279 lines
28 KiB
XML
279 lines
28 KiB
XML
<document id="91E008FE791BA2ED9C974EDE3D1AB55A" ID-CLB-Dataset="35142" ID-DOI="http://dx.doi.org/10.3897/zookeys.339.4903" ID-GBIF-Dataset="f5e2c4d6-742c-4235-bf0e-b97e1ec678d2" ID-PMC="PMC3800827" ID-Pensoft-Pub="1313-2970-339-1" ID-PubMed="24146585" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-339-1" ModsDocOrigin="ZooKeys 339" ModsDocTitle="Taxonomic revision of the genus Carasobarbus Karaman, 1971 (Actinopterygii, Cyprinidae)" checkinTime="1451246844427" checkinUser="pensoft" docAuthor="Borkenhagen, Kai & Krupp, Friedhelm" docDate="2013" docId="172EB0233D61E16171EFC009AC764B55" docLanguage="en" docName="ZooKeys 339: 1-53" docOrigin="ZooKeys 339" docSource="http://dx.doi.org/10.3897/zookeys.339.4903" docTitle="Carasobarbus kosswigi Ladiges 1960" docType="treatment" docVersion="5" lastPageNumber="29" masterDocId="FF8B0A687873FFFAFFC0C22FFFF3FFD2" masterDocTitle="Taxonomic revision of the genus Carasobarbus Karaman, 1971 (Actinopterygii, Cyprinidae)" masterLastPageNumber="53" masterPageNumber="1" pageNumber="27" updateTime="1732747224765" updateUser="ExternalLinkService">
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<mods:title id="E31F8F2875C7F7C9BB9CC043F1C108B7">Taxonomic revision of the genus Carasobarbus Karaman, 1971 (Actinopterygii, Cyprinidae)</mods:title>
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<mods:namePart id="4955D3DC7D4C06CCC22FCDF1B943EF28">Borkenhagen, Kai</mods:namePart>
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<mods:namePart id="3C1FC9BB799B87ABD8E6400C71CD5A2C">Krupp, Friedhelm</mods:namePart>
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<mods:date id="04F7166DA86E194C1BB35A6157A2467C">2013</mods:date>
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<mods:number id="9E206C2AEA71DF49EF2DE4E888388C74">339</mods:number>
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<mods:classification id="EB5A534322D3FC81CDA740527D1C3F5D">journal article</mods:classification>
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<mods:identifier id="1855E413B745A0F00BDC8AED5A87490C" type="DOI">http://dx.doi.org/10.3897/zookeys.339.4903</mods:identifier>
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<mods:identifier id="BE621AF0765424875FA4D8FF70FEFB73" type="Pensoft-Pub">1313-2970-339-1</mods:identifier>
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<treatment id="172EB0233D61E16171EFC009AC764B55" ID-GBIF-Taxon="152048583" LSID="urn:lsid:plazi:treatment:172EB0233D61E16171EFC009AC764B55" httpUri="http://treatment.plazi.org/id/172EB0233D61E16171EFC009AC764B55" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27" scope_family="Cyprinidae">
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<subSubSection id="9ACC0F3F0B26BB5761CE3CEE864479B2" pageId="26" pageNumber="27" type="nomenclature">
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<paragraph id="26FD855552EBC820CBC48E91E740C101" pageId="26" pageNumber="27">
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<taxonomicName id="AD66307D5AF18D7D4B2F3CF8B611BAA6" ID-CoL="R2ZX" LSID="http://species-id.net/wiki/Carasobarbus_kosswigi" authority="Ladiges, 1960" authorityName="Ladiges" authorityYear="1960" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus kosswigi" order="Cypriniformes" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="kosswigi">Carasobarbus kosswigi (Ladiges, 1960)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="A88F4DC79830ADC9EA391D74EBA8F8C4" pageId="26" pageNumber="27" type="reference_group">
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<paragraph id="CB916B99769B8F16B2B81A3706C5C126" pageId="26" pageNumber="27">
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<taxonomicName id="D517B800EB360FB46B0907AA560D8239" family="Cyprinidae" genus="Cyclocheilichthys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocheilichthys kosswigi" order="Cypriniformes" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="kosswigi">Cyclocheilichthys kosswigi</taxonomicName>
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<bibRefCitation id="98F2ADF560639F85D62410B773641E82" author="Ladiges, W" journalOrPublisher="Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut" pageId="45" pageNumber="46" pagination="105 - 150" title="Suesswasserfische der Tuerkei. I. Teil Cyprinidae." volume="58" year="1960">Ladiges 1960</bibRefCitation>
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: 135.
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</paragraph>
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</subSubSection>
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<subSubSection id="9908A28E8CA36A64B761F004AF2D82B3" pageId="26" pageNumber="27" type="material">
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<paragraph id="38B3C44FD5F294F05011A4FFE77CEAD5" pageId="26" pageNumber="27">Material.</paragraph>
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<paragraph id="D1DFB5377E146D5BC158CA44D2C20A0B" pageId="26" pageNumber="27">
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Type material. Holotype of
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<taxonomicName id="80D41CB6BCD6BD57C72D8A5082E1E1E4" family="Cyprinidae" genus="Cyclocheilichthys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocheilichthys kosswigi" order="Cypriniformes" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="kosswigi">Cyclocheilichthys kosswigi</taxonomicName>
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: ZMH H 1148, Turkey, Batman
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<normalizedToken id="32ECD6029D0F453E07BFD6550E419165" originalValue="Çayı">Cayi</normalizedToken>
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[
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<geoCoordinate id="22823C90C7965AE002F31B2AF2009EF7" direction="north" orientation="latitude" precision="15" value="37.787777">37°47'16"N</geoCoordinate>
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,
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<geoCoordinate id="0127555C28C545927515B06F3E0EAEF7" direction="east" orientation="longitude" precision="15" value="41.014168">41°0'51"E</geoCoordinate>
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], C. Kosswig, IV.1939.
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</paragraph>
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<paragraph id="0AB462AA32CFCF53411E8CBB4858CAD3" pageId="26" pageNumber="27">
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Non-type material. Tigris-Euphrates system. NMW 90369, 1, Turkey, Batman
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<normalizedToken id="A005068822F7FD529DD4107073FA31D4" originalValue="Çayı">Cayi</normalizedToken>
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near Baschkaja [
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<geoCoordinate id="2E1B695766FB2E7FF2E0428AA75B266A" direction="north" orientation="latitude" precision="15" value="37.8875">37°53'15"N</geoCoordinate>
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,
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<geoCoordinate id="9007773F72AD345E8D7B39EE0CE7580F" direction="east" orientation="longitude" precision="15" value="41.13222">41°7'56"E</geoCoordinate>
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], V. Pietschmann, 15 Jul 1914. - NMW 90805, 1, Turkey,
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<normalizedToken id="15AF9B05C0E8FD13CEC915BB0C48B0B6" originalValue="Gökçesu">Goekcesu</normalizedToken>
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<normalizedToken id="B03AADFCC95BDDB4938069354E8673DA" originalValue="Çayı">Cayi</normalizedToken>
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(
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<geoCoordinate id="117285305895F2CC8F530F520B54DBE7" direction="north" orientation="latitude" precision="925" value="37.75">37°45'N</geoCoordinate>
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,
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<geoCoordinate id="A0242242B4C48B2DC939F29A9F5687FE" direction="east" orientation="longitude" precision="925" value="41.75">41°45'E</geoCoordinate>
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), 26 Sep 1985. - ZMH 9548, 2, Turkey,
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<normalizedToken id="B3C58EFF7289503398BCA84415EBB2CC" originalValue="Ceylanpınar">Ceylanpinar</normalizedToken>
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[
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<geoCoordinate id="0538D20055B2E09B9AB1421782916AC9" direction="north" orientation="latitude" precision="15" value="36.84722">36°50'50"N</geoCoordinate>
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,
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<geoCoordinate id="2AE57EB1F436F3EBEA718F17C7F65413" direction="east" orientation="longitude" precision="15" value="40.05">40°3'0"E</geoCoordinate>
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]. - SMF 33119, 1, Syria, Nahr al
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<normalizedToken id="AB2559785D42E99D983747458C42BB7C" originalValue="Khābūr">Khabūr</normalizedToken>
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at Al
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<normalizedToken id="487E063DC4BEC36375FACF70680AF980" originalValue="Ḩasakah">Ḩasakah</normalizedToken>
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[
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||
<geoCoordinate id="A4FB4F7425778CCB841B2CD07C8BB982" direction="north" orientation="latitude" precision="15" value="36.5025">36°30'9"N</geoCoordinate>
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,
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||
<geoCoordinate id="91137F2EEBD69CEA9023869951AA03AD" direction="east" orientation="longitude" precision="15" value="40.74778">40°44'52"E</geoCoordinate>
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||
], F. Krupp. - SMF 30172, 1, Syria, Nahr al
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||
<normalizedToken id="DC8A55017C5A1CC396E19B0F43188CEA" originalValue="Khābūr">Khabūr</normalizedToken>
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near Tall
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<normalizedToken id="79A12A93204F6A4525564D060EA17DAA" originalValue="Budayrī">Budayrī</normalizedToken>
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(
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<geoCoordinate id="237311558BC10A401AF3BA7795B990CB" direction="north" orientation="latitude" precision="925" value="36.4">36°24'N</geoCoordinate>
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,
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<geoCoordinate id="75A781858987CD5EE0373501004AF9F0" direction="east" orientation="longitude" precision="925" value="40.866665">40°52'E</geoCoordinate>
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), F. Krupp, 2-4 Nov 1986. - SMF 30173, 1, Syria, Nahr al
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<normalizedToken id="4C3846367B68C5CF181E6F2BDF7442E0" originalValue="Khābūr">Khabūr</normalizedToken>
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near
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<normalizedToken id="7142C74F7B059BE7026F86A750F8D162" originalValue="Nahāb">Nahab</normalizedToken>
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(
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<geoCoordinate id="6BCACAEAF51590DDCF694B9816BB67E3" direction="north" orientation="latitude" precision="925" value="36.383335">36°23'N</geoCoordinate>
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,
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<geoCoordinate id="C58D648868AA3ED7FE8711F22E470485" direction="east" orientation="longitude" precision="925" value="40.833332">40°50'E</geoCoordinate>
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), F. Krupp, 23-27 May 1989. - SMF 30174, 1, Syria, Nahr al
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||
<normalizedToken id="6A7006292CE9EB92767F4938799867F2" originalValue="Khābūr">Khabūr</normalizedToken>
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near
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||
<normalizedToken id="3DE8A2348357DB423CD9F32BF2BECDCF" originalValue="Nahāb">Nahab</normalizedToken>
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(
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||
<geoCoordinate id="AFE50F4A5F85297D0004E7F75084D5CA" direction="north" orientation="latitude" precision="925" value="36.383335">36°23'N</geoCoordinate>
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,
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||
<geoCoordinate id="6C7692ECC97FBBBBD0E97CF2875E8F4F" direction="east" orientation="longitude" precision="925" value="40.833332">40°50'E</geoCoordinate>
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||
), F. Krupp, 28
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||
<normalizedToken id="03632EEE1172DAC5CEF0E33DC7CEEACE" originalValue="Sep–">Sep-</normalizedToken>
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||
8 Oct 1988. - CMNFI 79-0290, 2, Iran,
|
||
<normalizedToken id="54752085A836520F282F4216379A67EA" originalValue="Qaşr-e">Qasr-e</normalizedToken>
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||
<normalizedToken id="3EEFDACB65BFC453BC40B3A313444BA8" originalValue="Shīrīn">Shīrīn</normalizedToken>
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||
(
|
||
<geoCoordinate id="E3B1C6BA009FDEF1D42C451F55812779" direction="north" orientation="latitude" precision="925" value="34.516666">34°31'N</geoCoordinate>
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||
,
|
||
<geoCoordinate id="369F831F859A82C0903A9E7D603A3D2B" direction="east" orientation="longitude" precision="925" value="45.583332">45°35'E</geoCoordinate>
|
||
). - CMNFI 79-0289, 1, Iran, 25-30 km from
|
||
<normalizedToken id="91989401A091776CB478F80E14448991" originalValue="Qaşr-e">Qasr-e</normalizedToken>
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||
<normalizedToken id="007F1ECF5D9EC6A73C04FE16FD062917" originalValue="Shīrīn">Shīrīn</normalizedToken>
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(
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||
<geoCoordinate id="A56B59092854F3F123B322048FC89B1A" direction="north" orientation="latitude" precision="925" value="34.466667">34°28'N</geoCoordinate>
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||
,
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||
<geoCoordinate id="5F972C248145EF66A663DE232B2365A3" direction="east" orientation="longitude" precision="925" value="45.866665">45°52'E</geoCoordinate>
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||
). - BMNH 1974.2.22:1292-1296, 4; BMNH 1974.2.22:1281, 1, Iraq, Euphrates at
|
||
<normalizedToken id="BDD814408936C1944EB3BC6EADEAC038" originalValue="Ḩadīthah">Ḩadīthah</normalizedToken>
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[
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||
<geoCoordinate id="12EA0CE6E0E29E22EB61779AB43CC047" direction="north" orientation="latitude" precision="15" value="34.139725">34°8'23"N</geoCoordinate>
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||
,
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||
<geoCoordinate id="4E0FEF6D19EEE76F596D6DA289A6F7AC" direction="east" orientation="longitude" precision="15" value="42.378056">42°22'41"E</geoCoordinate>
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||
], 19 Oct 1953. - CMNFI 79-0275, 1, Iran,
|
||
<normalizedToken id="A34734FF27BB4937E2DCBA3263937A94" originalValue="Rūdkhāneh-ye">Rūdkhaneh-ye</normalizedToken>
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||
<normalizedToken id="9D6FB8020B6E7D9D2BAB9F9C95485C20" originalValue="Kashgān">Kashgan</normalizedToken>
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||
, 2 km from
|
||
<normalizedToken id="B2335E06DC61969BE920CACFE84A25D3" originalValue="Ma‘mūlān">Ma'mūlan</normalizedToken>
|
||
(
|
||
<geoCoordinate id="0BEA1C39CBC1C606CE7DFE6E92BD7D40" direction="north" orientation="latitude" precision="925" value="33.416668">33°25'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="FCA177B3118D8D5DC38068FE7B3D4161" direction="east" orientation="longitude" precision="925" value="47.966667">47°58'E</geoCoordinate>
|
||
). - SMF 33129, 3, Iran,
|
||
<normalizedToken id="57A8A6E6485EA7A2EB3EECF3D592F3DC" originalValue="Rūdkhāneh-ye">Rūdkhaneh-ye</normalizedToken>
|
||
Karkheh at Pol-e Dokhtar (
|
||
<geoCoordinate id="6156FD85BB884BFB25558942434E3927" direction="north" orientation="latitude" precision="15" value="33.16">33°9'36"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="8F9A0CD37715B7E997F40B9B28FF1CB8" direction="east" orientation="longitude" precision="15" value="47.72">47°43'12"E</geoCoordinate>
|
||
), N. Alwan et al., 3 Mar 2008. - ZM-CBSU 4153, 1; ZM-CBSU 4154, 1, Iran,
|
||
<normalizedToken id="B0BF45C43FC93BA0FD686CB0870F2276" originalValue="Rūdkhāneh-ye">Rūdkhaneh-ye</normalizedToken>
|
||
Dez at
|
||
<normalizedToken id="84A331066470F601AEFC8AEAB14A4648" originalValue="Dezfūl">Dezfūl</normalizedToken>
|
||
[
|
||
<geoCoordinate id="719062398ADB523418CDB4A53040AB38" direction="north" orientation="latitude" precision="15" value="32.3825">32°22'57"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="97C6DCA60BD6AE7E355A7A5BCE430143" direction="east" orientation="longitude" precision="15" value="48.401947">48°24'7"E</geoCoordinate>
|
||
], F. Bossaghzadeh, 8 Jun 2005.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2A97C598CEE45A41E223CEE2BBC78CAA" pageId="26" pageNumber="27" type="diagnosis">
|
||
<paragraph id="445D96E43AF3063C184262294FA40163" pageId="26" pageNumber="27">Diagnosis.</paragraph>
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||
<paragraph id="AA4070625856B6500DBD17B83D38E729" pageId="26" pageNumber="27">Two pairs of barbels; 32 to 38 scales in the lateral line, usually 14 to 16 scales around the least circumference of the caudal peduncle; last unbranched dorsal-fin ray markedly longer than head; mouth narrow, lower lip spatulate and median lobe present.</paragraph>
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||
</subSubSection>
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||
<subSubSection id="E923E0C6DAA61B16AA3B7EFD580707A9" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27" type="description">
|
||
<paragraph id="2FE1629AFEC75D54CD5D6A7E305148F1" pageId="26" pageNumber="27">Description.</paragraph>
|
||
<paragraph id="0AE2ED8D4D2E818BAE0CEC6ABAB9A6E9" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
|
||
Body moderately high, laterally compressed and without a nuchal hump. The greatest body depth is at the point of the origin of the dorsal fin. The ventral profile of the head is straight, its dorsal profile has a slight to pronounced hump
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||
<pageBreakToken id="645BBD9CB8190A95E0F5EF915E52BA58" pageId="27" pageNumber="28" start="start">near</pageBreakToken>
|
||
the nostrils (Figs 21, 22). The head is short and narrow. The mouth is inferior. The maximum body depth is bigger than the head length (Fig. 12). The lips are comparatively thick and the lower jaw is narrow with a sharp horny sheath and a median lobe. The two pairs of barbels (Table 2) are stout and the anterior pair is quite long. The eyes are rather high in the middle of the head and rather small. The morphometric characters are summarised in Table 1.
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||
</paragraph>
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||
<caption id="5E476D03FABDC85F3B3A29A1EE295A8E" pageId="27" pageNumber="28">
|
||
<paragraph id="DB16BE135E6AFFB0065B63BE533D268E" pageId="27" pageNumber="28">
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Figure 21.
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<taxonomicName id="CA0CB1CE4A9B580A3A4F5671EBF502DD" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus kosswigi" order="Cypriniformes" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="kosswigi">Carasobarbus kosswigi</taxonomicName>
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, holotype (ZMH 1148) from Batman
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<normalizedToken id="05C4EB260F4DA5B5AE1E86E56E8A4561" originalValue="Çayı">Cayi</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="8C53C85A45033C95B2D495DEE5FED8DA" pageId="27" pageNumber="28">
|
||
<paragraph id="07C7884E46899D2F94EE4B6017A124B0" pageId="27" pageNumber="28">
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Figure 22.
|
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<taxonomicName id="3C6FA9CD8A3C3C7D59A0B96B1D941ED7" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus kosswigi" order="Cypriniformes" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="kosswigi">Carasobarbus kosswigi</taxonomicName>
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, live specimen from
|
||
<normalizedToken id="BDA1E15792447756A9B23EAD6066DDC2" originalValue="Rūdkhāneh-ye">Rūdkhaneh-ye</normalizedToken>
|
||
Karkheh.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="71AF82763B32F81D32D5C6C6C2348E2F" pageId="27" pageNumber="28">The dorsal fin is long and usually has four unbranched and nine or 10 branched rays (Table 3). The last unbranched ray is long and well ossified; only the tip is flexible. It is considerably longer than the head (Fig. 4). The anal fin usually has three unbranched rays and six branched rays (Table 4). Its base is long. The bases of the dorsal and anal fin have a sheath of scales.</paragraph>
|
||
<paragraph id="502E8F2AA66AB7E7BF7F0E1FF33D8FAA" pageId="27" pageNumber="28">There are 32 to 38 scales in the lateral line (Table 5), 5.5 to seven scales above the lateral line (Table 6), 4.5 to 6.5 scales below the lateral line (Table 7) and (12) 14 to 16 scales around the least circumference of the caudal peduncle (Table 8). The scales are shown in Fig. 5.</paragraph>
|
||
<paragraph id="7D6A38A4D590F14343E92EE6CAEDE08B" pageId="28" pageNumber="29">
|
||
<pageBreakToken id="8E092FE8B7C7EB90DB3F5C8B71A1783C" pageId="28" pageNumber="29" start="start">The</pageBreakToken>
|
||
pharyngeal teeth count is 2.3.5-5.3.2 in seven specimens, 2.3.5- in one specimen and -4.3.2 in one specimen. The pharyngeal teeth are hooked at their tips (Fig. 6).
|
||
</paragraph>
|
||
<paragraph id="63D6FF7F3D5832A957456905DCE57EB5" pageId="28" pageNumber="29">Live specimens are silvery. The back is darker than the belly, which is almost white (Fig. 22). Fixed specimens are yellow-brown and some have a darker back.</paragraph>
|
||
<paragraph id="4C76AE920B5DE46FC347C7B3B0F8EC77" pageId="28" pageNumber="29">
|
||
<taxonomicName id="75D092CDAC63C672BF3B458CC9BF611D" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus kosswigi" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="kosswigi">Carasobarbus kosswigi</taxonomicName>
|
||
differs from all congeners, except
|
||
<taxonomicName id="8D2EAE2162F150AC281B73A83036FD7E" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus sublimus" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="sublimus">Carasobarbus sublimus</taxonomicName>
|
||
, by having a spatulate lower jaw with a median lobe on the lower lip vs. a crescent-shaped lower jaw and a lower lip without median lobe. It differs from
|
||
<taxonomicName id="9E04354F03191A4DC2680119814B4E68" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus sublimus" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="sublimus">Carasobarbus sublimus</taxonomicName>
|
||
by having 32 to 38 scales in the lateral line vs. 27 to 29 and modally 14 scales around the least circumference of the caudal peduncle vs. 12 and by having a longer and more ossified last unbranched ray in the dorsal fin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="857D1C5377EA559EDBEE811E0B3694E7" pageId="28" pageNumber="29" type="distribution">
|
||
<paragraph id="8FAFC59AD9DD47F519B45A9F2639F92B" pageId="28" pageNumber="29">Distribution.</paragraph>
|
||
<paragraph id="1389310C3B3CEAC0E725DE0B8DE474F6" pageId="28" pageNumber="29">
|
||
<taxonomicName id="227247F18F7122A63B65BE7EBFBC82A7" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus kosswigi" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="kosswigi">Carasobarbus kosswigi</taxonomicName>
|
||
occurs in the Euphrates-Tigris system (Fig. 7).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A09D22A6525539B2A9E56B40A464DDD5" pageId="28" pageNumber="29" type="habitats and biology">
|
||
<paragraph id="5B146523D467899EF323069C0835E27F" pageId="28" pageNumber="29">Habitats and biology.</paragraph>
|
||
<paragraph id="F0720F565103832A8A9DF701A84257D1" pageId="28" pageNumber="29">
|
||
<taxonomicName id="A1525E5D33857F8FC7034F82D8E0B377" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus kosswigi" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="kosswigi">Carasobarbus kosswigi</taxonomicName>
|
||
is rare, inhabits fast-flowing reaches of rivers and feeds on small animals (
|
||
<bibRefCitation id="5D789B7E306108F7B18D8CA00A932290" author="Krupp, F" editor="Kuehne, H" journalOrPublisher="Harrassowitz-Verlag, Wiesbaden" pageId="45" pageNumber="46" pagination="41 - 51" title="Die Fischfauna des Nahr al-Ḫabūr, Nordost-Syrien." year="2008">Krupp and Schneider 2008</bibRefCitation>
|
||
). The maximum length is about 150 mm SL and this species has no economic importance (
|
||
<bibRefCitation id="A2A5F60B1733871EA9946271B27BE529" author="Krupp, F" editor="Kuehne, H" journalOrPublisher="Harrassowitz-Verlag, Wiesbaden" pageId="45" pageNumber="46" pagination="41 - 51" title="Die Fischfauna des Nahr al-Ḫabūr, Nordost-Syrien." year="2008">Krupp and Schneider 2008</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0224C50982BE7CF5515499165B29C951" pageId="28" pageNumber="29" type="conservation status">
|
||
<paragraph id="620CCDB0CBE4C45E78E73484E2F4D2C2" pageId="28" pageNumber="29">Conservation status.</paragraph>
|
||
<paragraph id="4F356BC6E48B7507FC7664A12CB65C1E" pageId="28" pageNumber="29">Little information is available, but because this species is dependent on fast-flowing water, it is probably impacted by the construction of dams.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F4DCB8BFFF7F6345CA97FABFC80B3EBD" pageId="28" pageNumber="29" type="discussion">
|
||
<paragraph id="8F020EC22A41641687E9615955F7CE0E" pageId="28" pageNumber="29">Discussion.</paragraph>
|
||
<paragraph id="6C8084753DCBC32A84D75334AF07B397" pageId="28" pageNumber="29">
|
||
<taxonomicName id="D5C33EA3CB4D9807FA9DCECC0FE2F87A" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus kosswigi" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="kosswigi">Carasobarbus kosswigi</taxonomicName>
|
||
was described from the Batman
|
||
<normalizedToken id="323EFCFC9ACD335FEF9108DF9CCD267D" originalValue="Çayı">Cayi</normalizedToken>
|
||
and placed in the genus
|
||
<taxonomicName id="6D4DB4B4E9D6E73EC4780316D156C993" family="Cyprinidae" genus="Cyclocheilichthys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocheilichthys" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">Cyclocheilichthys</taxonomicName>
|
||
(
|
||
<bibRefCitation id="DED944FD09F9BCB40A09329870CCC336" author="Ladiges, W" journalOrPublisher="Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut" pageId="45" pageNumber="46" pagination="105 - 150" title="Suesswasserfische der Tuerkei. I. Teil Cyprinidae." volume="58" year="1960">Ladiges 1960</bibRefCitation>
|
||
). Karaman erected the new genus
|
||
<taxonomicName id="F244B5DD3461AC048FF6C9D5EE3DD16B" family="Cyprinidae" genus="Kosswigobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kosswigobarbus" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">Kosswigobarbus</taxonomicName>
|
||
for this species (
|
||
<bibRefCitation id="571A80B3585863EBA3B22607C492F1D1" author="Karaman, MS" journalOrPublisher="Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut" pageId="44" pageNumber="45" pagination="175 - 254" title="Suesswasserfische der Tuerkei. 8. Teil: Revision der Barben Europas, Vorderasiens und Nordafrikas." volume="67" year="1971">Karaman 1971</bibRefCitation>
|
||
). Coad gave a detailed re-description of this species and transferred it to the genus
|
||
<taxonomicName id="BDE203A98F4A41D0D782C938017F640D" family="Cyprinidae" genus="Barbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Barbus" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">Barbus</taxonomicName>
|
||
(
|
||
<bibRefCitation id="141AF74C7A2F126103B4F8A541CA15CA" author="Coad, BW" journalOrPublisher="Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut" pageId="42" pageNumber="43" pagination="263 - 265" title="A re-description and generic re-assignment of Kosswigobarbus kosswigi (Ladiges, 1960), a cyprinid fish from Turkey and Iran." volume="79" year="1982">Coad 1982</bibRefCitation>
|
||
).
|
||
<taxonomicName id="F7E7C5031DF76C3C42080279970B5756" family="Cyprinidae" genus="Kosswigobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kosswigobarbus" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">Kosswigobarbus</taxonomicName>
|
||
was revalidated (
|
||
<bibRefCitation id="B8779B4FE3DD51B3D850A7DC9BE325CB" author="Ekmekci, FG" journalOrPublisher="Folia Zoologica" pageId="43" pageNumber="44" pagination="87 - 96" title="A revision of the generic position of Barynotus (Systomus) verhoeffi, and the validity of the genera Carasobarbus, Kosswigobarbus and Mesopotamichthys (Pisces, Cyprinidae)." volume="47" year="1998">
|
||
<normalizedToken id="1A50C4FBDA98B5733BEFC4BA2B7ED869" originalValue="Ekmekçi">Ekmekci</normalizedToken>
|
||
and Banarescu 1998
|
||
</bibRefCitation>
|
||
) and sometimes used as a subgenus of
|
||
<taxonomicName id="781E5CE5A2BC06A3474068593B9F97CD" family="Cyprinidae" genus="Barbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Barbus" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">Barbus</taxonomicName>
|
||
(
|
||
<bibRefCitation id="451FC79BD649475CF275AB90F25839E2" author="Tsigenopoulos, CS" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="46" pageNumber="47" pagination="851 - 856" title="Phylogenetic relationships of hexaploid large-sized barbs (genus Labeobarbus, Cyprinidae) based on mtDNA data." url="10.1016/j.ympev.2010.02.006" volume="56" year="2010">Tsigenopoulos et al. 2010</bibRefCitation>
|
||
). Later the species was placed in
|
||
<taxonomicName id="3E2D702C694B4D9E16B106438A7B6758" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">Carasobarbus</taxonomicName>
|
||
(
|
||
<bibRefCitation id="179F6FC6B7C650AE034617B69CD4ECC0" author="Borkenhagen, K" journalOrPublisher="Environmental Biology of Fishes" pageId="42" pageNumber="43" pagination="327 - 335" title="The molecular systematics of the Carasobarbus species from Iran and adjacent areas, with comments on Carasobarbus albus (Heckel, 1843)." url="10.1007/s10641-011-9787-1" volume="91" year="2011">Borkenhagen et al. 2011</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="9659E079AE176D7ACBBEBC5E580180FD" pageId="28" pageNumber="29">
|
||
<taxonomicName id="D56985702A0523D1E6A992EE545FE1EB" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus kosswigi" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="kosswigi">Carasobarbus kosswigi</taxonomicName>
|
||
is paraphyletic with respect to
|
||
<taxonomicName id="6B543C0502C6D5B56EB110AB7969A195" family="Cyprinidae" genus="Carasobarbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carasobarbus sublimus" order="Cypriniformes" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="sublimus">Carasobarbus sublimus</taxonomicName>
|
||
(
|
||
<bibRefCitation id="C41F12A35CF8F5B7FACFA21F3989F100" author="Borkenhagen, K" journalOrPublisher="Environmental Biology of Fishes" pageId="42" pageNumber="43" pagination="327 - 335" title="The molecular systematics of the Carasobarbus species from Iran and adjacent areas, with comments on Carasobarbus albus (Heckel, 1843)." url="10.1007/s10641-011-9787-1" volume="91" year="2011">Borkenhagen et al. 2011</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |