treatments-xml/data/2E/EA/71/2EEA7109CB2454DD9C0A9C30C75088C3.xml

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<mods:title id="9C30834815E48D529D9F8C00C93B4FBE">Viola shiweii, a new species of Viola (Violaceae) from karst forest in Guizhou, China</mods:title>
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<mods:namePart id="9D869B9C6F2D332DBD596555D653B788">Li, Xiao-Chen</mods:namePart>
<mods:nameIdentifier id="62EE8806CC488C864577515C19271485" type="ORCID">https://orcid.org/0000-0003-3885-328X</mods:nameIdentifier>
<mods:affiliation id="6D57CEF90DCD7C2FE112F607034115E7">Eastern China Conservation Center for Wild Endangered Plant Resources, Shanghai 201602, China &amp; Shanghai Chenshan Botanical Garden, Shanghai 201602, China</mods:affiliation>
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<mods:namePart id="5D8E558DC7BADC95C3C01597499121BA">Wang, Zheng-Wei</mods:namePart>
<mods:affiliation id="8D1333D65BAAE5BE7E1D3D955E1A394E">Eastern China Conservation Center for Wild Endangered Plant Resources, Shanghai 201602, China &amp; Shanghai Chenshan Botanical Garden, Shanghai 201602, China</mods:affiliation>
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<mods:namePart id="874A5AD6CC5E7D09F0032CC739A640E8">Wang, Qi</mods:namePart>
<mods:affiliation id="1C4CFFEF287F23B75EAD38F9D9158AEE">Eastern China Conservation Center for Wild Endangered Plant Resources, Shanghai 201602, China &amp; Shanghai Chenshan Botanical Garden, Shanghai 201602, China</mods:affiliation>
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<mods:namePart id="0E03DCC61C09E7E5E1CA2385A5C92408">Ge, Bin-Jie</mods:namePart>
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<mods:affiliation id="949AB5038459D645E80327E469E1BE1E">Eastern China Conservation Center for Wild Endangered Plant Resources, Shanghai 201602, China &amp; Shanghai Chenshan Botanical Garden, Shanghai 201602, China</mods:affiliation>
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<mods:namePart id="0C2852F61CA3DD9708DA2F034D9C6D69">Chen, Bin</mods:namePart>
<mods:nameIdentifier id="C870D02303A735882D18000AEFE7E595" type="ORCID">https://orcid.org/0000-0003-1822-4486</mods:nameIdentifier>
<mods:affiliation id="3FC912225BACDF20BFC17F3AF8074E04">Eastern China Conservation Center for Wild Endangered Plant Resources, Shanghai 201602, China &amp; Shanghai Chenshan Botanical Garden, Shanghai 201602, China</mods:affiliation>
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<mods:namePart id="EDCC125478CA328DED5088BE0D94E450">Yu, Ping</mods:namePart>
<mods:affiliation id="7462B893D01C48B617DE08A44A9B2617">Guizhou Maolan Karst Forest, Libo 558400, China</mods:affiliation>
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<mods:namePart id="B7166C971C7A67AC7780C7071902424C">Zhong, Xin</mods:namePart>
<mods:affiliation id="EA831A6C9EC5BC61E10A18D79AEC8839">Eastern China Conservation Center for Wild Endangered Plant Resources, Shanghai 201602, China &amp; Shanghai Chenshan Botanical Garden, Shanghai 201602, China</mods:affiliation>
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<mods:date id="E0AEC70330A7B2D9AA82FBFE76601BFB">2022</mods:date>
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<treatment id="2EEA7109CB2454DD9C0A9C30C75088C3" ID-GBIF-Taxon="195348451" LSID="urn:lsid:plazi:treatment:2EEA7109CB2454DD9C0A9C30C75088C3" httpUri="http://treatment.plazi.org/id/2EEA7109CB2454DD9C0A9C30C75088C3" lastPageNumber="63" pageId="0" pageNumber="63">
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<paragraph id="ABF67F2247A9C6C8A905E0CAF9F397C9" pageId="0" pageNumber="63">
<taxonomicName id="03BD4C6B844F80989140232E8082A5E9" ID-CoL="992J2" LSID="2EEA7109-CB24-54DD-9C0A-9C30C75088C3" authority="Xiao C. Li &amp; Z. W. Wang" authorityName="Xiao C. Li &amp; Z. W. Wang" authorityYear="2022" class="Magnoliopsida" family="Violaceae" genus="Viola" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Viola shiweii" order="Malpighiales" pageId="0" pageNumber="63" phylum="Tracheophyta" rank="species" species="shiweii" status="sp. nov.">Viola shiweii Xiao C. Li &amp; Z. W. Wang</taxonomicName>
<taxonomicNameLabel id="D121EA57CF3E803AFC23E00189A42A3C" pageId="0" pageNumber="63">sp. nov.</taxonomicNameLabel>
</paragraph>
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<paragraph id="E837F34F01B11846FD98C973A8A81CC9" pageId="0" pageNumber="63">
<figureCitation id="0F0107C3C415C7E4A40941B07342F677" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Viola shiweii sp. nov. A flower side view B flower front view C habit D stipule E capsule F stamens and pistil." figureDoi="10.3897/phytokeys.196.83176.figure2" httpUri="https://binary.pensoft.net/fig/683701" pageId="0" pageNumber="63">Figs 2</figureCitation>
<figureCitation id="852E7100E3A0849A7F66B0B0740F7311" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Viola shiweii sp. nov. A petals, sepals, stamens and pistils B, G flower in front view and detail of a longitudinal section in side C, H basal leaf adaxially and abaxially D cleistogamous flowers E capsule and seeds F margin teeth I stipule J leaves on stolons." figureDoi="10.3897/phytokeys.196.83176.figure3" httpUri="https://binary.pensoft.net/fig/683702" pageId="0" pageNumber="63">, 3</figureCitation>
<figureCitation id="E00187C16F92947853B9FBA7A58957B4" captionStart="Figure A1" captionStartId="F7" captionText="Figure A 1. Holotype: LXC 00927 (CSH 0182173)." figureDoi="10.3897/phytokeys.196.83176.figurea1" httpUri="https://binary.pensoft.net/fig/683706" pageId="0" pageNumber="63">: Figs A1</figureCitation>
<figureCitation id="B733C0818A8D3161232696261C696877" captionStart="Figure A2" captionStartId="F8" captionText="Figure A 2. Isotype: LXC 00927 (CSH 0182170)." figureDoi="10.3897/phytokeys.196.83176.figurea2" httpUri="https://binary.pensoft.net/fig/683707" pageId="0" pageNumber="63">, A2</figureCitation>
<figureCitation id="C3162D03F1BF59F41A43FC66DEF16EBA" captionStart="Figure A3" captionStartId="F9" captionText="Figure A 3. Isotype: LXC 00927 (CSH 0182171)." figureDoi="10.3897/phytokeys.196.83176.figurea3" httpUri="https://binary.pensoft.net/fig/683708" pageId="0" pageNumber="63">, A3</figureCitation>
<figureCitation id="CA250FB40B6071BEB850274D65142BB3" captionStart="Figure A4" captionStartId="F10" captionText="Figure A 4. Isotype: LXC 00927 (CSH 0182172)." figureDoi="10.3897/phytokeys.196.83176.figurea4" httpUri="https://binary.pensoft.net/fig/683709" pageId="0" pageNumber="63">, A4</figureCitation>
<figureCitation id="496D6EB01AA0E0B3F40E32573D2BC427" captionStart="Figure A5" captionStartId="F11" captionText="Figure A 5. Paratype: LXC 02320 (CSH 0189181)." figureDoi="10.3897/phytokeys.196.83176.figurea5" httpUri="https://binary.pensoft.net/fig/683710" pageId="0" pageNumber="63">, A5</figureCitation>
<figureCitation id="8B3E46845388B3D09D5173AE7D539469" captionStart="Figure A6" captionStartId="F12" captionText="Figure A 6. Paratype: LXC 02322 (CSH 0189180)." figureDoi="10.3897/phytokeys.196.83176.figurea6" httpUri="https://binary.pensoft.net/fig/683711" pageId="0" pageNumber="63">, A6</figureCitation>
<figureCitation id="22CA1F149E86CD2972834CE4D2C46BEE" captionStart="Figure A7" captionStartId="F13" captionText="Figure A 7. Paratype: LXC 02323 (CSH 0189177)." figureDoi="10.3897/phytokeys.196.83176.figurea7" httpUri="https://binary.pensoft.net/fig/683712" pageId="0" pageNumber="63">, A7</figureCitation>
<figureCitation id="2E96EF45589D05CA40B3891F11C4080A" captionStart="Figure A8" captionStartId="F14" captionText="Figure A 8. Paratype: LXC 02323 (CSH 0189178)." figureDoi="10.3897/phytokeys.196.83176.figurea8" httpUri="https://binary.pensoft.net/fig/683713" pageId="0" pageNumber="63">, A8</figureCitation>
<figureCitation id="56F16E454273C01BEBF033B52212459F" captionStart="Figure A9" captionStartId="F15" captionText="Figure A 9. Paratype: LXC 02323 (CSH 0189179)." figureDoi="10.3897/phytokeys.196.83176.figurea9" httpUri="https://binary.pensoft.net/fig/683714" pageId="0" pageNumber="63">, A9</figureCitation>
<figureCitation id="38D72BA51DB6CDCB6CFB706F04D6E9DE" captionStart="Figure A10" captionStartId="F16" captionText="Figure A 10. Paratype: LXC 02324 (CSH 0189176)." figureDoi="10.3897/phytokeys.196.83176.figurea10" httpUri="https://binary.pensoft.net/fig/683715" pageId="0" pageNumber="63">, A10</figureCitation>
<figureCitation id="D0EC09972D95385AA89F050F7A811265" captionStart="Figure A11" captionStartId="F17" captionText="Figure A 11. Paratype: LXC 02325 (CSH 0189174)." figureDoi="10.3897/phytokeys.196.83176.figurea11" httpUri="https://binary.pensoft.net/fig/683716" pageId="0" pageNumber="63">, A11</figureCitation>
<figureCitation id="856DFEF9EC476FC58436F1B680235548" captionStart="Figure A12" captionStartId="F18" captionText="Figure A 12. Paratype: LXC 02325 (CSH 0189175)." figureDoi="10.3897/phytokeys.196.83176.figurea12" httpUri="https://binary.pensoft.net/fig/683717" pageId="0" pageNumber="63">, A12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A158590ACCD49F07C221AC57752B59DB" pageId="0" pageNumber="63" type="materials_examined">
<paragraph id="6E1F0EF6CDA06A570043514E3B8D3B55" pageId="0" pageNumber="63">
<typeStatus id="7C981335029DB7D07138E037B56741AE">Type</typeStatus>
.
</paragraph>
<paragraph id="9C9C1B89F5732759A9CA61BF0DC65692" pageId="0" pageNumber="63">
<materialsCitation id="69A2E9BED0FEDD4ADD668A2D32C10BE6" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="China" location="Shanghai" specimenCount="1" stateProvince="Shanghai" typeStatus="holotype">
<emphasis id="592A3BC591F2B3BCCCC6766B173B3C8C" bold="true" pageId="0" pageNumber="63">
<collectingCountry id="6BDFA05ECB6F383F3FB525CD2179E94F" name="China">China</collectingCountry>
.
</emphasis>
<collectingRegion id="30200214B191D1D24170286657333EC7" country="China" name="Shanghai">Shanghai</collectingRegion>
Chenshan Botanical Garden, cultivated plants collected from
</materialsCitation>
<materialsCitation id="F6FCCCE309F44D219DFBA60A63F0A16E" accessionNumber="LXC00927" collectingDate="2019-11-09" country="China" county="Qiannan Buyi" elevation="867" latitude="25.277529" location="Libo county" longLatPrecision="1" longitude="107.91743" municipality="Miao Autonomous Prefecture" specimenCount="1" stateProvince="Guizhou" typeStatus="Holotype">
<collectingRegion id="3FE0EAA18C91DD9EB8539906D19C3F13" country="China" name="Guizhou">Guizhou</collectingRegion>
, 
<collectingCounty id="28EBAB57112DB68D99EADAB41E784365">Qiannan Buyi</collectingCounty>
and 
<collectingMunicipality id="5AC42E3B2972E4480FA2023BB7F08514">Miao Autonomous Prefecture</collectingMunicipality>
(黔南布依族苗族自治区), 
<location id="944DE565FD9587C2CA3B2121FCDAFFA6" LSID="urn:lsid:plazi:treatment:2EEA7109CB2454DD9C0A9C30C75088C3:944DE565FD9587C2CA3B2121FCDAFFA6" country="China" county="Qiannan Buyi" latitude="25.277529" longLatPrecision="1" longitude="107.91743" municipality="Miao Autonomous Prefecture" name="Libo county" stateProvince="Guizhou">Libo county</location>
(荔波县), 
<location id="9236FBCDB72914B6E25AD7C86566C428" LSID="urn:lsid:plazi:treatment:2EEA7109CB2454DD9C0A9C30C75088C3:9236FBCDB72914B6E25AD7C86566C428" country="China" county="Qiannan Buyi" latitude="25.277529" longLatPrecision="1" longitude="107.91743" municipality="Miao Autonomous Prefecture" name="Maolan National Nature Reserve" stateProvince="Guizhou">Maolan National Nature Reserve</location>
(茂兰国家级自然保护区), on the rocks along the karst forest margin, 
<geoCoordinate id="A71996D99441B1B49B60EF6672F80071" degrees="25" direction="north" minutes="16" orientation="latitude" precision="1" seconds="39.1039" value="25.277529">25°16'39.1039&quot;N</geoCoordinate>
, 
<geoCoordinate id="3FB9B9B080B086D0A436C7BD971B9992" degrees="107" direction="east" minutes="55" orientation="longitude" precision="1" seconds="2.7598" value="107.91743">107°55'2.7598&quot;E</geoCoordinate>
, 
<quantity id="B2EDD059DD1D8B0A29E004EBA940127E" metricMagnitude="2" metricUnit="m" metricValue="8.67" unit="m" value="867.0">
<elevation id="1F632195CDBE4D5C91302DDC17C82993" metricMagnitude="2" metricUnit="m" metricValue="8.67" unit="m" value="867.0">867 m</elevation>
</quantity>
elevation, 
<collectingDate id="3C863F90FF9BB9179575F5EDA4D83451" value="2019-11-09">9 Nov 2019</collectingDate>
, Wang Zheng-wei and Li Xiao-chen, 
<accessionNumber id="0179CB471245B51CE6F435E1BEAAC3F1" httpUri="http://www.ncbi.nlm.nih.gov/protein/LXC00927">LXC00927</accessionNumber>
<emphasis id="32C2B084F1F25FB24FD52C596565D801" bold="true" italics="true" pageId="0" pageNumber="63">
<typeStatus id="2F3C0F05FFC739848FAADAFA874642F1">Holotype</typeStatus>
</emphasis>
: CSH0182173 (CSH!); 
<emphasis id="970F4D29980ED76E365C6F6180F3C8B0" bold="true" italics="true" pageId="0" pageNumber="63">isotypes</emphasis>
: ZJFC!, CSFI!, IBSC
</materialsCitation>
!.
</paragraph>
<caption id="F34F7B4581650CB891134C525926ACD3" doi="10.3897/phytokeys.196.83176.figure2" httpUri="https://binary.pensoft.net/fig/683701" pageId="0" pageNumber="63" start="Figure 2" startId="F2">
<paragraph id="B1A18ABE1E312C8B592EF64B8DF5AA19" pageId="0" pageNumber="63">
<emphasis id="17493B174D2350034FCE698C80EB3D1F" bold="true" pageId="0" pageNumber="63">Figure 2.</emphasis>
<taxonomicName id="1F7CC4D22A76362D85ECF910C9DDE62E" authorityName="Xiao C. Li &amp; Z. W. Wang" authorityYear="2022" class="Magnoliopsida" family="Violaceae" genus="Viola" kingdom="Plantae" lsidName="Viola shiweii" order="Malpighiales" pageId="0" pageNumber="63" phylum="Tracheophyta" rank="species" species="shiweii">
<emphasis id="5B28D0FA0A6B4B4FAF69AC460F46DDCE" italics="true" pageId="0" pageNumber="63">Viola shiweii</emphasis>
</taxonomicName>
sp. nov. 
<emphasis id="AC6BF1EC3D8FFDACF609E2B5DF43DF44" bold="true" pageId="0" pageNumber="63">A</emphasis>
flower side view 
<emphasis id="3E9FFE10D82A07B88015B3442D19CB79" bold="true" pageId="0" pageNumber="63">B</emphasis>
flower front view 
<emphasis id="18389E6607F329AF4E80B80EB265B14D" bold="true" pageId="0" pageNumber="63">C</emphasis>
habit 
<emphasis id="11F7A0D4712B9A9D8B94E8628ED0473A" bold="true" pageId="0" pageNumber="63">D</emphasis>
stipule 
<emphasis id="9BB077ADF81D5F844E8E1F7DD5D48858" bold="true" pageId="0" pageNumber="63">E</emphasis>
capsule 
<emphasis id="FF8572515BBC124FF360A0B8F662F532" bold="true" pageId="0" pageNumber="63">F</emphasis>
stamens and pistil.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="0C1CCC2B4BD513153204D41971E7CEC0" pageId="0" pageNumber="63" type="diagnosis">
<paragraph id="B2CFCDA9EAAEED3426EB4F0E9842F9E9" pageId="0" pageNumber="63">Diagnosis.</paragraph>
<paragraph id="19EE4D10EF5EB5E665503344577FBA85" pageId="0" pageNumber="63">
The new species is morphologically most similar to 
<taxonomicName id="8469EB434B00FF0C8865580082E1F502" class="Magnoliopsida" family="Violaceae" genus="Viola" kingdom="Plantae" lsidName="V. kwangtungensis" order="Malpighiales" pageId="0" pageNumber="63" phylum="Tracheophyta" rank="species" species="kwangtungensis">
<emphasis id="BE89F7514B606E49BB826BF06F80BCDD" italics="true" pageId="0" pageNumber="63">V. kwangtungensis</emphasis>
</taxonomicName>
Melchior in its glabrous lateral petals and stoloniferous habit, but can be distinguished by its obtuse teeth along the leaf margin, its abaxially pale green leaf blade and its broader and entire sepals with a distinct basal appendage.
</paragraph>
</subSubSection>
<subSubSection id="9E8D531872C41D67FF7708AA02AD2D9E" pageId="0" pageNumber="63" type="description">
<paragraph id="7C2063E3AEDCB0D68F6AF26E78A09D53" pageId="0" pageNumber="63">Description.</paragraph>
<paragraph id="940948C2D6B86E3618E2D32DCF1FFB9A" pageId="0" pageNumber="63">
Perennial herb, acaulescent, with stolons. Rhizome short, straight or oblique, densely noded, usually covered by brown remains of stipules. 
<emphasis id="9648CFEF6D0FD84CA0244461C6F98D2A" bold="true" pageId="0" pageNumber="63">Stipules</emphasis>
free, brown, broadly lanceolate, 5-10 mm long, margin long fimbriate-dentate, lobes remotely dentate. 
<emphasis id="80A36B580BCD4E03B9CA375C04E9088B" bold="true" pageId="0" pageNumber="63">Basal leaves</emphasis>
glabrous, slightly grooved, with petioles stout, petioles unequal in length; blade thick leathery, ovate or suborbicular, 15-30 
<normalizedToken id="B8C611B0DD04EA8B54AB5C273F5B1F82" originalValue="×">x</normalizedToken>
15-20 mm, base deep cordate, apex usually obtuse, abaxially green, scabrous, abaxially greyish-green, mid-vein distinct above, glabrous on both surfaces, margin glandular-serrate or shallowly glandular-crenate, slightly wavy, teeth obtuse at the apex; 
<emphasis id="85EB32AF37FF804146844F13831C999F" bold="true" pageId="0" pageNumber="63">stolon leaves</emphasis>
scattered, smaller. Pedicel equal to or longer than petiole, two bracts narrowly lanceolate, at the middle or lower part of the pedicel. 
<emphasis id="51B942832A79D86197EC6AAEEC60401F" bold="true" pageId="0" pageNumber="63">Sepals</emphasis>
5, ca. 6 mm long and 2 mm wide, lanceolate, glabrous, margin narrowly membranous, apex somewhat acute, base distinctly decurrent, apex obtuse or shallowly dentate. Flower 1.5-2.5 cm in diameter, 
<emphasis id="BFAD1157B16958081E3A65C6CEDBAAE3" bold="true" pageId="0" pageNumber="63">petals</emphasis>
5, white, posterior and lateral ones obovate, ca. 1.2 cm 
<normalizedToken id="79CF94D9027D48FFADA1F5E82D8C6C12" originalValue="×">x</normalizedToken>
5 mm, narrow at the base, lateral petals purplish near the middle, glabrous, anterior petal shorter, ca. 10 mm (spur included) long, oblong, purple-veined, apex rounded, obtuse, spur saccate, 2-3 mm long and 1.5 mm wide. 
<emphasis id="18B8CDD0DC511F422DE0F1B6B55559FE" bold="true" pageId="0" pageNumber="63">Style</emphasis>
clavate, base geniculate, stigmas flattened on top, narrowly margined on lateral sides and abaxially, shortly beaked ventrally. 
<emphasis id="0BFF5B551925B21A8358D1623F8A4214" bold="true" pageId="0" pageNumber="63">Capsule</emphasis>
ellipsoid, valves carinate, ca. 10 mm long and 2.5 mm wide, glabrous. 
<emphasis id="E4560CD4C6824B24EAB4AF4542F96360" bold="true" pageId="0" pageNumber="63">Seeds</emphasis>
black, ca. 2 mm long and 1 mm in diameter.
</paragraph>
<caption id="9F4011AEB8D16648CA444029F18F1FE9" doi="10.3897/phytokeys.196.83176.figure3" httpUri="https://binary.pensoft.net/fig/683702" pageId="0" pageNumber="63" start="Figure 3" startId="F3">
<paragraph id="EE72CF80043F04A5D7AC3BCAD72FC758" pageId="0" pageNumber="63">
<emphasis id="CD075435572DF1A3134DD852DDFAD3B5" bold="true" pageId="0" pageNumber="63">Figure 3.</emphasis>
<taxonomicName id="BC66C34E700EBF75D560DA60591DEFBA" authorityName="Xiao C. Li &amp; Z. W. Wang" authorityYear="2022" class="Magnoliopsida" family="Violaceae" genus="Viola" kingdom="Plantae" lsidName="Viola shiweii" order="Malpighiales" pageId="0" pageNumber="63" phylum="Tracheophyta" rank="species" species="shiweii">
<emphasis id="3944FEF2EE3E9BE5EB4214A172F69AC9" italics="true" pageId="0" pageNumber="63">Viola shiweii</emphasis>
</taxonomicName>
sp. nov. 
<emphasis id="4B4D1A78CD38D7B07692FA5154C61EB4" bold="true" pageId="0" pageNumber="63">A</emphasis>
petals, sepals, stamens and pistils 
<emphasis id="3D92C4A7C2C9BC4C7AAE8F88B06ED7C9" bold="true" pageId="0" pageNumber="63">B, G</emphasis>
flower in front view and detail of a longitudinal section in side 
<emphasis id="650C00D0046FDE565693E4D2C96215BF" bold="true" pageId="0" pageNumber="63">C, H</emphasis>
basal leaf adaxially and abaxially 
<emphasis id="397E225E7AD606AC5D202F2693FFAD00" bold="true" pageId="0" pageNumber="63">D</emphasis>
cleistogamous flowers 
<emphasis id="CCC93B49E65ABAEF4F5CB7C7E2152B3F" bold="true" pageId="0" pageNumber="63">E</emphasis>
capsule and seeds 
<emphasis id="059C5497EA557D693764AD8785A362E0" bold="true" pageId="0" pageNumber="63">F</emphasis>
margin teeth 
<emphasis id="80C9540E846C445CE5E2D52B75CBC05B" bold="true" pageId="0" pageNumber="63">I</emphasis>
stipule 
<emphasis id="C2D788852AA045EC6C5E066AE260E7EC" bold="true" pageId="0" pageNumber="63">J</emphasis>
leaves on stolons.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="2EC7E06AA81FEA172DE3D15793A5D21E" pageId="0" pageNumber="63" type="distribution">
<paragraph id="2A37E06F1604B07006D19E354D141E11" pageId="0" pageNumber="63">Distribution and habitat.</paragraph>
<paragraph id="F2418A6BFA931B889BDE17E62B509874" pageId="0" pageNumber="63">The species was observed to grow on dry and partially shaded limestone, around the karst forest edge, at 700-900 m elevation.</paragraph>
</subSubSection>
<subSubSection id="E7CD86E8AE93403C178EB7941E9006A6" pageId="0" pageNumber="63" type="materials_examined">
<paragraph id="853B0DECA5D36B5C7CB8D63EB42A1D6B" pageId="0" pageNumber="63">Additional specimens examined.</paragraph>
<paragraph id="C28C3DEA1D0C6C563BD3BF335B0A2B2D" pageId="0" pageNumber="63">
<materialsCitation id="D44ACFD16DB1DAC2CD18762E393F1000" collectingDate="2008-07-24" collectorName="Zhang Dai-Gui" country="China" county="Libo County" location="Maolan National Nature Reserve" municipality="Qiannan Autonomous Prefecture" specimenCount="1" stateProvince="Guizhou">
<emphasis id="6877CA00FC9D77E0E81CDCC2303F65B7" bold="true" pageId="0" pageNumber="63">
<collectingCountry id="6DEF0D0B2D15D91040C444735E23E840" name="China">China</collectingCountry>
</emphasis>
, 
<collectingRegion id="4915CDD37859E5B0EBE872E5464644CC" country="China" name="Guizhou">Guizhou</collectingRegion>
, 
<collectingMunicipality id="A3AEB9B28D96EB507A5EB20BABF56C12">Qiannan Autonomous Prefecture</collectingMunicipality>
, 
<collectingCounty id="F0C805263B9C45C108E2399B5003AFE1">Libo County</collectingCounty>
, 
<location id="C83D4654BC724287B5A6647FCF012449" LSID="urn:lsid:plazi:treatment:2EEA7109CB2454DD9C0A9C30C75088C3:C83D4654BC724287B5A6647FCF012449" country="China" county="Libo County" municipality="Qiannan Autonomous Prefecture" name="Maolan National Nature Reserve" stateProvince="Guizhou">Maolan National Nature Reserve</location>
, karst forest, 
<collectingDate id="2173E2510C916AA916D18AE655BC16D3" value="2008-07-24">24 Jul 2008</collectingDate>
, 
<collectorName id="D13121946F5757D5BA97B51B6B7006E6">Zhang Dai-Gui</collectorName>
080724077 (JIU!)
</materialsCitation>
; 
<materialsCitation id="D2910998BEC61732AF7E520E3C3EAAE2" accessionNumber="LXC02320, LXC02322, LXC02323, LXC02324, LXC02325" collectingDate="2021-11-21" collectorName="Li Xiao-chen, Wang Zheng-wei, Wei Hong-jin" country="China" county="Libo County" location="Miao Autonomous Prefecture" municipality="Qiannan Buyi" specimenCount="1" stateProvince="Guizhou">
<emphasis id="B61B2AD574D541C7C0E53E9B29E7FBD9" bold="true" pageId="0" pageNumber="63">
<collectingCountry id="F4629F2E2A808658793F53F8643E1067" name="China">China</collectingCountry>
</emphasis>
, 
<collectingRegion id="FA27E7C08643DBFA561934CE681E48EA" country="China" name="Guizhou">Guizhou</collectingRegion>
, 
<collectingMunicipality id="76F228350A911A155A2489FD327A2792">Qiannan Buyi</collectingMunicipality>
and 
<location id="5895D11DC4DFF138226E80761E649A9A" LSID="urn:lsid:plazi:treatment:2EEA7109CB2454DD9C0A9C30C75088C3:5895D11DC4DFF138226E80761E649A9A" country="China" county="Libo County" municipality="Qiannan Buyi" name="Miao Autonomous Prefecture" stateProvince="Guizhou">Miao Autonomous Prefecture</location>
, 
<collectingCounty id="149117E2F781A1A2E6E70D5931363386">Libo County</collectingCounty>
, 
<location id="0F3926F7D00133D2EF5372820F3B0178" LSID="urn:lsid:plazi:treatment:2EEA7109CB2454DD9C0A9C30C75088C3:0F3926F7D00133D2EF5372820F3B0178" country="China" county="Libo County" municipality="Qiannan Buyi" name="Maolan National Nature Reserve" stateProvince="Guizhou">Maolan National Nature Reserve</location>
, 
<collectingDate id="6DEE460B45789BA58532F420078D9ADC" value="2021-11-21">21 Nov 2021</collectingDate>
, 
<collectorName id="C1068891440767A6258BA3922291FB93">Li Xiao-chen</collectorName>
, 
<collectorName id="C3243D830D2B8AA778EC527ECA22B358">Wang Zheng-wei</collectorName>
&amp; 
<collectorName id="652EC77073696515721C3DD8E0B833C9">Wei Hong-jin</collectorName>
, 
<accessionNumber id="5535E16095E064C0A87ED41D77BF834B" httpUri="http://www.ncbi.nlm.nih.gov/protein/LXC02320">LXC02320</accessionNumber>
(CSH!), 
<accessionNumber id="72C2A504C20F6CEDBA7453C98EFEB739" httpUri="http://www.ncbi.nlm.nih.gov/protein/LXC02322">LXC02322</accessionNumber>
(CSH!), 
<accessionNumber id="29235C40C4E4C19A543D1FEAD47E1E84" httpUri="http://www.ncbi.nlm.nih.gov/protein/LXC02323">LXC02323</accessionNumber>
(CSH!), 
<accessionNumber id="12F7CAA0A83E00861CBB5F9F78AF90AE" httpUri="http://www.ncbi.nlm.nih.gov/protein/LXC02324">LXC02324</accessionNumber>
(CSH!), 
<accessionNumber id="5AC0C042964B2FE37E7A6D3ED0C7E0AA" httpUri="http://www.ncbi.nlm.nih.gov/protein/LXC02325">LXC02325</accessionNumber>
(CSH!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C822C7C7F5CCE162428867A128200379" pageId="0" pageNumber="63" type="phenology">
<paragraph id="639EE04FAA05505619DA87380CD41CDC" pageId="0" pageNumber="63">Phenology.</paragraph>
<paragraph id="21F057A0884BD794A414004A7C7D2C50" pageId="0" pageNumber="63">Cultivated plants flower in September-March, fruiting in September.</paragraph>
</subSubSection>
<subSubSection id="7E6DE711668B8AD49C9466835334DB71" pageId="0" pageNumber="63" type="etymology">
<paragraph id="F1B1A5C234A306F2B8E8E88D2AA9ABA6" pageId="0" pageNumber="63">Etymology.</paragraph>
<paragraph id="19BF468DAB5F03E2BAF9744A86E543B0" pageId="0" pageNumber="63">
The specific name epithet &quot; 
<emphasis id="E95AE73C491CEF30F068C05160DF4A44" italics="true" pageId="0" pageNumber="63">shiweii</emphasis>
&quot; was proposed in memory of Deng Shi-wei (191?-1936), who dedicated his life to the exploration of the flora of Guizhou. The Chinese name is given as 
<normalizedToken id="63A94C589173D231B6321B47F492386D" originalValue="“世纬堇菜”">&quot;世纬堇菜&quot;</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection id="FC4ECE826AD0B03E2D52CC26CF559026" pageId="0" pageNumber="63" type="conservation status">
<paragraph id="797D0050AD6EEBA8101856FE7F67842E" pageId="0" pageNumber="63">Conservation status.</paragraph>
<paragraph id="5EEE226788DE9795C2EC430787EDEC2F" pageId="0" pageNumber="63">
Only two populations of 
<taxonomicName id="BB11C7CCEEB01CDDCDE269F486D2C216" class="Magnoliopsida" family="Violaceae" genus="Viola" kingdom="Plantae" lsidName="V. shiweii" order="Malpighiales" pageId="0" pageNumber="63" phylum="Tracheophyta" rank="species" species="shiweii">
<emphasis id="AF3BC542430B76CE7F2DD97AE388A5BB" italics="true" pageId="0" pageNumber="63">V. shiweii</emphasis>
</taxonomicName>
are currently known from Maolan National Nature Reserve, Libo County, in an area of the karst formation across Guizhou and Guangxi (Fig. 
<figureCitation id="4096590362477846CB388DB7513D6C55" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitat and distribution of Viola shiweii." figureDoi="10.3897/phytokeys.196.83176.figure4" httpUri="https://binary.pensoft.net/fig/683703" pageId="0" pageNumber="63">4</figureCitation>
). This species is represented by no more than 200 large and mature individuals. Due to its rarity, the low number of individuals and habitat vulnerability, 
<taxonomicName id="62FC8CA7D6068C4AC520300536971B77" class="Magnoliopsida" family="Violaceae" genus="Viola" kingdom="Plantae" lsidName="V. shiweii" order="Malpighiales" pageId="0" pageNumber="63" phylum="Tracheophyta" rank="species" species="shiweii">
<emphasis id="37BA39F6F2233401C7A32CB5B87A6E51" italics="true" pageId="0" pageNumber="63">V. shiweii</emphasis>
</taxonomicName>
is considered to be Critically Endangered (CR, B1), according to the 
<bibRefCitation id="3FE9B20A1A0594E076141149F0F5AF71" author="IUCN" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="63" publicationUrl="http://cmsdocs.s3.amazonaws.com/RedListGuidelines.pdf" refId="B19" refString="IUCN, 2019. Guidelines for Using the IUCN Red List Categories and Criteria. Version 14. Prepared by the Standards and Petitions Committee. http://cmsdocs.s3.amazonaws.com/RedListGuidelines.pdf" title="Guidelines for Using the IUCN Red List Categories and Criteria. Version 14. Prepared by the Standards and Petitions Committee." url="http://cmsdocs.s3.amazonaws.com/RedListGuidelines.pdf" year="2019">IUCN (2019)</bibRefCitation>
.
</paragraph>
<caption id="9D0ED50262D7F2F2D5DB867BCE6E2432" doi="10.3897/phytokeys.196.83176.figure4" httpUri="https://binary.pensoft.net/fig/683703" pageId="0" pageNumber="63" start="Figure 4" startId="F4">
<paragraph id="913424B0F47CBFD9AD047C41F16D4B61" pageId="0" pageNumber="63">
<emphasis id="5B4342AEEBC2051638E6EFC75D46C9B8" bold="true" pageId="0" pageNumber="63">Figure 4.</emphasis>
Habitat and distribution of 
<taxonomicName id="38058E907DAC62384C664A5345F3709E" authorityName="Xiao C. Li &amp; Z. W. Wang" authorityYear="2022" class="Magnoliopsida" family="Violaceae" genus="Viola" kingdom="Plantae" lsidName="Viola shiweii" order="Malpighiales" pageId="0" pageNumber="63" phylum="Tracheophyta" rank="species" species="shiweii">
<emphasis id="0DC301B7470FFF07F7644B70D7673725" italics="true" pageId="0" pageNumber="63">Viola shiweii</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>