treatments-xml/data/37/FB/26/37FB26D6CA345F2EB479DF84D78FF828.xml

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<document ID-CLB-Dataset="4509" ID-DOI="http://dx.doi.org/10.3897/zookeys.1058.68628" ID-GBIF-Dataset="4477866e-bbd0-4938-86ed-bd708b00559c" ID-PMC="PMC8421324" ID-Pensoft-Pub="1313-2970-1058-1" ID-Pensoft-UUID="61FB9E86BEE45968AAC868B21AF7ED86" ID-PubMed="34557053" ID-ZooBank="654932353DDB4E1B8848EAB69F2C20FD" ModsDocID="1313-2970-1058-1" checkinTime="1630349293316" checkinUser="pensoft" docAuthor="Vagalinski, Boyan &amp; Golovatch, Sergei I." docDate="2021" docId="37FB26D6CA345F2EB479DF84D78FF828" docLanguage="en" docName="ZooKeys 1058: 1-127" docOrigin="ZooKeys 1058" docPubDate="2021-08-30" docSource="http://dx.doi.org/10.3897/zookeys.1058.68628" docTitle="Megaphyllum (Megaphyllum) spathulatum" docType="treatment" docVersion="7" id="61FB9E86BEE45968AAC868B21AF7ED86" lastPageNumber="1" masterDocId="61FB9E86BEE45968AAC868B21AF7ED86" masterDocTitle="The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae)" masterLastPageNumber="127" masterPageNumber="1" pageNumber="1" updateTime="1732926145943" updateUser="ExternalLinkService">
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<mods:title id="CA57FF420131BB6BE7EA28C5DFA5E40E">The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae)</mods:title>
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<mods:namePart id="C6A997CEF987D0EF3A39647B077559AE">Vagalinski, Boyan</mods:namePart>
<mods:affiliation id="815BBAAEBCB22E56BD988C7BC186E532">Institute of Biodiversity and Ecosystem Research at the Bulgarian Academy of Sciences, 2 Yurii Gagarin Street, 1113, Sofia, Bulgaria</mods:affiliation>
<mods:nameIdentifier id="C7FF1DEB2CC4A87477282B3AF8BEA724" type="email">boyan_vagalinski@excite.com</mods:nameIdentifier>
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<mods:namePart id="DC93799F2473FFE625DBB92EAA27165A">Golovatch, Sergei I.</mods:namePart>
<mods:affiliation id="32B786C629F3980498F25A1F136FB700">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia</mods:affiliation>
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<mods:date id="621F1324198C3584E463310C89969F80">2021</mods:date>
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<mods:number id="14840022A64DD2D96BB60A691EF0B1ED">1058</mods:number>
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<mods:identifier id="0BAB4F6C4CBB1DA8E916231EFABB112D" type="DOI">http://dx.doi.org/10.3897/zookeys.1058.68628</mods:identifier>
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<treatment id="37FB26D6CA345F2EB479DF84D78FF828" ID-DOI="http://doi.org/10.5281/zenodo.7019381" ID-GBIF-Taxon="185371679" ID-Zenodo-Dep="7019381" LSID="urn:lsid:plazi:treatment:37FB26D6CA345F2EB479DF84D78FF828" httpUri="http://treatment.plazi.org/id/37FB26D6CA345F2EB479DF84D78FF828" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Diplopoda" scope_family="Julidae" scope_order="Julida" scope_subGenus="Megaphyllum" scope_tribe="Brachyiulini">
<subSubSection id="797FDB2912CF8365E7E28B7F2A33B88F" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="2295D3E4DFCFB4FA0CB7A88520BD0AC7" pageId="0" pageNumber="1">
<taxonomicName id="237B763976594B5C238325FB23120CE1" ID-CoL="8KR84" LSID="37FB26D6-CA34-5F2E-B479-DF84D78FF828" authority="(Lohmander, 1936)" baseAuthorityName="Lohmander" baseAuthorityYear="1936" class="Diplopoda" family="Julidae" genus="Megaphyllum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megaphyllum (Megaphyllum) spathulatum" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="spathulatum" subGenus="Megaphyllum">Megaphyllum (Megaphyllum) spathulatum (Lohmander, 1936)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="7DAA24880058C0B1BA2B9DAD1C806EFA" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="DC860F517E507AD06F60F41BF00DF5C8" pageId="0" pageNumber="1">
<taxonomicName id="2DF12B217D0189BD379D1A4747DF3CE9" authorityName="Lohmander" authorityYear="1936" class="Diplopoda" family="Julidae" genus="Chromatoiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chromatoiulus (Chromatoiulus) spathulatus" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="spathulatus" subGenus="Chromatoiulus">Chromatoiulus (Chromatoiulus) spathulatus</taxonomicName>
Lohmander, 1936: 104-109, figs 80-84.
</paragraph>
<paragraph id="570FA9782115D0B89606E36EF4710021" pageId="0" pageNumber="1">
<taxonomicName id="F1C01C0663195E14FBA12D9C64D30915" authorityName="Lohmander" authorityYear="1936" class="Diplopoda" family="Julidae" genus="Chromatoiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chromatoiulus (Chromatoiulus) spathulatus" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="spathulatus" subGenus="Chromatoiulus">Chromatoiulus (Chromatoiulus) spathulatus</taxonomicName>
:
<bibRefCitation id="4ADB7E9FBA1803C5D6583504369C737D" author="Attems, C" journalOrPublisher="Annalen des Naturhistorischen Museums in Wien" pageId="0" pageNumber="1" pagination="294 - 327" refId="B9" refString="Attems, C, 1940. Beitraege zur Kenntnis der Iuliden. Annalen des Naturhistorischen Museums in Wien 50: 294 - 327" title="Beitraege zur Kenntnis der Iuliden." volume="50" year="1940">Attems 1940</bibRefCitation>
: 306.
</paragraph>
<paragraph id="4E1E52370B64646C7B1327A1F17B8D5E" pageId="0" pageNumber="1">
<taxonomicName id="3D75EEA216938EC661E65722678056EC" class="Diplopoda" family="Julidae" genus="Chromatoiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chromatoiulus spathulus" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="spathulus">Chromatoiulus spathulus</taxonomicName>
(sic!):
<bibRefCitation id="9E6DB4F11B4B70F362A53E7DD3201B4E" author="Lang, J" journalOrPublisher="Zoologicheskii zhurnal" pageId="0" pageNumber="1" pagination="1790 - 1796" refId="B54" refString="Lang, J, 1959. [To the knowledge of millipedes (Diplopoda) of the USSR territory]. Zoologicheskii zhurnal 38 (12): 1790 - 1796" title="[To the knowledge of millipedes (Diplopoda) of the USSR territory]." volume="38" year="1959">Lang 1959</bibRefCitation>
: 1791.
</paragraph>
<paragraph id="6559F9CFC5AB9CFDB63554A179F0663B" pageId="0" pageNumber="1">
<taxonomicName id="63B69BD6B75A77354F760397C605BFB4" baseAuthorityName="Lohmander" baseAuthorityYear="1936" class="Diplopoda" family="Julidae" genus="Megaphyllum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megaphyllum spathulatum" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="spathulatum">Megaphyllum spathulatum</taxonomicName>
:
<bibRefCitation id="FF33DF1E28941DB0A881203BAB4AB9D0" author="Golovatch, SI" editor="Minelli, A" journalOrPublisher="E. J. Brill, Leiden" pageId="0" pageNumber="1" pagination="361 - 366" refId="B37" refString="Golovatch, SI, 1990b. On the distribution and faunogenesis of Crimean millipedes (Diplopoda). In: Minelli, A, Ed., Proceedings of the 7th International Congress of Myriapodology. E.J. Brill, Leiden: 361 - 366" title="On the distribution and faunogenesis of Crimean millipedes (Diplopoda)." volumeTitle="Proceedings of the 7 th International Congress of Myriapodology." year="1990 b">Golovatch 1990b</bibRefCitation>
: 364; 1992: 381;
<bibRefCitation id="B8AE96B88A027C4F9976E8DF12CB992B" DOI="https://doi.org/10.11646/zootaxa.3741.1.2" author="Lazanyi, E" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="055 - 100" refId="B57" refString="Lazanyi, E, Vagalinski, B, 2013. Redefinition of the millipede subgenus Megaphyllum sensu stricto Verhoeff, 1894 and neotype designation for Megaphyllum austriacum (Latzel, 1884) (Myriapoda: Diplopoda: Julida: Julidae). Zootaxa 3741 (1): 055 - 100, DOI: https://doi.org/10.11646/zootaxa.3741.1.2" title="Redefinition of the millipede subgenus Megaphyllum sensu stricto Verhoeff, 1894 and neotype designation for Megaphyllum austriacum (Latzel, 1884) (Myriapoda: Diplopoda: Julida: Julidae)." url="https://doi.org/10.11646/zootaxa.3741.1.2" volume="3741" year="2013">
<normalizedToken id="828BE6FBB17A00446F95D5FB7CC59D32" originalValue="Lazányi">Lazanyi</normalizedToken>
and Vagalinski 2013
</bibRefCitation>
: 87, fig. 18b-d;
<bibRefCitation id="28A0EE33B5E796697A4AA729752B49A6" DOI="https://doi.org/10.3897/zookeys.741.20042" author="Kokhia, MS" editor="Stoev, P" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="35 - 48" refId="B52" refString="Kokhia, MS, Golovatch, SI, 2018. A checklist of the millipedes of Georgia, Caucasus (Diplopoda). In: Stoev, P, Edgecombe, GD, Eds., Proceedings of the 17th International Congress of Myriapodology, Krabi, Thailand. ZooKeys 741: 35 - 48, DOI: https://doi.org/10.3897/zookeys.741.20042" title="A checklist of the millipedes of Georgia, Caucasus (Diplopoda)." url="https://doi.org/10.3897/zookeys.741.20042" volume="741" volumeTitle="Proceedings of the 17 th International Congress of Myriapodology, Krabi, Thailand." year="2018">Kokhia and Golovatch 2018</bibRefCitation>
: 41;
<bibRefCitation id="B148341FA59C495716F1ACD5F442B538" DOI="https://doi.org/10.3897/zookeys.930.47490" author="Kokhia, MS" editor="Korsos, Z" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="199 - 219" refId="B53" refString="Kokhia, MS, Golovatch, SI, 2020. Diversity and distribution of the millipedes (Diplopoda) of Georgia, Caucasus. In: Korsos, Z, Danyi, L, Eds., Proceedings of the 18th International Congress of Myriapodology, Budapest, Hungary. ZooKeys 930: 199 - 219, DOI: https://doi.org/10.3897/zookeys.930.47490" title="Diversity and distribution of the millipedes (Diplopoda) of Georgia, Caucasus." url="https://doi.org/10.3897/zookeys.930.47490" volume="930" volumeTitle="Proceedings of the 18 th International Congress of Myriapodology, Budapest, Hungary." year="2020">2020</bibRefCitation>
: 206.
</paragraph>
</subSubSection>
<subSubSection id="75AD91B0506637F99C086DC001D7CEA1" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="3FB42D6E39FA83ADC24C06D50E00A401" pageId="0" pageNumber="1">Material examined</paragraph>
<paragraph id="8BCF878F388B75985A5F3562BCF98A7E" pageId="0" pageNumber="1">
<emphasis id="824304C48F3C09963F00E74D7C54C113" bold="true" pageId="0" pageNumber="1">
(SMNG):
<materialsCitation id="B3C4B9A6AC5E65E8B30979EE7EAAB84F" collectingDate="2012-08-24" collectorName="F. Walther" country="Russia" county="Republic" elevation="430" latitude="44.2844" location="Dakhovskaya Canyon" longLatPrecision="6" longitude="40.1778" municipality="Kamennomostkiy" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Adygea">
<collectingCountry id="4BBAD6A5E8A163215BE8EF68FDBEAC89" name="Russia">Russia</collectingCountry>
: RU12-86,
<specimenCount id="1B163CEB9803A5B6FEAB2B471BFD6197" type="male">1 ♂</specimenCount>
,
<specimenCount id="D857D021D676D1C25E78BE72031D5476" type="female">1 ♀</specimenCount>
,
<collectingCounty id="63F718A6EE60660F333A75418485A1C5">Republic</collectingCounty>
of
<collectingRegion id="644C2315CE876E44F06FC8AD2D4DA599" country="Russia" name="Adygeya">Adygea</collectingRegion>
,
<collectingMunicipality id="597FDD588C84FD4AFC06D39BA94426D9">Kamennomostkiy</collectingMunicipality>
, northern end of
<location id="FEAF5A480CD513A656364CB50366F5D6" LSID="urn:lsid:plazi:treatment:37FB26D6CA345F2EB479DF84D78FF828:FEAF5A480CD513A656364CB50366F5D6" country="Russia" county="Republic" latitude="44.2844" longLatPrecision="6" longitude="40.1778" municipality="Kamennomostkiy" name="Dakhovskaya Canyon" stateProvince="Adygea">Dakhovskaya Canyon</location>
near quarry, limestone cliffs,
<geoCoordinate id="7EF179896EA3B1A55C8D03DCAAADE8CB" degrees="44.2844" direction="north" orientation="latitude" precision="5" value="44.2844">44.2844°N</geoCoordinate>
,
<geoCoordinate id="2CB5C2FF088DEE3FB9D72554D5EEB048" degrees="40.1778" direction="east" orientation="longitude" precision="5" value="40.1778">40.1778°E</geoCoordinate>
,
<elevation id="8E06875A4330060C8E58D05C6D114F9D" metricMagnitude="2" metricUnit="m" metricValue="4.3" unit="m" value="430.0">
<quantity id="07AECE7694C008B630876ADDA4FDA421" metricMagnitude="2" metricUnit="m" metricValue="4.3" unit="m" value="430.0">430 m</quantity>
a.s.l.
</elevation>
,
<collectingDate id="A19F0E5CD34FC69413D2362AB3696BF7" value="2012-08-24">24.VIII.2012</collectingDate>
,
<collectorName id="02B27C870B64CCE352F74E2114709C82">F. Walther</collectorName>
leg.
</materialsCitation>
</emphasis>
;
<materialsCitation id="51DBF8BE8E27C6C776FC33DF7D8E5D3D" collectingDate="2012-08-13" collectorName="F. Walther" country="Russia" elevation="800" latitude="44.2439" location="Dakhovskaya" longLatPrecision="6" longitude="40.1758" specimenCount="4" specimenCount-juvenile="1" specimenCount-male="3">
RU12-051,
<specimenCount id="D26ADAB829B1C8A88493A0190CF7E973" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="C2D404FE3E32032D2B8F82FDB7237665" type="generic">1 juv.</specimenCount>
,
<location id="FA3F1BCBFAEDAFB6AD39E507812DECCF" LSID="urn:lsid:plazi:treatment:37FB26D6CA345F2EB479DF84D78FF828:FA3F1BCBFAEDAFB6AD39E507812DECCF" country="Russia" latitude="44.2439" longLatPrecision="6" longitude="40.1758" name="Dakhovskaya">Dakhovskaya</location>
, limestone cliffs NW of,
<geoCoordinate id="5E380C3C557ADF2B8864EF768D782940" degrees="44.2439" direction="north" orientation="latitude" precision="5" value="44.2439">44.2439°N</geoCoordinate>
,
<geoCoordinate id="C9B645E99D5F558091CEAABCC09A0CEC" degrees="40.1758" direction="east" orientation="longitude" precision="5" value="40.1758">40.1758°E</geoCoordinate>
,
<elevation id="25C2EC920BDE3967DA415746292C1C9C" metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">
<quantity id="9891ADC381C53BD2CCBAE84972789DA2" metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">800 m</quantity>
a.s.l.
</elevation>
,
<collectingDate id="B4F0180153BC6A027F0C3110284DA831" value="2012-08-13">13.VIII.2012</collectingDate>
,
<collectorName id="3663F7E29D4700A5EA149DF997D83632">F. Walther</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="909BD6E7478391FFD8E4A64EB1EBBE0D" collectingDate="2004-05-21" collectorName="K. Voigtlaender" country="Russia" location="Mount Shibaba" municipality="Republic" specimenCount="1" specimenCount-male="1" stateProvince="Adygea">
33579,
<specimenCount id="96F6A0D93A9CBCC058EA847F917979AC" type="male">1 ♂</specimenCount>
,
<collectingMunicipality id="A6D02AA4F65424364EC23894D260BDB9">Republic</collectingMunicipality>
of
<collectingRegion id="B6A2E3D65C2643C62744270FB1C9A6D1" country="Russia" name="Adygeya">Adygea</collectingRegion>
,
<location id="056A029EB6C7A6285087DF27E3119399" LSID="urn:lsid:plazi:treatment:37FB26D6CA345F2EB479DF84D78FF828:056A029EB6C7A6285087DF27E3119399" country="Russia" municipality="Republic" name="Mount Shibaba" stateProvince="Adygea">Mount Shibaba</location>
,
<collectingDate id="E5BC669C99957B890453A95AF90C9EEE" value="2004-05-21">21.V.2004</collectingDate>
,
<collectorName id="BAB651037E93FA149593706363FF9BAE">
K.
<normalizedToken id="BFDA311FEEE6F0095EA32BCD7A241A7A" originalValue="Voigtländer">Voigtlaender</normalizedToken>
</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D834D0B65C170C84DFACFF6571CD10AD" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="3AFD7E13C2E919B74AB383E7FA39DC76" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="6FBCA371976920B45C72293993929707" pageId="0" pageNumber="1">
Differs from its only congener known from the Caucasus,
<taxonomicName id="E4454A1F41E6A83374B73EE22F53CA93" genus="M." lsidName="M." pageId="0" pageNumber="1" rank="genus">
<emphasis id="59269CDDA92FE86ABEB76D3AE650E033" italics="true" pageId="0" pageNumber="1">M.</emphasis>
</taxonomicName>
(
<emphasis id="07C38844720519118D6D942A0596CA5E" italics="true" pageId="0" pageNumber="1">s. str.</emphasis>
)
<taxonomicName id="01735D019A4A12744D8189406A1BDE6C" baseAuthorityName="Verhoeff" baseAuthorityYear="1901" class="Diplopoda" family="Julidae" genus="Megaphyllum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megaphyllum hercules" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hercules">
<emphasis id="751CEE9CCE69E59D4CD6F896A2716129" italics="true" pageId="0" pageNumber="1">Megaphyllum hercules</emphasis>
</taxonomicName>
, mainly by the colour pattern: body with a light yellow to ochre dorsum divided by a black axial line, vs. body uniformly dark with an orange to dark red mid-dorsal line in
<taxonomicName id="4B25CB2E205FF64264A323129689AB59" lsidName="M. hercules" pageId="0" pageNumber="1" rank="species" species="hercules">
<emphasis id="3EB4E674A30C1A151BA029F28F702946" italics="true" pageId="0" pageNumber="1">M. hercules</emphasis>
</taxonomicName>
; and by the promere significantly tapering all the way to a narrowly rounded apex, vs. the same having mostly parallel side margins, ending with a flat apex in
<taxonomicName id="9EBD13ED66DBC130B1769899B0CB176C" lsidName="M. hercules" pageId="0" pageNumber="1" rank="species" species="hercules">
<emphasis id="0482EDE2C7F758B81C3C6EFFDC2B4049" italics="true" pageId="0" pageNumber="1">M. hercules</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="7D019948E1684B1CE78FA5BF71E542EE" pageId="0" pageNumber="1" type="previous records from the caucasus">
<paragraph id="BC984DD475E181D7172E1B38A5C4AAB0" pageId="0" pageNumber="1">Previous records from the Caucasus.</paragraph>
<paragraph id="98EE18F067387A5C8DBABDFF06008A95" pageId="0" pageNumber="1">
Western Caucasus (unspecified type locality) (
<bibRefCitation id="21BD1575C70CCDC5480415C80D86EE09" author="Lohmander, H" journalOrPublisher="Zoologischer Anzeiger" pageId="0" pageNumber="1" refId="B63" refString="Lohmander, H, 1936. Ueber die Diplopoden des Kaukasusgebietes. Goeteborgs Kungliga Vetenskaps- och Vitterhets-Samhaelles Handlingar, Ser. 5B, 5(1): 1-196." title="Ueber die Diplopoden des Kaukasusgebietes. Goeteborgs Kungliga Vetenskaps- och Vitterhets-Samhaelles Handlingar, Ser. 5 B, 5 (1): 1 - 196." year="1936">Lohmander 1936</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9A4E1D7D0591B1B2CC1E11BA79F83BBF" pageId="0" pageNumber="1" type="distribution">
<paragraph id="1214F59FB9D4433A8D0223EF1D07F736" pageId="0" pageNumber="1">General distribution.</paragraph>
<paragraph id="B490D9F8AAD532A861ED06AE493551F8" pageId="0" pageNumber="1">WCIS?</paragraph>
</subSubSection>
<subSubSection id="55CF7621558A8FAB68BB6C1685115CDC" pageId="0" pageNumber="1" type="remarks">
<paragraph id="66F19A6EDC2600EF74EF763AA484A8D7" pageId="0" pageNumber="1">Remark.</paragraph>
<paragraph id="AC8D320FC3286534FF4ADAD823C2CE1A" pageId="0" pageNumber="1">This species seems to be a narrow local endemic of the northwestern foothills of the Caucasus Major, showing preferences for limestone terrain.</paragraph>
</subSubSection>
</treatment>
</document>