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<document id="74C6F4152D663BA1AAEF39B1F8C571A0" ID-DOI="10.11646/zootaxa.5306.3.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="8063106" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1687332062705" checkinUser="plazi" docAuthor="Sherwood, Danniella, Jocqué, Rudy, Henrard, Arnaud &amp; Fowler, Liza" docDate="2023" docId="2E0387F8B8761B2DFF0D7409FF039090" docLanguage="en" docName="zootaxa.5306.3.8.pdf" docOrigin="Zootaxa 5306 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5306.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Howaia mogera" docType="treatment" docVersion="3" lastPageNumber="400" masterDocId="D23AFF80B8741B2EFF9A771AFFDE9044" masterDocTitle="The scaffold web spider Nesticus helenensis Hubert, 1977, a junior synonym of Howaia mogera (Yaginuma, 1972) rest. comb., with revalidation of Howaia Lehtinen &amp; Saaristo, 1980 (Araneae: Nesticidae)" masterLastPageNumber="400" masterPageNumber="397" pageNumber="399" updateTime="1687368369211" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="19C7936463CF293864FCEF281C2641C7">The scaffold web spider Nesticus helenensis Hubert, 1977, a junior synonym of Howaia mogera (Yaginuma, 1972) rest. comb., with revalidation of Howaia Lehtinen &amp; Saaristo, 1980 (Araneae: Nesticidae)</mods:title>
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<mods:namePart id="51E5B8125F100BA64176D1282E7535FE">Sherwood, Danniella</mods:namePart>
<mods:affiliation id="1B12B087A0240F63605E02325491E1BA">Arachnology Research Association, London, United Kingdom. &amp; Fundación Ariguanabo, San Antonio de Los Baños, Cuba. &amp; IUCN Species Survival Commission, Mid-Atlantic Islands Invertebrate Specialist Group.</mods:affiliation>
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<mods:namePart id="6740F3455052888E46F05095418415A6">Jocqué, Rudy</mods:namePart>
<mods:affiliation id="40314A744873336895F54D5E28E9536A">Royal Museum for Central Africa, Tervuren, Belgium.</mods:affiliation>
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<mods:namePart id="4BB7CBC6416B56C6037361AD52348684">Henrard, Arnaud</mods:namePart>
<mods:affiliation id="433F57C5A825A175CB77BFADF2A95B01">Royal Museum for Central Africa, Tervuren, Belgium.</mods:affiliation>
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<mods:namePart id="9CF2F51BA2125B2A12ECCE72EDCDC915">Fowler, Liza</mods:namePart>
<mods:affiliation id="F1AAF9A596DB255F47A17151F8626F20">IUCN Species Survival Commission, Mid-Atlantic Islands Invertebrate Specialist Group. &amp; Saint Helena National Trust, Jamestown, Saint Helena.</mods:affiliation>
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<paragraph id="A61536EEB8761B2CFF0D7409FD129369" blockId="2.[151,1178,787,997]" box="[151,716,787,813]" pageId="2" pageNumber="399">
<heading id="FD5D8182B8761B2CFF0D7409FD129369" bold="true" box="[151,716,787,813]" fontSize="11" level="1" pageId="2" pageNumber="399" reason="1">
<emphasis id="94DEEAFCB8761B2CFF0D7409FD129369" bold="true" box="[151,716,787,813]" pageId="2" pageNumber="399">
<taxonomicName id="61AA4D6DB8761B2CFF0D7409FDE59369" authority="(Yaginuma, 1972)" baseAuthorityName="Yaginuma" baseAuthorityYear="1972" box="[151,571,787,813]" class="Arachnida" family="Nesticidae" genus="Howaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="mogera">
<emphasis id="94DEEAFCB8761B2CFF0D7409FE869369" bold="true" box="[151,344,787,813]" italics="true" pageId="2" pageNumber="399">Howaia mogera</emphasis>
(
<bibRefCitation id="C23B4B1FB8761B2CFEFD7409FDED9369" author="Yaginuma, T." box="[359,563,787,813]" pageId="2" pageNumber="399" pagination="619 - 622" refId="ref3090" refString="Yaginuma, T. (1972) Revision of the short-legged nesticid spiders of Japan. Bulletin of the National Museum of Nature and Science Tokyo, 15, 619 - 622." type="journal article" year="1972">Yaginuma, 1972</bibRefCitation>
)
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rest. comb.
</emphasis>
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</paragraph>
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<subSubSection id="EEB06565B8761B2CFF0D742EFDEA93A1" pageId="2" pageNumber="399" type="reference_group">
<paragraph id="A61536EEB8761B2CFF0D742EFCBC930F" blockId="2.[151,1178,787,997]" box="[151,866,819,846]" pageId="2" pageNumber="399">
<treatmentCitationGroup id="86BA11C0B8761B2CFF0D742EFCBC930F" box="[151,866,819,846]" pageId="2" pageNumber="399">
<treatmentCitation id="270B10FFB8761B2CFF0D742EFDE99308" author="Yaginuma, T." box="[151,567,819,844]" page="390" pageId="2" pageNumber="399" year="1970">
<taxonomicName id="61AA4D6DB8761B2CFF0D742EFDE99308" authority=": Yaginuma, 1970: 390" authorityName="Yaginuma" authorityPageNumber="390" authorityYear="1970" box="[151,567,819,844]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="terrestris">
<emphasis id="94DEEAFCB8761B2CFF0D742EFE91930F" box="[151,335,820,843]" italics="true" pageId="2" pageNumber="399">Nesticus terrestris</emphasis>
:
<bibRefCitation id="C23B4B1FB8761B2CFEC67429FDE99308" author="Yaginuma, T." box="[348,567,819,844]" pageId="2" pageNumber="399" pagination="385 - 394" refId="ref3056" refString="Yaginuma, T. (1970) Two new species of small nesticid spiders of Japan. Bulletin of the National Museum of Nature and Science Tokyo, 13, 385 - 394." type="journal article" year="1970">Yaginuma, 1970: 390</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 7 (♁, misidentification).
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</paragraph>
<paragraph id="A61536EEB8761B2CFF0D7449FD789329" blockId="2.[151,1178,787,997]" box="[151,678,850,877]" pageId="2" pageNumber="399">
<treatmentCitationGroup id="86BA11C0B8761B2CFF0D7449FD789329" box="[151,678,850,877]" pageId="2" pageNumber="399">
<treatmentCitation id="270B10FFB8761B2CFF0D7449FDFC932E" author="Yaginuma, T." box="[151,546,850,874]" page="621" pageId="2" pageNumber="399" year="1972">
<taxonomicName id="61AA4D6DB8761B2CFF0D7449FDFC932E" authority="Yaginuma, 1972: 621" authorityName="Yaginuma" authorityPageNumber="621" authorityYear="1972" box="[151,546,850,874]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="mogera">
<emphasis id="94DEEAFCB8761B2CFF0D7449FE9E932E" box="[151,320,851,874]" italics="true" pageId="2" pageNumber="399">Nesticus mogera</emphasis>
<bibRefCitation id="C23B4B1FB8761B2CFEDC7448FDFC932E" author="Yaginuma, T." box="[326,546,850,874]" pageId="2" pageNumber="399" pagination="619 - 622" refId="ref3090" refString="Yaginuma, T. (1972) Revision of the short-legged nesticid spiders of Japan. Bulletin of the National Museum of Nature and Science Tokyo, 15, 619 - 622." type="journal article" year="1972">Yaginuma, 1972: 621</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 1 (♁
<specimenCount id="B0ACFD67B8761B2CFD1F744EFD499329" box="[645,663,852,877]" pageId="2" pageNumber="399" type="female"></specimenCount>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A61536EEB8761B2CFF0D7468FCF793CD" blockId="2.[151,1178,787,997]" box="[151,809,881,907]" pageId="2" pageNumber="399">
<treatmentCitationGroup id="86BA11C0B8761B2CFF0D7468FCF793CD" box="[151,809,881,907]" pageId="2" pageNumber="399">
<treatmentCitation id="270B10FFB8761B2CFF0D7468FDC093CD" author="Hubert, M." box="[151,542,881,905]" page="153" pageId="2" pageNumber="399" year="1977">
<taxonomicName id="61AA4D6DB8761B2CFF0D7468FDC093CD" authority="Hubert, 1977: 153" authorityName="Hubert" authorityPageNumber="153" authorityYear="1977" box="[151,542,881,905]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="helenensis">
<emphasis id="94DEEAFCB8761B2CFF0D7468FE8393CD" box="[151,349,881,905]" italics="true" pageId="2" pageNumber="399">Nesticus helenensis</emphasis>
<bibRefCitation id="C23B4B1FB8761B2CFEF9746BFDC093CD" author="Hubert, M." box="[355,542,881,905]" pageId="2" pageNumber="399" pagination="153 - 156" refId="ref2435" refString="Hubert, M. (1977) Fam. Nesticidae. In: La faune terrestre de l'ile de Sainte-Helene IV. Annales, Musee Royal de l'Afrique Centrale, Sciences zoologiques (Zool. - Ser. 8 °), 220, 153 - 156." type="journal article" year="1977">Hubert, 1977: 153</bibRefCitation>
</taxonomicName>
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, figs. 67ah. (♁
<specimenCount id="B0ACFD67B8761B2CFD587468FD0D93CF" box="[706,723,882,907]" pageId="2" pageNumber="399" type="female"></specimenCount>
)
<emphasis id="94DEEAFCB8761B2CFD79746BFCF793CD" bold="true" box="[739,809,881,905]" pageId="2" pageNumber="399">n. syn.</emphasis>
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</paragraph>
<paragraph id="A61536EEB8761B2CFF0D748AFB4493EC" blockId="2.[151,1178,787,997]" box="[151,1178,911,938]" pageId="2" pageNumber="399">
<treatmentCitationGroup id="86BA11C0B8761B2CFF0D748AFB4493EC" box="[151,1178,911,938]" pageId="2" pageNumber="399">
<treatmentCitation id="270B10FFB8761B2CFF0D748AFD5993E3" author="Lehtinen, P. T. &amp; Saaristo, M. I." box="[151,647,911,936]" page="53" pageId="2" pageNumber="399" year="1980">
<taxonomicName id="61AA4D6DB8761B2CFF0D748AFDBC93EC" authority=": Lehtinen &amp; Saaristo (1980)" authorityName="Lehtinen &amp; Saaristo" authorityYear="1980" box="[151,610,911,936]" class="Arachnida" family="Nesticidae" genus="Howaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="mogera">
<emphasis id="94DEEAFCB8761B2CFF0D748AFEE593E3" box="[151,315,912,935]" italics="true" pageId="2" pageNumber="399">Howaia mogera</emphasis>
:
<bibRefCitation id="C23B4B1FB8761B2CFED37495FDBC93EC" author="Lehtinen, P. T. &amp; Saaristo, M. I." box="[329,610,911,936]" pageId="2" pageNumber="399" pagination="47 - 66" refId="ref2591" refString="Lehtinen, P. T. &amp; Saaristo, M. I. (1980) Spiders of the Oriental-Australian region. II. Nesticidae. Annales Zoologici Fennici, 17, 47 - 66." type="journal article" year="1980">Lehtinen &amp; Saaristo (1980)</bibRefCitation>
</taxonomicName>
: 53
</treatmentCitation>
, figs. 79, 2223, 29b (♁
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, transfer from
<taxonomicName id="61AA4D6DB8761B2CFBAC748AFB5293E3" authorityName=": Yaginuma" authorityYear="1972" box="[1078,1164,912,935]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="399" phylum="Arthropoda" rank="genus">
<emphasis id="94DEEAFCB8761B2CFBAC748AFB5293E3" box="[1078,1164,912,935]" italics="true" pageId="2" pageNumber="399">Nesticus</emphasis>
</taxonomicName>
).
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</paragraph>
<paragraph id="A61536EEB8761B2CFF0D74B5FB829382" blockId="2.[151,1178,787,997]" box="[151,1116,942,969]" pageId="2" pageNumber="399">
<treatmentCitationGroup id="86BA11C0B8761B2CFF0D74B5FB829382" box="[151,1116,942,969]" pageId="2" pageNumber="399">
<treatmentCitation id="270B10FFB8761B2CFF0D74B5FD4F9382" author="Marusik, Y. M. &amp; Guseinov, E." box="[151,657,942,966]" page="38" pageId="2" pageNumber="399" year="2003">
<taxonomicName id="61AA4D6DB8761B2CFF0D74B5FDB39382" authority=": Marusik &amp; Guseinov (2003)" authorityName=": Marusik &amp; Guseinov" authorityYear="2003" box="[151,621,942,966]" class="Arachnida" family="Nesticidae" genus="Howaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="mogera">
<emphasis id="94DEEAFCB8761B2CFF0D74B5FEE59382" box="[151,315,943,966]" italics="true" pageId="2" pageNumber="399">Howaia mogera</emphasis>
:
<bibRefCitation id="C23B4B1FB8761B2CFED374B4FDB39382" author="Marusik, Y. M. &amp; Guseinov, E." box="[329,621,942,966]" pageId="2" pageNumber="399" pagination="29 - 46" refId="ref2745" refString="Marusik, Y. M. &amp; Guseinov, E. (2003) Spiders (Arachnida: Aranei) of Azerbaijan. 1. New family and genus records. Arthropoda Selecta, 12 (1), 29 - 46." type="journal article" year="2003">Marusik &amp; Guseinov (2003)</bibRefCitation>
</taxonomicName>
: 38
</treatmentCitation>
, figs. 1721 (♁
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, not explicitly revalidated).
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</paragraph>
<paragraph id="A61536EEB8761B2CFF0D74D7FDEA93A1" blockId="2.[151,1178,787,997]" box="[151,564,973,997]" pageId="2" pageNumber="399">
For full synonymy list, see
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.
</paragraph>
</subSubSection>
<subSubSection id="EEB06565B8761B2CFF0D730FFD2F95D1" pageId="2" pageNumber="399" type="materials_examined">
<paragraph id="A61536EEB8761B2CFF0D730FFD2F95D1" blockId="2.[151,1437,1045,2041]" pageId="2" pageNumber="399">
<emphasis id="94DEEAFCB8761B2CFF0D730FFEA79469" bold="true" box="[151,377,1045,1069]" inLineHeading="true" pageId="2" pageNumber="399" reason="1">Material examined.</emphasis>
Type material of
<taxonomicName id="61AA4D6DB8761B2CFDDE730CFCC69469" authorityName="Hubert" authorityYear="1977" box="[580,792,1045,1069]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="helenensis">
<emphasis id="94DEEAFCB8761B2CFDDE730CFCC69469" box="[580,792,1045,1069]" italics="true" pageId="2" pageNumber="399">Nesticus helenensis</emphasis>
</taxonomicName>
:
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,
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<materialsCitation id="16C23CB3B8761B2CFC2C730FFEDC9431" collectingDate="1965-12-14" collectingDateMax="1965-12-15" collectingDateMin="1965-12-14" collectionCode="BE, RMCA" collectorName="P. Basilewsky &amp; P. L. G. Benoit &amp; N. Leleup" country="Saint Helena, Ascension and Tristan da Cunha" elevation="305" location="Ara." municipality="Fisher's Valley" pageId="2" pageNumber="399" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="7911884CB8761B2CFC2C730FFBCC9469" box="[950,1042,1045,1069]" pageId="2" pageNumber="399">paratype</typeStatus>
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(
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_
<collectionCode id="C0BBAE2BB8761B2CFBE9730FFB189469" box="[1139,1222,1045,1069]" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="2" pageNumber="399" type="Museum">RMCA</collectionCode>
_ARA.
<location id="A3756035B8761B2CFA88730FFA9E9469" LSID="urn:lsid:plazi:treatment:2E0387F8B8761B2DFF0D7409FF039090:A3756035B8761B2CFA88730FFA9E9469" box="[1298,1344,1045,1069]" country="Saint Helena, Ascension and Tristan da Cunha" municipality="Fisher's Valley" name="Ara." pageId="2" pageNumber="399">Ara.</location>
129084),
<collectingMunicipality id="4671AC94B8761B2CFF0D7323FEE99415" box="[151,311,1081,1105]" pageId="2" pageNumber="399">Fisher's Valley</collectingMunicipality>
,
<collectingCountry id="DEBD767EB8761B2CFEDF7323FE0A9415" box="[325,468,1081,1105]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="2" pageNumber="399">Saint Helena</collectingCountry>
,
<quantity id="61529B0BB8761B2CFE787323FDE69415" box="[482,568,1081,1105]" metricMagnitude="2" metricUnit="m" metricValue="3.048" pageId="2" pageNumber="399" unit="ft" value="1000.0">
<elevation id="2D87D1DDB8761B2CFE787323FDE69415" box="[482,568,1081,1105]" metricMagnitude="2" metricUnit="m" metricValue="3.048" pageId="2" pageNumber="399" unit="ft" value="1000.0">1,000ft.</elevation>
</quantity>
,
<date id="D214102EB8761B2CFDDF7323FD169415" box="[581,712,1081,1105]" pageId="2" pageNumber="399" value="1965-12-14">
<collectingDate id="C250E9C6B8761B2CFDDF7323FD169415" box="[581,712,1081,1105]" pageId="2" pageNumber="399" value="1965-12-14">14.XII.1965</collectingDate>
</date>
<date id="D214102EB8761B2CFD4D7323FC879415" box="[727,857,1081,1105]" pageId="2" pageNumber="399" value="1965-12-15">
<collectingDate id="C250E9C6B8761B2CFD4D7323FC879415" box="[727,857,1081,1105]" pageId="2" pageNumber="399" value="1965-12-15">15.XII.1965</collectingDate>
</date>
, coll.
<collectorName id="0B5F5338B8761B2CFC3A7323FBE69415" box="[928,1080,1081,1105]" pageId="2" pageNumber="399">P. Basilewsky</collectorName>
,
<collectorName id="0B5F5338B8761B2CFBDC7323FB309415" box="[1094,1262,1081,1105]" pageId="2" pageNumber="399">P. L. G. Benoit</collectorName>
and
<collectorName id="0B5F5338B8761B2CFAB27323FA469415" box="[1320,1432,1081,1105]" pageId="2" pageNumber="399">N. Leleup</collectorName>
, examined
</materialsCitation>
;
<materialsCitation id="16C23CB3B8761B2CFE967347FDC594DD" collectingDate="1967-01-26" collectionCode="BE, RMCA" collectorName="J. Decelle &amp; N. Leleup" country="Saint Helena, Ascension and Tristan da Cunha" elevation="305" location="Ara." municipality="Fisher's Valley" pageId="2" pageNumber="399" specimenCount="1" typeStatus="paratype">
<typeStatus id="7911884CB8761B2CFE967347FEB69431" box="[268,360,1117,1141]" pageId="2" pageNumber="399">paratype</typeStatus>
♁ [allotype] (
<collectionCode id="C0BBAE2BB8761B2CFE647347FDFF9431" box="[510,545,1117,1141]" pageId="2" pageNumber="399">BE</collectionCode>
_
<collectionCode id="C0BBAE2BB8761B2CFDB57347FD5C9431" box="[559,642,1117,1141]" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="2" pageNumber="399" type="Museum">RMCA</collectionCode>
_ARA.
<location id="A3756035B8761B2CFD547347FD229431" LSID="urn:lsid:plazi:treatment:2E0387F8B8761B2DFF0D7409FF039090:A3756035B8761B2CFD547347FD229431" box="[718,764,1117,1141]" country="Saint Helena, Ascension and Tristan da Cunha" municipality="Fisher's Valley" name="Ara." pageId="2" pageNumber="399">Ara.</location>
133336),
<collectingMunicipality id="4671AC94B8761B2CFCC47347FC259431" box="[862,1019,1117,1141]" pageId="2" pageNumber="399">Fisher's Valley</collectingMunicipality>
,
<collectingCountry id="DEBD767EB8761B2CFB9F7347FB4E9431" box="[1029,1168,1117,1141]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="2" pageNumber="399">Saint Helena</collectingCountry>
,
<quantity id="61529B0BB8761B2CFB017347FB2F9431" box="[1179,1265,1117,1141]" metricMagnitude="2" metricUnit="m" metricValue="3.048" pageId="2" pageNumber="399" unit="ft" value="1000.0">
<elevation id="2D87D1DDB8761B2CFB017347FB2F9431" box="[1179,1265,1117,1141]" metricMagnitude="2" metricUnit="m" metricValue="3.048" pageId="2" pageNumber="399" unit="ft" value="1000.0">1,000ft.</elevation>
</quantity>
,
<date id="D214102EB8761B2CFB607347FABD9431" box="[1274,1379,1117,1141]" pageId="2" pageNumber="399" value="1967-01-26">
<collectingDate id="C250E9C6B8761B2CFB607347FABD9431" box="[1274,1379,1117,1141]" pageId="2" pageNumber="399" value="1967-01-26">26.I.1967</collectingDate>
</date>
, coll.
<collectorName id="0B5F5338B8761B2CFF0D739BFEDF94DD" box="[151,257,1153,1177]" pageId="2" pageNumber="399">J. Decelle</collectorName>
and
<collectorName id="0B5F5338B8761B2CFEAC739BFE7B94DD" box="[310,421,1153,1177]" pageId="2" pageNumber="399">N. Leleup</collectorName>
, examined
</materialsCitation>
;
<materialsCitation id="16C23CB3B8761B2CFDBC739BFAE994F9" collectingDate="1966-01-06" collectionCode="BE, RMCA" collectorName="P. Basilewsky &amp; P. L. G. Benoit &amp; N. Leleup" country="Saint Helena, Ascension and Tristan da Cunha" elevation="305" location="Ara." municipality="The Briars" pageId="2" pageNumber="399" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="7911884CB8761B2CFDBC739BFD5C94DD" box="[550,642,1153,1177]" pageId="2" pageNumber="399">paratype</typeStatus>
<specimenCount id="B0ACFD67B8761B2CFD10739BFD4094DC" box="[650,670,1153,1176]" pageId="2" pageNumber="399" type="female"></specimenCount>
(
<collectionCode id="C0BBAE2BB8761B2CFD36739BFD1194DD" box="[684,719,1153,1177]" pageId="2" pageNumber="399">BE</collectionCode>
_
<collectionCode id="C0BBAE2BB8761B2CFD47739BFCEE94DD" box="[733,816,1153,1177]" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="2" pageNumber="399" type="Museum">RMCA</collectionCode>
_ARA.
<location id="A3756035B8761B2CFCE7739BFC7594DD" LSID="urn:lsid:plazi:treatment:2E0387F8B8761B2DFF0D7409FF039090:A3756035B8761B2CFCE7739BFC7594DD" box="[893,939,1153,1177]" country="Saint Helena, Ascension and Tristan da Cunha" municipality="The Briars" name="Ara." pageId="2" pageNumber="399">Ara.</location>
129205),
<collectingMunicipality id="4671AC94B8761B2CFB97739BFBA194DD" box="[1037,1151,1153,1177]" pageId="2" pageNumber="399">The Briars</collectingMunicipality>
[listed as Prosperous Bay, Bryan's Rock in
<bibRefCitation id="C23B4B1FB8761B2CFEDC73BFFE3F94F9" author="Hubert, M." box="[326,481,1189,1213]" pageId="2" pageNumber="399" pagination="153 - 156" refId="ref2435" refString="Hubert, M. (1977) Fam. Nesticidae. In: La faune terrestre de l'ile de Sainte-Helene IV. Annales, Musee Royal de l'Afrique Centrale, Sciences zoologiques (Zool. - Ser. 8 °), 220, 153 - 156." type="journal article" year="1977">Hubert (1977)</bibRefCitation>
],
<quantity id="61529B0BB8761B2CFE6873BFFD9694F9" box="[498,584,1189,1213]" metricMagnitude="2" metricUnit="m" metricValue="3.048" pageId="2" pageNumber="399" unit="ft" value="1000.0">
<elevation id="2D87D1DDB8761B2CFE6873BFFD9694F9" box="[498,584,1189,1213]" metricMagnitude="2" metricUnit="m" metricValue="3.048" pageId="2" pageNumber="399" unit="ft" value="1000.0">1,000ft.</elevation>
</quantity>
,
<date id="D214102EB8761B2CFDCB73BFFD7294F9" box="[593,684,1189,1213]" pageId="2" pageNumber="399" value="1966-01-06">
<collectingDate id="C250E9C6B8761B2CFDCB73BFFD7294F9" box="[593,684,1189,1213]" pageId="2" pageNumber="399" value="1966-01-06">6.I.1966</collectingDate>
</date>
, coll.
<collectorName id="0B5F5338B8761B2CFD7173BFFCA194F9" box="[747,895,1189,1213]" pageId="2" pageNumber="399">P. Basilewsky</collectorName>
,
<collectorName id="0B5F5338B8761B2CFC1273BFFBFA94F9" box="[904,1060,1189,1213]" pageId="2" pageNumber="399">P. L. G. Benoit</collectorName>
and
<collectorName id="0B5F5338B8761B2CFBCC73BFFB1C94F9" box="[1110,1218,1189,1213]" pageId="2" pageNumber="399">N. Leleup</collectorName>
, examined
</materialsCitation>
;
<materialsCitation id="16C23CB3B8761B2CFADA73BFFDDE9541" collectingDate="1965-11-14" collectionCode="BE, RMCA" collectorName="P. Basilewsky &amp; P. L. G. Benoit &amp; N. Leleup" country="Saint Helena, Ascension and Tristan da Cunha" county="Bryan's Rock" elevation="305" location="Ara." municipality="Prosperous Bay" pageId="2" pageNumber="399" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="7911884CB8761B2CFADA73BFFA4294F9" box="[1344,1436,1189,1213]" pageId="2" pageNumber="399">paratype</typeStatus>
<specimenCount id="B0ACFD67B8761B2CFF0D73D3FF7594A4" box="[151,171,1225,1248]" pageId="2" pageNumber="399" type="female"></specimenCount>
(
<collectionCode id="C0BBAE2BB8761B2CFF2673D3FF0194A5" box="[188,223,1225,1249]" pageId="2" pageNumber="399">BE</collectionCode>
_
<collectionCode id="C0BBAE2BB8761B2CFF7773D3FE9E94A5" box="[237,320,1225,1249]" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="2" pageNumber="399" type="Museum">RMCA</collectionCode>
_ARA.
<location id="A3756035B8761B2CFE1773D3FE6594A5" LSID="urn:lsid:plazi:treatment:2E0387F8B8761B2DFF0D7409FF039090:A3756035B8761B2CFE1773D3FE6594A5" box="[397,443,1225,1249]" country="Saint Helena, Ascension and Tristan da Cunha" county="Bryan's Rock" municipality="Prosperous Bay" name="Ara." pageId="2" pageNumber="399">Ara.</location>
129183),
<collectingMunicipality id="4671AC94B8761B2CFDBA73D3FD1294A5" box="[544,716,1225,1249]" pageId="2" pageNumber="399">Prosperous Bay</collectingMunicipality>
,
<collectingCounty id="4F744E62B8761B2CFD4373D3FCB094A5" box="[729,878,1225,1249]" pageId="2" pageNumber="399">Bryan's Rock</collectingCounty>
,
<quantity id="61529B0BB8761B2CFCE173D3FC0C94A5" box="[891,978,1225,1249]" metricMagnitude="2" metricUnit="m" metricValue="3.048" pageId="2" pageNumber="399" unit="ft" value="1000.0">
<elevation id="2D87D1DDB8761B2CFCE173D3FC0C94A5" box="[891,978,1225,1249]" metricMagnitude="2" metricUnit="m" metricValue="3.048" pageId="2" pageNumber="399" unit="ft" value="1000.0">1,000ft.</elevation>
</quantity>
,
<date id="D214102EB8761B2CFC4473D3FB8594A5" box="[990,1115,1225,1249]" pageId="2" pageNumber="399" value="1965-11-14">
<collectingDate id="C250E9C6B8761B2CFC4473D3FB8594A5" box="[990,1115,1225,1249]" pageId="2" pageNumber="399" value="1965-11-14">14.XI.1965</collectingDate>
</date>
, coll.
<collectorName id="0B5F5338B8761B2CFB3A73D3FAE694A5" box="[1184,1336,1225,1249]" pageId="2" pageNumber="399">P. Basilewsky</collectorName>
,
<collectorName id="0B5F5338B8761B2CFADC73D3FF009541" pageId="2" pageNumber="399">P. L. G. Benoit</collectorName>
and
<collectorName id="0B5F5338B8761B2CFE8D73F7FE569541" box="[279,392,1261,1285]" pageId="2" pageNumber="399">N. Leleup</collectorName>
, examined
</materialsCitation>
;
<materialsCitation id="16C23CB3B8761B2CFD9773F7FC74956D" collectingDate="1966-01-06" collectionCode="BE, RMCA" collectorName="P. Basilewsky &amp; P. L. G. Benoit &amp; N. Leleup" country="Saint Helena, Ascension and Tristan da Cunha" elevation="305" location="Ara." municipality="Fisher's Valley" pageId="2" pageNumber="399" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="7911884CB8761B2CFD9773F7FDB79541" box="[525,617,1261,1285]" pageId="2" pageNumber="399">paratype</typeStatus>
<specimenCount id="B0ACFD67B8761B2CFDE873F7FD589540" box="[626,646,1261,1284]" pageId="2" pageNumber="399" type="female"></specimenCount>
(
<collectionCode id="C0BBAE2BB8761B2CFD0D73F7FD649541" box="[663,698,1261,1285]" pageId="2" pageNumber="399">BE</collectionCode>
_
<collectionCode id="C0BBAE2BB8761B2CFD5273F7FCC59541" box="[712,795,1261,1285]" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="2" pageNumber="399" type="Museum">RMCA</collectionCode>
_ARA.
<location id="A3756035B8761B2CFCF273F7FC489541" LSID="urn:lsid:plazi:treatment:2E0387F8B8761B2DFF0D7409FF039090:A3756035B8761B2CFCF273F7FC489541" box="[872,918,1261,1285]" country="Saint Helena, Ascension and Tristan da Cunha" municipality="Fisher's Valley" name="Ara." pageId="2" pageNumber="399">Ara.</location>
129171),
<collectingMunicipality id="4671AC94B8761B2CFC6073F7FB449541" box="[1018,1178,1261,1285]" pageId="2" pageNumber="399">Fisher's Valley</collectingMunicipality>
,
<collectingCountry id="DEBD767EB8761B2CFB3D73F7FAEB9541" box="[1191,1333,1261,1285]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="2" pageNumber="399">Saint Helena</collectingCountry>
,
<quantity id="61529B0BB8761B2CFAD973F7FA479541" box="[1347,1433,1261,1285]" metricMagnitude="2" metricUnit="m" metricValue="3.048" pageId="2" pageNumber="399" unit="ft" value="1000.0">
<elevation id="2D87D1DDB8761B2CFAD973F7FA479541" box="[1347,1433,1261,1285]" metricMagnitude="2" metricUnit="m" metricValue="3.048" pageId="2" pageNumber="399" unit="ft" value="1000.0">1,000ft.</elevation>
</quantity>
,
<date id="D214102EB8761B2CFF0D720BFF2C956D" box="[151,242,1297,1321]" pageId="2" pageNumber="399" value="1966-01-06">
<collectingDate id="C250E9C6B8761B2CFF0D720BFF2C956D" box="[151,242,1297,1321]" pageId="2" pageNumber="399" value="1966-01-06">6.I.1966</collectingDate>
</date>
, coll.
<collectorName id="0B5F5338B8761B2CFEA3720BFE0F956D" box="[313,465,1297,1321]" pageId="2" pageNumber="399">P. Basilewsky</collectorName>
,
<collectorName id="0B5F5338B8761B2CFE45720BFD59956D" box="[479,647,1297,1321]" pageId="2" pageNumber="399">P. L. G. Benoit</collectorName>
and
<collectorName id="0B5F5338B8761B2CFD5B720BFCEF956D" box="[705,817,1297,1321]" pageId="2" pageNumber="399">N. Leleup</collectorName>
, examined
</materialsCitation>
;
<materialsCitation id="16C23CB3B8761B2CFC22720BFA469509" collectingDate="1965-12-19" collectionCode="BE, RMCA" collectorName="P. Basilewsky &amp; P. L. G. Benoit &amp; N. Leleup" country="Saint Helena, Ascension and Tristan da Cunha" elevation="305" location="Ara." municipality="Fisher's Valley" pageId="2" pageNumber="399" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="7911884CB8761B2CFC22720BFBCA956D" box="[952,1044,1297,1321]" pageId="2" pageNumber="399">paratype</typeStatus>
<specimenCount id="B0ACFD67B8761B2CFB87720BFBEF956C" box="[1053,1073,1297,1320]" pageId="2" pageNumber="399" type="female"></specimenCount>
(
<collectionCode id="C0BBAE2BB8761B2CFBD8720BFBBB956D" box="[1090,1125,1297,1321]" pageId="2" pageNumber="399">BE</collectionCode>
_
<collectionCode id="C0BBAE2BB8761B2CFBE9720BFB18956D" box="[1139,1222,1297,1321]" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="2" pageNumber="399" type="Museum">RMCA</collectionCode>
_ARA.
<location id="A3756035B8761B2CFA89720BFA9F956D" LSID="urn:lsid:plazi:treatment:2E0387F8B8761B2DFF0D7409FF039090:A3756035B8761B2CFA89720BFA9F956D" box="[1299,1345,1297,1321]" country="Saint Helena, Ascension and Tristan da Cunha" municipality="Fisher's Valley" name="Ara." pageId="2" pageNumber="399">Ara.</location>
129425),
<collectingMunicipality id="4671AC94B8761B2CFF0D722FFEE79509" box="[151,313,1333,1357]" pageId="2" pageNumber="399">Fisher's Valley</collectingMunicipality>
,
<collectingCountry id="DEBD767EB8761B2CFED2722FFE069509" box="[328,472,1333,1357]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="2" pageNumber="399">Saint Helena</collectingCountry>
,
<quantity id="61529B0BB8761B2CFE7D722FFDE09509" box="[487,574,1333,1357]" metricMagnitude="2" metricUnit="m" metricValue="3.048" pageId="2" pageNumber="399" unit="ft" value="1000.0">
<elevation id="2D87D1DDB8761B2CFE7D722FFDE09509" box="[487,574,1333,1357]" metricMagnitude="2" metricUnit="m" metricValue="3.048" pageId="2" pageNumber="399" unit="ft" value="1000.0">1,000ft.</elevation>
</quantity>
,
<date id="D214102EB8761B2CFDD6722FFD0F9509" box="[588,721,1333,1357]" pageId="2" pageNumber="399" value="1965-12-19">
<collectingDate id="C250E9C6B8761B2CFDD6722FFD0F9509" box="[588,721,1333,1357]" pageId="2" pageNumber="399" value="1965-12-19">19.XII.1965</collectingDate>
</date>
, coll.
<collectorName id="0B5F5338B8761B2CFC80722FFC6A9509" box="[794,948,1333,1357]" pageId="2" pageNumber="399">P. Basilewsky</collectorName>
,
<collectorName id="0B5F5338B8761B2CFC59722FFBB19509" box="[963,1135,1333,1357]" pageId="2" pageNumber="399">P. L. G. Benoit</collectorName>
and
<collectorName id="0B5F5338B8761B2CFB36722FFAC09509" box="[1196,1310,1333,1357]" pageId="2" pageNumber="399">N. Leleup</collectorName>
, examined
</materialsCitation>
;
<materialsCitation id="16C23CB3B8761B2CFF0D7243FD3295D1" collectingDate="1965-12-14" collectingDateMax="1965-12-15" collectingDateMin="1965-12-14" collectionCode="BE, RMCA" collectorName="P. Basilewsky &amp; P. L. G. Benoit &amp; N. Leleup" country="Saint Helena, Ascension and Tristan da Cunha" elevation="305" location="Ara." municipality="Fisher's Valley" pageId="2" pageNumber="399" specimenCount="1" typeStatus="paratype">
<typeStatus id="7911884CB8761B2CFF0D7243FF2D9535" box="[151,243,1369,1393]" pageId="2" pageNumber="399">paratype</typeStatus>
imm. ♁ (
<collectionCode id="C0BBAE2BB8761B2CFECD7243FEA49535" box="[343,378,1369,1393]" pageId="2" pageNumber="399">BE</collectionCode>
_
<collectionCode id="C0BBAE2BB8761B2CFE1D7243FE049535" box="[391,474,1369,1393]" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="2" pageNumber="399" type="Museum">RMCA</collectionCode>
_ARA.
<location id="A3756035B8761B2CFDBD7243FD8B9535" LSID="urn:lsid:plazi:treatment:2E0387F8B8761B2DFF0D7409FF039090:A3756035B8761B2CFDBD7243FD8B9535" box="[551,597,1369,1393]" country="Saint Helena, Ascension and Tristan da Cunha" municipality="Fisher's Valley" name="Ara." pageId="2" pageNumber="399">Ara.</location>
135896),
<collectingMunicipality id="4671AC94B8761B2CFD2C7243FC8C9535" box="[694,850,1369,1393]" pageId="2" pageNumber="399">Fisher's Valley</collectingMunicipality>
,
<collectingCountry id="DEBD767EB8761B2CFCC67243FC389535" box="[860,998,1369,1393]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="2" pageNumber="399">Saint Helena</collectingCountry>
,
<quantity id="61529B0BB8761B2CFC6A7243FB989535" box="[1008,1094,1369,1393]" metricMagnitude="2" metricUnit="m" metricValue="3.048" pageId="2" pageNumber="399" unit="ft" value="1000.0">
<elevation id="2D87D1DDB8761B2CFC6A7243FB989535" box="[1008,1094,1369,1393]" metricMagnitude="2" metricUnit="m" metricValue="3.048" pageId="2" pageNumber="399" unit="ft" value="1000.0">1,000ft.</elevation>
</quantity>
,
<date id="D214102EB8761B2CFBD57243FB0C9535" box="[1103,1234,1369,1393]" pageId="2" pageNumber="399" value="1965-12-14">
<collectingDate id="C250E9C6B8761B2CFBD57243FB0C9535" box="[1103,1234,1369,1393]" pageId="2" pageNumber="399" value="1965-12-14">14.XII.1965</collectingDate>
</date>
<date id="D214102EB8761B2CFB787243FABD9535" box="[1250,1379,1369,1393]" pageId="2" pageNumber="399" value="1965-12-15">
<collectingDate id="C250E9C6B8761B2CFB787243FABD9535" box="[1250,1379,1369,1393]" pageId="2" pageNumber="399" value="1965-12-15">15.XII.1965</collectingDate>
</date>
, coll.
<collectorName id="0B5F5338B8761B2CFF0D7267FEF295D1" box="[151,300,1405,1429]" pageId="2" pageNumber="399">P. Basilewsky</collectorName>
,
<collectorName id="0B5F5338B8761B2CFEAD7267FE0B95D1" box="[311,469,1405,1429]" pageId="2" pageNumber="399">P. L. G. Benoit</collectorName>
and
<collectorName id="0B5F5338B8761B2CFD937267FDA995D1" box="[521,631,1405,1429]" pageId="2" pageNumber="399">N. Leleup</collectorName>
, examined
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EEB06565B8761B2CFF5D72BBFE659629" pageId="2" pageNumber="399" type="diagnosis">
<paragraph id="A61536EEB8761B2CFF5D72BBFE659629" blockId="2.[151,1437,1045,2041]" pageId="2" pageNumber="399">
<emphasis id="94DEEAFCB8761B2CFF5D72BBFEE295FD" bold="true" box="[199,316,1441,1465]" inLineHeading="true" pageId="2" pageNumber="399" reason="1">Diagnosis.</emphasis>
(Adapted from
<bibRefCitation id="C23B4B1FB8761B2CFE7D72BBFD5395FD" author="Lin, Y. C. &amp; Ballarin, F. &amp; Li, S. Q." box="[487,653,1441,1465]" pageId="2" pageNumber="399" pagination="1 - 168" refId="ref2627" refString="Lin, Y. C., Ballarin, F. &amp; Li, S. Q. (2016) A survey of the spider family Nesticidae (Arachnida, Araneae) in Asia and Madagascar, with the description of forty-three new species. ZooKeys, 627, 1 - 168. https: // doi. org / 10.3897 / zookeys. 627.8629" type="journal article" year="2016">
Lin
<emphasis id="94DEEAFCB8761B2CFD8972B8FD9795FD" box="[531,585,1441,1465]" italics="true" pageId="2" pageNumber="399">et al.</emphasis>
, 2016
</bibRefCitation>
)
<taxonomicName id="61AA4D6DB8761B2CFD0172B8FC9695FD" authorityName=": Marusik &amp; Guseinov" authorityYear="2003" box="[667,840,1442,1465]" class="Arachnida" family="Nesticidae" genus="Howaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="mogera">
<emphasis id="94DEEAFCB8761B2CFD0172B8FC9695FD" box="[667,840,1442,1465]" italics="true" pageId="2" pageNumber="399">Howaia mogera</emphasis>
</taxonomicName>
can be distinguished from most known male congeners based on the thin ventral apophysis, and blunted paracymbial process (
<emphasis id="94DEEAFCB8761B2CFC3572DCFC1B9599" box="[943,965,1478,1501]" italics="true" pageId="2" pageNumber="399">vs</emphasis>
. ventral apophysis wider and paracymbial process not blunted in most known congeners, except
<taxonomicName id="61AA4D6DB8761B2CFD5772F0FC5D9645" authorityName="Lin, Ballarin &amp; Li" authorityYear="2016" baseAuthorityName="Lin, Ballarin &amp; Li" baseAuthorityYear="2016" box="[717,899,1513,1537]" class="Arachnida" family="Nesticidae" genus="Howaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="rongtangensis">
<emphasis id="94DEEAFCB8761B2CFD5772F0FD3E9645" box="[717,736,1514,1537]" italics="true" pageId="2" pageNumber="399">H</emphasis>
.
<emphasis id="94DEEAFCB8761B2CFD7772F0FC5D9645" box="[749,899,1514,1537]" italics="true" pageId="2" pageNumber="399">rongtangensis</emphasis>
</taxonomicName>
). Females can be distinguished by the short, robust and conspicuously protruding scape (
<emphasis id="94DEEAFCB8761B2CFDAE7114FD949661" box="[564,586,1550,1573]" italics="true" pageId="2" pageNumber="399">vs</emphasis>
. scape not short, robust and conspicuously protruding in known congeners).
<taxonomicName id="61AA4D6DB8761B2CFF0D7128FE9B960D" authorityName=": Marusik &amp; Guseinov" authorityYear="2003" box="[151,325,1586,1609]" class="Arachnida" family="Nesticidae" genus="Howaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="mogera">
<emphasis id="94DEEAFCB8761B2CFF0D7128FE9B960D" box="[151,325,1586,1609]" italics="true" pageId="2" pageNumber="399">Howaia mogera</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="61AA4D6DB8761B2CFDA67128FD2A960D" authorityName="Lin, Ballarin &amp; Li" authorityYear="2016" baseAuthorityName="Lin, Ballarin &amp; Li" baseAuthorityYear="2016" box="[572,756,1586,1609]" class="Arachnida" family="Nesticidae" genus="Howaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="rongtangensis">
<emphasis id="94DEEAFCB8761B2CFDA67128FD2A960D" box="[572,756,1586,1609]" italics="true" pageId="2" pageNumber="399">H. rongtangensis</emphasis>
</taxonomicName>
in both sexes by the presence of eyes (
<emphasis id="94DEEAFCB8761B2CFB3C7128FB62960D" box="[1190,1212,1586,1609]" italics="true" pageId="2" pageNumber="399">vs</emphasis>
. eyes absent in both sexes of
<taxonomicName id="61AA4D6DB8761B2CFF6E714CFE759629" authorityName="Lin, Ballarin &amp; Li" authorityYear="2016" baseAuthorityName="Lin, Ballarin &amp; Li" baseAuthorityYear="2016" box="[244,427,1622,1645]" class="Arachnida" family="Nesticidae" genus="Howaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="rongtangensis">
<emphasis id="94DEEAFCB8761B2CFF6E714CFE759629" box="[244,427,1622,1645]" italics="true" pageId="2" pageNumber="399">H. rongtangensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="EEB06565B8761B2CFF5D7163FCA497BD" pageId="2" pageNumber="399" type="description">
<paragraph id="A61536EEB8761B2CFF5D7163FE179765" blockId="2.[151,1437,1045,2041]" pageId="2" pageNumber="399">
<emphasis id="94DEEAFCB8761B2CFF5D7163FE8C96D5" bold="true" box="[199,338,1657,1681]" inLineHeading="true" pageId="2" pageNumber="399" reason="1">Description.</emphasis>
For detailed description, see
<bibRefCitation id="C23B4B1FB8761B2CFD0E7163FC9596D5" author="Lin, Y. C. &amp; Ballarin, F. &amp; Li, S. Q." box="[660,843,1657,1681]" pageId="2" pageNumber="399" pagination="1 - 168" refId="ref2627" refString="Lin, Y. C., Ballarin, F. &amp; Li, S. Q. (2016) A survey of the spider family Nesticidae (Arachnida, Araneae) in Asia and Madagascar, with the description of forty-three new species. ZooKeys, 627, 1 - 168. https: // doi. org / 10.3897 / zookeys. 627.8629" type="journal article" year="2016">
Lin
<emphasis id="94DEEAFCB8761B2CFD5B7160FD2796D5" box="[705,761,1657,1681]" italics="true" pageId="2" pageNumber="399">et al.</emphasis>
(2016)
</bibRefCitation>
. However, it is important to state that all recent works illustrating and/or redescribing
<taxonomicName id="61AA4D6DB8761B2CFE717184FD8096F1" authorityName="Sherwood &amp; Jocqué &amp; Henrard &amp; Fowler" authorityYear="2023" baseAuthorityName="Sherwood &amp; Jocqué &amp; Henrard &amp; Fowler" baseAuthorityYear="1972" box="[491,606,1694,1717]" class="Arachnida" family="Nesticidae" genus="Howaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="mogera" status="n. comb.">
<emphasis id="94DEEAFCB8761B2CFE717184FD8096F1" box="[491,606,1694,1717]" italics="true" pageId="2" pageNumber="399">H. mogera</emphasis>
</taxonomicName>
<emphasis id="94DEEAFCB8761B2CFDFD7187FD1796F1" bold="true" box="[615,713,1693,1717]" pageId="2" pageNumber="399">
<taxonomicNameLabel id="8FED5787B8761B2CFDFD7187FD1796F1" box="[615,713,1693,1717]" pageId="2" pageNumber="399" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
have focused on non-Japanese material (e.g.,
<collectingCountry id="DEBD767EB8761B2CFB277187FAEC96F1" box="[1213,1330,1693,1717]" name="Azerbaijan" pageId="2" pageNumber="399">Azerbaijan</collectingCountry>
(Marusik &amp; Guisenov, 2003),
<collectingCountry id="DEBD767EB8761B2CFEEB71DBFE6F969D" box="[369,433,1729,1753]" name="China" pageId="2" pageNumber="399">China</collectingCountry>
(
<bibRefCitation id="C23B4B1FB8761B2CFE2571DBFD4F969D" author="Gong, M. X. &amp; Zhu, C. D." box="[447,657,1729,1753]" pageId="2" pageNumber="399" pagination="62" refId="ref2357" refString="Gong, M. X. &amp; Zhu, C. D. (1982) [Description of Nesticus mogera Yaginuma (Araneae: Nesticidae) from China]. Journal of the Bethune Medical University, 8, 62." type="journal article" year="1982">Gong &amp; Zhu, 1982</bibRefCitation>
;
<bibRefCitation id="C23B4B1FB8761B2CFD0671DBFC9C969D" author="Liu, J. &amp; Li, S. Q." box="[668,834,1729,1753]" pageId="2" pageNumber="399" pagination="501 - 547" refId="ref2693" refString="Liu, J. &amp; Li, S. Q. (2013) New cave-dwelling spiders of the family Nesticidae (Arachnida, Araneae) from China. Zootaxa, 3613 (6), 501 - 547. https: // doi. org / 10.11646 / zootaxa. 3613.6.1" type="journal article" year="2013">Liu &amp; Li, 2013</bibRefCitation>
),
<collectingCountry id="DEBD767EB8761B2CFCCF71DBFCA4969D" box="[853,890,1729,1753]" name="Fiji" pageId="2" pageNumber="399">Fiji</collectingCountry>
(
<bibRefCitation id="C23B4B1FB8761B2CFC1271DBFB77969D" author="Lehtinen, P. T. &amp; Saaristo, M. I." box="[904,1193,1729,1753]" pageId="2" pageNumber="399" pagination="47 - 66" refId="ref2591" refString="Lehtinen, P. T. &amp; Saaristo, M. I. (1980) Spiders of the Oriental-Australian region. II. Nesticidae. Annales Zoologici Fennici, 17, 47 - 66." type="journal article" year="1980">Lehtinen &amp; Saaristo, 1980</bibRefCitation>
), the Hawaiian Islands (
<bibRefCitation id="C23B4B1FB8761B2CFF0471FFFEE796B9" author="Gertsch, W. J." box="[158,313,1765,1789]" pageId="2" pageNumber="399" pagination="1 - 91" refId="ref2316" refString="Gertsch, W. J. (1984) The spider family Nesticidae (Araneae) in North America, Central America, and the West Indies. Bulletin of the Texas Memorial Museum, 31, 1 - 91." type="journal article" year="1984">Gertsch, 1984</bibRefCitation>
),
<collectingCountry id="DEBD767EB8761B2CFED171FFFEA296B9" box="[331,380,1765,1789]" name="Italy" pageId="2" pageNumber="399">Italy</collectingCountry>
(
<bibRefCitation id="C23B4B1FB8761B2CFE1071FFFD8796B9" author="Pantini, P. &amp; Bonelli, D. &amp; Bonacci, T." box="[394,601,1765,1789]" pageId="2" pageNumber="399" pagination="25 - 32" refId="ref2856" refString="Pantini, P., Bonelli, D. &amp; Bonacci, T. (2020) I ragni epigei (Arachnida, Araneae) di un ambiente litoraneo della Calabria. Rivista del Museo Civico di Scienze Naturali &quot; Enrico Caffi &quot;, Bergamo, 32, 25 - 32." type="journal article" year="2020">
Pantini
<emphasis id="94DEEAFCB8761B2CFE4471FCFDCD96B9" box="[478,531,1765,1789]" italics="true" pageId="2" pageNumber="399">et al.</emphasis>
, 2020
</bibRefCitation>
),
<collectingCountry id="DEBD767EB8761B2CFDF171FFFD7296B8" box="[619,684,1765,1788]" name="South Korea" pageId="2" pageNumber="399">Korea</collectingCountry>
(
<bibRefCitation id="C23B4B1FB8761B2CFD2071FFFCB296B9" author="Kim, J. P. &amp; Yoo, Y. S. &amp; Lee, Y. B." box="[698,876,1765,1789]" pageId="2" pageNumber="399" pagination="183 - 187" refId="ref2540" refString="Kim, J. P., Yoo, Y. S. &amp; Lee, Y. B. (1999) A newly recorded species of the genus Nesticella (Aranea, Nesticidae) in Korea. Korean Journal of Systematic Zoology, 15, 183 - 187." type="journal article" year="1999">
Kim
<emphasis id="94DEEAFCB8761B2CFD6871FCFCF996B9" box="[754,807,1765,1789]" italics="true" pageId="2" pageNumber="399">et al.</emphasis>
, 1999
</bibRefCitation>
;
<bibRefCitation id="C23B4B1FB8761B2CFCED71FFFBEA96B9" author="Kim, S. T. &amp; Lee, S. Y." box="[887,1076,1765,1789]" pageId="2" pageNumber="399" pagination="1 - 79" refId="ref2483" refString="Kim, S. T. &amp; Lee, S. Y. (2018) Spiders IV. Arthropoda: Arachnida: Araneae: Atypidae, Scytodidae, Nesticidae, Anapidae, Ctenidae, Hahniidae, Miturgidae, Liocranidae, Trochanteriidae. Invertebrate Fauna of Korea, 21 (45), 1 - 79." type="journal article" year="2018">Kim &amp; Lee, 2018</bibRefCitation>
) and
<collectingCountry id="DEBD767EB8761B2CFBF571FFFB2696B9" box="[1135,1272,1765,1789]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="2" pageNumber="399">Saint Helena</collectingCountry>
(
<bibRefCitation id="C23B4B1FB8761B2CFA9F71FFFA4B96B9" author="Hubert, M." box="[1285,1429,1765,1789]" pageId="2" pageNumber="399" pagination="153 - 156" refId="ref2435" refString="Hubert, M. (1977) Fam. Nesticidae. In: La faune terrestre de l'ile de Sainte-Helene IV. Annales, Musee Royal de l'Afrique Centrale, Sciences zoologiques (Zool. - Ser. 8 °), 220, 153 - 156." type="journal article" year="1977">Hubert, 1977</bibRefCitation>
) [the latter as
<taxonomicName id="61AA4D6DB8761B2CFEB87010FE6E9765" authorityName="Hubert" authorityYear="1977" box="[290,432,1801,1825]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="helenensis">
<emphasis id="94DEEAFCB8761B2CFEB87010FE6E9765" box="[290,432,1801,1825]" italics="true" pageId="2" pageNumber="399">N. helenensis</emphasis>
</taxonomicName>
]).
</paragraph>
<paragraph id="A61536EEB8761B2CFF5D7037FCA497BD" blockId="2.[151,1437,1045,2041]" pageId="2" pageNumber="399">
<emphasis id="94DEEAFCB8761B2CFF5D7037FE069701" bold="true" box="[199,472,1837,1861]" inLineHeading="true" pageId="2" pageNumber="399" reason="1">Rationale for synonymy.</emphasis>
The palp morphology of the
<typeStatus id="7911884CB8761B2CFC997037FC819701" box="[771,863,1837,1861]" pageId="2" pageNumber="399">paratype</typeStatus>
male of
<taxonomicName id="61AA4D6DB8761B2CFC2C7034FB5A9701" authorityName="Hubert" authorityYear="1977" box="[950,1156,1837,1861]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="helenensis">
<emphasis id="94DEEAFCB8761B2CFC2C7034FB5A9701" box="[950,1156,1837,1861]" italics="true" pageId="2" pageNumber="399">Nesticus helenensis</emphasis>
</taxonomicName>
(
<figureCitation id="3E912A6BB8761B2CFB0B7037FB2F9701" box="[1169,1265,1837,1861]" captionStart="FIGURES 16" captionStartId="1.[152,270,1903,1927]" captionTargetBox="[151,1436,260,1879]" captionTargetId="figure-50@1.[151,1436,260,1879]" captionTargetPageId="1" captionText="FIGURES 16. Holotype female (BE_RMCA_ARA.Ara.129084) and paratype male (BE_RMCA_ARA.Ara.133336) of Nesticus helenensis Hubert, 1977 [= Howaia mogera (Yaginuma, 1972) n. syn.]. 1, Male habitus, dorsal. 2, Male palp, ventral. 3, Female habitus, dorsal. 4, Epigynum, ventral. 5, Male palp, retrolateral (modified from Hubert, 1977). 6, Epigynum, ventral (modified from Hubert, 1977)." figureDoi="http://doi.org/10.5281/zenodo.8063108" httpUri="https://zenodo.org/record/8063108/files/figure.png" pageId="2" pageNumber="399">Figs. 2, 5</figureCitation>
) is in agreement with all prior published illustrations of the male palp of
<taxonomicName id="61AA4D6DB8761B2CFC977048FC62972D" authorityName="Sherwood &amp; Jocqué &amp; Henrard &amp; Fowler" authorityYear="2023" baseAuthorityName="Sherwood &amp; Jocqué &amp; Henrard &amp; Fowler" baseAuthorityYear="1972" box="[781,956,1874,1897]" class="Arachnida" family="Nesticidae" genus="Howaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="mogera">
<emphasis id="94DEEAFCB8761B2CFC977048FC62972D" box="[781,956,1874,1897]" italics="true" pageId="2" pageNumber="399">Howaia mogera</emphasis>
</taxonomicName>
. Similarly, the epigynal morphology of the
<typeStatus id="7911884CB8761B2CFF0D706FFF2A97C9" box="[151,244,1909,1933]" pageId="2" pageNumber="399">holotype</typeStatus>
(
<figureCitation id="3E912A6BB8761B2CFE9E706FFEB497C9" box="[260,362,1909,1933]" captionStart="FIGURES 16" captionStartId="1.[152,270,1903,1927]" captionTargetBox="[151,1436,260,1879]" captionTargetId="figure-50@1.[151,1436,260,1879]" captionTargetPageId="1" captionText="FIGURES 16. Holotype female (BE_RMCA_ARA.Ara.129084) and paratype male (BE_RMCA_ARA.Ara.133336) of Nesticus helenensis Hubert, 1977 [= Howaia mogera (Yaginuma, 1972) n. syn.]. 1, Male habitus, dorsal. 2, Male palp, ventral. 3, Female habitus, dorsal. 4, Epigynum, ventral. 5, Male palp, retrolateral (modified from Hubert, 1977). 6, Epigynum, ventral (modified from Hubert, 1977)." figureDoi="http://doi.org/10.5281/zenodo.8063108" httpUri="https://zenodo.org/record/8063108/files/figure.png" pageId="2" pageNumber="399">Figs. 4, 6</figureCitation>
) fits with previously published illustrations of
<taxonomicName id="61AA4D6DB8761B2CFCF9706CFC0B97C9" authorityName="Sherwood &amp; Jocqué &amp; Henrard &amp; Fowler" authorityYear="2023" baseAuthorityName="Sherwood &amp; Jocqué &amp; Henrard &amp; Fowler" baseAuthorityYear="1972" box="[867,981,1910,1933]" class="Arachnida" family="Nesticidae" genus="Howaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="mogera">
<emphasis id="94DEEAFCB8761B2CFCF9706CFC0B97C9" box="[867,981,1910,1933]" italics="true" pageId="2" pageNumber="399">H. mogera</emphasis>
</taxonomicName>
. The habitus (
<figureCitation id="3E912A6BB8761B2CFBEA706FFB0897C9" box="[1136,1238,1909,1933]" captionStart="FIGURES 16" captionStartId="1.[152,270,1903,1927]" captionTargetBox="[151,1436,260,1879]" captionTargetId="figure-50@1.[151,1436,260,1879]" captionTargetPageId="1" captionText="FIGURES 16. Holotype female (BE_RMCA_ARA.Ara.129084) and paratype male (BE_RMCA_ARA.Ara.133336) of Nesticus helenensis Hubert, 1977 [= Howaia mogera (Yaginuma, 1972) n. syn.]. 1, Male habitus, dorsal. 2, Male palp, ventral. 3, Female habitus, dorsal. 4, Epigynum, ventral. 5, Male palp, retrolateral (modified from Hubert, 1977). 6, Epigynum, ventral (modified from Hubert, 1977)." figureDoi="http://doi.org/10.5281/zenodo.8063108" httpUri="https://zenodo.org/record/8063108/files/figure.png" pageId="2" pageNumber="399">Figs. 1, 3</figureCitation>
) is also congruent with that of preserved specimens of
<taxonomicName id="61AA4D6DB8761B2CFDBA7080FD4D97F5" authorityName="Sherwood &amp; Jocqué &amp; Henrard &amp; Fowler" authorityYear="2023" baseAuthorityName="Sherwood &amp; Jocqué &amp; Henrard &amp; Fowler" baseAuthorityYear="1972" box="[544,659,1946,1969]" class="Arachnida" family="Nesticidae" genus="Howaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="mogera">
<emphasis id="94DEEAFCB8761B2CFDBA7080FD4D97F5" box="[544,659,1946,1969]" italics="true" pageId="2" pageNumber="399">H. mogera</emphasis>
</taxonomicName>
(DS pers. obs.). The potential for synonymy was already recognised by
<bibRefCitation id="C23B4B1FB8761B2CFF0D70A7FE169791" author="Lehtinen, P. T. &amp; Saaristo, M. I." box="[151,456,1981,2005]" pageId="2" pageNumber="399" pagination="47 - 66" refId="ref2591" refString="Lehtinen, P. T. &amp; Saaristo, M. I. (1980) Spiders of the Oriental-Australian region. II. Nesticidae. Annales Zoologici Fennici, 17, 47 - 66." type="journal article" year="1980">Lehtinen &amp; Saaristo (1980)</bibRefCitation>
, but no formal synonymy was made. We hereby formally make the synonymy:
<taxonomicName id="61AA4D6DB8761B2CFAD870A4FE4797BD" authority="Hubert, 1977" authorityName="Hubert" authorityYear="1977" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="helenensis">
<emphasis id="94DEEAFCB8761B2CFAD870A4FED897BD" italics="true" pageId="2" pageNumber="399">Nesticus helenensis</emphasis>
<bibRefCitation id="C23B4B1FB8761B2CFE9770FBFE4797BD" author="Hubert, M." box="[269,409,2017,2041]" pageId="2" pageNumber="399" pagination="153 - 156" refId="ref2435" refString="Hubert, M. (1977) Fam. Nesticidae. In: La faune terrestre de l'ile de Sainte-Helene IV. Annales, Musee Royal de l'Afrique Centrale, Sciences zoologiques (Zool. - Ser. 8 °), 220, 153 - 156." type="journal article" year="1977">Hubert, 1977</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="61AA4D6DB8761B2CFE2C70F8FCF597BD" authority="(Yaginuma, 1972)" baseAuthorityName="Yaginuma" baseAuthorityYear="1972" box="[438,811,2017,2041]" class="Arachnida" family="Nesticidae" genus="Howaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="mogera" status="n. syn.">
<emphasis id="94DEEAFCB8761B2CFE2C70F8FDBC97BD" box="[438,610,2018,2041]" italics="true" pageId="2" pageNumber="399">Howaia mogera</emphasis>
(
<bibRefCitation id="C23B4B1FB8761B2CFDEA70FBFCFD97BD" author="Yaginuma, T." box="[624,803,2017,2041]" pageId="2" pageNumber="399" pagination="619 - 622" refId="ref3090" refString="Yaginuma, T. (1972) Revision of the short-legged nesticid spiders of Japan. Bulletin of the National Museum of Nature and Science Tokyo, 15, 619 - 622." type="journal article" year="1972">Yaginuma, 1972</bibRefCitation>
)
</taxonomicName>
<emphasis id="94DEEAFCB8761B2CFCAB70FBFCA497BD" bold="true" box="[817,890,2017,2041]" pageId="2" pageNumber="399">
<taxonomicNameLabel id="8FED5787B8761B2CFCAB70FBFCA497BD" box="[817,890,2017,2041]" pageId="2" pageNumber="399">n. syn.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="EEB06565B8771B2DFF5D778DFF039090" pageId="3" pageNumber="400" type="distribution">
<paragraph id="A61536EEB8771B2DFF5D778DFF039090" blockId="3.[151,1436,151,212]" pageId="3" pageNumber="400">
<emphasis id="94DEEAFCB8771B2DFF5D778DFE8490EB" bold="true" box="[199,346,151,175]" inLineHeading="true" pageId="3" pageNumber="400" reason="1">Distribution.</emphasis>
<collectingCountry id="DEBD767EB8771B2DFEFE778DFDC390EB" box="[356,541,151,175]" name="United Kingdom" pageId="3" pageNumber="400">United Kingdom</collectingCountry>
to
<collectingCountry id="DEBD767EB8771B2DFDD37782FD5690EB" box="[585,648,152,175]" name="Japan" pageId="3" pageNumber="400">Japan</collectingCountry>
, introduced to Hawaii and
<collectingCountry id="DEBD767EB8771B2DFC25778DFB9390EB" box="[959,1101,151,175]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="3" pageNumber="400">Saint Helena</collectingCountry>
(
<bibRefCitation id="C23B4B1FB8771B2DFBC5778DFA9390F4" author="Nentwig, W. &amp; Blick, T. &amp; Bosmans, R. &amp; Gloor, D. &amp; Hanggi, A. &amp; Kropf, C." box="[1119,1357,151,176]" pageId="3" pageNumber="400" refId="ref2788" refString="Nentwig, W., Blick, T., Bosmans, R., Gloor, D., Hanggi, A. &amp; Kropf, C. (2023) Spiders of Europe. Version 05.2023. Available from: https: // www. araneae. nmbe. ch (accessed 7 May 2023) https: // doi. org / 10.24436 / 1" type="url" year="2023">
Nentwig
<emphasis id="94DEEAFCB8771B2DFB527782FADD90EB" box="[1224,1283,151,175]" italics="true" pageId="3" pageNumber="400">et al.</emphasis>
, 2023
</bibRefCitation>
;
<bibRefCitation id="C23B4B1FB8771B2DFAC67782FA4290F4" author="WSC" box="[1372,1436,152,176]" pageId="3" pageNumber="400" refId="ref2908" refString="WSC (2023) World Spider Catalog. Version 24. Natural History Museum Bern, Bern. Available from: http: // wsc. nmbe. ch (accessed 27 March 2023) https: // doi. org / 10.24436 / 2" type="url" year="2023">WSC,</bibRefCitation>
2023).
</paragraph>
</subSubSection>
</treatment>
</document>