treatments-xml/data/0B/24/F8/0B24F87CFFBEFFB1FB29B5C6CAA62016.xml
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<document ID-DOI="http://doi.org/10.5281/zenodo.4391208" ID-GBIF-Dataset="39c9ce8d-d95d-467e-9590-ab184d952201" ID-GBIF-Taxon="159879543" ID-Zenodo-Dep="4391208" checkinTime="1562074625476" checkinUser="jeremy" docAuthor="DeMarmels, Jurg" docDate="1992" docId="0B24F87CFFBEFFB1FB29B5C6CAA62016" docLanguage="en" docName="De Marmels - 1992 - Caballitos del diablo (Odonata) de las Sierras de Tapirapeco y Unturán, en el extremo Sur de Venezuela.pdf.imf" docOrigin="Acta Biologica Venezuelica 14 (1)" docStyle="DocumentStyle{}" docTitle="Metaleptobasis fernandezi Racenis 1955" docType="treatment" docVersion="12" lastPageId="7" lastPageNumber="63" masterDocId="F71D8004FFB8FFB6FFC7B11BC92F2226" masterDocTitle="Dragonflies (Odonata) From The Sierras Of Tapirapeco And Unturan, In The Extreme South Of Venezuela" masterLastPageNumber="78" masterPageNumber="57" pageId="6" pageNumber="63" updateTime="1632324435586" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Dragonflies (Odonata) From The Sierras Of Tapirapeco And Unturan, In The Extreme South Of Venezuela</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>DeMarmels, Jurg</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Acta Biologica Venezuelica</mods:title>
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<mods:part>
<mods:date>1992</mods:date>
<mods:detail type="volume">
<mods:number>14</mods:number>
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<mods:number>1</mods:number>
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<mods:identifier type="DOI">10.5281/zenodo.3266063</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4391208" ID-GBIF-Taxon="159879543" ID-Zenodo-Dep="4391208" LSID="urn:lsid:plazi:treatment:0B24F87CFFBEFFB1FB29B5C6CAA62016" httpUri="http://treatment.plazi.org/id/0B24F87CFFBEFFB1FB29B5C6CAA62016" lastPageId="7" lastPageNumber="63" pageId="6" pageNumber="63">
<subSubSection box="[1262,2063,1245,1283]" pageId="6" pageNumber="63" type="nomenclature">
<paragraph blockId="6.[1257,2220,417,3268]" box="[1262,2063,1245,1283]" pageId="6" pageNumber="63">
<taxonomicName authorityName="Racenis" authorityYear="1955" box="[1262,2063,1245,1283]" class="Insecta" family="Coenagrionidae" genus="Metaleptobasis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="fernandezi">
<emphasis box="[1262,1538,1245,1283]" italics="true" pageId="6" pageNumber="63">Metaleptobasis</emphasis>
<emphasis box="[1567,1764,1245,1283]" pageId="6" pageNumber="63">fernandezi</emphasis>
Racenis. 1955
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection box="[1300,1560,1297,1334]" pageId="6" pageNumber="63" type="description">
<paragraph blockId="6.[1257,2220,417,3268]" box="[1300,1560,1297,1334]" pageId="6" pageNumber="63">
<figureCitation box="[1300,1560,1297,1334]" captionStart="Plate V" captionStartId="17.[207,304,2852,2886]" captionTargetBox="[158,2205,437,2815]" captionTargetPageId="17" captionText="Plate V: Figures 31-35. Leptobasis yanomami sp.n. (31) abdominal segment 10 with anal appendages, holotype male, left lateral view; (32) same, dorsal view; (33) penis, paratype, left lateral view; (34) same, ventral view; (35) hind margin of pronotum, paratype female, dorsal view. Figure 36. Leptobasis mammiiuris, female (Venezuela, Bolivar, Canaima), hind margin of pronotum, dorsal view. Figures 37-40. Metaieptobasis fernandezi. (37) female, hind margin of pronotum, dorsal view; (38) male, abdominal segment 10 with anal appendages, dorsal view; (39) same, left lateral view; (40) male, left mesothoracic horn, left lateral view. Figures 41-45. Metaleptobasis mauritia (41) male (Trinidad, Valencia), abdominal sEgment 10 with anal appendages, dorsal view; (42) same, left lateral view; (43) left mesothoracic horn, left lateral view; (44) female (Venezuela, Amazonas, San Juan de Manapiare), hind margin of pronotum, dorsal view; (45) left mesothoracic horn, left lateral view, same specimen. Figure 46. Metaleptobasis manicaria, female (Venezuela, Delta Amacuro, Jotacuay), hind margin of pronotum, dorsal view." httpUri="https://zenodo.org/record/3266075/files/figure.png" pageId="6" pageNumber="63">Figures 37-40</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="63" type="materials_examined">
<paragraph blockId="6.[1257,2220,417,3268]" pageId="6" pageNumber="63">
Material. (6♂, 1♀).
<materialsCitation ID-GBIF-Occurrence="2433522117" box="[1718,2144,1361,1399]" collectingDate="1989-02-14" collectingDateMax="1989-03-06" collectingDateMin="1989-02-14" collectionCode="MIZA" collectorName="J. DeMarmels &amp; A. Chacon" country="Venezuela" elevation="160" latitude="2.036111" location="Base Camp and surroundings, on the upper Rio Mavaca, left bank" longitude="-65.113884" pageId="6" pageNumber="63" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Amazonas">
<specimenCount box="[1718,1764,1361,1390]" pageId="6" pageNumber="63" type="male">4♂</specimenCount>
,
<specimenCount box="[1812,1860,1362,1391]" pageId="6" pageNumber="63" type="female">1♀</specimenCount>
, Base Camp
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2433522149" box="[1261,1497,1412,1452]" collectingDate="1989-02-23" collectingDateMax="1989-02-25" collectingDateMin="1989-02-23" collectionCode="MIZA" collectorName="J. DeMarmels &amp; A. Chacon" country="Venezuela" elevation="500" latitude="1.7194445" location="Siapa 2 (upper Rio Siapa)" longitude="-64.530556" pageId="6" pageNumber="63" specimenCount="2" specimenCount-male="2" stateProvince="Amazonas">
<specimenCount box="[1261,1304,1412,1443]" pageId="6" pageNumber="63" type="male">2♂</specimenCount>
, Siapa 2.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="64" pageId="6" pageNumber="63" type="discussion">
<paragraph blockId="6.[1257,2220,417,3268]" pageId="6" pageNumber="63">
Remarks.
<taxonomicName authority="Calvert" authorityName="Calvert" authorityYear="1907" box="[1504,1950,1481,1519]" class="Insecta" family="Coenagrionidae" genus="Metaleptobasis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis box="[1504,1785,1481,1519]" italics="true" pageId="6" pageNumber="63">Metaleptobasis</emphasis>
Calvert
</taxonomicName>
is a difficult genus, chiefly because the amplitude of intraspe. cific variability of most species is unknown.
<emphasis box="[2165,2208,1587,1624]" italics="true" pageId="6" pageNumber="63">M</emphasis>
.
<emphasis box="[1260,1455,1639,1677]" pageId="6" pageNumber="63">fernandezi</emphasis>
was described from a single female (
<bibRefCitation author="RACENIS, J." box="[1276,1544,1692,1729]" journalOrPublisher="Boletin del Museo de Ciencias Naturales, Caracas" pageId="6" pageNumber="63" pagination="3 - 20" part="1" refId="ref10621" refString="RACENIS, J. 1955 - El genero Metaleptobasis (Odonata: Coenagrionidae) en Venezuela. Boletin del Museo de Ciencias Naturales, Caracas, 1 (1): 3 - 20." title="El genero Metaleptobasis (Odonata: Coenagrionidae) en Venezuela" type="journal article" year="1955">Racenis, 1955</bibRefCitation>
). The prothoracic hind lobe of the present female (
<figureCitation box="[1584,1724,1744,1781]" captionStart="Plate V" captionStartId="17.[207,304,2852,2886]" captionTargetBox="[158,2205,437,2815]" captionTargetPageId="17" captionText="Plate V: Figures 31-35. Leptobasis yanomami sp.n. (31) abdominal segment 10 with anal appendages, holotype male, left lateral view; (32) same, dorsal view; (33) penis, paratype, left lateral view; (34) same, ventral view; (35) hind margin of pronotum, paratype female, dorsal view. Figure 36. Leptobasis mammiiuris, female (Venezuela, Bolivar, Canaima), hind margin of pronotum, dorsal view. Figures 37-40. Metaieptobasis fernandezi. (37) female, hind margin of pronotum, dorsal view; (38) male, abdominal segment 10 with anal appendages, dorsal view; (39) same, left lateral view; (40) male, left mesothoracic horn, left lateral view. Figures 41-45. Metaleptobasis mauritia (41) male (Trinidad, Valencia), abdominal sEgment 10 with anal appendages, dorsal view; (42) same, left lateral view; (43) left mesothoracic horn, left lateral view; (44) female (Venezuela, Amazonas, San Juan de Manapiare), hind margin of pronotum, dorsal view; (45) left mesothoracic horn, left lateral view, same specimen. Figure 46. Metaleptobasis manicaria, female (Venezuela, Delta Amacuro, Jotacuay), hind margin of pronotum, dorsal view." httpUri="https://zenodo.org/record/3266075/files/figure.png" pageId="6" pageNumber="63">Fig. 37</figureCitation>
) corresponds well with the
<typeStatus box="[1339,1501,1797,1834]" pageId="6" pageNumber="63">holotype</typeStatus>
and with the figure given by Geijs. kes (1932, sub
<emphasis box="[1558,1749,1850,1888]" italics="true" pageId="6" pageNumber="63">
<taxonomicName authorityName="Williamson" authorityYear="1915" box="[1558,1743,1850,1888]" class="Insecta" family="Coenagrionidae" genus="Metaleptobasis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="manicaria">manicaria</taxonomicName>
,
</emphasis>
see
<bibRefCitation author="RACENIS, J." box="[1844,2111,1850,1887]" journalOrPublisher="Boletin del Museo de Ciencias Naturales, Caracas" pageId="6" pageNumber="63" pagination="3 - 20" part="1" refId="ref10621" refString="RACENIS, J. 1955 - El genero Metaleptobasis (Odonata: Coenagrionidae) en Venezuela. Boletin del Museo de Ciencias Naturales, Caracas, 1 (1): 3 - 20." title="El genero Metaleptobasis (Odonata: Coenagrionidae) en Venezuela" type="journal article" year="1955">Racenis, 1955</bibRefCitation>
). The mesothoracic horns are reduced to mere tuber. cles. The thoracic middorsal stripe is slightly broader in the female from Base Camp than in the
<typeStatus box="[1341,1425,2062,2099]" pageId="6" pageNumber="63">type</typeStatus>
. With some doubts I am referring also the sixmales to
<taxonomicName authorityName="Racenis" authorityYear="1955" box="[1572,1771,2115,2153]" class="Insecta" family="Coenagrionidae" genus="Metaleptobasis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="fernandezi">iernandezi</taxonomicName>
. The male of this species has not so far been described. The present specimens are exceedingly similar to
<taxonomicName authority="Calvert" authorityName="Calvert" authorityYear="1907" box="[1989,2038,2220,2257]" class="Insecta" family="Coenagrionidae" genus="Metaleptobasis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis box="[1989,2032,2220,2257]" italics="true" pageId="6" pageNumber="63">M</emphasis>
.
</taxonomicName>
<taxonomicName authority="Williamson" authorityName="Williamson" authorityYear="1915" class="Insecta" family="Coenagrionidae" genus="Metaleptobasis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="mauritia">
<emphasis box="[2062,2218,2220,2258]" pageId="6" pageNumber="63">mauritia</emphasis>
Williamson
</taxonomicName>
, but differ from that species (only 1♂ compared: Valencia, Trinidad,
<date box="[1863,2086,2325,2362]" pageId="6" pageNumber="63" value="1965-04-07">7 Apr 1965</date>
, T. W. Donnelly leg.) in the superior anal appendages:
<taxonomicName authorityName="Williamson" authorityYear="1915" box="[1264,1484,2431,2469]" class="Insecta" family="Coenagrionidae" genus="Metaleptobasis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="mauritia">In mauritia</taxonomicName>
they are slightly narrower and less sharply angled internally in the apical third; the tip is clearly marked- off from the internal margin, and is reddish (
<figureCitation box="[1678,1897,2590,2627]" captionStart="Plate V" captionStartId="17.[207,304,2852,2886]" captionTargetBox="[158,2205,437,2815]" captionTargetPageId="17" captionText="Plate V: Figures 31-35. Leptobasis yanomami sp.n. (31) abdominal segment 10 with anal appendages, holotype male, left lateral view; (32) same, dorsal view; (33) penis, paratype, left lateral view; (34) same, ventral view; (35) hind margin of pronotum, paratype female, dorsal view. Figure 36. Leptobasis mammiiuris, female (Venezuela, Bolivar, Canaima), hind margin of pronotum, dorsal view. Figures 37-40. Metaieptobasis fernandezi. (37) female, hind margin of pronotum, dorsal view; (38) male, abdominal segment 10 with anal appendages, dorsal view; (39) same, left lateral view; (40) male, left mesothoracic horn, left lateral view. Figures 41-45. Metaleptobasis mauritia (41) male (Trinidad, Valencia), abdominal sEgment 10 with anal appendages, dorsal view; (42) same, left lateral view; (43) left mesothoracic horn, left lateral view; (44) female (Venezuela, Amazonas, San Juan de Manapiare), hind margin of pronotum, dorsal view; (45) left mesothoracic horn, left lateral view, same specimen. Figure 46. Metaleptobasis manicaria, female (Venezuela, Delta Amacuro, Jotacuay), hind margin of pronotum, dorsal view." httpUri="https://zenodo.org/record/3266075/files/figure.png" pageId="6" pageNumber="63">Figs. 41-42</figureCitation>
). In
<taxonomicName authorityName="Racenis" authorityYear="1955" box="[2022,2217,2590,2628]" class="Insecta" family="Coenagrionidae" genus="Metaleptobasis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="fernandezi">
<emphasis box="[2022,2217,2590,2628]" italics="true" pageId="6" pageNumber="63">fernandezi</emphasis>
</taxonomicName>
the superior appendages are slightly broader and more angled internally in the apical third; the transition between internal margin and tip is smooth, the tip dark brown (
<figureCitation box="[1877,2102,2801,2838]" captionStart="Plate V" captionStartId="17.[207,304,2852,2886]" captionTargetBox="[158,2205,437,2815]" captionTargetPageId="17" captionText="Plate V: Figures 31-35. Leptobasis yanomami sp.n. (31) abdominal segment 10 with anal appendages, holotype male, left lateral view; (32) same, dorsal view; (33) penis, paratype, left lateral view; (34) same, ventral view; (35) hind margin of pronotum, paratype female, dorsal view. Figure 36. Leptobasis mammiiuris, female (Venezuela, Bolivar, Canaima), hind margin of pronotum, dorsal view. Figures 37-40. Metaieptobasis fernandezi. (37) female, hind margin of pronotum, dorsal view; (38) male, abdominal segment 10 with anal appendages, dorsal view; (39) same, left lateral view; (40) male, left mesothoracic horn, left lateral view. Figures 41-45. Metaleptobasis mauritia (41) male (Trinidad, Valencia), abdominal sEgment 10 with anal appendages, dorsal view; (42) same, left lateral view; (43) left mesothoracic horn, left lateral view; (44) female (Venezuela, Amazonas, San Juan de Manapiare), hind margin of pronotum, dorsal view; (45) left mesothoracic horn, left lateral view, same specimen. Figure 46. Metaleptobasis manicaria, female (Venezuela, Delta Amacuro, Jotacuay), hind margin of pronotum, dorsal view." httpUri="https://zenodo.org/record/3266075/files/figure.png" pageId="6" pageNumber="63">Figs. 38-39</figureCitation>
). The hind border of the pronotum has a metallic green spot in the middle in male
<taxonomicName authorityName="Williamson" authorityYear="1915" box="[1797,1958,2907,2945]" class="Insecta" family="Coenagrionidae" genus="Metaleptobasis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="mauritia">mauritia</taxonomicName>
, which is absent in
<emphasis box="[1407,1613,2960,2998]" italics="true" pageId="6" pageNumber="63">
<taxonomicName authorityName="Racenis" authorityYear="1955" box="[1407,1606,2960,2998]" class="Insecta" family="Coenagrionidae" genus="Metaleptobasis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="fernandezi">fernandezi</taxonomicName>
.
</emphasis>
The mesothoracic horns are similarly shaped in both species (males) (
<figureCitation captionStart="Plate V" captionStartId="17.[207,304,2852,2886]" captionTargetBox="[158,2205,437,2815]" captionTargetPageId="17" captionText="Plate V: Figures 31-35. Leptobasis yanomami sp.n. (31) abdominal segment 10 with anal appendages, holotype male, left lateral view; (32) same, dorsal view; (33) penis, paratype, left lateral view; (34) same, ventral view; (35) hind margin of pronotum, paratype female, dorsal view. Figure 36. Leptobasis mammiiuris, female (Venezuela, Bolivar, Canaima), hind margin of pronotum, dorsal view. Figures 37-40. Metaieptobasis fernandezi. (37) female, hind margin of pronotum, dorsal view; (38) male, abdominal segment 10 with anal appendages, dorsal view; (39) same, left lateral view; (40) male, left mesothoracic horn, left lateral view. Figures 41-45. Metaleptobasis mauritia (41) male (Trinidad, Valencia), abdominal sEgment 10 with anal appendages, dorsal view; (42) same, left lateral view; (43) left mesothoracic horn, left lateral view; (44) female (Venezuela, Amazonas, San Juan de Manapiare), hind margin of pronotum, dorsal view; (45) left mesothoracic horn, left lateral view, same specimen. Figure 46. Metaleptobasis manicaria, female (Venezuela, Delta Amacuro, Jotacuay), hind margin of pronotum, dorsal view." httpUri="https://zenodo.org/record/3266075/files/figure.png" pageId="6" pageNumber="63">Figs. 40. 43</figureCitation>
).
</paragraph>
<paragraph blockId="6.[1257,2220,417,3268]" lastBlockId="7.[203,1166,417,3269]" lastPageId="7" lastPageNumber="64" pageId="6" pageNumber="63">
Female
<taxonomicName box="[1456,1615,3125,3163]" class="Insecta" family="Calopterygidae" genus="Iridictyon" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="mauritia">
<emphasis box="[1456,1615,3125,3163]" italics="true" pageId="6" pageNumber="63">mauritia</emphasis>
</taxonomicName>
have a differently shaped prothoracic hind lobe (
<figureCitation box="[1648,1782,3178,3215]" captionStart="Plate V" captionStartId="17.[207,304,2852,2886]" captionTargetBox="[158,2205,437,2815]" captionTargetPageId="17" captionText="Plate V: Figures 31-35. Leptobasis yanomami sp.n. (31) abdominal segment 10 with anal appendages, holotype male, left lateral view; (32) same, dorsal view; (33) penis, paratype, left lateral view; (34) same, ventral view; (35) hind margin of pronotum, paratype female, dorsal view. Figure 36. Leptobasis mammiiuris, female (Venezuela, Bolivar, Canaima), hind margin of pronotum, dorsal view. Figures 37-40. Metaieptobasis fernandezi. (37) female, hind margin of pronotum, dorsal view; (38) male, abdominal segment 10 with anal appendages, dorsal view; (39) same, left lateral view; (40) male, left mesothoracic horn, left lateral view. Figures 41-45. Metaleptobasis mauritia (41) male (Trinidad, Valencia), abdominal sEgment 10 with anal appendages, dorsal view; (42) same, left lateral view; (43) left mesothoracic horn, left lateral view; (44) female (Venezuela, Amazonas, San Juan de Manapiare), hind margin of pronotum, dorsal view; (45) left mesothoracic horn, left lateral view, same specimen. Figure 46. Metaleptobasis manicaria, female (Venezuela, Delta Amacuro, Jotacuay), hind margin of pronotum, dorsal view." httpUri="https://zenodo.org/record/3266075/files/figure.png" pageId="6" pageNumber="63">Fig. 44</figureCitation>
) and developed mesothoracic horns (
<figureCitation box="[1571,1711,3231,3268]" captionStart="Plate V" captionStartId="17.[207,304,2852,2886]" captionTargetBox="[158,2205,437,2815]" captionTargetPageId="17" captionText="Plate V: Figures 31-35. Leptobasis yanomami sp.n. (31) abdominal segment 10 with anal appendages, holotype male, left lateral view; (32) same, dorsal view; (33) penis, paratype, left lateral view; (34) same, ventral view; (35) hind margin of pronotum, paratype female, dorsal view. Figure 36. Leptobasis mammiiuris, female (Venezuela, Bolivar, Canaima), hind margin of pronotum, dorsal view. Figures 37-40. Metaieptobasis fernandezi. (37) female, hind margin of pronotum, dorsal view; (38) male, abdominal segment 10 with anal appendages, dorsal view; (39) same, left lateral view; (40) male, left mesothoracic horn, left lateral view. Figures 41-45. Metaleptobasis mauritia (41) male (Trinidad, Valencia), abdominal sEgment 10 with anal appendages, dorsal view; (42) same, left lateral view; (43) left mesothoracic horn, left lateral view; (44) female (Venezuela, Amazonas, San Juan de Manapiare), hind margin of pronotum, dorsal view; (45) left mesothoracic horn, left lateral view, same specimen. Figure 46. Metaleptobasis manicaria, female (Venezuela, Delta Amacuro, Jotacuay), hind margin of pronotum, dorsal view." httpUri="https://zenodo.org/record/3266075/files/figure.png" pageId="6" pageNumber="63">Fig. 45</figureCitation>
). The specimens from Base Camp were collected together with
<taxonomicName authorityName="Williamson" authorityYear="1915" class="Insecta" family="Coenagrionidae" genus="Metaleptobasis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="brysonima">
<emphasis box="[938,983,417,454]" italics="true" pageId="7" pageNumber="64">M</emphasis>
.
<emphasis box="[1008,1156,417,454]" pageId="7" pageNumber="64">
<emphasis box="[205,261,469,508]" italics="true" pageId="7" pageNumber="64">brysonima</emphasis>
</emphasis>
</taxonomicName>
<emphasis box="[260,266,469,508]" italics="true" pageId="7" pageNumber="64">,</emphasis>
but the two males from Siapa 2 were taken in a (then) dry hollow in the forest.
</paragraph>
</subSubSection>
</treatment>
</document>