170 lines
16 KiB
XML
170 lines
16 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.4391168" ID-GBIF-Dataset="39c9ce8d-d95d-467e-9590-ab184d952201" ID-GBIF-Taxon="159879578" ID-Zenodo-Dep="4391168" checkinTime="1562074625476" checkinUser="jeremy" docAuthor="DeMarmels, Jurg" docDate="1992" docId="0B24F87CFFBBFFB2FB3EB747C8CF24C4" docLanguage="en" docName="De Marmels - 1992 - Caballitos del diablo (Odonata) de las Sierras de Tapirapeco y Unturán, en el extremo Sur de Venezuela.pdf.imf" docOrigin="Acta Biologica Venezuelica 14 (1)" docStyle="DocumentStyle{}" docTitle="Lestes debellardi DeMarmels 1992, sp. n." docType="treatment" docVersion="11" lastPageId="4" lastPageNumber="61" masterDocId="F71D8004FFB8FFB6FFC7B11BC92F2226" masterDocTitle="Dragonflies (Odonata) From The Sierras Of Tapirapeco And Unturan, In The Extreme South Of Venezuela" masterLastPageNumber="78" masterPageNumber="57" pageId="3" pageNumber="60" updateTime="1632324435586" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Dragonflies (Odonata) From The Sierras Of Tapirapeco And Unturan, In The Extreme South Of Venezuela</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>DeMarmels, Jurg</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Acta Biologica Venezuelica</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>1992</mods:date>
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<mods:detail type="volume">
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<mods:number>14</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>57</mods:start>
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<mods:end>78</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5281/zenodo.3266063</mods:identifier>
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<mods:identifier type="GBIF-Dataset">39c9ce8d-d95d-467e-9590-ab184d952201</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3266063</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4391168" ID-GBIF-Taxon="159879578" ID-Zenodo-Dep="4391168" LSID="urn:lsid:plazi:treatment:0B24F87CFFBBFFB2FB3EB747C8CF24C4" httpUri="http://treatment.plazi.org/id/0B24F87CFFBBFFB2FB3EB747C8CF24C4" lastPageId="4" lastPageNumber="61" pageId="3" pageNumber="60">
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<subSubSection box="[1273,1708,1628,1665]" pageId="3" pageNumber="60" type="nomenclature">
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<paragraph blockId="3.[1271,2235,414,3269]" box="[1273,1708,1628,1665]" pageId="3" pageNumber="60">
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<taxonomicName authority="DeMarmels, 1992" authorityName="DeMarmels" authorityYear="1992" box="[1273,1592,1628,1665]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="debellardi" status="sp. n.">
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<emphasis box="[1273,1592,1628,1665]" italics="true" pageId="3" pageNumber="60">Lestes debellardi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1609,1708,1628,1665]" pageId="3" pageNumber="60" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection box="[1320,1580,1680,1717]" pageId="3" pageNumber="60" type="description">
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<paragraph blockId="3.[1271,2235,414,3269]" box="[1320,1580,1680,1717]" pageId="3" pageNumber="60">
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<figureCitation box="[1320,1580,1680,1717]" captionStart="plate III" captionStartId="15.[224,319,3031,3065]" captionTargetBox="[242,2213,424,2984]" captionTargetPageId="15" captionText="plate III: Figures 15-19. Lestes debellardi sp.n. (15) holotype male, pectoral color pattern, ventral view; (16) segment 10 with anal appendages, same specimen, left lateral view; (17) same, dorsal view; (18) penis of paratype, right lateral view; (19) same, ventral view. Figures 20-22. Epipleoneura lamina. (20) penis, right lateral view. (21) same, ventral view; (22) hind margin of pronotum of female taken in tandem, dorsal view. Figure 22a. Neoneura denticulata (?). hind margin of pronotum (female), dorsal view. Figures 23-24. Neoneura desana, female taken in tandem. (23) color pattern of head, dorsal view; (24) hind margin of pronotum, dorsal view" httpUri="https://zenodo.org/record/3266071/files/figure.png" pageId="3" pageNumber="60">Figures 15-19</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="60" type="materials_examined">
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<paragraph blockId="3.[1271,2235,414,3269]" pageId="3" pageNumber="60">
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<materialsCitation ID-GBIF-Occurrence="2433522096" collectingDate="1989-03-01" collectingDateMax="1989-03-06" collectingDateMin="1989-03-01" collectionCode="MIZA" collectorName="J. DeMarmels" country="Venezuela" elevation="160" latitude="2.036111" location="upper Rio Mavaca" longitude="-65.113884" pageId="3" pageNumber="60" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas" typeStatus="holotype">
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<specimenCount box="[1319,1414,1740,1777]" pageId="3" pageNumber="60" type="male">Male</specimenCount>
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(
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<typeStatus box="[1460,1627,1740,1777]" pageId="3" pageNumber="60">holotype</typeStatus>
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).
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<collectingCountry box="[1677,1868,1740,1777]" name="Venezuela" pageId="3" pageNumber="60" value="Venezuela">Venezuela</collectingCountry>
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,
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<collectingRegion box="[1908,2223,1740,1777]" pageId="3" pageNumber="60">Amazonas State</collectingRegion>
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, upper
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<collectingMunicipality box="[1407,1645,1793,1830]" pageId="3" pageNumber="60">Rio Mavaca</collectingMunicipality>
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,
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<elevation box="[1677,1802,1793,1830]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="60" unit="m" value="160.0">
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<quantity box="[1677,1802,1793,1830]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="60" unit="m" value="160.0">160 m</quantity>
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</elevation>
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,
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<geoCoordinate box="[1835,2071,1793,1830]" direction="north" orientation="latitude" pageId="3" pageNumber="60" precision="15" value="2.036111">02°02'10" N</geoCoordinate>
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, 65°06'- 50"W,
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<collectingDate box="[1412,1677,1846,1883]" pageId="3" pageNumber="60" value="1989-03-01" valueMax="1989-03-06" valueMin="1989-03-01">
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<date box="[1412,1677,1846,1883]" pageId="3" pageNumber="60" value="1989-03-01" valueMax="1989-03-06" valueMin="1989-03-01">1-6 Mar 1989</date>
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</collectingDate>
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,
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<collectorName box="[1700,1970,1846,1883]" pageId="3" pageNumber="60">J. DeMarmels</collectorName>
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leg
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</materialsCitation>
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.
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</paragraph>
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<paragraph blockId="3.[1271,2235,414,3269]" pageId="3" pageNumber="60">
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Material (3♂, including
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<typeStatus box="[1814,1975,1906,1943]" pageId="3" pageNumber="60">holotype</typeStatus>
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and
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<typeStatus pageId="3" pageNumber="60">paratypes</typeStatus>
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).
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<materialsCitation ID-GBIF-Occurrence="2433522110" collectingDate="1982-11-01" collectingDateMax="1982-11-03" collectingDateMin="1982-11-01" collectionCode="MIZA" collectorName="E. Rubio & T. Borrego" country="Venezuela" location="km 85 road El Dorado Santa Elena de Uairen" pageId="3" pageNumber="60" specimenCount="1" specimenCount-male="1" stateProvince="Bolivar" typeStatus="paratype">
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<specimenCount box="[1381,1425,1957,1988]" pageId="3" pageNumber="60" type="male">1♂</specimenCount>
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.
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<typeStatus box="[1463,1630,1959,1996]" pageId="3" pageNumber="60">paratype</typeStatus>
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,
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<locationDeviation box="[1654,2094,1959,1996]" pageId="3" pageNumber="60">km 85 road El Dorado</locationDeviation>
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- Santa Elena de
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<collectingMunicipality box="[1459,1588,2012,2049]" pageId="3" pageNumber="60">Uairen</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:0B24F87CFFBBFFB2FB3EB747C8CF24C4:86521FB1FFBBFFB5F988B6C7CFF62A27" box="[1615,1753,2012,2049]" country="Venezuela" municipality="Uairen" name="Bolivar" pageId="3" pageNumber="60">Bolivar</location>
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,
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<collectingDate box="[1780,2042,2012,2049]" pageId="3" pageNumber="60" value="1982-11-01" valueMax="1982-11-03" valueMin="1982-11-01">
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<date box="[1780,2042,2012,2049]" pageId="3" pageNumber="60" value="1982-11-01" valueMax="1982-11-03" valueMin="1982-11-01">1-3 Nov 1982</date>
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</collectingDate>
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,
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<collectorName pageId="3" pageNumber="60">E. Rubio & T. Borrego</collectorName>
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leg.
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</materialsCitation>
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:
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<materialsCitation ID-GBIF-Occurrence="2433522079" collectingDate="1985-05-22" collectionCode="MIZA" collectorName="F. Fernandez" country="Venezuela" location="km 35 road El Dorado Santa Elena de Uairen" pageId="3" pageNumber="60" specimenCount="1" specimenCount-male="1" stateProvince="Bolivar" typeStatus="paratype">
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<specimenCount box="[1669,1712,2064,2095]" pageId="3" pageNumber="60" type="male">1♂</specimenCount>
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,
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<typeStatus box="[1765,1933,2065,2102]" pageId="3" pageNumber="60">paratype</typeStatus>
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,
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<locationDeviation pageId="3" pageNumber="60">km 33 (same road</locationDeviation>
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),
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<collectingDate box="[1406,1666,2118,2155]" pageId="3" pageNumber="60" value="1985-05-22">
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<date box="[1406,1666,2118,2155]" pageId="3" pageNumber="60" value="1985-05-22">22 May 1985</date>
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</collectingDate>
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,
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<collectorName box="[1692,1941,2118,2155]" pageId="3" pageNumber="60">F. Fernandez</collectorName>
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leg
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="61" pageId="3" pageNumber="60" type="description">
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<paragraph blockId="3.[1271,2235,414,3269]" lastBlockId="4.[183,1147,411,2498]" lastPageId="4" lastPageNumber="61" pageId="3" pageNumber="60">
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Male (
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<typeStatus box="[1448,1615,2178,2215]" pageId="3" pageNumber="60">holotype</typeStatus>
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). Labium and rear of head pale, labrum and mandibles sky blue, top of head black with a metallic green luster. Compound eyes (in life) blue above, yellow green below, Pronotum laterally blue, dorsally predominantly brown. Mesepisternum brassy black (brown along median carina), with a strongly obscured green stripe along humeral suture, this stripe less than half as broad as dark portion of mesepisternum. Thorax laterally sky blue, pectoral color pattern as in
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<figureCitation box="[1397,1539,2705,2742]" captionStart="plate III" captionStartId="15.[224,319,3031,3065]" captionTargetBox="[242,2213,424,2984]" captionTargetPageId="15" captionText="plate III: Figures 15-19. Lestes debellardi sp.n. (15) holotype male, pectoral color pattern, ventral view; (16) segment 10 with anal appendages, same specimen, left lateral view; (17) same, dorsal view; (18) penis of paratype, right lateral view; (19) same, ventral view. Figures 20-22. Epipleoneura lamina. (20) penis, right lateral view. (21) same, ventral view; (22) hind margin of pronotum of female taken in tandem, dorsal view. Figure 22a. Neoneura denticulata (?). hind margin of pronotum (female), dorsal view. Figures 23-24. Neoneura desana, female taken in tandem. (23) color pattern of head, dorsal view; (24) hind margin of pronotum, dorsal view" httpUri="https://zenodo.org/record/3266071/files/figure.png" pageId="3" pageNumber="60">Fig. 15</figureCitation>
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. Coxae blue. femora brown with a black external line; tibiae pale blue externally, creamy internally with black longitudinal line. Wings hyaline. venation black, pterostigma dark brown. Three wings with 13 px, left hind wing 14. R3 branching between third and fourth px (fore wings), between second and third (hind wings). Abdomen dorsally brassy black, segments 1-4 and 8 laterally sky blue; segments 3-7 with basal ring of same color, otherwise brown laterally. Segment 9 black with a characteristic pruinescent white basal mark on top; segment 10 black. Anal appendages black, the superiors without
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<emphasis box="[1094,1136,464,500]" italics="true" pageId="4" pageNumber="61">in</emphasis>
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· ternal basal spine; inferiors about half as long as superiors, in their apical portion wiht a brush of stiff yellow hairs (
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<figureCitation box="[549,767,623,659]" captionStart="plate III" captionStartId="15.[224,319,3031,3065]" captionTargetBox="[242,2213,424,2984]" captionTargetPageId="15" captionText="plate III: Figures 15-19. Lestes debellardi sp.n. (15) holotype male, pectoral color pattern, ventral view; (16) segment 10 with anal appendages, same specimen, left lateral view; (17) same, dorsal view; (18) penis of paratype, right lateral view; (19) same, ventral view. Figures 20-22. Epipleoneura lamina. (20) penis, right lateral view. (21) same, ventral view; (22) hind margin of pronotum of female taken in tandem, dorsal view. Figure 22a. Neoneura denticulata (?). hind margin of pronotum (female), dorsal view. Figures 23-24. Neoneura desana, female taken in tandem. (23) color pattern of head, dorsal view; (24) hind margin of pronotum, dorsal view" httpUri="https://zenodo.org/record/3266071/files/figure.png" pageId="4" pageNumber="61">Figs. 16-17</figureCitation>
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).
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</paragraph>
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<paragraph blockId="4.[183,1147,411,2498]" pageId="4" pageNumber="61">Measurements. Total length (excl. anal appendages) 46.6; abdomen (excl. app.) 37.0; anal appendages 1.4; hind wing 24.5.</paragraph>
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<paragraph blockId="4.[183,1147,411,2498]" pageId="4" pageNumber="61">
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The
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<typeStatus box="[321,503,858,894]" pageId="4" pageNumber="61">paratypes</typeStatus>
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are similar to the
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<typeStatus box="[891,1055,858,894]" pageId="4" pageNumber="61">holotype</typeStatus>
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, but with their color less well preserved. Fore wings with 12-15 px, hind wings with 12-14. R, branching slightly before to considerably after fourth px. Size similar, the male from km 33 some· what smaller (total length 44). Penis of this male illustrated in
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<figureCitation box="[451,757,1175,1211]" captionStart="plate III" captionStartId="15.[224,319,3031,3065]" captionTargetBox="[242,2213,424,2984]" captionTargetPageId="15" captionText="plate III: Figures 15-19. Lestes debellardi sp.n. (15) holotype male, pectoral color pattern, ventral view; (16) segment 10 with anal appendages, same specimen, left lateral view; (17) same, dorsal view; (18) penis of paratype, right lateral view; (19) same, ventral view. Figures 20-22. Epipleoneura lamina. (20) penis, right lateral view. (21) same, ventral view; (22) hind margin of pronotum of female taken in tandem, dorsal view. Figure 22a. Neoneura denticulata (?). hind margin of pronotum (female), dorsal view. Figures 23-24. Neoneura desana, female taken in tandem. (23) color pattern of head, dorsal view; (24) hind margin of pronotum, dorsal view" httpUri="https://zenodo.org/record/3266071/files/figure.png" pageId="4" pageNumber="61">Figs. 18 and 19</figureCitation>
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.
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</paragraph>
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<paragraph blockId="4.[183,1147,411,2498]" box="[232,581,1239,1275]" pageId="4" pageNumber="61">Female. Unknown</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="61" type="discussion">
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<paragraph blockId="4.[183,1147,411,2498]" pageId="4" pageNumber="61">
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Remarks.
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<taxonomicName authorityName="DeMarmels" authorityYear="1992" box="[427,748,1303,1340]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="debellardi">
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<emphasis box="[427,748,1303,1340]" italics="true" pageId="4" pageNumber="61">Lestes debellardi</emphasis>
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</taxonomicName>
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is similar to
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<taxonomicName authority="Hagen in Selys" authorityName="Hagen in Selys" authorityYear="1862" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="pictus">
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<emphasis italics="true" pageId="4" pageNumber="61">L. pictus</emphasis>
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Hagen in Selys
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</taxonomicName>
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and
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<taxonomicName authority="Sjostedt." authorityName="Sjostedt." box="[678,1051,1355,1392]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="falcifer">
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<emphasis box="[678,869,1355,1392]" italics="true" pageId="4" pageNumber="61">L. falcifer</emphasis>
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Sjostedt.
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</taxonomicName>
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The latter two have, however, differently shaped inferior appendages and a diverging color pattern. The
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<typeStatus box="[188,349,1516,1552]" pageId="4" pageNumber="61">holotype</typeStatus>
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was taken in the forest, away from water, together with some
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<taxonomicName box="[569,755,1567,1604]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="falcifer">
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<emphasis box="[569,755,1567,1604]" italics="true" pageId="4" pageNumber="61">L. falcifer</emphasis>
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</taxonomicName>
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. I dedicate this species to Dr. Eugenio de Bellard Pietri, Caracas, the organizer of the expedition. who allowed me to take part in it.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |