treatments-xml/data/01/4B/E7/014BE744FFE2A82CFF4B45AA5BCDFDA3.xml

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<mods:title id="F5A92CB898C35F9BD29FA47750F1E53C">On the genus Tenagogonus Stål (Hemiptera: Heteroptera: Gerridae) from India with a description of a new species</mods:title>
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<mods:namePart id="AFE7629390B57808C9375C5C7D7A8F61">Jehamalar, E. Eyarin</mods:namePart>
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<mods:namePart id="B31CD2AB9D2DD2FB2F759CC849F5D76A">Chandra, Kailash</mods:namePart>
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<paragraph id="895D5652FFE2A825FF4B45AA5DBFFF68" blockId="1.[151,765,221,279]" box="[151,765,221,247]" pageId="1" pageNumber="379">
<heading id="D215E13EFFE2A825FF4B45AA5DBFFF68" bold="true" box="[151,765,221,247]" fontSize="11" level="1" pageId="1" pageNumber="379" reason="1">
<taxonomicName id="4EE22DD1FFE2A825FF4B45AA5DBFFF68" ID-CoL="7C35C" authority="Hungerford &amp; Matsuda" authorityName="Hungerford &amp; Matsuda" box="[151,765,221,247]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="379" phylum="Arthropoda" rank="species" species="ceylonensis">
<emphasis id="BB968A40FFE2A825FF4B45AA5DBFFF68" bold="true" box="[151,765,221,247]" pageId="1" pageNumber="379">
<emphasis id="BB968A40FFE2A825FF4B45AA5E88FF68" bold="true" box="[151,458,221,247]" italics="true" pageId="1" pageNumber="379">Tenagogonus ceylonensis</emphasis>
Hungerford &amp; Matsuda
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</paragraph>
<paragraph id="895D5652FFE2A825FF4B45885EE5FE88" blockId="1.[151,765,221,279]" box="[151,423,255,279]" pageId="1" pageNumber="379">
<figureCitation id="11D94AD7FFE2A825FF4B45885FA3FE88" box="[151,225,255,279]" captionStart="FIGURE 1. A – H, T" captionStartId="2.[151,250,1919,1942]" captionTargetBox="[218,1385,188,1872]" captionTargetId="figure@2.[218,1386,157,1909]" captionTargetPageId="2" captionText="FIGURE 1. A – H, T. venkataramani sp. nov.: A, male (holotype), dorsal view; B, same, ventral view; C, female (paratype), dorsal view; D, male (holotype), left lateral view; E, macropterous female (paratype), dorsal view; F, apterous female (paratype), left lateral view; G, apterous female (paratype), terminal abdominal segments, ventral view; H, male (holotype), genital segments, left lateral view; I, T. ceylonensis, apterous male genital segments, left lateral view; J, T. nicobarensis, same." httpUri="https://zenodo.org/record/219885/files/figure.png" pageId="1" pageNumber="379">
Figs. 
<date id="FD5C7092FFE2A825FF0945885FA3FE88" box="[213,225,255,279]" pageId="1" pageNumber="379">1</date>
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I, 2A, D, G, J, 3A
</paragraph>
<paragraph id="895D5652FFE2A825FF4B443E5C47FEC2" blockId="1.[151,773,328,350]" box="[151,773,328,350]" pageId="1" pageNumber="379">
<taxonomicName id="4EE22DD1FFE2A825FF4B443E5C43FEC2" authority="Hungerford &amp; Matsuda, 1962: 141" authorityName="Hungerford &amp; Matsuda" authorityPageNumber="141" authorityYear="1962" box="[151,769,328,350]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="379" phylum="Arthropoda" rank="species" species="ceylonensis">
<emphasis id="BB968A40FFE2A825FF4B443E5EDAFEC1" box="[151,408,328,350]" italics="true" pageId="1" pageNumber="379">Tenagogonus ceylonensis</emphasis>
Hungerford &amp; Matsuda, 1962: 141
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.
</paragraph>
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<subSubSection id="C1F805D9FFE2A825FF4B44FD5F9AFE75" pageId="1" pageNumber="379" type="materials_examined">
<paragraph id="895D5652FFE2A825FF4B44FD5F9AFE75" blockId="1.[151,1436,394,922]" pageId="1" pageNumber="379">
<emphasis id="BB968A40FFE2A825FF4B44FD5EC8FE3C" bold="true" box="[151,394,394,419]" pageId="1" pageNumber="379">Material examined.</emphasis>
<collectingCountry id="F1F516C2FFE2A825FE4644FC5EACFE3C" box="[410,494,395,419]" name="India" pageId="1" pageNumber="379">INDIA</collectingCountry>
: Tamil Nadu, Kanyakumari District, Cherukole Village, 
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[
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], 
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, 
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, 
<date id="FD5C7092FFE2A825FE2044D95D2FFE59" box="[508,621,430,454]" pageId="1" pageNumber="379" value="2011-05-20">20.v.2011</date>
, Coll: E. Eyarin Jehamalar, 8 3 (apt.), 1 Ƥ (apt.) (ZSI Reg. No. 2720/ H15).
</paragraph>
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<subSubSection id="C1F805D9FFE2A825FF1544815D9EFC95" pageId="1" pageNumber="379" type="diagnosis">
<paragraph id="895D5652FFE2A825FF1544815D9EFC95" blockId="1.[151,1436,394,922]" pageId="1" pageNumber="379">
<emphasis id="BB968A40FFE2A825FF1544815E01FD90" bold="true" box="[201,323,502,527]" pageId="1" pageNumber="379">Diagnosis.</emphasis>
Apterous form: 
<collectingCountry id="F1F516C2FFE2A825FDD944815D7CFD91" box="[517,574,502,526]" name="Malta" pageId="1" pageNumber="379">Male</collectingCountry>
mean length 4.49, range 4.40–4.59 (n=5), mean width 1.6, range 1.49–1.6 (n=5), and female length 5.27 (n=1) excluding lateral processes of seventh segment (= connexival spines), width 1.98 (n=1); colour: Ochraceous; head with four stripes, central pair of stripes united at base and slightly pale; each fourth antennal segment longer than other segments; antero-sublateral region of anterior pronotal lobe with short broad band on each side; pronotal lobe with median brown stripe; mesopleuron with three brown longitudinal stripes and white fascia near each side of pronotal lobe; male devoid of connexival spines and females with long connexival spines, converging on dry specimen (
<figureCitation id="11D94AD7FFE2A825FD1F47B95C4AFD79" box="[707,776,718,742]" captionStart="FIGURE 2. A, D, G, J, T" captionStartId="3.[151,250,1826,1849]" captionTargetBox="[259,1322,205,1749]" captionTargetId="figure@3.[242,1344,193,1790]" captionTargetPageId="3" captionText="FIGURE 2. A, D, G, J, T. ceylonensis: A, pygophore, parameres and endosoma, ventral view; D, endosoma, right lateral view; G, terminal abdominal segment of female showing connexival spines, dorsal view; J, male genital segments, ventral view. B, E, H, K, T. nicobarensis: B, pygophore, parameres and endosoma, ventral view; E, endosoma, right lateral view; H, terminal abdominal segment of female showing connexival spines, dorsal view; K, male genital segments, ventral view. C, F, I, L, T. venkataramani: C, pygophore, parameres and endosoma, ventral view; F, endosoma, right lateral view; I, terminal abdominal segment of female showing connexival spines, dorsal view; L, male genital segments, ventral view (ds—dorsal sclerite, en—endosoma, ls—lateral sclerite, pa—paramere, pg—pygophore, vs—ventral sclerite)." httpUri="https://zenodo.org/record/219886/files/figure.png" pageId="1" pageNumber="379">Fig. 2</figureCitation>
G); posterior margin of eighth sternum of male bisinuated (
<figureCitation id="11D94AD7FFE2A825FF4347855FA8FC95" box="[159,234,754,778]" captionStart="FIGURE 1. A – H, T" captionStartId="2.[151,250,1919,1942]" captionTargetBox="[218,1385,188,1872]" captionTargetId="figure@2.[218,1386,157,1909]" captionTargetPageId="2" captionText="FIGURE 1. A – H, T. venkataramani sp. nov.: A, male (holotype), dorsal view; B, same, ventral view; C, female (paratype), dorsal view; D, male (holotype), left lateral view; E, macropterous female (paratype), dorsal view; F, apterous female (paratype), left lateral view; G, apterous female (paratype), terminal abdominal segments, ventral view; H, male (holotype), genital segments, left lateral view; I, T. ceylonensis, apterous male genital segments, left lateral view; J, T. nicobarensis, same." httpUri="https://zenodo.org/record/219885/files/figure.png" pageId="1" pageNumber="379">
Figs. 
<date id="FD5C7092FFE2A825FF0247855FA8FC95" box="[222,234,754,778]" pageId="1" pageNumber="379">1</date>
</figureCitation>
I, 2J); endosomal sclerites as in 
<figureCitation id="11D94AD7FFE2A825FD8F47855D9AFC95" box="[595,728,754,778]" captionStart="FIGURE 2. A, D, G, J, T" captionStartId="3.[151,250,1826,1849]" captionTargetBox="[259,1322,205,1749]" captionTargetId="figure@3.[242,1344,193,1790]" captionTargetPageId="3" captionText="FIGURE 2. A, D, G, J, T. ceylonensis: A, pygophore, parameres and endosoma, ventral view; D, endosoma, right lateral view; G, terminal abdominal segment of female showing connexival spines, dorsal view; J, male genital segments, ventral view. B, E, H, K, T. nicobarensis: B, pygophore, parameres and endosoma, ventral view; E, endosoma, right lateral view; H, terminal abdominal segment of female showing connexival spines, dorsal view; K, male genital segments, ventral view. C, F, I, L, T. venkataramani: C, pygophore, parameres and endosoma, ventral view; F, endosoma, right lateral view; I, terminal abdominal segment of female showing connexival spines, dorsal view; L, male genital segments, ventral view (ds—dorsal sclerite, en—endosoma, ls—lateral sclerite, pa—paramere, pg—pygophore, vs—ventral sclerite)." httpUri="https://zenodo.org/record/219886/files/figure.png" pageId="1" pageNumber="379">Figs. 2A, D</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C1F805D9FFE2A825FF1546615C8FFCB1" box="[201,973,790,815]" pageId="1" pageNumber="379" type="distribution">
<paragraph id="895D5652FFE2A825FF1546615C8FFCB1" blockId="1.[151,1436,394,922]" box="[201,973,790,815]" pageId="1" pageNumber="379">
<emphasis id="BB968A40FFE2A825FF1546615E20FCB0" bold="true" box="[201,354,790,815]" pageId="1" pageNumber="379">Distribution.</emphasis>
<collectingCountry id="F1F516C2FFE2A825FEB546615EE1FCB1" box="[361,419,790,814]" name="India" pageId="1" pageNumber="379">India</collectingCountry>
(Karnataka, Kerala, and Tamil Nadu), 
<collectingCountry id="F1F516C2FFE2A825FC8446615C8BFCB1" box="[856,969,790,814]" name="Sri Lanka" pageId="1" pageNumber="379">Sri Lanka</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C1F805D9FFE2A825FF15464C5EEDFBB5" pageId="1" pageNumber="379" type="discussion">
<paragraph id="895D5652FFE2A825FF15464C5C9BFC05" blockId="1.[151,1436,394,922]" pageId="1" pageNumber="379">
<emphasis id="BB968A40FFE2A825FF15464C5E7FFCCC" bold="true" box="[201,317,827,851]" pageId="1" pageNumber="379">Remarks.</emphasis>
In the original description of the species, the venter of the mesonotum was described as nearly black, but the specimens studied in this study have the venter pale yellow and clothed with minute brown hairs. These specimens represent the first record for this species for Tamil Nadu.
</paragraph>
<paragraph id="895D5652FFE2A825FF4B46985EEDFBB5" blockId="1.[151,573,1006,1066]" pageId="1" pageNumber="379">
<taxonomicName id="4EE22DD1FFE2A825FF4B46985F8DFBB5" authority="Andersen Figs." authorityName="Andersen Figs." class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="379" phylum="Arthropoda" rank="species" species="nicobarensis">
<emphasis id="BB968A40FFE2A825FF4B46985E88FB98" bold="true" box="[151,458,1006,1031]" italics="true" pageId="1" pageNumber="379">Tenagogonus nicobarensis</emphasis>
Andersen 
<figureCitation id="11D94AD7FFE2A825FF4B41655F8DFBB5" box="[151,207,1042,1066]" captionStart="FIGURE 1. A – H, T" captionStartId="2.[151,250,1919,1942]" captionTargetBox="[218,1385,188,1872]" captionTargetId="figure@2.[218,1386,157,1909]" captionTargetPageId="2" captionText="FIGURE 1. A – H, T. venkataramani sp. nov.: A, male (holotype), dorsal view; B, same, ventral view; C, female (paratype), dorsal view; D, male (holotype), left lateral view; E, macropterous female (paratype), dorsal view; F, apterous female (paratype), left lateral view; G, apterous female (paratype), terminal abdominal segments, ventral view; H, male (holotype), genital segments, left lateral view; I, T. ceylonensis, apterous male genital segments, left lateral view; J, T. nicobarensis, same." httpUri="https://zenodo.org/record/219885/files/figure.png" pageId="1" pageNumber="379">Figs.</figureCitation>
</taxonomicName>
1J, 2B, E, H, K, 3B
</paragraph>
</subSubSection>
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<paragraph id="895D5652FFE2A825FF4B412C5DC6FBEF" blockId="1.[151,644,1114,1136]" box="[151,644,1114,1136]" pageId="1" pageNumber="379">
<taxonomicName id="4EE22DD1FFE2A825FF4B412C5DC3FBEF" authority="Andersen, 1964: 322" authorityName="Andersen" authorityPageNumber="322" authorityYear="1964" box="[151,641,1114,1136]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="379" phylum="Arthropoda" rank="species" species="nicobarensis">
<emphasis id="BB968A40FFE2A825FF4B412C5EE1FBEF" box="[151,419,1114,1136]" italics="true" pageId="1" pageNumber="379">Tenagogonus nicobarensis</emphasis>
Andersen, 1964: 322
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C1F805D9FFE2A825FF4B41EA5CBEFA4A" pageId="1" pageNumber="379" type="materials_examined">
<paragraph id="895D5652FFE2A825FF4B41EA5CBEFA4A" blockId="1.[151,1436,1181,1926]" pageId="1" pageNumber="379">
<emphasis id="BB968A40FFE2A825FF4B41EA5EC5FB29" bold="true" box="[151,391,1181,1206]" pageId="1" pageNumber="379">Material examined.</emphasis>
<collectingCountry id="F1F516C2FFE2A825FE4841E95EAAFB29" box="[404,488,1182,1206]" name="India" pageId="1" pageNumber="379">INDIA</collectingCountry>
: Andaman and 
<collectingCountry id="F1F516C2FFE2A825FD7541EA5C20FB2A" box="[681,866,1181,1205]" name="India" pageId="1" pageNumber="379">Nicobar Islands</collectingCountry>
, South Andaman: Manjery, 
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[
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], 
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, 
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, 
<date id="FD5C7092FFE2A825FE0941B65D01FB46" box="[469,579,1217,1241]" pageId="1" pageNumber="379" value="2012-03-03">3.iii.2012</date>
, 1 3 (mac.), 3 Ƥ (mac.); Wandoor, 
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[57.0 m], 
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, 
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, 
<date id="FD5C7092FFE2A825FEF541925ED4FB62" box="[297,406,1253,1277]" pageId="1" pageNumber="379" value="2012-03-05">5.iii.2012</date>
, 3 3 (apt.), 6 Ƥ (apt.); Humphrygunj, 
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[89.0 m], 
<geoCoordinate id="ECD63095FFE2A825FB9B41915BA5FB62" box="[1095,1255,1253,1277]" direction="north" orientation="latitude" pageId="1" pageNumber="379" precision="1" value="12.418616">N 11°85.117’</geoCoordinate>
, 
<geoCoordinate id="ECD63095FFE2A825FB2541915AD5FB62" box="[1273,1431,1253,1277]" direction="east" orientation="longitude" pageId="1" pageNumber="379" precision="1" value="92.65755">E 92°39.453’</geoCoordinate>
, 
<date id="FD5C7092FFE2A825FF4B407E5E47FABE" box="[151,261,1289,1313]" pageId="1" pageNumber="379" value="2012-03-06">6.iii.2012</date>
, 3 3 (apt.), 3 3 (mac.), 2 Ƥ (apt.), 2 Ƥ (mac.); Mongulton, 
<quantity id="4E1AFBB7FFE2A825FC76407E5C9DFABE" box="[938,991,1289,1313]" metricMagnitude="1" metricUnit="m" metricValue="1.40208" pageId="1" pageNumber="379" unit="ft" value="46.0">46 ft</quantity>
[14.0 m], 
<geoCoordinate id="ECD63095FFE2A825FB8B407D5BB1FABE" box="[1111,1267,1289,1313]" direction="north" orientation="latitude" pageId="1" pageNumber="379" precision="1" value="11.580167">N 11°34.810’</geoCoordinate>
, 
<geoCoordinate id="ECD63095FFE2A825FB23407D5AD5FABE" box="[1279,1431,1289,1313]" direction="east" orientation="longitude" pageId="1" pageNumber="379" precision="1" value="92.65495">E 92°39.297’</geoCoordinate>
, 
<date id="FD5C7092FFE2A825FF4B405A5E47FADA" box="[151,261,1325,1349]" pageId="1" pageNumber="379" value="2012-03-06">6.iii.2012</date>
, 3 3 (apt.), 4 3 (mac.), 10 Ƥ (apt.); Guptapara, 
<quantity id="4E1AFBB7FFE2A825FCFC405A5C17FADA" box="[800,853,1325,1349]" metricMagnitude="1" metricUnit="m" metricValue="2.19456" pageId="1" pageNumber="379" unit="ft" value="72.0">72 ft</quantity>
[
<quantity id="4E1AFBB7FFE2A825FCBE405A5CF5FADA" box="[866,951,1325,1349]" metricMagnitude="1" metricUnit="m" metricValue="2.19" pageId="1" pageNumber="379" unit="m" value="21.9">21.9 m</quantity>
], 
<geoCoordinate id="ECD63095FFE2A825FC1740595B27FADA" box="[971,1125,1325,1349]" direction="north" orientation="latitude" pageId="1" pageNumber="379" precision="1" value="11.574133">N 11°34.448’</geoCoordinate>
, 
<geoCoordinate id="ECD63095FFE2A825FBAD40595A4AFADA" box="[1137,1288,1325,1349]" direction="east" orientation="longitude" pageId="1" pageNumber="379" precision="1" value="92.66615">E 92°39.969’</geoCoordinate>
, 
<date id="FD5C7092FFE2A825FAC9405A5AC0FADA" box="[1301,1410,1325,1349]" pageId="1" pageNumber="379" value="2012-03-06">6.iii.2012</date>
, 1 m (mac.); Bambooflat, 
<quantity id="4E1AFBB7FFE2A825FE4140265E9DFAF6" box="[413,479,1361,1385]" metricMagnitude="1" metricUnit="m" metricValue="6.18744" pageId="1" pageNumber="379" unit="ft" value="203.0">203 ft</quantity>
[
<quantity id="4E1AFBB7FFE2A825FE3040265D03FAF6" box="[492,577,1361,1385]" metricMagnitude="1" metricUnit="m" metricValue="6.1899999999999995" pageId="1" pageNumber="379" unit="m" value="61.9">61.9 m</quantity>
], 
<geoCoordinate id="ECD63095FFE2A825FD8840255DACFAF6" box="[596,750,1361,1385]" direction="north" orientation="latitude" pageId="1" pageNumber="379" precision="1" value="11.719017">N 11°43.141’</geoCoordinate>
, 
<geoCoordinate id="ECD63095FFE2A825FD2540255CD3FAF6" box="[761,913,1361,1385]" direction="east" orientation="longitude" pageId="1" pageNumber="379" precision="1" value="92.71962">E 92°43.177’</geoCoordinate>
, 
<date id="FD5C7092FFE2A825FC4040265B48FAF6" box="[924,1034,1361,1385]" pageId="1" pageNumber="379" value="2012-03-07">7.iii.2012</date>
, 3 3 (apt.), 4 Ƥ (apt.); Shoalbay, 
<quantity id="4E1AFBB7FFE2A825FA5C40265FEAFA12" metricMagnitude="1" metricUnit="m" metricValue="2.83464" pageId="1" pageNumber="379" unit="ft" value="93.0">93 ft</quantity>
[
<quantity id="4E1AFBB7FFE2A825FF6A40025E49FA12" box="[182,267,1397,1421]" metricMagnitude="1" metricUnit="m" metricValue="2.83" pageId="1" pageNumber="379" unit="m" value="28.3">28.3 m</quantity>
], 
<geoCoordinate id="ECD63095FFE2A825FEC340015EF8FA12" box="[287,442,1397,1421]" direction="north" orientation="latitude" pageId="1" pageNumber="379" precision="1" value="11.82575">N 11°49.545’</geoCoordinate>
, 
<geoCoordinate id="ECD63095FFE2A825FE1940015D1CFA12" box="[453,606,1397,1421]" direction="east" orientation="longitude" pageId="1" pageNumber="379" precision="1" value="92.7273">E 92°43.638’</geoCoordinate>
, 
<date id="FD5C7092FFE2A825FDB640025DA4FA12" box="[618,742,1397,1421]" pageId="1" pageNumber="379" value="2012-03-14">14.iii.2012</date>
, 6 3 (apt.), 3 3 (mac.), 3 Ƥ (apt.), 1 Ƥ (mac.). All specimens were collected from streams of 
<collectingCountry id="F1F516C2FFE2A825FE2440ED5D08FA2D" box="[504,586,1434,1458]" name="India" pageId="1" pageNumber="379">INDIA</collectingCountry>
, Andaman &amp; 
<collectingCountry id="F1F516C2FFE2A825FD3A40EE5CDBFA2E" box="[742,921,1433,1457]" name="India" pageId="1" pageNumber="379">Nicobar Islands</collectingCountry>
, South Andaman, by E. Eyarin Jehamalar and all specimens are deposited in ZSI Kolkata, Reg. No. 2693/H15 to 2699/H15.
</paragraph>
</subSubSection>
<subSubSection id="C1F805D9FFE2A825FF1B40965CB5F886" pageId="1" pageNumber="379" type="diagnosis">
<paragraph id="895D5652FFE2A825FF1B40965CB5F886" blockId="1.[151,1436,1181,1926]" pageId="1" pageNumber="379">
<emphasis id="BB968A40FFE2A825FF1B40965E03FA65" bold="true" box="[199,321,1505,1530]" pageId="1" pageNumber="379">Diagnosis.</emphasis>
Apterous form: 
<collectingCountry id="F1F516C2FFE2A825FDDC40965D7BFA66" box="[512,569,1505,1529]" name="Malta" pageId="1" pageNumber="379">Male</collectingCountry>
mean length 6.38, range 5.40–7.57 (n=5), mean width 2.01, range 1.60–2.5 (n=5), and female mean length 7.07, range 6.75–7.43 (n=5), mean width 2.46, range 2.17–2.67 (n=5); colour yellowish brown; head with four black stripes; mesopleuron with two black stripes, outer one broad; lateral region of mesosternum with interrupted black band; male devoid of lateral processes of seventh sternum (= connexival spines) and female with long connexival processes (
<figureCitation id="11D94AD7FFE2A825FD2943065C7EF916" box="[757,828,1649,1673]" captionStart="FIGURE 2. A, D, G, J, T" captionStartId="3.[151,250,1826,1849]" captionTargetBox="[259,1322,205,1749]" captionTargetId="figure@3.[242,1344,193,1790]" captionTargetPageId="3" captionText="FIGURE 2. A, D, G, J, T. ceylonensis: A, pygophore, parameres and endosoma, ventral view; D, endosoma, right lateral view; G, terminal abdominal segment of female showing connexival spines, dorsal view; J, male genital segments, ventral view. B, E, H, K, T. nicobarensis: B, pygophore, parameres and endosoma, ventral view; E, endosoma, right lateral view; H, terminal abdominal segment of female showing connexival spines, dorsal view; K, male genital segments, ventral view. C, F, I, L, T. venkataramani: C, pygophore, parameres and endosoma, ventral view; F, endosoma, right lateral view; I, terminal abdominal segment of female showing connexival spines, dorsal view; L, male genital segments, ventral view (ds—dorsal sclerite, en—endosoma, ls—lateral sclerite, pa—paramere, pg—pygophore, vs—ventral sclerite)." httpUri="https://zenodo.org/record/219886/files/figure.png" pageId="1" pageNumber="379">Fig. 2</figureCitation>
H, 3B); seventh abdominal sternum of male with Vshaped impression on its posterior margin (
<figureCitation id="11D94AD7FFE2A825FDA443E25D84F932" box="[632,710,1685,1709]" captionStart="FIGURE 2. A, D, G, J, T" captionStartId="3.[151,250,1826,1849]" captionTargetBox="[259,1322,205,1749]" captionTargetId="figure@3.[242,1344,193,1790]" captionTargetPageId="3" captionText="FIGURE 2. A, D, G, J, T. ceylonensis: A, pygophore, parameres and endosoma, ventral view; D, endosoma, right lateral view; G, terminal abdominal segment of female showing connexival spines, dorsal view; J, male genital segments, ventral view. B, E, H, K, T. nicobarensis: B, pygophore, parameres and endosoma, ventral view; E, endosoma, right lateral view; H, terminal abdominal segment of female showing connexival spines, dorsal view; K, male genital segments, ventral view. C, F, I, L, T. venkataramani: C, pygophore, parameres and endosoma, ventral view; F, endosoma, right lateral view; I, terminal abdominal segment of female showing connexival spines, dorsal view; L, male genital segments, ventral view (ds—dorsal sclerite, en—endosoma, ls—lateral sclerite, pa—paramere, pg—pygophore, vs—ventral sclerite)." httpUri="https://zenodo.org/record/219886/files/figure.png" pageId="1" pageNumber="379">Figs. 2</figureCitation>
K); ninth sternum of male with two sheath-like structures leaving a broad median longitudinal depression widened anteriorly; endosomal sclerites as in 
<figureCitation id="11D94AD7FFE2A825FBAC43CE5B96F94E" box="[1136,1236,1721,1745]" captionStart="FIGURE 2. A, D, G, J, T" captionStartId="3.[151,250,1826,1849]" captionTargetBox="[259,1322,205,1749]" captionTargetId="figure@3.[242,1344,193,1790]" captionTargetPageId="3" captionText="FIGURE 2. A, D, G, J, T. ceylonensis: A, pygophore, parameres and endosoma, ventral view; D, endosoma, right lateral view; G, terminal abdominal segment of female showing connexival spines, dorsal view; J, male genital segments, ventral view. B, E, H, K, T. nicobarensis: B, pygophore, parameres and endosoma, ventral view; E, endosoma, right lateral view; H, terminal abdominal segment of female showing connexival spines, dorsal view; K, male genital segments, ventral view. C, F, I, L, T. venkataramani: C, pygophore, parameres and endosoma, ventral view; F, endosoma, right lateral view; I, terminal abdominal segment of female showing connexival spines, dorsal view; L, male genital segments, ventral view (ds—dorsal sclerite, en—endosoma, ls—lateral sclerite, pa—paramere, pg—pygophore, vs—ventral sclerite)." httpUri="https://zenodo.org/record/219886/files/figure.png" pageId="1" pageNumber="379">Figs. 2B</figureCitation>
, E; macropterous form: 
<collectingCountry id="F1F516C2FFE2A825FF0143AA5E54F96A" box="[221,278,1757,1781]" name="Malta" pageId="1" pageNumber="379">Male</collectingCountry>
mean length 7.38, range 6.59–8.32 (n=4), mean width 2.10, range 1.77–2.44 (n=4), and female mean length 7.52, range 7.26–7.91 (n=4), mean width 2.48, range 2.34–2.72 (n=4).
</paragraph>
</subSubSection>
<subSubSection id="C1F805D9FFE2A825FF1B42525DEAF8A2" box="[199,680,1829,1854]" pageId="1" pageNumber="379" type="distribution">
<paragraph id="895D5652FFE2A825FF1B42525DEAF8A2" blockId="1.[151,1436,1181,1926]" box="[199,680,1829,1854]" pageId="1" pageNumber="379">
<emphasis id="BB968A40FFE2A825FF1B42525E22F8A1" bold="true" box="[199,352,1829,1854]" pageId="1" pageNumber="379">Distribution.</emphasis>
Andaman &amp; 
<collectingCountry id="F1F516C2FFE2A825FE2442525DEAF8A2" box="[504,680,1829,1853]" name="India" pageId="1" pageNumber="379">Nicobar Islands</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="C1F805D9FFE2A821FF1B423D5E9EFF4B" lastPageId="5" lastPageNumber="383" pageId="1" pageNumber="379" type="discussion">
<paragraph id="895D5652FFE2A825FF1B423D5A6FF81A" blockId="1.[151,1436,1181,1926]" pageId="1" pageNumber="379">
<emphasis id="BB968A40FFE2A825FF1B423D5E79F8FD" bold="true" box="[199,315,1866,1890]" pageId="1" pageNumber="379">Remarks.</emphasis>
This species has been reported from the Andaman Islands and a few 
<collectingCountry id="F1F516C2FFE2A825FB9B423E5BE3F8FE" box="[1095,1185,1865,1889]" name="India" pageId="1" pageNumber="379">Nicobar</collectingCountry>
groups of Islands, but during our extensive surveys, 
<taxonomicName id="4EE22DD1FFE2A825FE3B42195DD3F81A" box="[487,657,1901,1925]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="379" phylum="Arthropoda" rank="species" species="nicobarensis">
<emphasis id="BB968A40FFE2A825FE3B42195DD3F81A" box="[487,657,1901,1925]" italics="true" pageId="1" pageNumber="379">T. nicobarensis</emphasis>
</taxonomicName>
has not been encountered by us from Great 
<collectingCountry id="F1F516C2FFE2A825FB5C421A5B98F81A" box="[1152,1242,1901,1925]" name="India" pageId="1" pageNumber="379">Nicobar</collectingCountry>
<collectingCountry id="F1F516C2FFE2A825FB3D421A5A6BF81A" box="[1249,1321,1901,1925]" name="Iceland" pageId="1" pageNumber="379">Island</collectingCountry>
.
</paragraph>
<caption id="DD9D06DAFFE1A826FF4B42085AD4F86E" httpUri="https://zenodo.org/record/219885/files/figure.png" pageId="2" pageNumber="380" targetBox="[218,1385,188,1872]" targetPageId="2">
<paragraph id="895D5652FFE1A826FF4B42085AD4F86E" blockId="2.[151,1436,1919,2033]" pageId="2" pageNumber="380">
<emphasis id="BB968A40FFE1A826FF4B42085E56F80A" bold="true" box="[151,276,1919,1942]" pageId="2" pageNumber="380">FIGURE 1.</emphasis>
A–H, 
<taxonomicName id="4EE22DD1FFE1A826FE8042F65D4DF80A" box="[348,527,1919,1942]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="380" phylum="Arthropoda" rank="species" species="venkataramani" status="sp. nov.">
<emphasis id="BB968A40FFE1A826FE8042F65D4DF80A" box="[348,527,1919,1942]" italics="true" pageId="2" pageNumber="380">T. venkataramani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0A5373BFFE1A826FDCB42F75D2BF80A" box="[535,617,1920,1941]" pageId="2" pageNumber="380" rank="species">
<emphasis id="BB968A40FFE1A826FDCB42F75D2BF80A" bold="true" box="[535,617,1920,1941]" pageId="2" pageNumber="380">sp. nov.</emphasis>
</taxonomicNameLabel>
: A, male (holotype), dorsal view; B, same, ventral view; C, female (paratype), dorsal view; D, male (holotype), left lateral view; E, macropterous female (paratype), dorsal view; F, apterous female (paratype), left lateral view; G, apterous female (paratype), terminal abdominal segments, ventral view; H, male (holotype), genital segments, left lateral view; I, 
<taxonomicName id="4EE22DD1FFE1A826FDD242AB5DD9F86E" box="[526,667,2011,2033]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="380" phylum="Arthropoda" rank="species" species="ceylonensis">
<emphasis id="BB968A40FFE1A826FDD242AB5DD9F86E" box="[526,667,2011,2033]" italics="true" pageId="2" pageNumber="380">T. ceylonensis</emphasis>
</taxonomicName>
, apterous male genital segments, left lateral view; J, 
<taxonomicName id="4EE22DD1FFE1A826FB6942AB5A0CF86E" box="[1205,1358,2011,2033]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="380" phylum="Arthropoda" rank="species" species="nicobarensis">
<emphasis id="BB968A40FFE1A826FB6942AB5A0CF86E" box="[1205,1358,2011,2033]" italics="true" pageId="2" pageNumber="380">T. nicobarensis</emphasis>
</taxonomicName>
, same.
</paragraph>
</caption>
<caption id="DD9D06DAFFE0A827FF4B42555B5CF86F" httpUri="https://zenodo.org/record/219886/files/figure.png" pageId="3" pageNumber="381" targetBox="[259,1322,205,1749]" targetPageId="3">
<paragraph id="895D5652FFE0A827FF4B42555B5CF86F" blockId="3.[151,1436,1826,2033]" pageId="3" pageNumber="381">
<emphasis id="BB968A40FFE0A827FF4B42555E51F8A7" bold="true" box="[151,275,1826,1849]" pageId="3" pageNumber="381">FIGURE 2.</emphasis>
A, D, G, J, 
<taxonomicName id="4EE22DD1FFE0A827FE5642535D5AF8A6" box="[394,536,1827,1849]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="381" phylum="Arthropoda" rank="species" species="ceylonensis">
<emphasis id="BB968A40FFE0A827FE5642535D5AF8A6" box="[394,536,1827,1849]" italics="true" pageId="3" pageNumber="381">T. ceylonensis</emphasis>
</taxonomicName>
: A, pygophore, parameres and endosoma, ventral view; D, endosoma, right lateral view; G, terminal abdominal segment of female showing connexival spines, dorsal view; J, male genital segments, ventral view. B, E, H, K, 
<taxonomicName id="4EE22DD1FFE0A827FF0742165E35F8E9" box="[219,375,1888,1910]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="381" phylum="Arthropoda" rank="species" species="nicobarensis">
<emphasis id="BB968A40FFE0A827FF0742165E35F8E9" box="[219,375,1888,1910]" italics="true" pageId="3" pageNumber="381">T. nicobarensis</emphasis>
</taxonomicName>
: B, pygophore, parameres and endosoma, ventral view; E, endosoma, right lateral view; H, terminal abdominal segment of female showing connexival spines, dorsal view; K, male genital segments, ventral view. C, F, I, L, 
<taxonomicName id="4EE22DD1FFE0A827FA5542F75E6DF82C" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="381" phylum="Arthropoda" rank="species" species="venkataramani">
<emphasis id="BB968A40FFE0A827FA5542F75E6DF82C" italics="true" pageId="3" pageNumber="381">T. venkataramani</emphasis>
</taxonomicName>
: C, pygophore, parameres and endosoma, ventral view; F, endosoma, right lateral view; I, terminal abdominal segment of female showing connexival spines, dorsal view; L, male genital segments, ventral view (ds—dorsal sclerite, en—endosoma, ls—lateral sclerite, pa—paramere, pg—pygophore, vs—ventral sclerite).
</paragraph>
</caption>
<caption id="DD9D06DAFFE7A820FF4B42E85D1FF86C" httpUri="https://zenodo.org/record/219887/files/figure.png" pageId="4" pageNumber="382" targetBox="[157,1412,206,1932]" targetPageId="4">
<paragraph id="895D5652FFE7A820FF4B42E85D1FF86C" blockId="4.[151,1435,1951,2035]" pageId="4" pageNumber="382">
<emphasis id="BB968A40FFE7A820FF4B42E85E56F82A" bold="true" box="[151,276,1951,1974]" pageId="4" pageNumber="382">FIGURE 3.</emphasis>
A, B, C, Female terminal abdominal segments showing connexival spines, lateral view; A, 
<taxonomicName id="4EE22DD1FFE7A820FB1042D65A19F829" box="[1228,1371,1952,1974]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="382" phylum="Arthropoda" rank="species" species="ceylonensis">
<emphasis id="BB968A40FFE7A820FB1042D65A19F829" box="[1228,1371,1952,1974]" italics="true" pageId="4" pageNumber="382">T. ceylonensis</emphasis>
</taxonomicName>
; B, 
<taxonomicName id="4EE22DD1FFE7A820FA5542D65E55F84A" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="382" phylum="Arthropoda" rank="species" species="nicobarensis">
<emphasis id="BB968A40FFE7A820FA5542D65E55F84A" italics="true" pageId="4" pageNumber="382">T. nicobarensis</emphasis>
</taxonomicName>
; C, 
<emphasis id="BB968A40FFE7A820FE9742B75D45F84A" box="[331,519,1983,2005]" italics="true" pageId="4" pageNumber="382">
<taxonomicName id="4EE22DD1FFE7A820FE9742B75D41F84A" box="[331,515,1983,2005]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="382" phylum="Arthropoda" rank="species" species="venkataramani">T. venkataramani</taxonomicName>
.
</emphasis>
Distribution of 
<taxonomicName id="4EE22DD1FFE7A820FD6942B75C04F84A" box="[693,838,1983,2005]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="382" phylum="Arthropoda" rank="species" species="ceylonensis">
<emphasis id="BB968A40FFE7A820FD6942B75C04F84A" box="[693,838,1983,2005]" italics="true" pageId="4" pageNumber="382">T. ceylonensis</emphasis>
</taxonomicName>
. Round dots denote records from literature and the star denotes the present record from Tamil Nadu.
</paragraph>
</caption>
<paragraph id="895D5652FFE6A821FF4B45EF5E9EFF4B" blockId="5.[151,583,151,212]" pageId="5" pageNumber="383">
<emphasis id="BB968A40FFE6A821FF4B45EF5D05FF2F" bold="true" box="[151,583,151,176]" pageId="5" pageNumber="383">
<taxonomicName id="4EE22DD1FFE6A821FF4B45EF5EA4FF2F" box="[151,486,151,176]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="383" phylum="Arthropoda" rank="species" species="venkataramani" status="sp. nov.">
<emphasis id="BB968A40FFE6A821FF4B45EF5EA4FF2F" bold="true" box="[151,486,151,176]" italics="true" pageId="5" pageNumber="383">Tenagogonus venkataramani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0A5373BFFE6A821FE3145EF5D05FF2F" box="[493,583,152,176]" pageId="5" pageNumber="383" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="11D94AD7FFE6A821FF4345CB5E67FF4B" box="[159,293,188,212]" captionStart="FIGURE 1. A – H, T" captionStartId="2.[151,250,1919,1942]" captionTargetBox="[218,1385,188,1872]" captionTargetId="figure@2.[218,1386,157,1909]" captionTargetPageId="2" captionText="FIGURE 1. A – H, T. venkataramani sp. nov.: A, male (holotype), dorsal view; B, same, ventral view; C, female (paratype), dorsal view; D, male (holotype), left lateral view; E, macropterous female (paratype), dorsal view; F, apterous female (paratype), left lateral view; G, apterous female (paratype), terminal abdominal segments, ventral view; H, male (holotype), genital segments, left lateral view; I, T. ceylonensis, apterous male genital segments, left lateral view; J, T. nicobarensis, same." httpUri="https://zenodo.org/record/219885/files/figure.png" pageId="5" pageNumber="383">Figs. 1A–H</figureCitation>
, 
<figureCitation id="11D94AD7FFE6A821FEED45CB5E11FF4B" box="[305,339,188,212]" captionStart="FIGURE 2. A, D, G, J, T" captionStartId="3.[151,250,1826,1849]" captionTargetBox="[259,1322,205,1749]" captionTargetId="figure@3.[242,1344,193,1790]" captionTargetPageId="3" captionText="FIGURE 2. A, D, G, J, T. ceylonensis: A, pygophore, parameres and endosoma, ventral view; D, endosoma, right lateral view; G, terminal abdominal segment of female showing connexival spines, dorsal view; J, male genital segments, ventral view. B, E, H, K, T. nicobarensis: B, pygophore, parameres and endosoma, ventral view; E, endosoma, right lateral view; H, terminal abdominal segment of female showing connexival spines, dorsal view; K, male genital segments, ventral view. C, F, I, L, T. venkataramani: C, pygophore, parameres and endosoma, ventral view; F, endosoma, right lateral view; I, terminal abdominal segment of female showing connexival spines, dorsal view; L, male genital segments, ventral view (ds—dorsal sclerite, en—endosoma, ls—lateral sclerite, pa—paramere, pg—pygophore, vs—ventral sclerite)." httpUri="https://zenodo.org/record/219886/files/figure.png" pageId="5" pageNumber="383">2C</figureCitation>
, F, I, L, 3C)
</paragraph>
</subSubSection>
<subSubSection id="C1F805D9FFE6A821FF4B44745E69FD6D" pageId="5" pageNumber="383" type="materials_examined">
<paragraph id="895D5652FFE6A821FF4B44745E4DFD18" blockId="5.[151,1437,259,2015]" pageId="5" pageNumber="383">
<emphasis id="BB968A40FFE6A821FF4B44745D46FE83" bold="true" box="[151,516,259,284]" pageId="5" pageNumber="383">
Material examined. 
<typeStatus id="5659E8F0FFE6A821FE4C44735EBDFE83" box="[400,511,260,284]" pageId="5" pageNumber="383" type="holotype">Holotype</typeStatus>
.
</emphasis>
3 (apt.): 
<collectingCountry id="F1F516C2FFE6A821FDA344735D93FE83" box="[639,721,260,284]" name="India" pageId="5" pageNumber="383">INDIA</collectingCountry>
, Andaman &amp; 
<collectingCountry id="F1F516C2FFE6A821FCA644735B70FE83" box="[890,1074,260,284]" name="India" pageId="5" pageNumber="383">Nicobar Islands</collectingCountry>
, Great 
<collectingCountry id="F1F516C2FFE6A821FB5744735BA7FE83" box="[1163,1253,260,284]" name="India" pageId="5" pageNumber="383">Nicobar</collectingCountry>
<collectingCountry id="F1F516C2FFE6A821FB2C44735A75FE83" box="[1264,1335,260,284]" name="Iceland" pageId="5" pageNumber="383">Island</collectingCountry>
, GNBR, Govind Nagar, Nature Trail, 
<quantity id="4E1AFBB7FFE6A821FE0444505D5BFEA0" box="[472,537,295,319]" metricMagnitude="2" metricUnit="m" metricValue="1.0607039999999999" pageId="5" pageNumber="383" unit="ft" value="348.0">348 ft</quantity>
[
<quantity id="4E1AFBB7FFE6A821FDFA44505DCAFEA0" box="[550,648,295,319]" metricMagnitude="2" metricUnit="m" metricValue="1.061" pageId="5" pageNumber="383" unit="m" value="106.1">106.1 m</quantity>
], 
<geoCoordinate id="ECD63095FFE6A821FD47445F5C74FEA0" box="[667,822,295,319]" direction="north" orientation="latitude" pageId="5" pageNumber="383" precision="1" value="7.0033665">N 07°00.202’</geoCoordinate>
, 
<geoCoordinate id="ECD63095FFE6A821FC9D445F5CA4FEA0" box="[833,998,295,319]" direction="east" orientation="longitude" pageId="5" pageNumber="383" precision="1" value="93.88035">E 093°52.821’</geoCoordinate>
, 
<date id="FD5C7092FFE6A821FC2D44505B29FEA0" box="[1009,1131,295,319]" pageId="5" pageNumber="383" value="2010-11-10">10.xi.2010</date>
, Coll: E. Eyarin Jehamalar. 
<emphasis id="BB968A40FFE6A821FF4B443B5E54FEFB" bold="true" box="[151,278,332,356]" pageId="5" pageNumber="383">
<typeStatus id="5659E8F0FFE6A821FF4B443B5E53FEFB" box="[151,273,332,356]" pageId="5" pageNumber="383" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Same data as for 
<typeStatus id="5659E8F0FFE6A821FE00443B5D03FEFB" box="[476,577,332,356]" pageId="5" pageNumber="383" type="holotype">holotype</typeStatus>
, 1 Ƥ (apt.). Govind Nagar, East West Road, 
<quantity id="4E1AFBB7FFE6A821FBF2443B5B32FEFB" box="[1070,1136,332,356]" metricMagnitude="1" metricUnit="m" metricValue="8.29056" pageId="5" pageNumber="383" unit="ft" value="272.0">272 ft</quantity>
[
<quantity id="4E1AFBB7FFE6A821FBA2443B5B90FEFB" box="[1150,1234,332,356]" metricMagnitude="1" metricUnit="m" metricValue="8.29" pageId="5" pageNumber="383" unit="m" value="82.9">82.9 m</quantity>
], 
<geoCoordinate id="ECD63095FFE6A821FB39443A5A3DFEFB" box="[1253,1407,332,356]" direction="north" orientation="latitude" pageId="5" pageNumber="383" precision="1" value="7.0018">N 07°00.108’</geoCoordinate>
, 
<geoCoordinate id="ECD63095FFE6A821FA57443A5E66FE18" direction="east" orientation="longitude" pageId="5" pageNumber="383" precision="1" value="93.88129">E 093°52.877’</geoCoordinate>
, 
<date id="FD5C7092FFE6A821FEE944185EE3FE18" box="[309,417,367,391]" pageId="5" pageNumber="383" value="2010-11-06">6.xi.2010</date>
, 5 3 (apt.), 2 Ƥ (apt.); Johinder Nagar, Chingen Village, 
<date id="FD5C7092FFE6A821FB8A44185B80FE18" box="[1110,1218,367,391]" pageId="5" pageNumber="383" value="2010-11-08">8.xi.2010</date>
, 38 ft [
<quantity id="4E1AFBB7FFE6A821FAC244185A37FE18" box="[1310,1397,367,391]" metricMagnitude="1" metricUnit="m" metricValue="1.16" pageId="5" pageNumber="383" unit="m" value="11.6">11.6 m</quantity>
] 
<geoCoordinate id="ECD63095FFE6A821FA5444075E54FE33" direction="north" orientation="latitude" pageId="5" pageNumber="383" precision="1" value="6.9745502">N 06°58.473’</geoCoordinate>
, 
<geoCoordinate id="ECD63095FFE6A821FEF444E25E96FE33" box="[296,468,404,428]" direction="east" orientation="longitude" pageId="5" pageNumber="383" precision="1" value="93.9258">E 093°55.548’</geoCoordinate>
, 12 3 (apt.), 2 Ƥ (mac.), 3 Ƥ (apt.); Laxmi Nagar, 
<geoCoordinate id="ECD63095FFE6A821FB8944E25BAAFE33" box="[1109,1256,404,428]" direction="north" orientation="latitude" pageId="5" pageNumber="383" precision="1" value="6.9005666">N 6°54.034’</geoCoordinate>
, 
<geoCoordinate id="ECD63095FFE6A821FB2544E25ADAFE33" box="[1273,1432,404,428]" direction="east" orientation="longitude" pageId="5" pageNumber="383" precision="1" value="93.89118">E 93°53.471’</geoCoordinate>
, 
<date id="FD5C7092FFE6A821FF4B44C05E50FE50" box="[151,274,439,463]" pageId="5" pageNumber="383" value="2012-03-23">23.iii.2012</date>
, 3 3 (apt.); Sastri Nagar, 
<geoCoordinate id="ECD63095FFE6A821FD9844CF5D96FE50" box="[580,724,439,463]" direction="north" orientation="latitude" pageId="5" pageNumber="383" precision="1" value="6.8058667">N 6°48.352’</geoCoordinate>
, E 93°53.297, 
<date id="FD5C7092FFE6A821FC5F44C05CBDFE50" box="[899,1023,439,463]" pageId="5" pageNumber="383" value="2012-03-24">24.iii.2012</date>
, 6 3 (apt.), 4 Ƥ (apt.), 1 Ƥ (mac.); Gandhi Nagar, 
<geoCoordinate id="ECD63095FFE6A821FE9644AA5E9EFE6B" box="[330,476,476,500]" direction="north" orientation="latitude" pageId="5" pageNumber="383" precision="1" value="6.836917">N 6°50.215’</geoCoordinate>
, 
<geoCoordinate id="ECD63095FFE6A821FE3744AA5DCAFE6B" box="[491,648,476,500]" direction="east" orientation="longitude" pageId="5" pageNumber="383" precision="1" value="93.88635">E 93°53.181’</geoCoordinate>
, 
<date id="FD5C7092FFE6A821FD4B44AB5C51FE6B" box="[663,787,476,500]" pageId="5" pageNumber="383" value="2012-03-25">25.iii.2012</date>
, 4 3 (apt.), 2 Ƥ (apt.); Magar Nalah, 
<geoCoordinate id="ECD63095FFE6A821FB0044AA5A39FE6B" box="[1244,1403,476,500]" direction="north" orientation="latitude" pageId="5" pageNumber="383" precision="1" value="6.9886665">N 06°59.320’</geoCoordinate>
, 
<geoCoordinate id="ECD63095FFE6A821FA5744AA5E67FD85" direction="east" orientation="longitude" pageId="5" pageNumber="383" precision="1" value="93.90963">E 093°54.578’</geoCoordinate>
, 
<date id="FD5C7092FFE6A821FEEF47755EEDFD85" box="[307,431,514,538]" pageId="5" pageNumber="383" value="2012-03-28">28.iii.2012</date>
, 1 3 (apt.); East West Road 19th Km, Galathea tributary, 
<geoCoordinate id="ECD63095FFE6A821FB9F47745BA2FD85" box="[1091,1248,514,538]" direction="north" orientation="latitude" pageId="5" pageNumber="383" precision="1" value="6.9666166">N 06°57.997’</geoCoordinate>
, 
<geoCoordinate id="ECD63095FFE6A821FB3347745AD5FD85" box="[1263,1431,514,538]" direction="east" orientation="longitude" pageId="5" pageNumber="383" precision="1" value="93.82575">E 093°49.545’</geoCoordinate>
, 
<date id="FD5C7092FFE6A821FF4B47505E43FDA0" box="[151,257,551,575]" pageId="5" pageNumber="383" value="2012-04-03">3.iv.2012</date>
, 2 3 (apt.), 1 Ƥ (apt.); Afra Bay, 
<geoCoordinate id="ECD63095FFE6A821FD4E475F5C72FDA0" box="[658,816,551,575]" direction="north" orientation="latitude" pageId="5" pageNumber="383" precision="1" value="8.517333">N 07°91.040’</geoCoordinate>
, 
<geoCoordinate id="ECD63095FFE6A821FCE3475F5C98FDA0" box="[831,986,551,575]" direction="east" orientation="longitude" pageId="5" pageNumber="383" precision="1" value="93.7364">E 93°44.184’</geoCoordinate>
, 
<date id="FD5C7092FFE6A821FC3547505B20FDA0" box="[1001,1122,551,575]" pageId="5" pageNumber="383" value="2012-04-15">15.iv.2012</date>
, 9 3 (apt.), 2 Ƥ (apt.). All specimens were collected from streams of 
<collectingCountry id="F1F516C2FFE6A821FD5A473C5D9AFDFC" box="[646,728,587,611]" name="India" pageId="5" pageNumber="383">INDIA</collectingCountry>
, Andaman &amp; 
<collectingCountry id="F1F516C2FFE6A821FCA0473D5B70FDFD" box="[892,1074,586,610]" name="India" pageId="5" pageNumber="383">Nicobar Islands</collectingCountry>
, 
<collectingCountry id="F1F516C2FFE6A821FB9D473D5BDFFDFD" box="[1089,1181,586,610]" name="India" pageId="5" pageNumber="383">Nicobar</collectingCountry>
, GNBR, by E. Eyarin Jehamalar.
</paragraph>
<paragraph id="895D5652FFE6A821FF1B47E55E69FD6D" blockId="5.[151,1437,259,2015]" pageId="5" pageNumber="383">
<emphasis id="BB968A40FFE6A821FF1B47E55E0DFD34" bold="true" box="[199,335,658,683]" pageId="5" pageNumber="383">Repository.</emphasis>
The 
<typeStatus id="5659E8F0FFE6A821FE5547E45EFBFD34" box="[393,441,659,683]" pageId="5" pageNumber="383">type</typeStatus>
specimens are deposited in the Central Entomological Laboratory of Zoological Survey of 
<collectingCountry id="F1F516C2FFE6A821FF6A47C05FB0FD50" box="[182,242,695,719]" name="India" pageId="5" pageNumber="383">India</collectingCountry>
, New Alipore, Kolkata, West Bengal, 
<collectingCountry id="F1F516C2FFE6A821FD7947C05DA0FD50" box="[677,738,695,719]" name="India" pageId="5" pageNumber="383">India</collectingCountry>
. 
<typeStatus id="5659E8F0FFE6A821FD3247C05C14FD50" box="[750,854,695,719]" pageId="5" pageNumber="383" type="holotype">Holotype</typeStatus>
Reg. No. 2748/H15, 
<typeStatus id="5659E8F0FFE6A821FB9647CF5BF5FD4F" box="[1098,1207,696,720]" pageId="5" pageNumber="383" type="paratype">Paratypes</typeStatus>
Reg. No. 2749/H15 to 2757/H15.
</paragraph>
</subSubSection>
<subSubSection id="C1F805D9FFE6A821FF1B47885D91FCA5" pageId="5" pageNumber="383" type="etymology">
<paragraph id="895D5652FFE6A821FF1B47885D91FCA5" blockId="5.[151,1437,259,2015]" pageId="5" pageNumber="383">
<emphasis id="BB968A40FFE6A821FF1B47885E0CFC88" bold="true" box="[199,334,767,791]" pageId="5" pageNumber="383">Etymology.</emphasis>
<taxonomicName id="4EE22DD1FFE6A821FE8646775DE1FC88" box="[346,675,767,791]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="383" phylum="Arthropoda" rank="species" species="venkataramani" status="sp. nov.">
<emphasis id="BB968A40FFE6A821FE8646775DE1FC88" box="[346,675,767,791]" italics="true" pageId="5" pageNumber="383">Tenagogonus venkataramani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0A5373BFFE6A821FD7346775C49FC88" box="[687,779,768,791]" pageId="5" pageNumber="383" rank="species">sp. nov.</taxonomicNameLabel>
is named in honour of Dr. K. Venkataraman, Director, Zoological Survey of 
<collectingCountry id="F1F516C2FFE6A821FE5046555E8AFCA5" box="[396,456,802,826]" name="India" pageId="5" pageNumber="383">India</collectingCountry>
, New Alipore, Kolkata.
</paragraph>
</subSubSection>
<subSubSection id="C1F805D9FFE6A821FF1B46315D9FFC55" pageId="5" pageNumber="383" type="diagnosis">
<paragraph id="895D5652FFE6A821FF1B46315D9FFC55" blockId="5.[151,1437,259,2015]" pageId="5" pageNumber="383">
<emphasis id="BB968A40FFE6A821FF1B46315E03FCC0" bold="true" box="[199,321,838,863]" pageId="5" pageNumber="383">Diagnosis.</emphasis>
<taxonomicName id="4EE22DD1FFE6A821FE94463F5DC9FCC0" box="[328,651,839,863]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="383" phylum="Arthropoda" rank="species" species="venkataramani" status="sp. nov.">
<emphasis id="BB968A40FFE6A821FE94463F5DC9FCC0" box="[328,651,839,863]" italics="true" pageId="5" pageNumber="383">Tenagogonus venkataramani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0A5373BFFE6A821FD4D46305DA9FCC0" box="[657,747,839,863]" pageId="5" pageNumber="383" rank="species">
<emphasis id="BB968A40FFE6A821FD4D46305DA9FCC0" bold="true" box="[657,747,839,863]" pageId="5" pageNumber="383">sp. nov.</emphasis>
</taxonomicNameLabel>
can easily be identified by the presence of an almost parallelsided, broad deep groove ventrally in the middle of the first genital segment (eighth abdominal segment) and the contiguous longitudinal setal tufts (
<figureCitation id="11D94AD7FFE6A821FDFC46F85DC3FC38" box="[544,641,911,935]" captionStart="FIGURE 1. A – H, T" captionStartId="2.[151,250,1919,1942]" captionTargetBox="[218,1385,188,1872]" captionTargetId="figure@2.[218,1386,157,1909]" captionTargetPageId="2" captionText="FIGURE 1. A – H, T. venkataramani sp. nov.: A, male (holotype), dorsal view; B, same, ventral view; C, female (paratype), dorsal view; D, male (holotype), left lateral view; E, macropterous female (paratype), dorsal view; F, apterous female (paratype), left lateral view; G, apterous female (paratype), terminal abdominal segments, ventral view; H, male (holotype), genital segments, left lateral view; I, T. ceylonensis, apterous male genital segments, left lateral view; J, T. nicobarensis, same." httpUri="https://zenodo.org/record/219885/files/figure.png" pageId="5" pageNumber="383">Figs. 1B</figureCitation>
; 2L). The seventh sternum of the female has a median apical projection and flap-like apical connexival projections laterally.
</paragraph>
</subSubSection>
<subSubSection id="C1F805D9FFE6A823FF1546A15D88F8B9" lastPageId="7" lastPageNumber="385" pageId="5" pageNumber="383" type="description">
<paragraph id="895D5652FFE6A821FF1546A15D53FB8D" blockId="5.[151,1437,259,2015]" pageId="5" pageNumber="383">
<emphasis id="BB968A40FFE6A821FF1546A15D55FC70" bold="true" box="[201,535,982,1007]" pageId="5" pageNumber="383">Description. Apterous male</emphasis>
(
<typeStatus id="5659E8F0FFE6A821FDF646A05DCDFC70" box="[554,655,983,1007]" pageId="5" pageNumber="383" type="holotype">holotype</typeStatus>
) (
<figureCitation id="11D94AD7FFE6A821FD7646A05C1EFC70" box="[682,860,983,1007]" captionStart="FIGURE 1. A – H, T" captionStartId="2.[151,250,1919,1942]" captionTargetBox="[218,1385,188,1872]" captionTargetId="figure@2.[218,1386,157,1909]" captionTargetPageId="2" captionText="FIGURE 1. A – H, T. venkataramani sp. nov.: A, male (holotype), dorsal view; B, same, ventral view; C, female (paratype), dorsal view; D, male (holotype), left lateral view; E, macropterous female (paratype), dorsal view; F, apterous female (paratype), left lateral view; G, apterous female (paratype), terminal abdominal segments, ventral view; H, male (holotype), genital segments, left lateral view; I, T. ceylonensis, apterous male genital segments, left lateral view; J, T. nicobarensis, same." httpUri="https://zenodo.org/record/219885/files/figure.png" pageId="5" pageNumber="383">Figs. 1A, B, D</figureCitation>
) Body length 5.82, width (across mesoacetabula) 1.93. Body narrower than female.
</paragraph>
<paragraph id="895D5652FFE6A821FF1B41695B6AFBC5" blockId="5.[151,1437,259,2015]" pageId="5" pageNumber="383">
<emphasis id="BB968A40FFE6A821FF1B41695E62FBA8" bold="true" box="[199,288,1054,1079]" pageId="5" pageNumber="383">Colour.</emphasis>
Yellowish brown with piceous to black stripes; antennae and legs fuscous; apices of femora, tibiae and tarsi dark; omphalium and surrounding area pale red; body venter pinkish yellow.
</paragraph>
<paragraph id="895D5652FFE6A821FF4B41105E3FFBE0" blockId="5.[151,1437,259,2015]" box="[151,381,1127,1151]" pageId="5" pageNumber="383">Structural characters</paragraph>
<paragraph id="895D5652FFE6A821FF1B41FC5B3AFAAD" blockId="5.[151,1437,259,2015]" pageId="5" pageNumber="383">
<emphasis id="BB968A40FFE6A821FF1B41FC5E4FFB3C" bold="true" box="[199,269,1163,1187]" pageId="5" pageNumber="383">Head.</emphasis>
Length: Head 0.94, antennal segments one to four 1.78, 1.55, 2.01, 3.01; rostral segments one to four 0.31, 0.16, 1.64, 0.34. Width: Head 1.29; head width between eyes 0.69. Head with pair of lateral shorter stripes and central longer V-shaped stripe, no stripes uniting anteriorly (
<figureCitation id="11D94AD7FFE6A821FCA941A55C92FB75" box="[885,976,1234,1258]" captionStart="FIGURE 1. A – H, T" captionStartId="2.[151,250,1919,1942]" captionTargetBox="[218,1385,188,1872]" captionTargetId="figure@2.[218,1386,157,1909]" captionTargetPageId="2" captionText="FIGURE 1. A – H, T. venkataramani sp. nov.: A, male (holotype), dorsal view; B, same, ventral view; C, female (paratype), dorsal view; D, male (holotype), left lateral view; E, macropterous female (paratype), dorsal view; F, apterous female (paratype), left lateral view; G, apterous female (paratype), terminal abdominal segments, ventral view; H, male (holotype), genital segments, left lateral view; I, T. ceylonensis, apterous male genital segments, left lateral view; J, T. nicobarensis, same." httpUri="https://zenodo.org/record/219885/files/figure.png" pageId="5" pageNumber="383">Fig. 1A</figureCitation>
). Subapex medially with pair of piceous spots covered by brown hue. Antennae much longer than body. Rostrum reaching to middle of mesosternum (
<figureCitation id="11D94AD7FFE6A821FAB241805FFBFAAD" captionStart="FIGURE 1. A – H, T" captionStartId="2.[151,250,1919,1942]" captionTargetBox="[218,1385,188,1872]" captionTargetId="figure@2.[218,1386,157,1909]" captionTargetPageId="2" captionText="FIGURE 1. A – H, T. venkataramani sp. nov.: A, male (holotype), dorsal view; B, same, ventral view; C, female (paratype), dorsal view; D, male (holotype), left lateral view; E, macropterous female (paratype), dorsal view; F, apterous female (paratype), left lateral view; G, apterous female (paratype), terminal abdominal segments, ventral view; H, male (holotype), genital segments, left lateral view; I, T. ceylonensis, apterous male genital segments, left lateral view; J, T. nicobarensis, same." httpUri="https://zenodo.org/record/219885/files/figure.png" pageId="5" pageNumber="383">Fig. 1B</figureCitation>
); apical flap of third rostral segment and fourth rostral segment fully black and shiny.
</paragraph>
<paragraph id="895D5652FFE6A821FF1B40485C1CF898" blockId="5.[151,1437,259,2015]" pageId="5" pageNumber="383">
<emphasis id="BB968A40FFE6A821FF1B40485E64FAC8" bold="true" box="[199,294,1343,1367]" pageId="5" pageNumber="383">Thorax.</emphasis>
Length: Anterior pronotal lobe 0.61; pronotal lobe 2.52; prosternum 0.58; mesosternum 2.41, metasternum 0.45; fore femora 2.39; fore tibia 2.14; fore tarsus I 0.19; fore tarsus II 0.26; mid femora 6.18; mid tibia 4.33; mid tarsus I 1.98; mid tarsus II 0.43; hind femora 6.08; hind tibia 2.74; hind tarsus I 0.51; hind tarsus II 0.35. Width: Pronotal lobe 0.96; fore femora 0.24, mid femora 0.17, hind femora 0.13. Pronotum and mesonotum with median and marginal stripes; mesopleuron with distinct stripe very close to pronotal lobe and obscure stripe on middle of mesopleuron; black stripe on sublateral region of mesosternum originating near base of fore trochanter and reaching near anterior region of mesoacetabular groove and not interrupted medially (
<figureCitation id="11D94AD7FFE6A821FAEC43605ACEF9B0" box="[1328,1420,1559,1583]" captionStart="FIGURE 1. A – H, T" captionStartId="2.[151,250,1919,1942]" captionTargetBox="[218,1385,188,1872]" captionTargetId="figure@2.[218,1386,157,1909]" captionTargetPageId="2" captionText="FIGURE 1. A – H, T. venkataramani sp. nov.: A, male (holotype), dorsal view; B, same, ventral view; C, female (paratype), dorsal view; D, male (holotype), left lateral view; E, macropterous female (paratype), dorsal view; F, apterous female (paratype), left lateral view; G, apterous female (paratype), terminal abdominal segments, ventral view; H, male (holotype), genital segments, left lateral view; I, T. ceylonensis, apterous male genital segments, left lateral view; J, T. nicobarensis, same." httpUri="https://zenodo.org/record/219885/files/figure.png" pageId="5" pageNumber="383">Fig. 1D</figureCitation>
); piceous spot on mesoacetabula open anteriorly; indistinct stripe on sublateral region of mesonotum brown; metacetabulum near to connexivum with curved piceous stripe; posterior margin of metasternum slightly sinuated laterally and convexed at middle; omphalium protruding just above posterior margin of metasternum, surrounded with pale reddish hue; stripe on metanotum not reaching posterior margin; W-shaped marking between metanotum and first abdominal tergite black. 
<emphasis id="BB968A40FFE6A821FDD143BB5D0BF97C" box="[525,585,1740,1763]" italics="true" pageId="5" pageNumber="383">Legs.</emphasis>
Fore femora and fore tibiae slightly curved; apex of each middle femur with small spine; apices of hind coxae near fifth abdominal sternum.
</paragraph>
<paragraph id="895D5652FFE6A822FF1B42655F8AFEA0" blockId="5.[151,1437,259,2015]" lastBlockId="6.[151,1437,151,932]" lastPageId="6" lastPageNumber="384" pageId="5" pageNumber="383">
<emphasis id="BB968A40FFE6A821FF1B42655E01F8B4" bold="true" box="[199,323,1810,1835]" pageId="5" pageNumber="383">Abdomen.</emphasis>
Nine abdominal tergites visible from above; length of tergites one to nine, respectively, 0.34, 0.22, 0.15, 0.14, 0.12, 0.18, 0.39, 0.27, 0.09; abdominal sternal length two to nine, respectively, 0.12, 0.08, 0.08, 0.08, 0.10, 0.27, 0.24, 0.44. Abdominal tergites one to two with broad piceous band medially enclosing yellow spot; tergite three with pair of narrow piceous stripes; tergites four to seven each with pair of small piceous spots anteriorly; tergite seven with pair of longitudinal linear streaks laterally near connexiva; intersegmental regions of connexiva each with piceous spot; abdominal sternum eight with tuft of brownish yellow setae on each side of the median longitudinal groove. (
<figureCitation id="11D94AD7FFE5A822FE3445E05D33FF30" box="[488,625,151,175]" captionStart="FIGURE 1. A – H, T" captionStartId="2.[151,250,1919,1942]" captionTargetBox="[218,1385,188,1872]" captionTargetId="figure@2.[218,1386,157,1909]" captionTargetPageId="2" captionText="FIGURE 1. A – H, T. venkataramani sp. nov.: A, male (holotype), dorsal view; B, same, ventral view; C, female (paratype), dorsal view; D, male (holotype), left lateral view; E, macropterous female (paratype), dorsal view; F, apterous female (paratype), left lateral view; G, apterous female (paratype), terminal abdominal segments, ventral view; H, male (holotype), genital segments, left lateral view; I, T. ceylonensis, apterous male genital segments, left lateral view; J, T. nicobarensis, same." httpUri="https://zenodo.org/record/219885/files/figure.png" pageId="6" pageNumber="384">Figs. 1B, H</figureCitation>
; 2L); posterior margin of eighth abdominal segment slightly trisinuate; anterior region of ninth abdominal sternum fuscous, shiny and downwardly sloped; connexiva not produced apically; parameres very small and oval; reticulation seen on endosoma; dorsal sclerite of endosoma curved, pair of lateral sclerites encircling dorsal region of endosoma (
<figureCitation id="11D94AD7FFE5A822FD2F44735C74FE83" box="[755,822,260,284]" captionStart="FIGURE 2. A, D, G, J, T" captionStartId="3.[151,250,1826,1849]" captionTargetBox="[259,1322,205,1749]" captionTargetId="figure@3.[242,1344,193,1790]" captionTargetPageId="3" captionText="FIGURE 2. A, D, G, J, T. ceylonensis: A, pygophore, parameres and endosoma, ventral view; D, endosoma, right lateral view; G, terminal abdominal segment of female showing connexival spines, dorsal view; J, male genital segments, ventral view. B, E, H, K, T. nicobarensis: B, pygophore, parameres and endosoma, ventral view; E, endosoma, right lateral view; H, terminal abdominal segment of female showing connexival spines, dorsal view; K, male genital segments, ventral view. C, F, I, L, T. venkataramani: C, pygophore, parameres and endosoma, ventral view; F, endosoma, right lateral view; I, terminal abdominal segment of female showing connexival spines, dorsal view; L, male genital segments, ventral view (ds—dorsal sclerite, en—endosoma, ls—lateral sclerite, pa—paramere, pg—pygophore, vs—ventral sclerite)." httpUri="https://zenodo.org/record/219886/files/figure.png" pageId="6" pageNumber="384">Fig. 2</figureCitation>
F); ventral sclerite broad and excavated anteriorly (
<figureCitation id="11D94AD7FFE5A822FAB344735FF9FEA0" captionStart="FIGURE 2. A, D, G, J, T" captionStartId="3.[151,250,1826,1849]" captionTargetBox="[259,1322,205,1749]" captionTargetId="figure@3.[242,1344,193,1790]" captionTargetPageId="3" captionText="FIGURE 2. A, D, G, J, T. ceylonensis: A, pygophore, parameres and endosoma, ventral view; D, endosoma, right lateral view; G, terminal abdominal segment of female showing connexival spines, dorsal view; J, male genital segments, ventral view. B, E, H, K, T. nicobarensis: B, pygophore, parameres and endosoma, ventral view; E, endosoma, right lateral view; H, terminal abdominal segment of female showing connexival spines, dorsal view; K, male genital segments, ventral view. C, F, I, L, T. venkataramani: C, pygophore, parameres and endosoma, ventral view; F, endosoma, right lateral view; I, terminal abdominal segment of female showing connexival spines, dorsal view; L, male genital segments, ventral view (ds—dorsal sclerite, en—endosoma, ls—lateral sclerite, pa—paramere, pg—pygophore, vs—ventral sclerite)." httpUri="https://zenodo.org/record/219886/files/figure.png" pageId="6" pageNumber="384">Fig. 2C</figureCitation>
).
</paragraph>
<paragraph id="895D5652FFE5A822FF1B443C5DCFFE18" blockId="6.[151,1437,151,932]" pageId="6" pageNumber="384">
<emphasis id="BB968A40FFE5A822FF1B443C5ECDFEFB" bold="true" box="[199,399,331,356]" pageId="6" pageNumber="384">Apterous female</emphasis>
(
<typeStatus id="5659E8F0FFE5A822FE7D443B5D50FEFB" box="[417,530,332,356]" pageId="6" pageNumber="384" type="paratype">Paratypes</typeStatus>
) (
<figureCitation id="11D94AD7FFE5A822FDF0443B5DF7FEFB" box="[556,693,332,356]" captionStart="FIGURE 1. A – H, T" captionStartId="2.[151,250,1919,1942]" captionTargetBox="[218,1385,188,1872]" captionTargetId="figure@2.[218,1386,157,1909]" captionTargetPageId="2" captionText="FIGURE 1. A – H, T. venkataramani sp. nov.: A, male (holotype), dorsal view; B, same, ventral view; C, female (paratype), dorsal view; D, male (holotype), left lateral view; E, macropterous female (paratype), dorsal view; F, apterous female (paratype), left lateral view; G, apterous female (paratype), terminal abdominal segments, ventral view; H, male (holotype), genital segments, left lateral view; I, T. ceylonensis, apterous male genital segments, left lateral view; J, T. nicobarensis, same." httpUri="https://zenodo.org/record/219885/files/figure.png" pageId="6" pageNumber="384">Figs. 1C, F</figureCitation>
). Mean body length 6.52, range 5.88–7.25 (n=5), width (across mesoacetabula) 2.41, range 2.13–2.76 (n=5).
</paragraph>
<paragraph id="895D5652FFE5A822FF1B44E45C19FE6B" blockId="6.[151,1437,151,932]" pageId="6" pageNumber="384">
<emphasis id="BB968A40FFE5A822FF1B44E45E62FE33" bold="true" box="[199,288,403,428]" pageId="6" pageNumber="384">Colour.</emphasis>
Similar to apterous male except the colour pattern of mesopleura and mesoacetabulum. The stripe on middle of the mesopleuron thick, broad, and connected with the lateral stripe of pronotum. The short stripes on the dorsal side of mesoacetabulum is enclosed a small yellow spot.
</paragraph>
<paragraph id="895D5652FFE5A822FF1B44885C47FD88" blockId="6.[151,1437,151,932]" box="[199,773,511,535]" pageId="6" pageNumber="384">Structural characters (range is given in parenthesis)</paragraph>
<paragraph id="895D5652FFE5A822FF1B47535D44FD38" blockId="6.[151,1437,151,932]" pageId="6" pageNumber="384">
<emphasis id="BB968A40FFE5A822FF1B47535E4FFDA3" bold="true" box="[199,269,548,572]" pageId="6" pageNumber="384">Head.</emphasis>
Mean length (n=5): Head 1.0 (0.93–1.03), antennal segments one to four, respectively, 1.57(1.45–1.72), 1.38 (1.23–1.47), 1.76 (1.57–1.91), 2.68 (2.41–2.85); rostral segments three and four, respectively, 1.82 (1.37–1.82) and 0.42 (0.36–0.49). Mean width (n=5): Head 1.30 (1.23–1.43), width between eyes 0.65 (0.56–0.73). Other characters as for 
<typeStatus id="5659E8F0FFE5A822FE4047F85D40FD38" box="[412,514,655,679]" pageId="6" pageNumber="384" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="895D5652FFE5A822FF1B47C35C98FC3B" blockId="6.[151,1437,151,932]" pageId="6" pageNumber="384">
<emphasis id="BB968A40FFE5A822FF1B47C35E64FD53" bold="true" box="[199,294,692,716]" pageId="6" pageNumber="384">Thorax.</emphasis>
Mean length (n=5): Anterior pronotal lobe 0.66 (0.60–0.68); pronotal lobe 2.67 (2.43–2.96); prosternum 0.52 (0.45–0.58); mesosternum 2.51(2.28–3.01), metasternum 0.55 (0.47–0.61); maximum width of pronotal lobe 1.09 (0.95–1.25). Mesopleuron with two distinct stripes, one very close to pronotal lobe and another on middle of mesopleuron anteriorly attached with contiguous stripe; large piceous spot on mesoacetabula enclosing oblique yellow to orange spot; posterior margin of metasternum not strongly bisinuate; fore femora and fore tibiae straight; apices of middle femora without spines; hind coxae surpassing third abdominal sternum; all other characters similar to those of 
<typeStatus id="5659E8F0FFE5A822FDFE46FB5DCAFC3B" box="[546,648,908,932]" pageId="6" pageNumber="384" type="holotype">holotype</typeStatus>
; leg measurements in 
<tableCitation id="C46063E9FFE5A822FC5C46FB5C91FC3B" box="[896,979,908,932]" captionStart="TABLE 1" captionStartId="6.[151,235,980,1001]" captionTargetBox="[156,1363,1029,1313]" captionTargetPageId="6" captionText="TABLE 1. Mean length of legs (in mm. n = 5), range is given in parenthesis." httpUri="http://table.plazi.org/id/DD9D06DAFFE5A822FF4B46A35CD7FC76" pageId="6" pageNumber="384" tableUuid="DD9D06DAFFE5A822FF4B46A35CD7FC76">Table 1</tableCitation>
.
</paragraph>
<caption id="DD9D06DAFFE5A822FF4B46A35CD7FC76" ID-Table-UUID="DD9D06DAFFE5A822FF4B46A35CD7FC76" box="[151,917,979,1002]" httpUri="http://table.plazi.org/id/DD9D06DAFFE5A822FF4B46A35CD7FC76" pageId="6" pageNumber="384" targetBox="[156,1363,1029,1313]" targetIsTable="true" targetPageId="6">
<paragraph id="895D5652FFE5A822FF4B46A35CD7FC76" blockId="6.[151,917,979,1002]" box="[151,917,979,1002]" pageId="6" pageNumber="384">
<emphasis id="BB968A40FFE5A822FF4B46A35E46FC76" bold="true" box="[151,260,980,1001]" pageId="6" pageNumber="384">TABLE 1.</emphasis>
Mean length of legs (in mm. n=5), range is given in parenthesis.
</paragraph>
</caption>
<paragraph id="895D5652FFE5A822FF4041715A0FFABF" pageId="6" pageNumber="384">
<table id="FBE2A4F2FFE557DBFF4041725A11FABE" box="[156,1363,1029,1313]" gridcols="5" gridrows="5" pageId="6" pageNumber="384">
<tr id="37D25410FFE557DBFF4041725A11FB84" box="[156,1363,1029,1051]" gridrow="0" pageId="6" pageNumber="384">
<th id="74033D6CFFE557DBFF4041725FBCFB84" box="[156,254,1029,1051]" gridcol="0" gridrow="0" pageId="6" pageNumber="384">Paratypes</th>
<th id="74033D6CFFE557DBFE9E41725EE2FB84" box="[322,416,1029,1051]" gridcol="1" gridrow="0" pageId="6" pageNumber="384">Leg parts</th>
<th id="74033D6CFFE557DBFE3541725C27FB84" box="[489,869,1029,1051]" gridcol="2" gridrow="0" pageId="6" pageNumber="384">Femur Tibia</th>
<th id="74033D6CFFE557DBFC7C41725B0AFB84" box="[928,1096,1029,1051]" gridcol="3" gridrow="0" pageId="6" pageNumber="384">I Tarsal segment</th>
<th id="74033D6CFFE557DBFB7841725A11FB84" box="[1188,1363,1029,1051]" gridcol="4" gridrow="0" pageId="6" pageNumber="384">II Tarsal segment</th>
</tr>
<tr id="37D25410FFE557DBFF4041445A11FBE8" box="[156,1363,1075,1143]" gridrow="1" pageId="6" pageNumber="384">
<th id="74033D6CFFE557DBFF4041445FBCFBE8" box="[156,254,1075,1143]" gridcol="0" gridrow="1" pageId="6" pageNumber="384">Apterous males</th>
<td id="74033D6CFFE557DBFE9E41445EE2FBE8" box="[322,416,1075,1143]" gridcol="1" gridrow="1" pageId="6" pageNumber="384">Fore leg Mid leg</td>
<td id="74033D6CFFE557DBFE3541445C27FBE8" box="[489,869,1075,1143]" gridcol="2" gridrow="1" pageId="6" pageNumber="384">2.26 (2.18–2.39) 2.07 (1.98–2.14) 5.84 (5.59–6.18) 4.22 (3.99–4.35)</td>
<td id="74033D6CFFE557DBFC7C41445B0AFBE8" box="[928,1096,1075,1143]" gridcol="3" gridrow="1" pageId="6" pageNumber="384">0.20 (0.17–0.24) 1.75 (1.58–1.98)</td>
<td id="74033D6CFFE557DBFB7841445A11FBE8" box="[1188,1363,1075,1143]" gridcol="4" gridrow="1" pageId="6" pageNumber="384">0.34 (0.26–0.41) 0.43 (0.4–0.45)</td>
</tr>
<tr id="37D25410FFE557DBFF4041FD5A11FB3F" box="[156,1363,1162,1184]" gridrow="2" pageId="6" pageNumber="384" rowspan-0="1">
<td id="74033D6CFFE557DBFE9E41FD5EE2FB3F" box="[322,416,1162,1184]" gridcol="1" gridrow="2" pageId="6" pageNumber="384">Hind leg</td>
<td id="74033D6CFFE557DBFE3541FD5C27FB3F" box="[489,869,1162,1184]" gridcol="2" gridrow="2" pageId="6" pageNumber="384">5.81 (5.52–6.08) 2.60 (2.45–2.74)</td>
<td id="74033D6CFFE557DBFC7C41FD5B0AFB3F" box="[928,1096,1162,1184]" gridcol="3" gridrow="2" pageId="6" pageNumber="384">0.44 (0.4–0.51)</td>
<td id="74033D6CFFE557DBFB7841FD5A11FB3F" box="[1188,1363,1162,1184]" gridcol="4" gridrow="2" pageId="6" pageNumber="384">0.35 (0.31–0.36)</td>
</tr>
<tr id="37D25410FFE557DBFF4041C35A11FB68" box="[156,1363,1204,1271]" gridrow="3" pageId="6" pageNumber="384">
<th id="74033D6CFFE557DBFF4041C35FBCFB68" box="[156,254,1204,1271]" gridcol="0" gridrow="3" pageId="6" pageNumber="384">Apterous females</th>
<td id="74033D6CFFE557DBFE9E41C35EE2FB68" box="[322,416,1204,1271]" gridcol="1" gridrow="3" pageId="6" pageNumber="384">Fore leg Mid leg</td>
<td id="74033D6CFFE557DBFE3541C35C27FB68" box="[489,869,1204,1271]" gridcol="2" gridrow="3" pageId="6" pageNumber="384">2.60 (2.16–2.98) 2.32 (2.09–2.64) 6.47 (6–6.99) 4.89 (4.38–5.69)</td>
<td id="74033D6CFFE557DBFC7C41C35B0AFB68" box="[928,1096,1204,1271]" gridcol="3" gridrow="3" pageId="6" pageNumber="384">0.25 (0.21–0.30) 2.05 (1.76–2.5)</td>
<td id="74033D6CFFE557DBFB7841C35A11FB68" box="[1188,1363,1204,1271]" gridcol="4" gridrow="3" pageId="6" pageNumber="384">0.40 (0.28–0.45) 0.49 (0.46–0.51)</td>
</tr>
<tr id="37D25410FFE557DBFF40407D5A11FABE" box="[156,1363,1290,1313]" gridrow="4" pageId="6" pageNumber="384" rowspan-0="1">
<td id="74033D6CFFE557DBFE9E407D5EE2FABE" box="[322,416,1290,1313]" gridcol="1" gridrow="4" pageId="6" pageNumber="384">Hind leg</td>
<td id="74033D6CFFE557DBFE35407D5C27FABE" box="[489,869,1290,1313]" gridcol="2" gridrow="4" pageId="6" pageNumber="384">6.45 (5.91–7.06) 3.13 (2.89–3.42)</td>
<td id="74033D6CFFE557DBFC7C407D5B0AFABE" box="[928,1096,1290,1313]" gridcol="3" gridrow="4" pageId="6" pageNumber="384">0.58 (0.52–0.62)</td>
<td id="74033D6CFFE557DBFB78407D5A11FABE" box="[1188,1363,1290,1313]" gridcol="4" gridrow="4" pageId="6" pageNumber="384">0.42 (0.35–0.47)</td>
</tr>
</table>
</paragraph>
<paragraph id="895D5652FFE5A822FF1B40135E9AF95F" blockId="6.[151,1437,1380,2016]" pageId="6" pageNumber="384">
<emphasis id="BB968A40FFE5A822FF1B40135E01FAE2" bold="true" box="[199,323,1380,1405]" pageId="6" pageNumber="384">Abdomen.</emphasis>
Eight tergites visible from above; mean length of tergites one to eight, respectively, 0.39 (0.32–0.51), 0.26 (0.21–0.32), 0.24 (0.18–0.28), 0.22 (0.19–0.25), 0.22 (0.2–0.25), 0.28 (0.23–0.30), 0.43 (0.41–0.44) (n=5). Mean length of sternites two to seven, respectively, 0.24 (0.22–0.27), 0.23 (0.19–0.25), 0.24 (0.22–0.27), 0.25 (0.23–0.27), 0.31 (0.26–0.34), 0.67 (0.62–0.72) (n=5). Tergites one to three with broad piceous band medially enclosing yellow spot, intersegmental region between tergites four and five with pair of piceous spots; anterior region of tergite six with pair of round piceous stripes, tergite seven laterally near connexiva with thick piceous stripe on each side; connexiva each with broad piceous stripe; posterior margin of tergite seven slightly concave; venter of abdomen paler; posterior margin of seventh sternum strongly bisinuate, with median triangular projection; connexiva apically produced into flap-like projections (
<figureCitation id="11D94AD7FFE5A822FC2143F25B1FF902" box="[1021,1117,1669,1693]" captionStart="FIGURE 1. A – H, T" captionStartId="2.[151,250,1919,1942]" captionTargetBox="[218,1385,188,1872]" captionTargetId="figure@2.[218,1386,157,1909]" captionTargetPageId="2" captionText="FIGURE 1. A – H, T. venkataramani sp. nov.: A, male (holotype), dorsal view; B, same, ventral view; C, female (paratype), dorsal view; D, male (holotype), left lateral view; E, macropterous female (paratype), dorsal view; F, apterous female (paratype), left lateral view; G, apterous female (paratype), terminal abdominal segments, ventral view; H, male (holotype), genital segments, left lateral view; I, T. ceylonensis, apterous male genital segments, left lateral view; J, T. nicobarensis, same." httpUri="https://zenodo.org/record/219885/files/figure.png" pageId="6" pageNumber="384">Figs. 1G</figureCitation>
, 
<date id="FD5C7092FFE5A822FBB643F25BC3F902" box="[1130,1153,1669,1693]" pageId="6" pageNumber="384">
<figureCitation id="11D94AD7FFE5A822FBB643F25B3AF902" box="[1130,1144,1669,1693]" captionStart="FIGURE 2. A, D, G, J, T" captionStartId="3.[151,250,1826,1849]" captionTargetBox="[259,1322,205,1749]" captionTargetId="figure@3.[242,1344,193,1790]" captionTargetPageId="3" captionText="FIGURE 2. A, D, G, J, T. ceylonensis: A, pygophore, parameres and endosoma, ventral view; D, endosoma, right lateral view; G, terminal abdominal segment of female showing connexival spines, dorsal view; J, male genital segments, ventral view. B, E, H, K, T. nicobarensis: B, pygophore, parameres and endosoma, ventral view; E, endosoma, right lateral view; H, terminal abdominal segment of female showing connexival spines, dorsal view; K, male genital segments, ventral view. C, F, I, L, T. venkataramani: C, pygophore, parameres and endosoma, ventral view; F, endosoma, right lateral view; I, terminal abdominal segment of female showing connexival spines, dorsal view; L, male genital segments, ventral view (ds—dorsal sclerite, en—endosoma, ls—lateral sclerite, pa—paramere, pg—pygophore, vs—ventral sclerite)." httpUri="https://zenodo.org/record/219886/files/figure.png" pageId="6" pageNumber="384">2</figureCitation>
I
</date>
&amp; 3C); genital segments well retracted into abdomen.
</paragraph>
<paragraph id="895D5652FFE5A822FF1B43BA5E24F897" blockId="6.[151,1437,1380,2016]" pageId="6" pageNumber="384">
<emphasis id="BB968A40FFE5A822FF1B43BA5E87F97A" bold="true" box="[199,453,1740,1765]" pageId="6" pageNumber="384">Macropterous female</emphasis>
(
<figureCitation id="11D94AD7FFE5A822FE0843BA5D6BF97A" box="[468,553,1741,1765]" captionStart="FIGURE 1. A – H, T" captionStartId="2.[151,250,1919,1942]" captionTargetBox="[218,1385,188,1872]" captionTargetId="figure@2.[218,1386,157,1909]" captionTargetPageId="2" captionText="FIGURE 1. A – H, T. venkataramani sp. nov.: A, male (holotype), dorsal view; B, same, ventral view; C, female (paratype), dorsal view; D, male (holotype), left lateral view; E, macropterous female (paratype), dorsal view; F, apterous female (paratype), left lateral view; G, apterous female (paratype), terminal abdominal segments, ventral view; H, male (holotype), genital segments, left lateral view; I, T. ceylonensis, apterous male genital segments, left lateral view; J, T. nicobarensis, same." httpUri="https://zenodo.org/record/219885/files/figure.png" pageId="6" pageNumber="384">Fig. 1E</figureCitation>
). Mean body length 6.71 (6.42–6.93) (n=3), width (across mesoacetabula) 2.27 (2.16–2.33) (n=3).
</paragraph>
<paragraph id="895D5652FFE5A822FF1B42635DD0F8B2" blockId="6.[151,1437,1380,2016]" box="[199,658,1812,1837]" pageId="6" pageNumber="384">
<emphasis id="BB968A40FFE5A822FF1B42635E62F8B2" bold="true" box="[199,288,1812,1837]" pageId="6" pageNumber="384">Colour.</emphasis>
Same as that of apterous female.
</paragraph>
<paragraph id="895D5652FFE5A822FF1B424F5EF6F8CF" blockId="6.[151,1437,1380,2016]" box="[199,436,1848,1872]" pageId="6" pageNumber="384">Structural Characters</paragraph>
<paragraph id="895D5652FFE5A822FF1B422A5DD4F87F" blockId="6.[151,1437,1380,2016]" pageId="6" pageNumber="384">
<emphasis id="BB968A40FFE5A822FF1B422A5E4FF8EA" bold="true" box="[199,269,1885,1909]" pageId="6" pageNumber="384">Head.</emphasis>
Mean length (n=3): Head 0.94 (0.80–1.03), antennal segments one to four, respectively, 1.70 (1.68–1.70), 1.38 (1.29–1.47), 1.76 (1.65–1.86), 2.70 (2.47–2.84); rostral segments three and four, respectively, 1.60 (1.55–1.66) and 0.44 (0.43–0.0.45). Mean width (n=3): Head 1.30 (1.27–1.31), width between eyes 0.67 (0.61–0.71). Other characters as for 
<typeStatus id="5659E8F0FFE5A822FDF142BF5DD1F87F" box="[557,659,1992,2016]" pageId="6" pageNumber="384" type="holotype">holotype</typeStatus>
.
</paragraph>
<caption id="DD9D06DAFFE4A823FF4B43575E30F9EB" httpUri="https://zenodo.org/record/219888/files/figure.png" pageId="7" pageNumber="385" targetBox="[161,1417,214,1530]" targetPageId="7">
<paragraph id="895D5652FFE4A823FF4B43575E30F9EB" blockId="7.[151,1436,1568,1652]" pageId="7" pageNumber="385">
<emphasis id="BB968A40FFE4A823FF4B43575E56F9A9" bold="true" box="[151,276,1568,1591]" pageId="7" pageNumber="385">FIGURE 4.</emphasis>
Distribution of 
<taxonomicName id="4EE22DD1FFE4A823FE6743555D17F9A8" box="[443,597,1569,1591]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="385" phylum="Arthropoda" rank="species" species="nicobarensis">
<emphasis id="BB968A40FFE4A823FE6743555D17F9A8" box="[443,597,1569,1591]" italics="true" pageId="7" pageNumber="385">T. nicobarensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4EE22DD1FFE4A823FD5643555C7EF9A8" box="[650,828,1569,1591]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="385" phylum="Arthropoda" rank="species" species="venkataramani" status="sp. nov.">
<emphasis id="BB968A40FFE4A823FD5643555C7EF9A8" box="[650,828,1569,1591]" italics="true" pageId="7" pageNumber="385">T. venkataramani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0A5373BFFE4A823FC9843565CD4F9A9" box="[836,918,1569,1590]" pageId="7" pageNumber="385" rank="species">
<emphasis id="BB968A40FFE4A823FC9843565CD4F9A9" bold="true" box="[836,918,1569,1590]" pageId="7" pageNumber="385">sp. nov.</emphasis>
</taxonomicNameLabel>
The triangles denote present records and the stars denote earlier records of 
<taxonomicName id="4EE22DD1FFE4A823FE4643365D76F9CA" box="[410,564,1599,1622]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="385" phylum="Arthropoda" rank="species" species="nicobarensis">
<emphasis id="BB968A40FFE4A823FE4643365D76F9CA" box="[410,564,1599,1622]" italics="true" pageId="7" pageNumber="385">T. nicobarensis</emphasis>
</taxonomicName>
from literature. The squares denote the distribution of 
<taxonomicName id="4EE22DD1FFE4A823FBB243365A63F9CA" box="[1134,1313,1599,1622]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="385" phylum="Arthropoda" rank="species" species="venkataramani" status="sp. nov.">
<emphasis id="BB968A40FFE4A823FBB243365A63F9CA" box="[1134,1313,1599,1622]" italics="true" pageId="7" pageNumber="385">T. venkataramani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0A5373BFFE4A823FAF443375A38F9CA" box="[1320,1402,1600,1621]" pageId="7" pageNumber="385" rank="species">
<emphasis id="BB968A40FFE4A823FAF443375A38F9CA" bold="true" box="[1320,1402,1600,1621]" pageId="7" pageNumber="385">sp. nov.</emphasis>
</taxonomicNameLabel>
on Great Nicobar Island.
</paragraph>
</caption>
<paragraph id="895D5652FFE4A823FF1B43D65D88F8B9" blockId="7.[151,1436,1697,1830]" pageId="7" pageNumber="385">
<emphasis id="BB968A40FFE4A823FF1B43D65E64F926" bold="true" box="[199,294,1697,1721]" pageId="7" pageNumber="385">Thorax.</emphasis>
Mean anterior pronotal lobe 0.56 (0.53–0.59); pronotal lobe length 2.58 (2.38–2.71) (n=3), width across humeral angle 1.71 (1.68–1.75) (n=3); hemelytral length 3.95 (3.83–4.12) (n=3), maximum width 1.09 (1.05–1.12) (n=3). Hemelytra cinerous brown; all other characters same as those of apterous 
<typeStatus id="5659E8F0FFE4A823FB74439D5A4EF89D" box="[1192,1292,1770,1794]" pageId="7" pageNumber="385" type="paratype">paratype</typeStatus>
. 
<emphasis id="BB968A40FFE4A823FF1B427B5E01F8BA" bold="true" box="[199,323,1804,1829]" pageId="7" pageNumber="385">Abdomen.</emphasis>
Same as that of apterous 
<typeStatus id="5659E8F0FFE4A823FDBE42795D84F8B9" box="[610,710,1806,1830]" pageId="7" pageNumber="385" type="paratype">paratype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C1F805D9FFE4A82CFF4B42075BCDFDA3" lastPageId="8" lastPageNumber="386" pageId="7" pageNumber="385" type="discussion">
<paragraph id="895D5652FFE4A823FF4B42075E57F816" blockId="7.[151,277,1904,1929]" box="[151,277,1904,1929]" pageId="7" pageNumber="385">
<emphasis id="BB968A40FFE4A823FF4B42075E57F816" bold="true" box="[151,277,1904,1929]" pageId="7" pageNumber="385">Discussion</emphasis>
</paragraph>
<paragraph id="895D5652FFE4A82CFF4B42CE5D4CFE33" blockId="7.[151,1436,1977,2037]" lastBlockId="8.[151,1437,151,572]" lastPageId="8" lastPageNumber="386" pageId="7" pageNumber="385">
The male of 
<taxonomicName id="4EE22DD1FFE4A823FEEE42CD5EB8F84E" box="[306,506,1977,2001]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="385" phylum="Arthropoda" rank="species" species="venkataramani" status="sp. nov.">
<emphasis id="BB968A40FFE4A823FEEE42CD5EB8F84E" box="[306,506,1977,2001]" italics="true" pageId="7" pageNumber="385">T. venkataramani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0A5373BFFE4A823FDD942CE5D21F84E" box="[517,611,1977,2001]" pageId="7" pageNumber="385" rank="species">
<emphasis id="BB968A40FFE4A823FDD942CE5D21F84E" bold="true" box="[517,611,1977,2001]" pageId="7" pageNumber="385">sp. nov.</emphasis>
</taxonomicNameLabel>
is closely related to 
<taxonomicName id="4EE22DD1FFE4A823FC8342CD5BE8F84E" authority="Andersen, T." authorityName="Andersen, T." box="[863,1194,1977,2001]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="385" phylum="Arthropoda" rank="species" species="nicobarensis">
<emphasis id="BB968A40FFE4A823FC8342CD5B4EF84E" box="[863,1036,1977,2001]" italics="true" pageId="7" pageNumber="385">T. nicobarensis</emphasis>
Andersen, 
<emphasis id="BB968A40FFE4A823FB4942CD5BE8F84E" box="[1173,1194,1978,2001]" italics="true" pageId="7" pageNumber="385">T.</emphasis>
</taxonomicName>
maai Hungerford &amp; Matsuda, and 
<taxonomicName id="4EE22DD1FFE4A823FEE942A95D3EF86A" authority="Chen &amp; Nieser" authorityName="Chen &amp; Nieser" box="[309,636,2013,2037]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="385" phylum="Arthropoda" rank="species" species="akanthinos">
<emphasis id="BB968A40FFE4A823FEE942A95E8EF86A" box="[309,460,2013,2037]" italics="true" pageId="7" pageNumber="385">T. akanthinos</emphasis>
Chen &amp; Nieser
</taxonomicName>
in having the broad deep median longitudinal depression on the venter of the first genital segment and the absence of setal tufts on the sides of the pygophore (ninth abdominal segment). However, this new species can be differentiated from 
<taxonomicName id="4EE22DD1FFEBA82CFD3245CA5CD5FF4B" box="[750,919,188,212]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="386" phylum="Arthropoda" rank="species" species="nicobarensis">
<emphasis id="BB968A40FFEBA82CFD3245CA5CD5FF4B" box="[750,919,188,212]" italics="true" pageId="8" pageNumber="386">T. nicobarensis</emphasis>
</taxonomicName>
by the absence of the V-shaped impression on abdominal sternite VII. The new species shares one character with 
<taxonomicName id="4EE22DD1FFEBA82CFC6E45965B0FFF67" box="[946,1101,224,248]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="386" phylum="Arthropoda" rank="species" species="akanthinos">
<emphasis id="BB968A40FFEBA82CFC6E45965B0FFF67" box="[946,1101,224,248]" italics="true" pageId="8" pageNumber="386">T. akanthinos</emphasis>
</taxonomicName>
and 
<taxonomicName id="4EE22DD1FFEBA82CFB5045965BA1FF67" box="[1164,1251,225,248]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="386" phylum="Arthropoda" rank="species" species="maai">
<emphasis id="BB968A40FFEBA82CFB5045965BA1FF67" box="[1164,1251,225,248]" italics="true" pageId="8" pageNumber="386">T. maai</emphasis>
</taxonomicName>
: the abdominal segment VIII has a broad median longitudinal groove. This groove is bordered by slightly diverging carinae in 
<emphasis id="BB968A40FFEBA82CFA5B44725E5BFEDF" italics="true" pageId="8" pageNumber="386">
<taxonomicName id="4EE22DD1FFEBA82CFA5B44725E57FEDF" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="386" phylum="Arthropoda" rank="species" species="akanthinos">T. akanthinos</taxonomicName>
,
</emphasis>
but in 
<taxonomicName id="4EE22DD1FFEBA82CFEB4445E5D69FEDF" box="[360,555,296,320]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="386" phylum="Arthropoda" rank="species" species="venkataramani" status="sp. nov.">
<emphasis id="BB968A40FFEBA82CFEB4445E5D69FEDF" box="[360,555,296,320]" italics="true" pageId="8" pageNumber="386">T. venkataramani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0A5373BFFEBA82CFDEF445F5DCEFEDF" box="[563,652,296,320]" pageId="8" pageNumber="386" rank="species">
<emphasis id="BB968A40FFEBA82CFDEF445F5DCEFEDF" bold="true" box="[563,652,296,320]" pageId="8" pageNumber="386">sp. nov.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4EE22DD1FFEBA82CFD1F445E5C57FEDF" box="[707,789,297,320]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="386" phylum="Arthropoda" rank="species" species="maai">
<emphasis id="BB968A40FFEBA82CFD1F445E5C57FEDF" box="[707,789,297,320]" italics="true" pageId="8" pageNumber="386">T. maai</emphasis>
</taxonomicName>
the carinae are straight. However, it can be differentiated with 
<taxonomicName id="4EE22DD1FFEBA82CFF13443A5E62FEFB" box="[207,288,333,356]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="386" phylum="Arthropoda" rank="species" species="maai">
<emphasis id="BB968A40FFEBA82CFF13443A5E62FEFB" box="[207,288,333,356]" italics="true" pageId="8" pageNumber="386">T. maai</emphasis>
</taxonomicName>
by the presence of the tuft of setae on the carinae. The females of 
<taxonomicName id="4EE22DD1FFEBA82CFBD8443A5B17FEFB" box="[1028,1109,333,356]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="386" phylum="Arthropoda" rank="species" species="maai">
<emphasis id="BB968A40FFEBA82CFBD8443A5B17FEFB" box="[1028,1109,333,356]" italics="true" pageId="8" pageNumber="386">T. maai</emphasis>
</taxonomicName>
and 
<taxonomicName id="4EE22DD1FFEBA82CFB50443A5A60FEFB" box="[1164,1314,332,356]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="386" phylum="Arthropoda" rank="species" species="akanthinos">
<emphasis id="BB968A40FFEBA82CFB50443A5A60FEFB" box="[1164,1314,332,356]" italics="true" pageId="8" pageNumber="386">T. akanthinos</emphasis>
</taxonomicName>
have short and pointed connexival spine, whereas these connexival spines are flap-like and blunt in 
<taxonomicName id="4EE22DD1FFEBA82CFB4044065A21FE17" box="[1180,1379,368,392]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="386" phylum="Arthropoda" rank="species" species="venkataramani">
<emphasis id="BB968A40FFEBA82CFB4044065A21FE17" box="[1180,1379,368,392]" italics="true" pageId="8" pageNumber="386">T. venkataramani</emphasis>
</taxonomicName>
, and long and blunt in T. 
<taxonomicName id="4EE22DD1FFEBA82CFEA544E35D45FE33" box="[377,519,404,428]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="386" phylum="Arthropoda" rank="species" species="nicobarensis">
<emphasis id="BB968A40FFEBA82CFEA544E35D45FE33" box="[377,519,404,428]" italics="true" pageId="8" pageNumber="386">nicobarensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="895D5652FFEBA82CFF1B44CF5BCDFDA3" blockId="8.[151,1437,151,572]" pageId="8" pageNumber="386">
A total of 
<specimenCount id="9FE49DDBFFEBA82CFEE144C05EA7FE50" box="[317,485,439,463]" pageId="8" pageNumber="386" type="generic">121 specimens</specimenCount>
from 17 localities of 3 species of 
<taxonomicName id="4EE22DD1FFEBA82CFCB044CE5B42FE4F" box="[876,1024,441,464]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="386" phylum="Arthropoda" rank="genus">
<emphasis id="BB968A40FFEBA82CFCB044CE5B42FE4F" box="[876,1024,441,464]" italics="true" pageId="8" pageNumber="386">Tenagogonus</emphasis>
</taxonomicName>
were examined in the present study (61 
<taxonomicName id="4EE22DD1FFEBA82CFF1844AA5EC5FE6B" box="[196,391,476,500]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="386" phylum="Arthropoda" rank="species" species="venkataramani" status="sp. nov.">
<emphasis id="BB968A40FFEBA82CFF1844AA5EC5FE6B" box="[196,391,476,500]" italics="true" pageId="8" pageNumber="386">T. venkataramani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0A5373BFFEBA82CFE5244AB5EAAFE6B" box="[398,488,476,500]" pageId="8" pageNumber="386" rank="species">
<emphasis id="BB968A40FFEBA82CFE5244AB5EAAFE6B" bold="true" box="[398,488,476,500]" pageId="8" pageNumber="386">sp. nov.</emphasis>
</taxonomicNameLabel>
, 51 
<taxonomicName id="4EE22DD1FFEBA82CFDC644AA5D86FE6B" box="[538,708,476,500]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="386" phylum="Arthropoda" rank="species" species="nicobarensis">
<emphasis id="BB968A40FFEBA82CFDC644AA5D86FE6B" box="[538,708,476,500]" italics="true" pageId="8" pageNumber="386">T. nicobarensis</emphasis>
</taxonomicName>
, 10 
<taxonomicName id="4EE22DD1FFEBA82CFD2944AA5CD0FE6B" box="[757,914,476,500]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="386" phylum="Arthropoda" rank="species" species="ceylonensis">
<emphasis id="BB968A40FFEBA82CFD2944AA5CD0FE6B" box="[757,914,476,500]" italics="true" pageId="8" pageNumber="386">T. ceylonensis</emphasis>
</taxonomicName>
). Of the three species, only a small number of 
<taxonomicName id="4EE22DD1FFEBA82CFF4B47765E7BFD87" box="[151,313,512,536]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="386" phylum="Arthropoda" rank="species" species="ceylonensis">
<emphasis id="BB968A40FFEBA82CFF4B47765E7BFD87" box="[151,313,512,536]" italics="true" pageId="8" pageNumber="386">T. ceylonensis</emphasis>
</taxonomicName>
were collected from a single site in an agricultural area (monoculture rubber plantation). The distribution maps of known Indian species of 
<taxonomicName id="4EE22DD1FFEBA82CFD4B47525C69FDA3" box="[663,811,549,572]" class="Insecta" family="Gerridae" genus="Tenagogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="386" phylum="Arthropoda" rank="genus">
<emphasis id="BB968A40FFEBA82CFD4B47525C69FDA3" box="[663,811,549,572]" italics="true" pageId="8" pageNumber="386">Tenagogonus</emphasis>
</taxonomicName>
are provided in 
<figureCitation id="11D94AD7FFEBA82CFC3F47535B6CFDA3" box="[995,1070,548,572]" captionStart="FIGURE 3. A, B, C" captionStartId="4.[151,250,1951,1974]" captionTargetBox="[157,1412,206,1932]" captionTargetId="figure@4.[151,1436,193,1938]" captionTargetPageId="4" captionText="FIGURE 3. A, B, C, Female terminal abdominal segments showing connexival spines, lateral view; A, T. ceylonensis; B, T. nicobarensis; C, T. venkataramani. Distribution of T. ceylonensis. Round dots denote records from literature and the star denotes the present record from Tamil Nadu." httpUri="https://zenodo.org/record/219887/files/figure.png" pageId="8" pageNumber="386">Figs. 3</figureCitation>
D and 4.
</paragraph>
</subSubSection>
</treatment>
</document>