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<paragraph id="A31536D9FFDB2E3BFF46FE71FD2485E9" blockId="11.[226,688,447,473]" box="[226,688,447,473]" pageId="11" pageNumber="12">
<heading id="F85D81B5FFDB2E3BFF46FE71FD2485E9" box="[226,688,447,473]" centered="true" level="2" pageId="11" pageNumber="12" reason="4">
Genus
<taxonomicName id="64AA4D5AFFDB2E3BFE91FE71FDDF85E9" ID-CoL="5TJF6" authority="Peters, 1871" authorityName="Peters" authorityYear="1871" box="[309,587,447,473]" class="Mammalia" family="Hipposideridae" genus="Doryrhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="91DEEACBFFDB2E3BFE91FE71FE2485E9" box="[309,432,447,473]" italics="true" pageId="11" pageNumber="12">Doryrhina</emphasis>
<bibRefCitation id="C73B4B28FFDB2E3BFE1DFE71FDDF85E9" author="PETERS, W." box="[441,587,447,473]" journalOrPublisher="Monatsberichte der Koniglich Preussischen Akademie der Wissenschaften" pageId="11" pageNumber="12" pagination="301 - 332" part="1871" refId="ref14019" refString="PETERS, W. 1871. Uber die Gattungen und Arten der Hufeisennase, Rhinolophi. Monatsberichte der Koniglich Preussischen Akademie der Wissenschaften, 1871: 301 - 332." title="Uber die Gattungen und Arten der Hufeisennase, Rhinolophi" type="journal article" year="1871">Peters, 1871</bibRefCitation>
</taxonomicName>
(p. 314)
</heading>
</paragraph>
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<subSubSection id="EBB06552FFDB2E3BFF65FDCDFF77860E" pageId="11" pageNumber="12" type="multiple">
<paragraph id="A31536D9FFDB2E3BFF65FDCDFF77860E" blockId="11.[151,763,515,574]" pageId="11" pageNumber="12">
New combination —
<taxonomicName id="64AA4D5AFFDB2E3BFE64FDCDFF4B860E" authority="(Peters, 1871)" baseAuthorityName="Peters" baseAuthorityYear="1871" class="Mammalia" family="Hipposideridae" genus="Doryrhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cyclops">
<emphasis id="91DEEACBFFDB2E3BFE64FDCDFD0F862D" box="[448,667,515,541]" italics="true" pageId="11" pageNumber="12">Doryrhina cyclops</emphasis>
(
<bibRefCitation id="C73B4B28FFDB2E3BFD0FFDCDFF42860E" author="PETERS, W." journalOrPublisher="Monatsberichte der Koniglich Preussischen Akademie der Wissenschaften" pageId="11" pageNumber="12" pagination="301 - 332" part="1871" refId="ref14019" refString="PETERS, W. 1871. Uber die Gattungen und Arten der Hufeisennase, Rhinolophi. Monatsberichte der Koniglich Preussischen Akademie der Wissenschaften, 1871: 301 - 332." title="Uber die Gattungen und Arten der Hufeisennase, Rhinolophi" type="journal article" year="1871">Peters, 1871</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="EBB06552FFDB2E3BFF33FDA6FE8786B2" box="[151,275,616,642]" pageId="11" pageNumber="12" type="multiple">
<paragraph id="A31536D9FFDB2E3BFF33FDA6FE8786B2" blockId="11.[151,763,616,810]" box="[151,275,616,642]" pageId="11" pageNumber="12">Synonyms</paragraph>
</subSubSection>
<subSubSection id="EBB06552FFDB2E3BFF33FD46FD94871A" pageId="11" pageNumber="12" type="reference_group">
<paragraph id="A31536D9FFDB2E3BFF33FD46FD8586AD" blockId="11.[151,763,616,810]" box="[151,529,648,669]" pageId="11" pageNumber="12">
<taxonomicName id="64AA4D5AFFDB2E3BFF33FD46FD9986AD" authority="Temminck, 1853" authorityName="Temminck" authorityYear="1853" box="[151,525,648,669]" class="Mammalia" family="Hipposideridae" genus="Phyllorrhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cyclops">
<emphasis id="91DEEACBFFDB2E3BFF33FD46FEF586AD" box="[151,353,648,669]" italics="true" pageId="11" pageNumber="12">Phyllorrhina cyclops</emphasis>
Temminck, 1853
</taxonomicName>
.
</paragraph>
<paragraph id="A31536D9FFDB2E3BFF33FD6AFDBA86E5" blockId="11.[151,763,616,810]" pageId="11" pageNumber="12">
<taxonomicName id="64AA4D5AFFDB2E3BFF33FD6AFD888689" authority="Temminck, 1853" authorityName="Temminck" authorityYear="1853" box="[151,540,676,697]" class="Mammalia" family="Hipposideridae" genus="Phyllorrhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cyclops">
<emphasis id="91DEEACBFFDB2E3BFF33FD6AFEF38689" box="[151,359,676,697]" italics="true" pageId="11" pageNumber="12">Phyllorrhina cyclops</emphasis>
Temminck, 1853
</taxonomicName>
=
<taxonomicName id="64AA4D5AFFDB2E3BFDE6FD6AFEE286E5" ID-CoL="5TJF9" authority="(Temminck, 1853)" baseAuthorityName="Temminck" baseAuthorityYear="1853" class="Mammalia" family="Hipposideridae" genus="Doryrhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cyclops">
<emphasis id="91DEEACBFFDB2E3BFDE6FD6AFD6F8689" box="[578,763,676,697]" italics="true" pageId="11" pageNumber="12">Doryrhina cyclops</emphasis>
(Temminck, 1853)
</taxonomicName>
, see
<bibRefCitation id="C73B4B28FFDB2E3BFE02FD0EFDBD86E5" author="PETERS, W." box="[422,553,704,725]" journalOrPublisher="Monatsberichte der Koniglich Preussischen Akademie der Wissenschaften" pageId="11" pageNumber="12" pagination="301 - 332" part="1871" refId="ref14019" refString="PETERS, W. 1871. Uber die Gattungen und Arten der Hufeisennase, Rhinolophi. Monatsberichte der Koniglich Preussischen Akademie der Wissenschaften, 1871: 301 - 332." title="Uber die Gattungen und Arten der Hufeisennase, Rhinolophi" type="journal article" year="1871">Peters (1871)</bibRefCitation>
.
</paragraph>
<paragraph id="A31536D9FFDB2E3BFF33FD13FD8A86C2" blockId="11.[151,763,616,810]" box="[151,542,733,754]" pageId="11" pageNumber="12">
<taxonomicName id="64AA4D5AFFDB2E3BFF33FD13FD8E86C2" authority="de Winton, 1897" authorityName="de Winton" authorityYear="1897" box="[151,538,733,754]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="micaceus">
<emphasis id="91DEEACBFFDB2E3BFF33FD13FEE486C2" box="[151,368,733,754]" italics="true" pageId="11" pageNumber="12">Rhinolophus micaceus</emphasis>
de Winton, 1897
</taxonomicName>
.
</paragraph>
<paragraph id="A31536D9FFDB2E3BFF33FD37FD80873E" blockId="11.[151,763,616,810]" box="[151,532,761,782]" pageId="11" pageNumber="12">
<taxonomicName id="64AA4D5AFFDB2E3BFF33FD37FD84873E" ID-CoL="3M36N" authority="de Winton, 1899" authorityName="de Winton" authorityYear="1899" box="[151,528,761,782]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cyclops">
<emphasis id="91DEEACBFFDB2E3BFF33FD37FEF2873E" box="[151,358,761,782]" italics="true" pageId="11" pageNumber="12">Hipposideros cyclops</emphasis>
de Winton, 1899
</taxonomicName>
.
</paragraph>
<paragraph id="A31536D9FFDB2E3BFF33FCDBFD94871A" blockId="11.[151,763,616,810]" box="[151,512,789,810]" pageId="11" pageNumber="12">
<taxonomicName id="64AA4D5AFFDB2E3BFF33FCDBFE68871A" authority="J. A. Allen, 1917" authorityName="J. A. Allen" authorityYear="1917" box="[151,508,789,810]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="langi">
<emphasis id="91DEEACBFFDB2E3BFF33FCDBFEDB871A" box="[151,335,789,810]" italics="true" pageId="11" pageNumber="12">Hipposideros langi</emphasis>
J. A. Allen, 1917
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="EBB06552FFDB2E3BFE9BFCA2FDC087B6" box="[319,596,876,902]" pageId="11" pageNumber="12" type="multiple">
<paragraph id="A31536D9FFDB2E3BFE9BFCA2FDC087B6" blockId="11.[319,596,876,902]" box="[319,596,876,902]" pageId="11" pageNumber="12">
<taxonomicName id="64AA4D5AFFDB2E3BFE9BFCA2FDC087B6" authority="Peters, 1871" authorityName="Peters" authorityYear="1871" box="[319,596,876,902]" class="Mammalia" family="Hipposideridae" genus="Doryrhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="91DEEACBFFDB2E3BFE9BFCA2FE2E87B6" box="[319,442,876,902]" italics="true" pageId="11" pageNumber="12">Doryrhina</emphasis>
<bibRefCitation id="C73B4B28FFDB2E3BFE66FCA2FDC087B6" author="PETERS, W." box="[450,596,876,902]" journalOrPublisher="Monatsberichte der Koniglich Preussischen Akademie der Wissenschaften" pageId="11" pageNumber="12" pagination="301 - 332" part="1871" refId="ref14019" refString="PETERS, W. 1871. Uber die Gattungen und Arten der Hufeisennase, Rhinolophi. Monatsberichte der Koniglich Preussischen Akademie der Wissenschaften, 1871: 301 - 332." title="Uber die Gattungen und Arten der Hufeisennase, Rhinolophi" type="journal article" year="1871">Peters, 1871</bibRefCitation>
</taxonomicName>
</paragraph>
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<subSubSection id="EBB06552FFDB2E3CFF33FC61FDA58120" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="description">
<paragraph id="A31536D9FFDB2E3BFF33FC61FDD587F9" blockId="11.[151,577,943,969]" box="[151,577,943,969]" pageId="11" pageNumber="12">
<emphasis id="91DEEACBFFDB2E3BFF33FC61FE5487F9" box="[151,448,943,969]" italics="true" pageId="11" pageNumber="12">Description of the Genus</emphasis>
<taxonomicName id="64AA4D5AFFDB2E3BFE6CFC61FDD587F9" authorityName="Peters" authorityYear="1871" box="[456,577,943,969]" class="Mammalia" family="Hipposideridae" genus="Doryrhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">Doryrhina</taxonomicName>
</paragraph>
<paragraph id="A31536D9FFDB2E3BFF33FC23FE588037" blockId="11.[151,764,1005,1973]" box="[151,460,1005,1031]" pageId="11" pageNumber="12">
<emphasis id="91DEEACBFFDB2E3BFF33FC23FE588037" box="[151,460,1005,1031]" italics="true" pageId="11" pageNumber="12">Morphological characters</emphasis>
</paragraph>
<paragraph id="A31536D9FFDB2E3BFF65FBC1FD1480C2" blockId="11.[151,764,1005,1973]" pageId="11" pageNumber="12">
<bibRefCitation id="C73B4B28FFDB2E3BFF65FBC1FEF28019" author="PETERS, W." box="[193,358,1039,1065]" journalOrPublisher="Monatsberichte der Koniglich Preussischen Akademie der Wissenschaften" pageId="11" pageNumber="12" pagination="301 - 332" part="1871" refId="ref14019" refString="PETERS, W. 1871. Uber die Gattungen und Arten der Hufeisennase, Rhinolophi. Monatsberichte der Koniglich Preussischen Akademie der Wissenschaften, 1871: 301 - 332." title="Uber die Gattungen und Arten der Hufeisennase, Rhinolophi" type="journal article" year="1871">Peters (1871)</bibRefCitation>
in his naming of the subgenus
<taxonomicName id="64AA4D5AFFDB2E3BFF33FBFEFE86807A" authorityName="Peters" authorityYear="1871" box="[151,274,1072,1098]" class="Mammalia" family="Hipposideridae" genus="Doryrhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="91DEEACBFFDB2E3BFF33FBFEFE86807A" box="[151,274,1072,1098]" italics="true" pageId="11" pageNumber="12">Doryrhina</emphasis>
</taxonomicName>
presented different characters associated with the noseleaf structure, and the
<typeStatus id="7C11887BFFDB2E3BFDE5FB9CFDE0805C" box="[577,628,1106,1132]" pageId="11" pageNumber="12">type</typeStatus>
species for the subgenus was
<taxonomicName id="64AA4D5AFFDB2E3BFED4FBBAFD6780BE" baseAuthorityName="Hill" baseAuthorityYear="1963" box="[368,755,1140,1166]" class="Mammalia" family="Hipposideridae" genus="Phyllorhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cyclops" subGenus="Doryrhina">
<emphasis id="91DEEACBFFDB2E3BFED4FBBAFE6880BE" box="[368,508,1140,1166]" italics="true" pageId="11" pageNumber="12">Phyllorhina</emphasis>
(
<emphasis id="91DEEACBFFDB2E3BFDAAFBBAFD1D80BE" box="[526,649,1140,1166]" italics="true" pageId="11" pageNumber="12">Doryrhina</emphasis>
)
<emphasis id="91DEEACBFFDB2E3BFD3FFBBAFD6780BE" box="[667,755,1140,1166]" italics="true" pageId="11" pageNumber="12">cyclops</emphasis>
</taxonomicName>
; a tropical African taxon (
<bibRefCitation id="C73B4B28FFDB2E3BFE61FB5BFDD8809F" author="FAHR, J." bookContentInfo="800 pp." box="[453,588,1173,1199]" editor="M. HAPPOLD &amp; D. C. D. HAPPOLD" journalOrPublisher="Bloomsbury Publishing, London" pageId="11" pageNumber="12" pagination="380 - 383" refId="ref12366" refString="FAHR, J. 2013. Hipposideros cyclops cyclops leaf-nosed bat. Pp. 380 - 383, in Mammals of Africa. Volume IV (M. HAPPOLD and D. C. D. HAPPOLD, eds.). Bloomsbury Publishing, London, 800 pp." title="Hipposideros cyclops cyclops leaf-nosed bat." type="book chapter" volumeTitle="Mammals of Africa. Volume IV" year="2013">Fahr, 2013</bibRefCitation>
). Here we elevate the subgenus
<taxonomicName id="64AA4D5AFFDB2E3BFEDEFB79FE6180E1" authorityName="Peters" authorityYear="1871" box="[378,501,1207,1233]" class="Mammalia" family="Hipposideridae" genus="Doryrhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="91DEEACBFFDB2E3BFEDEFB79FE6180E1" box="[378,501,1207,1233]" italics="true" pageId="11" pageNumber="12">Doryrhina</emphasis>
</taxonomicName>
to the level of genus for
<taxonomicName id="64AA4D5AFFDB2E3BFF65FB16FE8D80C2" baseAuthorityName="Hill" baseAuthorityYear="1963" box="[193,281,1240,1266]" class="Mammalia" family="Hipposideridae" genus="Phyllorhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cyclops">
<emphasis id="91DEEACBFFDB2E3BFF65FB16FE8D80C2" box="[193,281,1240,1266]" italics="true" pageId="11" pageNumber="12">cyclops</emphasis>
</taxonomicName>
and expand Peters diagnosis.
</paragraph>
<paragraph id="A31536D9FFDB2E3BFF65FB34FBC681A5" blockId="11.[151,764,1005,1973]" lastBlockId="11.[809,1422,177,1973]" pageId="11" pageNumber="12">
The single species we place herein in the genus
<taxonomicName id="64AA4D5AFFDB2E3BFF33FAD2FE868106" authorityName="Peters" authorityYear="1871" box="[151,274,1308,1334]" class="Mammalia" family="Hipposideridae" genus="Doryrhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="91DEEACBFFDB2E3BFF33FAD2FE868106" box="[151,274,1308,1334]" italics="true" pageId="11" pageNumber="12">Doryrhina</emphasis>
</taxonomicName>
,
<taxonomicName id="64AA4D5AFFDB2E3BFE85FAD2FE098106" baseAuthorityName="Hill" baseAuthorityYear="1963" box="[289,413,1308,1334]" class="Mammalia" family="Hipposideridae" genus="Phyllorhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cyclops">
<emphasis id="91DEEACBFFDB2E3BFE85FAD2FE098106" box="[289,413,1308,1334]" italics="true" pageId="11" pageNumber="12">D. cyclops</emphasis>
</taxonomicName>
, has a frontal sac on the forehead (
<bibRefCitation id="C73B4B28FFDB2E3BFF45FAF3FEF48167" author="HILL, J. E." box="[225,352,1341,1367]" journalOrPublisher="Bulletin of the British Museum (Natural History), Zoology" pageId="11" pageNumber="12" pagination="1 - 129" part="11" refId="ref13158" refString="HILL, J. E. 1963. A revision of the genus Hipposideros. Bulletin of the British Museum (Natural History), Zoology, 11: 1 - 129." title="A revision of the genus Hipposideros" type="journal article" year="1963">Hill, 1963</bibRefCitation>
;
<bibRefCitation id="C73B4B28FFDB2E3BFECBFAF3FE608167" author="FAHR, J." bookContentInfo="800 pp." box="[367,500,1341,1367]" editor="M. HAPPOLD &amp; D. C. D. HAPPOLD" journalOrPublisher="Bloomsbury Publishing, London" pageId="11" pageNumber="12" pagination="380 - 383" refId="ref12366" refString="FAHR, J. 2013. Hipposideros cyclops cyclops leaf-nosed bat. Pp. 380 - 383, in Mammals of Africa. Volume IV (M. HAPPOLD and D. C. D. HAPPOLD, eds.). Bloomsbury Publishing, London, 800 pp." title="Hipposideros cyclops cyclops leaf-nosed bat." type="book chapter" volumeTitle="Mammals of Africa. Volume IV" year="2013">Fahr, 2013</bibRefCitation>
), in a central position behind the posterior noseleaf, and opening as a relatively small vertical slit (
<figureCitation id="3B912A5CFFDB2E3BFE67FA4FFD9A81AB" box="[451,526,1409,1435]" captionStart="FIG" captionStartId="9.[151,164,1783,1804]" captionTargetBox="[163,1411,1288,1771]" captionTargetId="figure-14@9.[765,1091,1288,1736]" captionTargetPageId="9" captionText="FIG. 4. Morphological comparisons of the frontal sacs and noseleaves in A — Macronycteris commersonii (FMNH 213588, ♀), Madagascar, Province dAntsiranana, Réserve Spéciale dAnkarana [now Parc National], near Andrafiabe Cave, 31 October 2010; B — M. gigas (FMNH 128212, ♀), Senegal, Casmance, Diabane, 12 km SW of Adeane, 15 January 1983; C — M. vittatus (FMNH 192800, ♀, sequenced for Cyt-b), Tanzania, Pemba Island, Kaskazini Region, Micheweni District, Kilijini Village, 3 August 2006; and D — Doryrhina cyclops (FMNH 164973), Uganda, Masindi District, Budongo Forest, 25 June 1998. The form of M. cryptovalorona is similar to M. commersonii and no comparative specimen material was available for M. thomensis. Drawing by Velizar Simeonovski" figureDoi="http://doi.org/10.5281/zenodo.6822116" httpUri="https://zenodo.org/record/6822116/files/figure.png" pageId="11" pageNumber="12">Fig. 4</figureCitation>
). Within the slit are stiff white hairs that when everted form a distinct hair tuft (
<bibRefCitation id="C73B4B28FFDB2E3BFEB2FA0AFE4E81EE" author="ROSEVEAR, D. R." bookContentInfo="418 pp." box="[278,474,1476,1502]" journalOrPublisher="Trustees of the British Museum (Natural History), London" pageId="11" pageNumber="12" refId="ref14852" refString="ROSEVEAR, D. R. 1965. The bats of West Africa. Trustees of the British Museum (Natural History), London, 418 pp." title="The bats of West Africa" type="book" year="1965">Rosevear, 1965</bibRefCitation>
). The noseleaf of
<taxonomicName id="64AA4D5AFFDB2E3BFD6AFA0AFF7A8230" authorityName="Peters" authorityYear="1871" class="Mammalia" family="Hipposideridae" genus="Doryrhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="91DEEACBFFDB2E3BFD6AFA0AFF7A8230" italics="true" pageId="11" pageNumber="12">Doryrhina</emphasis>
</taxonomicName>
shows several particular aspects which differ from other hipposiderids, including
<taxonomicName id="64AA4D5AFFDB2E3BFDD8F9C9FF5F8273" authorityName="Gray" authorityYear="1866" class="Mammalia" family="Hipposideridae" genus="Macronycteris" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="91DEEACBFFDB2E3BFDD8F9C9FF5F8273" italics="true" pageId="11" pageNumber="12">Macronycteris</emphasis>
</taxonomicName>
. Members of the genus
<taxonomicName id="64AA4D5AFFDB2E3BFDA7F9E7FDEA8273" authorityName="Peters" authorityYear="1871" box="[515,638,1577,1603]" class="Mammalia" family="Hipposideridae" genus="Doryrhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="91DEEACBFFDB2E3BFDA7F9E7FDEA8273" box="[515,638,1577,1603]" italics="true" pageId="11" pageNumber="12">Doryrhina</emphasis>
</taxonomicName>
have two lateral fleshy leaflets (
<figureCitation id="3B912A5CFFDB2E3BFE3AF985FE738255" box="[414,487,1611,1637]" captionStart="FIG" captionStartId="9.[151,164,1783,1804]" captionTargetBox="[163,1411,1288,1771]" captionTargetId="figure-14@9.[765,1091,1288,1736]" captionTargetPageId="9" captionText="FIG. 4. Morphological comparisons of the frontal sacs and noseleaves in A — Macronycteris commersonii (FMNH 213588, ♀), Madagascar, Province dAntsiranana, Réserve Spéciale dAnkarana [now Parc National], near Andrafiabe Cave, 31 October 2010; B — M. gigas (FMNH 128212, ♀), Senegal, Casmance, Diabane, 12 km SW of Adeane, 15 January 1983; C — M. vittatus (FMNH 192800, ♀, sequenced for Cyt-b), Tanzania, Pemba Island, Kaskazini Region, Micheweni District, Kilijini Village, 3 August 2006; and D — Doryrhina cyclops (FMNH 164973), Uganda, Masindi District, Budongo Forest, 25 June 1998. The form of M. cryptovalorona is similar to M. commersonii and no comparative specimen material was available for M. thomensis. Drawing by Velizar Simeonovski" figureDoi="http://doi.org/10.5281/zenodo.6822116" httpUri="https://zenodo.org/record/6822116/files/figure.png" pageId="11" pageNumber="12">Fig. 4</figureCitation>
), a common configuration in
<taxonomicName id="64AA4D5AFFDB2E3BFF54F9A2FE1982B6" box="[240,397,1644,1670]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="91DEEACBFFDB2E3BFF54F9A2FE1982B6" box="[240,397,1644,1670]" italics="true" pageId="11" pageNumber="12">Hipposideros</emphasis>
</taxonomicName>
, although several members of this genus, as well as
<taxonomicName id="64AA4D5AFFDB2E3BFE66F940FDF88298" authorityName="Gray" authorityYear="1866" box="[450,620,1678,1704]" class="Mammalia" family="Hipposideridae" genus="Macronycteris" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="91DEEACBFFDB2E3BFE66F940FDF88298" box="[450,620,1678,1704]" italics="true" pageId="11" pageNumber="12">Macronycteris</emphasis>
</taxonomicName>
, have four leaflets (
<bibRefCitation id="C73B4B28FFDB2E3BFF5AF97EFE2A82FA" author="ANDERSEN, K." box="[254,446,1712,1738]" journalOrPublisher="Annals and Magazine of Natural History" pageId="11" pageNumber="12" pagination="497 - 507" part="7" refId="ref11715" refString="ANDERSEN, K. 1905. On Hipposideros diadema and its closest allies. Annals and Magazine of Natural History, 7, 16: 497 - 507." title="On Hipposideros diadema and its closest allies" type="journal article" year="1905">Andersen, 1905</bibRefCitation>
,
<bibRefCitation id="C73B4B28FFDB2E3BFE69F97EFD9F82FA" author="ANDERSEN, K." box="[461,523,1712,1738]" journalOrPublisher="Annals and Magazine of Natural History" pageId="11" pageNumber="12" pagination="35 - 48" part="7" refId="ref11744" refString="ANDERSEN, K. 1906. On the bats of the Hipposideros armiger and Commersoni types. Annals and Magazine of Natural History, 7, 17: 35 - 48." title="On the bats of the Hipposideros armiger and Commersoni types" type="journal article" year="1906">1906</bibRefCitation>
;
<bibRefCitation id="C73B4B28FFDB2E3BFDBDF97EFD6382FA" author="PAYNE, J. &amp; C. M. FRANCIS &amp; K. PHILIPPS" bookContentInfo="332 pp." box="[537,759,1712,1738]" journalOrPublisher="The Sabah Society, Kota Kinabalu" pageId="11" pageNumber="12" refId="ref13982" refString="PAYNE, J., C. M. FRANCIS, and K. PHILIPPS. 1985. A field guide to the mammals of Borneo. The Sabah Society, Kota Kinabalu, 332 pp." title="A field guide to the mammals of Borneo" type="book" year="1985">
Payne
<emphasis id="91DEEACBFFDB2E3BFDCFF97EFD3482FA" box="[619,672,1712,1738]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
., 1985
</bibRefCitation>
;
<bibRefCitation id="C73B4B28FFDB2E3BFF33F91FFED682DB" author="STRAHAN, R." bookContentInfo="756 pp." box="[151,322,1745,1771]" journalOrPublisher="Smithsonian Institution Press, Washington, D. C" pageId="11" pageNumber="12" refId="ref15176" refString="STRAHAN, R. 1995. Mammals of Australia. Smithsonian Institution Press, Washington, D. C., 756 pp." title="Mammals of Australia" type="book" year="1995">Strahan, 1995</bibRefCitation>
;
<bibRefCitation id="C73B4B28FFDB2E3BFEF6F91FFD8182DB" author="FLANNERY, T." bookContentInfo="568 pp." box="[338,533,1745,1771]" journalOrPublisher="Cornell University Press, Ithaca" pageId="11" pageNumber="12" refId="ref12416" refString="FLANNERY, T. 1995 a. Mammals of New Guinea. Cornell University Press, Ithaca, 568 pp." title="Mammals of New Guinea" type="book" year="1995">
Flannery, 1995
<emphasis id="91DEEACBFFDB2E3BFDA2F91FFD8182DB" box="[518,533,1745,1771]" italics="true" pageId="11" pageNumber="12">a</emphasis>
</bibRefCitation>
,
<bibRefCitation id="C73B4B28FFDB2E3BFD83F91FFDE582DB" author="FLANNERY, T." bookContentInfo="464 pp." box="[551,625,1745,1771]" journalOrPublisher="Cornell University Press, Ithaca" pageId="11" pageNumber="12" refId="ref12437" refString="FLANNERY, T. 1995 b. Mammals of the south-west Pacific and Moluccan islands. Cornell University Press, Ithaca, 464 pp." title="Mammals of the south-west Pacific and Moluccan islands" type="book" year="1995">
1995
<emphasis id="91DEEACBFFDB2E3BFDC6F91FFDE582DB" box="[610,625,1745,1771]" italics="true" pageId="11" pageNumber="12">b</emphasis>
</bibRefCitation>
;
<bibRefCitation id="C73B4B28FFDB2E3BFD20F91FFED8833D" author="BATES, P. J. J. &amp; D. L. HARRISON" bookContentInfo="258 pp." journalOrPublisher="Harrison Zoological Museum, Sevenoaks" pageId="11" pageNumber="12" refId="ref11818" refString="BATES, P. J. J., and D. L. HARRISON. 1997. Bats of the Indian Subcontinent. Harrison Zoological Museum, Sevenoaks, 258 pp." title="Bats of the Indian Subcontinent" type="book" year="1997">Bates and Harrison, 1997</bibRefCitation>
;
<bibRefCitation id="C73B4B28FFDB2E3BFEFDF93DFE69833D" author="FRANCIS, C. M." bookContentInfo="392 pp." box="[345,509,1779,1805]" journalOrPublisher="Princeton University Press, Princeton" pageId="11" pageNumber="12" refId="ref12539" refString="FRANCIS, C. M. 2008. A guide to the mammals of southeast Asia. Princeton University Press, Princeton, 392 pp." title="A guide to the mammals of southeast Asia" type="book" year="2008">Francis, 2008</bibRefCitation>
;
<bibRefCitation id="C73B4B28FFDB2E3BFDAEF93DFD60833D" author="THONG, V. D. &amp; S. J. PUECHMAILLE &amp; A. DENZINGER &amp; P. J. BATES &amp; C. DIETZ &amp; G. CSORBA &amp; P. SOISOOK &amp; E. C. TEELING &amp; S. MATSUMURA &amp; N. M. FUREY" box="[522,756,1779,1805]" journalOrPublisher="Mammal Review" pageId="11" pageNumber="12" pagination="166 - 192" part="42" refId="ref15390" refString="THONG, V. D., S. J. PUECHMAILLE, A. DENZINGER, P. J. BATES, C. DIETZ, G. CSORBA, P. SOISOOK, E. C. TEELING, S. MATSUMURA, and N. M. FUREY. 2012 a. Systematics of the Hipposideros turpis complex and a description of a new subspecies from Vietnam. Mammal Review, 42: 166 - 192." title="Systematics of the Hipposideros turpis complex and a description of a new subspecies from Vietnam" type="journal article" year="2012">
Thong
<emphasis id="91DEEACBFFDB2E3BFDFBF93DFD00833D" box="[607,660,1779,1805]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
., 2012
<emphasis id="91DEEACBFFDB2E3BFD41F93DFD60833D" box="[741,756,1779,1805]" italics="true" pageId="11" pageNumber="12">a</emphasis>
</bibRefCitation>
,
<bibRefCitation id="C73B4B28FFDB2E3BFF33F8DBFF75831F" author="THONG, V. D. &amp; S. J. PUECHMAILLE &amp; A. DENZINGER &amp; C. DIETZ &amp; G. CSORBA &amp; P. J. BATES &amp; E. C. TEELING &amp; H. - U. SCHNITZ- LER" box="[151,225,1813,1839]" journalOrPublisher="Journal of Mammalogy" pageId="11" pageNumber="12" pagination="1 - 11" part="93" refId="ref15470" refString="THONG, V. D., S. J. PUECHMAILLE, A. DENZINGER, C. DIETZ, G. CSORBA, P. J. BATES, E. C. TEELING, and H. - U. SCHNITZ- LER. 2012 b. A new species of Hipposideros (Chiroptera: Hipposideridae) from Vietnam. Journal of Mammalogy, 93: 1 - 11." title="A new species of Hipposideros (Chiroptera: Hipposideridae) from Vietnam" type="journal article" year="2012">
2012
<emphasis id="91DEEACBFFDB2E3BFF76F8DBFF75831F" box="[210,225,1813,1839]" italics="true" pageId="11" pageNumber="12">b</emphasis>
</bibRefCitation>
;
<bibRefCitation id="C73B4B28FFDB2E3BFF57F8DBFE26831F" author="HAPPOLD, M." bookContentInfo="800 pp." box="[243,434,1813,1839]" editor="M. HAPPOLD &amp; D. C. D. HAPPOLD" journalOrPublisher="Bloomsbury Publishing, London" pageId="11" pageNumber="12" pagination="367 - 371" refId="ref13058" refString="HAPPOLD, M. 2013 c. Genus Hipposideros. Old World leaf-nosed bats. Pp. 367 - 371, in Mammals of Africa. Volume IV (M. HAPPOLD and D. C. D. HAPPOLD, eds.). Bloomsbury Publishing, London, 800 pp." title="Genus Hipposideros. Old World leaf-nosed bats." type="book chapter" volumeTitle="Mammals of Africa. Volume IV" year="2013">
Happold, 2013
<emphasis id="91DEEACBFFDB2E3BFE01F8DBFE26831F" box="[421,434,1813,1839]" italics="true" pageId="11" pageNumber="12">c</emphasis>
</bibRefCitation>
). The 2nd lateral leaflet in
<taxonomicName id="64AA4D5AFFDB2E3BFF33F8F8FE808360" baseAuthorityName="Hill" baseAuthorityYear="1963" box="[151,276,1846,1872]" class="Mammalia" family="Hipposideridae" genus="Phyllorhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cyclops">
<emphasis id="91DEEACBFFDB2E3BFF33F8F8FE808360" box="[151,276,1846,1872]" italics="true" pageId="11" pageNumber="12">D. cyclops</emphasis>
</taxonomicName>
extends posteriorly and forms a continuous extension of the posterior leaf (
<figureCitation id="3B912A5CFFDB2E3BFD9AF896FD1C8342" box="[574,648,1880,1906]" captionStart="FIG" captionStartId="9.[151,164,1783,1804]" captionTargetBox="[163,1411,1288,1771]" captionTargetId="figure-14@9.[765,1091,1288,1736]" captionTargetPageId="9" captionText="FIG. 4. Morphological comparisons of the frontal sacs and noseleaves in A — Macronycteris commersonii (FMNH 213588, ♀), Madagascar, Province dAntsiranana, Réserve Spéciale dAnkarana [now Parc National], near Andrafiabe Cave, 31 October 2010; B — M. gigas (FMNH 128212, ♀), Senegal, Casmance, Diabane, 12 km SW of Adeane, 15 January 1983; C — M. vittatus (FMNH 192800, ♀, sequenced for Cyt-b), Tanzania, Pemba Island, Kaskazini Region, Micheweni District, Kilijini Village, 3 August 2006; and D — Doryrhina cyclops (FMNH 164973), Uganda, Masindi District, Budongo Forest, 25 June 1998. The form of M. cryptovalorona is similar to M. commersonii and no comparative specimen material was available for M. thomensis. Drawing by Velizar Simeonovski" figureDoi="http://doi.org/10.5281/zenodo.6822116" httpUri="https://zenodo.org/record/6822116/files/figure.png" pageId="11" pageNumber="12">Fig. 4</figureCitation>
). Further, this species has a club-shaped process projecting from the posterior portion of the noseleaf, which distinguishes it from all African members of the genus
<taxonomicName id="64AA4D5AFFDB2E3BFC25FF1AFBB584DE" box="[897,1057,212,238]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="91DEEACBFFDB2E3BFC25FF1AFBB584DE" box="[897,1057,212,238]" italics="true" pageId="11" pageNumber="12">Hipposideros</emphasis>
</taxonomicName>
, as well as
<taxonomicName id="64AA4D5AFFDB2E3BFB7CFF1AFA1184DE" authorityName="Gray" authorityYear="1866" box="[1240,1413,212,238]" class="Mammalia" family="Hipposideridae" genus="Macronycteris" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="91DEEACBFFDB2E3BFB7CFF1AFA1184DE" box="[1240,1413,212,238]" italics="true" pageId="11" pageNumber="12">Macronycteris</emphasis>
</taxonomicName>
. Other aspects of the noseleaf of
<taxonomicName id="64AA4D5AFFDB2E3BFB0DFF38FAB28520" baseAuthorityName="Hill" baseAuthorityYear="1963" box="[1193,1318,246,272]" class="Mammalia" family="Hipposideridae" genus="Phyllorhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cyclops">
<emphasis id="91DEEACBFFDB2E3BFB0DFF38FAB28520" box="[1193,1318,246,272]" italics="true" pageId="11" pageNumber="12">D. cyclops</emphasis>
</taxonomicName>
include: 1) anterior portion is broad and with a club-like structure commencing at anterior margin of lateral leaflet and extending posteriorly to middle portion of noseleaf, internarial septa form relatively large and sculpted structures that partially obscure the deep-set nasal passages, and distinct lappets surrounded the nasal passages; 2) middle portion of noseleaf is absent; 3) posterior portion with a vertical medial septum (that extends as the club-shaped process mentioned above) and two prominent vertical lateral septa on either side of the vertical medial septum, which divide the structure into 6 separate cells, the lateral-most cell merging with the 2nd lateral leaflet, and posterior margin is a thin structure with little expansion. The different aspects of the noseleaf described herein are unique to
<taxonomicName id="64AA4D5AFFDB2E3BFAB6FCD8FA198700" authorityName="Peters" authorityYear="1871" box="[1298,1421,790,816]" class="Mammalia" family="Hipposideridae" genus="Doryrhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="91DEEACBFFDB2E3BFAB6FCD8FA198700" box="[1298,1421,790,816]" italics="true" pageId="11" pageNumber="12">Doryrhina</emphasis>
</taxonomicName>
and not found in another genus of the family
<taxonomicName id="64AA4D5AFFDB2E3BFA9EFCF6FC078744" authorityName="Lydekker" authorityYear="1891" class="Mammalia" family="Hipposideridae" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="family">Hipposideridae</taxonomicName>
(
<bibRefCitation id="C73B4B28FFDB2E3BFC07FC94FBCA8744" author="ROSEVEAR, D. R." bookContentInfo="418 pp." box="[931,1118,858,884]" journalOrPublisher="Trustees of the British Museum (Natural History), London" pageId="11" pageNumber="12" refId="ref14852" refString="ROSEVEAR, D. R. 1965. The bats of West Africa. Trustees of the British Museum (Natural History), London, 418 pp." title="The bats of West Africa" type="book" year="1965">Rosevear, 1965</bibRefCitation>
;
<bibRefCitation id="C73B4B28FFDB2E3BFBCEFC94FAD68744" author="PAYNE, J. &amp; C. M. FRANCIS &amp; K. PHILIPPS" bookContentInfo="332 pp." box="[1130,1346,858,884]" journalOrPublisher="The Sabah Society, Kota Kinabalu" pageId="11" pageNumber="12" refId="ref13982" refString="PAYNE, J., C. M. FRANCIS, and K. PHILIPPS. 1985. A field guide to the mammals of Borneo. The Sabah Society, Kota Kinabalu, 332 pp." title="A field guide to the mammals of Borneo" type="book" year="1985">
Payne
<emphasis id="91DEEACBFFDB2E3BFB1EFC94FB7A8744" box="[1210,1262,858,884]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
., 1985
</bibRefCitation>
;
<bibRefCitation id="C73B4B28FFDB2E3BFAEBFC94FC2A87A6" author="FLANNERY, T." bookContentInfo="568 pp." journalOrPublisher="Cornell University Press, Ithaca" pageId="11" pageNumber="12" refId="ref12416" refString="FLANNERY, T. 1995 a. Mammals of New Guinea. Cornell University Press, Ithaca, 568 pp." title="Mammals of New Guinea" type="book" year="1995">
Flannery, 1995
<emphasis id="91DEEACBFFDB2E3BFC0BFCB2FC2A87A6" box="[943,958,892,918]" italics="true" pageId="11" pageNumber="12">a</emphasis>
</bibRefCitation>
,
<bibRefCitation id="C73B4B28FFDB2E3BFC73FCB2FBB587A6" author="FLANNERY, T." bookContentInfo="464 pp." box="[983,1057,892,918]" journalOrPublisher="Cornell University Press, Ithaca" pageId="11" pageNumber="12" refId="ref12437" refString="FLANNERY, T. 1995 b. Mammals of the south-west Pacific and Moluccan islands. Cornell University Press, Ithaca, 464 pp." title="Mammals of the south-west Pacific and Moluccan islands" type="book" year="1995">
1995
<emphasis id="91DEEACBFFDB2E3BFBB6FCB2FBB587A6" box="[1042,1057,892,918]" italics="true" pageId="11" pageNumber="12">b</emphasis>
</bibRefCitation>
;
<bibRefCitation id="C73B4B28FFDB2E3BFB9FFCB2FA1C87A6" author="BATES, P. J. J. &amp; D. L. HARRISON" bookContentInfo="258 pp." box="[1083,1416,892,918]" journalOrPublisher="Harrison Zoological Museum, Sevenoaks" pageId="11" pageNumber="12" refId="ref11818" refString="BATES, P. J. J., and D. L. HARRISON. 1997. Bats of the Indian Subcontinent. Harrison Zoological Museum, Sevenoaks, 258 pp." title="Bats of the Indian Subcontinent" type="book" year="1997">Bates and Harrison, 1997</bibRefCitation>
;
<bibRefCitation id="C73B4B28FFDB2E3BFC8DFC50FC408788" author="FRANCIS, C. M." bookContentInfo="392 pp." box="[809,980,926,952]" journalOrPublisher="Princeton University Press, Princeton" pageId="11" pageNumber="12" refId="ref12539" refString="FRANCIS, C. M. 2008. A guide to the mammals of southeast Asia. Princeton University Press, Princeton, 392 pp." title="A guide to the mammals of southeast Asia" type="book" year="2008">Francis, 2008</bibRefCitation>
;
<bibRefCitation id="C73B4B28FFDB2E3BFC4CFC50FA998788" author="MONADJEM, A. &amp; P. J. TAYLOR &amp; F. P. D. COTERILL &amp; M. C. SCHOEMAN" bookContentInfo="596 pp." box="[1000,1293,926,952]" journalOrPublisher="Wits University Press, Johannesburg" pageId="11" pageNumber="12" refId="ref13742" refString="MONADJEM, A., P. J. TAYLOR, F. P. D. COTERILL, and M. C. SCHOEMAN. 2010. Bats of central and southern Africa: a biogeographic and taxonomic synthesis. Wits University Press, Johannesburg, 596 pp." title="Bats of central and southern Africa: a biogeographic and taxonomic synthesis" type="book" year="2010">
Monadjem
<emphasis id="91DEEACBFFDB2E3BFBD2FC50FB258788" box="[1142,1201,926,952]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
., 2010
</bibRefCitation>
;
<bibRefCitation id="C73B4B28FFDB2E3BFA85FC50FCE487EA" author="HAPPOLD, M." bookContentInfo="800 pp." editor="M. HAPPOLD &amp; D. C. D. HAPPOLD" journalOrPublisher="Bloomsbury Publishing, London" pageId="11" pageNumber="12" pagination="367 - 371" refId="ref13058" refString="HAPPOLD, M. 2013 c. Genus Hipposideros. Old World leaf-nosed bats. Pp. 367 - 371, in Mammals of Africa. Volume IV (M. HAPPOLD and D. C. D. HAPPOLD, eds.). Bloomsbury Publishing, London, 800 pp." title="Genus Hipposideros. Old World leaf-nosed bats." type="book chapter" volumeTitle="Mammals of Africa. Volume IV" year="2013">
Happold, 2013
<emphasis id="91DEEACBFFDB2E3BFCC7FC0EFCE487EA" box="[867,880,960,986]" italics="true" pageId="11" pageNumber="12">c</emphasis>
</bibRefCitation>
). The exception is the African species
<taxonomicName id="64AA4D5AFFDB2E3BFAE1FC0EFC1887CC" authorityName="Eisentraut" authorityYear="1956" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="camerunensis">
<emphasis id="91DEEACBFFDB2E3BFAE1FC0EFC1887CC" italics="true" pageId="11" pageNumber="12">camerunensis</emphasis>
</taxonomicName>
, apparently closely related to
<taxonomicName id="64AA4D5AFFDB2E3BFB4BFC2CFCE5802E" authority="(Hill, 1963)" baseAuthorityName="Hill" baseAuthorityYear="1963" class="Mammalia" family="Hipposideridae" genus="Phyllorhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cyclops">
<emphasis id="91DEEACBFFDB2E3BFB4BFC2CFAD387CC" box="[1263,1351,994,1020]" italics="true" pageId="11" pageNumber="12">cyclops</emphasis>
(
<bibRefCitation id="C73B4B28FFDB2E3BFAF3FC2CFCFC802E" author="HILL, J. E." journalOrPublisher="Bulletin of the British Museum (Natural History), Zoology" pageId="11" pageNumber="12" pagination="1 - 129" part="11" refId="ref13158" refString="HILL, J. E. 1963. A revision of the genus Hipposideros. Bulletin of the British Museum (Natural History), Zoology, 11: 1 - 129." title="A revision of the genus Hipposideros" type="journal article" year="1963">Hill, 1963</bibRefCitation>
)
</taxonomicName>
. Further,
<bibRefCitation id="C73B4B28FFDB2E3BFC40FBCAFBE4802E" author="TATE, G. H. H." box="[996,1136,1028,1054]" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="11" pageNumber="12" pagination="353 - 393" part="73" refId="ref15200" refString="TATE, G. H. H. 1941. Results of the Archbold Expeditions No. 35. A review of the genus Hipposideros with special reference to Indo-Australian species. Bulletin of the American Museum of Natural History, 73: 353 - 393." title="Results of the Archbold Expeditions No. 35. A review of the genus Hipposideros with special reference to Indo-Australian species" type="journal article" year="1941">Tate (1941)</bibRefCitation>
,
<bibRefCitation id="C73B4B28FFDB2E3BFBDBFBCAFA91802E" author="HILL, J. E." box="[1151,1285,1028,1054]" journalOrPublisher="Bulletin of the British Museum (Natural History), Zoology" pageId="11" pageNumber="12" pagination="1 - 129" part="11" refId="ref13158" refString="HILL, J. E. 1963. A revision of the genus Hipposideros. Bulletin of the British Museum (Natural History), Zoology, 11: 1 - 129." title="A revision of the genus Hipposideros" type="journal article" year="1963">Hill (1963)</bibRefCitation>
and
<bibRefCitation id="C73B4B28FFDB2E3BFAE6FBCAFC268070" author="KOOPMAN, K. F." bookContentInfo="217 pp." editor="J. NIETHAMMER &amp; H. SCHLIEMANN &amp; D. STARCK" journalOrPublisher="Walter de Gruyter, Berlin" pageId="11" pageNumber="12" pagination="100 - 109" refId="ref13354" refString="KOOPMAN, K. F. 1994. Chiroptera: Systematics. Pp. 100 - 109, in Handbuch der Zoologie. Volume VII (J. NIETHAMMER, H. SCHLIEMANN, and D. STARCK, eds.). Walter de Gruyter, Berlin, vii + 217 pp." title="Chiroptera: Systematics." type="book chapter" volumeTitle="Handbuch der Zoologie. Volume VII" year="1994">Koopman (1994)</bibRefCitation>
based on anatomical characters considered the
<taxonomicName id="64AA4D5AFFDB2E3BFC00FB86FBBF8052" baseAuthorityName="Hill" baseAuthorityYear="1963" box="[932,1067,1096,1122]" class="Mammalia" family="Hipposideridae" genus="Phyllorhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cyclops">
<emphasis id="91DEEACBFFDB2E3BFC00FB86FBBF8052" box="[932,1067,1096,1122]" italics="true" pageId="11" pageNumber="12">H. cyclops</emphasis>
</taxonomicName>
group to be distinct from other groups in this genus and composed of the African species
<taxonomicName id="64AA4D5AFFDB2E3BFC56FB42FBE18096" baseAuthorityName="Hill" baseAuthorityYear="1963" box="[1010,1141,1164,1190]" class="Mammalia" family="Hipposideridae" genus="Phyllorhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cyclops">
<emphasis id="91DEEACBFFDB2E3BFC56FB42FBE18096" box="[1010,1141,1164,1190]" italics="true" pageId="11" pageNumber="12">H. cyclops</emphasis>
</taxonomicName>
and
<taxonomicName id="64AA4D5AFFDB2E3BFB1EFB42FA118096" authorityName="Eisentraut" authorityYear="1956" box="[1210,1413,1164,1190]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="camerunensis">
<emphasis id="91DEEACBFFDB2E3BFB1EFB42FA118096" box="[1210,1413,1164,1190]" italics="true" pageId="11" pageNumber="12">H. camerunensis</emphasis>
</taxonomicName>
, and the Australian-New Guinea species
<taxonomicName id="64AA4D5AFFDB2E3BFAB2FB60FCC680DA" baseAuthorityName="Thomas &amp; Doria" baseAuthorityYear="1886" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="muscinus">
<emphasis id="91DEEACBFFDB2E3BFAB2FB60FCC680DA" italics="true" pageId="11" pageNumber="12">H. muscinus</emphasis>
</taxonomicName>
,
<taxonomicName id="64AA4D5AFFDB2E3BFCC4FB1EFC6A80DA" authorityName="Thomas" authorityYear="1913" box="[864,1022,1232,1258]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="wollastoni">
<emphasis id="91DEEACBFFDB2E3BFCC4FB1EFC6A80DA" box="[864,1022,1232,1258]" italics="true" pageId="11" pageNumber="12">H. wollastoni</emphasis>
</taxonomicName>
,
<taxonomicName id="64AA4D5AFFDB2E3BFBA8FB1EFB4280DA" authorityName="Hill" authorityYear="1985" box="[1036,1238,1232,1258]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="corynophyllus">
<emphasis id="91DEEACBFFDB2E3BFBA8FB1EFB4280DA" box="[1036,1238,1232,1258]" italics="true" pageId="11" pageNumber="12">H. corynophyllus</emphasis>
</taxonomicName>
,
<taxonomicName id="64AA4D5AFFDB2E3BFB41FB1EFACF80DA" authorityName="Matschie" authorityYear="1903" box="[1253,1371,1232,1258]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="semoni">
<emphasis id="91DEEACBFFDB2E3BFB41FB1EFACF80DA" box="[1253,1371,1232,1258]" italics="true" pageId="11" pageNumber="12">H. semoni</emphasis>
</taxonomicName>
and
<taxonomicName id="64AA4D5AFFDB2E3BFC8DFB3DFC3A813D" authorityName="Thomas" authorityYear="1913" box="[809,942,1267,1293]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="stenotis">
<emphasis id="91DEEACBFFDB2E3BFC8DFB3DFC3A813D" box="[809,942,1267,1293]" italics="true" pageId="11" pageNumber="12">H. stenotis</emphasis>
</taxonomicName>
. With the exception of
<taxonomicName id="64AA4D5AFFDB2E3BFB47FB3DFAAF813D" baseAuthorityName="Hill" baseAuthorityYear="1963" box="[1251,1339,1267,1293]" class="Mammalia" family="Hipposideridae" genus="Phyllorhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cyclops">
<emphasis id="91DEEACBFFDB2E3BFB47FB3DFAAF813D" box="[1251,1339,1267,1293]" italics="true" pageId="11" pageNumber="12">cyclops</emphasis>
</taxonomicName>
, these other species are not represented in our molecular dataset and are in need of study to determine if they are best placed in the genus
<taxonomicName id="64AA4D5AFFDB2E3BFBD3FA97FB668143" authorityName="Peters" authorityYear="1871" box="[1143,1266,1369,1395]" class="Mammalia" family="Hipposideridae" genus="Doryrhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="91DEEACBFFDB2E3BFBD3FA97FB668143" box="[1143,1266,1369,1395]" italics="true" pageId="11" pageNumber="12">Doryrhina</emphasis>
</taxonomicName>
or should be retained in
<taxonomicName id="64AA4D5AFFDB2E3BFC0AFAB5FBDF81A5" box="[942,1099,1403,1429]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="91DEEACBFFDB2E3BFC0AFAB5FBDF81A5" box="[942,1099,1403,1429]" italics="true" pageId="11" pageNumber="12">Hipposideros</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="A31536D9FFDB2E3CFCF6FA53FF7E84DE" blockId="11.[809,1422,177,1973]" lastBlockId="12.[151,764,177,784]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
The separated ears of
<taxonomicName id="64AA4D5AFFDB2E3BFBC4FA53FB4B8187" baseAuthorityName="Hill" baseAuthorityYear="1963" box="[1120,1247,1437,1463]" class="Mammalia" family="Hipposideridae" genus="Phyllorhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cyclops">
<emphasis id="91DEEACBFFDB2E3BFBC4FA53FB4B8187" box="[1120,1247,1437,1463]" italics="true" pageId="11" pageNumber="12">D. cyclops</emphasis>
</taxonomicName>
are long, narrow and terminating with pointed tips. The average ear length in this species is
<quantity id="64529B3CFFDB2E3BFB20FA2FFB6581CB" box="[1156,1265,1505,1531]" metricMagnitude="-2" metricUnit="m" metricValue="3.35" pageId="11" pageNumber="12" unit="mm" value="33.5">33.5 mm</quantity>
(sexes combined,
<emphasis id="91DEEACBFFDB2E3BFCDDF9CDFC1C822D" box="[889,904,1539,1565]" italics="true" pageId="11" pageNumber="12">n</emphasis>
= 125) (
<bibRefCitation id="C73B4B28FFDB2E3BFC4FF9CDFBE4822D" author="FAHR, J." bookContentInfo="800 pp." box="[1003,1136,1539,1565]" editor="M. HAPPOLD &amp; D. C. D. HAPPOLD" journalOrPublisher="Bloomsbury Publishing, London" pageId="11" pageNumber="12" pagination="380 - 383" refId="ref12366" refString="FAHR, J. 2013. Hipposideros cyclops cyclops leaf-nosed bat. Pp. 380 - 383, in Mammals of Africa. Volume IV (M. HAPPOLD and D. C. D. HAPPOLD, eds.). Bloomsbury Publishing, London, 800 pp." title="Hipposideros cyclops cyclops leaf-nosed bat." type="book chapter" volumeTitle="Mammals of Africa. Volume IV" year="2013">Fahr, 2013</bibRefCitation>
). Antitragus not present. Other measurements of
<taxonomicName id="64AA4D5AFFDB2E3BFBE4F9EBFB2F820F" baseAuthorityName="Hill" baseAuthorityYear="1963" box="[1088,1211,1573,1599]" class="Mammalia" family="Hipposideridae" genus="Phyllorhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cyclops">
<emphasis id="91DEEACBFFDB2E3BFBE4F9EBFB2F820F" box="[1088,1211,1573,1599]" italics="true" pageId="11" pageNumber="12">D. cyclops</emphasis>
</taxonomicName>
, which show sexual dimorphism, include: mean forearm length,
<quantity id="64529B3CFFDB2E3BFAFEF989FCC382B3" metricMagnitude="-2" metricUnit="m" metricValue="6.529999999999999" pageId="11" pageNumber="12" unit="mm" value="65.3">65.3 mm</quantity>
(
<specimenCount id="B5ACFD50FFDB2E3BFCCEF9A7FC0D82B3" box="[874,921,1641,1667]" pageId="11" pageNumber="12" type="male">♂♂</specimenCount>
,
<emphasis id="91DEEACBFFDB2E3BFC03F9A7FC2282B3" box="[935,950,1641,1667]" italics="true" pageId="11" pageNumber="12">n</emphasis>
= 53) and
<quantity id="64529B3CFFDB2E3BFBE4F9A7FB3F82B3" box="[1088,1195,1641,1667]" metricMagnitude="-2" metricUnit="m" metricValue="6.83" pageId="11" pageNumber="12" unit="mm" value="68.3">68.3 mm</quantity>
(
<specimenCount id="B5ACFD50FFDB2E3BFB64F9A7FB7F82B3" box="[1216,1259,1641,1667]" pageId="11" pageNumber="12" type="female">♀♀</specimenCount>
,
<emphasis id="91DEEACBFFDB2E3BFB5FF9A7FA9E82B3" box="[1275,1290,1641,1667]" italics="true" pageId="11" pageNumber="12">n</emphasis>
= 45); tail length,
<quantity id="64529B3CFFDB2E3BFC20F945FC7B8295" box="[900,1007,1675,1701]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" pageId="11" pageNumber="12" unit="mm" value="26.5">26.5 mm</quantity>
(
<specimenCount id="B5ACFD50FFDB2E3BFBA7F945FBA68295" box="[1027,1074,1675,1701]" pageId="11" pageNumber="12" type="male">♂♂</specimenCount>
,
<emphasis id="91DEEACBFFDB2E3BFBE5F945FBC48295" box="[1089,1104,1675,1701]" italics="true" pageId="11" pageNumber="12">n</emphasis>
= 42) and
<quantity id="64529B3CFFDB2E3BFB7EF945FAD28295" box="[1242,1350,1675,1701]" metricMagnitude="-2" metricUnit="m" metricValue="2.92" pageId="11" pageNumber="12" unit="mm" value="29.2">29.2 mm</quantity>
(
<specimenCount id="B5ACFD50FFDB2E3BFAF8F945FA138295" box="[1372,1415,1675,1701]" pageId="11" pageNumber="12" type="female">♀♀</specimenCount>
,
<emphasis id="91DEEACBFFDB2E3BFC8DF963FCAC82F7" box="[809,824,1709,1735]" italics="true" pageId="11" pageNumber="12">n</emphasis>
= 77); and hindfoot length (with claws),
<quantity id="64529B3CFFDB2E3BFA80F963FA1A82F7" box="[1316,1422,1709,1735]" metricMagnitude="-2" metricUnit="m" metricValue="1.98" pageId="11" pageNumber="12" unit="mm" value="19.8">19.8 mm</quantity>
(
<specimenCount id="B5ACFD50FFDB2E3BFC96F901FCF582D9" box="[818,865,1743,1769]" pageId="11" pageNumber="12" type="male">♂♂</specimenCount>
,
<emphasis id="91DEEACBFFDB2E3BFCD4F901FCEB82D9" box="[880,895,1743,1769]" italics="true" pageId="11" pageNumber="12">n</emphasis>
= 41) and
<quantity id="64529B3CFFDB2E3BFBAFF901FBE382D9" box="[1035,1143,1743,1769]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" pageId="11" pageNumber="12" unit="mm" value="20.5">20.5 mm</quantity>
(
<specimenCount id="B5ACFD50FFDB2E3BFB29F901FB2C82D9" box="[1165,1208,1743,1769]" pageId="11" pageNumber="12" type="female">♀♀</specimenCount>
,
<emphasis id="91DEEACBFFDB2E3BFB6DF901FB4C82D9" box="[1225,1240,1743,1769]" italics="true" pageId="11" pageNumber="12">n</emphasis>
= 73) (
<bibRefCitation id="C73B4B28FFDB2E3BFA92F901FC74833B" author="DECHER, J. &amp; J. FAHR" journalOrPublisher="Mammalian Species" pageId="11" pageNumber="12" pagination="1 - 7" part="763" refId="ref12091" refString="DECHER, J. and J. FAHR. 2005. Hipposideros cyclops. Mammalian Species, 763: 1 - 7." title="Hipposideros cyclops" type="journal article" year="2005">Decher and Fahr, 2005</bibRefCitation>
).
<taxonomicName id="64AA4D5AFFDB2E3BFC50F93FFB5B833B" baseAuthorityName="Temminck" baseAuthorityYear="1853" box="[1012,1231,1777,1803]" class="Mammalia" family="Hipposideridae" genus="Doryrhina" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cyclops">
<emphasis id="91DEEACBFFDB2E3BFC50F93FFB5B833B" box="[1012,1231,1777,1803]" italics="true" pageId="11" pageNumber="12">Doryrhina cyclops</emphasis>
</taxonomicName>
has dense, long and woolly pelage, on the dorsum generally blackish-brown and often with a frosted tint, while the ventrum is lighter and not frosted. No rufus or orange colour morphs are known. The wing and interfemoral membranes are blackish-brown and skin on forearm, wing digits and tibia are paler reddishbrown.
</paragraph>
<paragraph id="A31536D9FFDC2E3CFF65FF38FEFD85CF" blockId="12.[151,764,177,784]" pageId="12" pageNumber="13">
The skull of
<taxonomicName id="64AA4D5AFFDC2E3CFEF3FF38FE478520" baseAuthorityName="Temminck" baseAuthorityYear="1853" box="[343,467,246,272]" class="Mammalia" family="Hipposideridae" genus="Doryrhina" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="cyclops">
<emphasis id="91DEEACBFFDC2E3CFEF3FF38FE478520" box="[343,467,246,272]" italics="true" pageId="12" pageNumber="13">D. cyclops</emphasis>
</taxonomicName>
, is proportionately large, with a lengthened braincase, and low sagittal crest. The internarial septum is not enlarged. Rostrum is distinctly broad. Zygomatic arches are slender and form the widest portion of the skull. Premaxillae posteriorly wide and in broad contact with the palate. Anterior palatal foramina enclosed. Cochleae greatly expanded.
</paragraph>
<paragraph id="A31536D9FFDC2E3CFF65FDC9FE758720" blockId="12.[151,764,177,784]" pageId="12" pageNumber="13">Dental formula — premolars 1/2, canines 1/1, premolars 2/2 and molars 3/3. The upper incisors and associated cusps are not well developed. The upper canines lack well-defined cusps, but have distinct cingula. The upper anterior premolar is reduced, visible in lateral view, and in contact with the canine and posterior premolar. Anterior lower premolar is distinctly small.</paragraph>
<paragraph id="A31536D9FFDC2E3CFF33FCF4FE238764" blockId="12.[151,764,826,1296]" box="[151,439,826,852]" pageId="12" pageNumber="13">
<emphasis id="91DEEACBFFDC2E3CFF33FCF4FE238764" box="[151,439,826,852]" italics="true" pageId="12" pageNumber="13">Karyological characters</emphasis>
</paragraph>
<paragraph id="A31536D9FFDC2E3CFF65FC92FDA58120" blockId="12.[151,764,826,1296]" pageId="12" pageNumber="13">
On the basis of karyological information from the literature, further evidence can be found to support the resurrection of
<taxonomicName id="64AA4D5AFFDC2E3CFE0AFC6EFDBD878A" authorityName="Peters" authorityYear="1871" box="[430,553,928,954]" class="Mammalia" family="Hipposideridae" genus="Doryrhina" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="91DEEACBFFDC2E3CFE0AFC6EFDBD878A" box="[430,553,928,954]" italics="true" pageId="12" pageNumber="13">Doryrhina</emphasis>
</taxonomicName>
for a species previously placed in the genus
<taxonomicName id="64AA4D5AFFDC2E3CFE58FC0DFD0D87ED" box="[508,665,963,989]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="91DEEACBFFDC2E3CFE58FC0DFD0D87ED" box="[508,665,963,989]" italics="true" pageId="12" pageNumber="13">Hipposideros</emphasis>
</taxonomicName>
,
<taxonomicName id="64AA4D5AFFDC2E3CFD0AFC0DFF4187CF" baseAuthorityName="Temminck" baseAuthorityYear="1853" class="Mammalia" family="Hipposideridae" genus="Doryrhina" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="cyclops">
<emphasis id="91DEEACBFFDC2E3CFD0AFC0DFF4187CF" italics="true" pageId="12" pageNumber="13">H. cyclops</emphasis>
</taxonomicName>
.
<taxonomicName id="64AA4D5AFFDC2E3CFF4AFC2BFE1F87CF" box="[238,395,997,1023]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="91DEEACBFFDC2E3CFF4AFC2BFE1F87CF" box="[238,395,997,1023]" italics="true" pageId="12" pageNumber="13">Hipposideros</emphasis>
</taxonomicName>
shows extreme karyological conservatism, with nearly all species examined to date possessing a 2
<emphasis id="91DEEACBFFDC2E3CFE24FBE7FE1B8073" box="[384,399,1065,1091]" italics="true" pageId="12" pageNumber="13">n</emphasis>
complement of 32 (
<bibRefCitation id="C73B4B28FFDC2E3CFD22FBE7FE9E8055" author="HARADA, M. &amp; M. MINEZAWA &amp; S. TAKADA &amp; S. YENBUTRA &amp; P. NUN- PAKDEE &amp; S. OHTANI" journalOrPublisher="Caryologia" pageId="12" pageNumber="13" pagination="269 - 278" part="35" refId="ref13111" refString="HARADA, M., M. MINEZAWA, S. TAKADA, S. YENBUTRA, P. NUN- PAKDEE, and S. OHTANI. 1982. Karyological analysis of 12 species of bats from Thailand. Caryologia, 35: 269 - 278." title="Karyological analysis of 12 species of bats from Thailand" type="journal article" year="1982">
Harada
<emphasis id="91DEEACBFFDC2E3CFD42FBE7FF3A8055" italics="true" pageId="12" pageNumber="13">et al</emphasis>
., 1982
</bibRefCitation>
;
<bibRefCitation id="C73B4B28FFDC2E3CFEBAFB85FD908055" author="HOOD, C. S. &amp; D. A. SCHLITTER &amp; J. I. GEORGUDAKI &amp; S. YENBUTRA &amp; R. J. BAKER" box="[286,516,1099,1125]" journalOrPublisher="Annals of Carnegie Museum" pageId="12" pageNumber="13" pagination="99 - 109" part="57" refId="ref13191" refString="HOOD, C. S., D. A. SCHLITTER, J. I. GEORGUDAKI, S. YENBUTRA, and R. J. BAKER. 1988. Chromosomal studies of bats (Mammalia: Chiroptera) from Thailand. Annals of Carnegie Museum, 57: 99 - 109." title="Chromosomal studies of bats (Mammalia: Chiroptera) from Thailand" type="journal article" year="1988">
Hood
<emphasis id="91DEEACBFFDC2E3CFECAFB85FE3D8055" box="[366,425,1099,1125]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
., 1988
</bibRefCitation>
;
<bibRefCitation id="C73B4B28FFDC2E3CFDBCFB85FF4180B7" author="RAUTENBACH, I. &amp; G. BRONNER &amp; D. SCHLITTER" journalOrPublisher="Koedoe" pageId="12" pageNumber="13" pagination="87 - 104" part="36" refId="ref14661" refString="RAUTENBACH, I., G. BRONNER, and D. SCHLITTER. 1993. Karyotypic data and attendant systematic implications for the bats of southern Africa. Koedoe, 36: 87 - 104." title="Karyotypic data and attendant systematic implications for the bats of southern Africa" type="journal article" year="1993">
Rautenbach
<emphasis id="91DEEACBFFDC2E3CFD16FB85FD798055" box="[690,749,1099,1125]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
., 1993
</bibRefCitation>
;
<bibRefCitation id="C73B4B28FFDC2E3CFF43FBA3FDF380B7" author="BOGDANOWICZ, W. &amp; R. D. OWEN" bookContentInfo="365 pp." box="[231,615,1133,1159]" editor="T. H. KUNZ &amp; P. A. RACEY" journalOrPublisher="Smithsonian Institution Press. Washington, D. C." pageId="12" pageNumber="13" pagination="27 - 42" refId="ref11965" refString="BOGDANOWICZ, W., and R. D. OWEN. 1998. In the Minotaur's labyrinth: phylogeny of the bat family Hipposideridae. Pp. 27 - 42, in Bat biology and conservation (T. H. KUNZ and P. A. RACEY, eds.). Smithsonian Institution Press. Washington, D. C., 365 pp." title="In the Minotaur's labyrinth: phylogeny of the bat family Hipposideridae" type="book chapter" volumeTitle="Bat biology and conservation" year="1998">Bogdanowicz and Owen, 1998</bibRefCitation>
;
<bibRefCitation id="C73B4B28FFDC2E3CFDDDFBA3FEBE8099" author="KOUBINOVA, D. &amp; K. SREEPADA &amp; P. KOUBEK &amp; J. ZIMA" journalOrPublisher="Acta Chiropterologica" pageId="12" pageNumber="13" pagination="393 - 400" part="12" refId="ref13408" refString="KOUBINOVA, D., K. SREEPADA, P. KOUBEK, and J. ZIMA. 2010. Karyotypic variation in rhinolophid and hipposiderid bats (Chiroptera: Rhinolophidae, Hipposideridae). Acta Chiropterologica, 12: 393 - 400." title="Karyotypic variation in rhinolophid and hipposiderid bats (Chiroptera: Rhinolophidae, Hipposideridae)" type="journal article" year="2010">
Koubínová
<emphasis id="91DEEACBFFDC2E3CFF33FB41FF5B8099" box="[151,207,1167,1193]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
., 2010
</bibRefCitation>
). However, in the case of
<taxonomicName id="64AA4D5AFFDC2E3CFDD0FB41FD608099" baseAuthorityName="Temminck" baseAuthorityYear="1853" box="[628,756,1167,1193]" class="Mammalia" family="Hipposideridae" genus="Doryrhina" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="cyclops">
<emphasis id="91DEEACBFFDC2E3CFDD0FB41FD608099" box="[628,756,1167,1193]" italics="true" pageId="12" pageNumber="13">H. cyclops</emphasis>
</taxonomicName>
, 2
<emphasis id="91DEEACBFFDC2E3CFF02FB7CFF2180FC" box="[166,181,1202,1228]" italics="true" pageId="12" pageNumber="13">n</emphasis>
= 36 (
<bibRefCitation id="C73B4B28FFDC2E3CFEA7FB7CFD8D80FC" author="KOUBINOVA, D. &amp; K. SREEPADA &amp; P. KOUBEK &amp; J. ZIMA" box="[259,537,1202,1228]" journalOrPublisher="Acta Chiropterologica" pageId="12" pageNumber="13" pagination="393 - 400" part="12" refId="ref13408" refString="KOUBINOVA, D., K. SREEPADA, P. KOUBEK, and J. ZIMA. 2010. Karyotypic variation in rhinolophid and hipposiderid bats (Chiroptera: Rhinolophidae, Hipposideridae). Acta Chiropterologica, 12: 393 - 400." title="Karyotypic variation in rhinolophid and hipposiderid bats (Chiroptera: Rhinolophidae, Hipposideridae)" type="journal article" year="2010">
Koubínová
<emphasis id="91DEEACBFFDC2E3CFE2BFB7CFE5080FC" box="[399,452,1202,1228]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
., 2010
</bibRefCitation>
), which is also different from
<taxonomicName id="64AA4D5AFFDC2E3CFE83FB1AFE4580DE" authorityName="Gray" authorityYear="1866" box="[295,465,1236,1262]" class="Mammalia" family="Hipposideridae" genus="Macronycteris" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="91DEEACBFFDC2E3CFE83FB1AFE4580DE" box="[295,465,1236,1262]" italics="true" pageId="12" pageNumber="13">Macronycteris</emphasis>
</taxonomicName>
, 2
<emphasis id="91DEEACBFFDC2E3CFE54FB1AFE6B80DE" box="[496,511,1236,1262]" italics="true" pageId="12" pageNumber="13">n</emphasis>
= 52 (
<bibRefCitation id="C73B4B28FFDC2E3CFDF5FB1AFE978120" author="RAUTENBACH, I. &amp; G. BRONNER &amp; D. SCHLITTER" journalOrPublisher="Koedoe" pageId="12" pageNumber="13" pagination="87 - 104" part="36" refId="ref14661" refString="RAUTENBACH, I., G. BRONNER, and D. SCHLITTER. 1993. Karyotypic data and attendant systematic implications for the bats of southern Africa. Koedoe, 36: 87 - 104." title="Karyotypic data and attendant systematic implications for the bats of southern Africa" type="journal article" year="1993">
Rautenbach
<emphasis id="91DEEACBFFDC2E3CFD42FB1AFF3A8120" italics="true" pageId="12" pageNumber="13">et al</emphasis>
., 1993
</bibRefCitation>
;
<bibRefCitation id="C73B4B28FFDC2E3CFEB4FB38FDB18120" author="KOUBINOVA, D. &amp; K. SREEPADA &amp; P. KOUBEK &amp; J. ZIMA" box="[272,549,1270,1296]" journalOrPublisher="Acta Chiropterologica" pageId="12" pageNumber="13" pagination="393 - 400" part="12" refId="ref13408" refString="KOUBINOVA, D., K. SREEPADA, P. KOUBEK, and J. ZIMA. 2010. Karyotypic variation in rhinolophid and hipposiderid bats (Chiroptera: Rhinolophidae, Hipposideridae). Acta Chiropterologica, 12: 393 - 400." title="Karyotypic variation in rhinolophid and hipposiderid bats (Chiroptera: Rhinolophidae, Hipposideridae)" type="journal article" year="2010">
Koubínová
<emphasis id="91DEEACBFFDC2E3CFE3EFB38FE5A8120" box="[410,462,1270,1296]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
., 2010
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>