="3.[151,250,1747,1771]" captionTargetBox="[227,1361,270,1700]" captionTargetId="figure-58@3.[227,1361,270,1721]" captionTargetPageId="3" captionText="FIGURE 2. Amphinemura beibeng sp. nov., male. a: terminalia, dorsal view; b: terminalia, ventral view; c: epiproct, dorsal view; d: epiproct, lateral view; e: right paraproct, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14613099" httpUri="https://zenodo.org/record/14613099/files/figure.png" pageId="6" pageNumber="191">Figs. 2a</figureCitation>
,
<figureCitation id="42DC2A4D402EFFF38C9520EFFE1EFA07" box="[482,510,1417,1443]" captionStart="FIGURE 4" captionStartId="5.[151,250,1824,1848]" captionTargetBox="[164,1423,192,1787]" captionTargetId="figure-18@5.[151,1436,179,1798]" captionTargetPageId="5" captionText="FIGURE 4. Male terminalia, cleared, Amphinemura beibeng sp. nov. (a–c), Amphinemura pentaspina sp. nov. (d–f). a, d: terminalia, dorsal view; b, e: terminalia, ventral view; c, f: terminalia, lateral view. ds: dorsal sclerite, il: inner lobe of paraproct, la: lateral arm, ml: median lobe of paraproct, ol: outer lobe of paraproct, vs: ventral sclerite," figureDoi="http://doi.org/10.5281/zenodo.14613105" httpUri="https://zenodo.org/record/14613105/files/figure.png" pageId="6" pageNumber="191">4a</figureCitation>
) whereas for
<taxonomicName id="1DE74D4B402EFFF38FEE20EFFCD3FA06" authorityName="Kong &amp; Lin &amp; Bozdoğan &amp; Yang &amp; Li" authorityYear="2025" box="[665,819,1417,1443]" class="Insecta" family="Nemouridae" genus="Amphinemura" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="pentaspina" status="sp. nov.">
<emphasis id="E893EADA402EFFF38FEE20EFFCD3FA06" box="[665,819,1417,1443]" italics="true" pageId="6" pageNumber="191">A. pentaspina</emphasis>
</taxonomicName>
<emphasis id="E893EADA402EFFF38E4D20EFFC74FA07" bold="true" box="[826,916,1417,1443]" pageId="6" pageNumber="191">
<taxonomicNameLabel id="F3A057A1402EFFF38E4D20EFFC74FA07" box="[826,916,1417,1443]" pageId="6" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the outer paraproct lobe is entirely fused with median lobe (
<figureCitation id="42DC2A4D402EFFF38C4420CAFE70FA63" box="[307,400,1452,1479]" captionStart="FIGURE 3" captionStartId="4.[151,250,1553,1577]" captionTargetBox="[198,1389,179,1503]" captionTargetId="figure-30@4.[198,1389,179,1528]" captionTargetPageId="4" captionText="FIGURE 3. Amphinemura pentaspina sp. nov., male. a: terminalia, dorsal view; b: terminalia, ventral view; c: epiproct, dorsal view; d: epiproct, lateral view; e: right paraproct, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14613101" httpUri="https://zenodo.org/record/14613101/files/figure.png" pageId="6" pageNumber="191">Figs. 3e</figureCitation>
,
<figureCitation id="42DC2A4D402EFFF38CE920CBFE5CFA63" box="[414,444,1453,1479]" captionStart="FIGURE 4" captionStartId="5.[151,250,1824,1848]" captionTargetBox="[164,1423,192,1787]" captionTargetId="figure-18@5.[151,1436,179,1798]" captionTargetPageId="5" captionText="FIGURE 4. Male terminalia, cleared, Amphinemura beibeng sp. nov. (a–c), Amphinemura pentaspina sp. nov. (d–f). a, d: terminalia, dorsal view; b, e: terminalia, ventral view; c, f: terminalia, lateral view. ds: dorsal sclerite, il: inner lobe of paraproct, la: lateral arm, ml: median lobe of paraproct, ol: outer lobe of paraproct, vs: ventral sclerite," figureDoi="http://doi.org/10.5281/zenodo.14613105" httpUri="https://zenodo.org/record/14613105/files/figure.png" pageId="6" pageNumber="191">4e</figureCitation>
). The two new species also resemble two Himalayan species of the group,
<taxonomicName id="1DE74D4B402EFFF3886F20CBFECAFA4E" authority="Aubert, 1967" authorityName="Aubert" authorityYear="1967" class="Insecta" family="Nemouridae" genus="Amphinemura" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="bomdilai">
<emphasis id="E893EADA402EFFF3886F20CBFA7DFA62" box="[1304,1437,1452,1479]" italics="true" pageId="6" pageNumber="191">A. bomdilai</emphasis>
<bibRefCitation id="BE764B39402EFFF38DE020B6FECAFA4E" author="Aubert, J." box="[151,298,1488,1514]" pageId="6" pageNumber="191" pagination="209 - 253" refId="ref2659" refString="Aubert, J. (1967) Les Nemouridae de l'Assam (Plecopteres). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 39 (3 - 4), 209 - 253." type="journal article" year="1967">Aubert, 1967</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="1DE74D4B402EFFF38C2820B7FDE9FA4E" authorityName="Aubert" authorityYear="1967" box="[351,521,1488,1515]" class="Insecta" family="Nemouridae" genus="Amphinemura" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="paraluteipes">
<emphasis id="E893EADA402EFFF38C2820B7FDE9FA4E" box="[351,521,1488,1515]" italics="true" pageId="6" pageNumber="191">A. paraluteipes</emphasis>
</taxonomicName>
, in having similarly curved ventral ridge of the epiproct, but they can be separated from them by details of paraproct as summarized in
<tableCitation id="97650373402EFFF38F972092FCD6F9AB" box="[736,822,1524,1551]" captionStart="TABLE 2" captionStartId="6.[151,239,1598,1622]" captionText="TABLE 2. comparison of paraproct characters the new species with similar species in Amphinemura luteipes group." pageId="6" pageNumber="191">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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