1771 lines
120 KiB
XML
1771 lines
120 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/herpetozoa.36.e104707" ID-Pensoft-Pub="2682-955X-36-95" ID-Pensoft-UUID="FCFA9C9B396D5C3E9A41FDD281753499" ID-ZooBank="C17C61D315384F32B934B28A37E1A376" ModsDocID="2682-955X-36-95" checkinTime="1682372474577" checkinUser="pensoft" docAuthor="Gruenwald, Christoph I., Montano-Ruvalcaba, Carlos, Jones, Jason M., Ahumada-Carrillo, Ivan, Gruenwald, Andre J., Zheng, Jiacheng, Strickland, Jason L. & Reyes-Velasco, Jacobo" docDate="2023" docId="01355F73A0DA52D5B33EDBE47A4A9AA0" docLanguage="en" docName="Herpetozoa 36: 95-111" docOrigin="Herpetozoa 36" docPubDate="2023-04-24" docSource="http://dx.doi.org/10.3897/herpetozoa.36.e104707" docTitle="Eleutherodactylus franzi sp. nov." docType="treatment" docVersion="2" id="FCFA9C9B396D5C3E9A41FDD281753499" lastPageNumber="95" masterDocId="FCFA9C9B396D5C3E9A41FDD281753499" masterDocTitle="A novel species of piping frog Eleutherodactylus (Anura, Eleutherodactylidae) from southern Mexico" masterLastPageNumber="111" masterPageNumber="95" pageNumber="95" updateTime="1682372776239" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>A novel species of piping frog Eleutherodactylus (Anura, Eleutherodactylidae) from southern Mexico</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Gruenwald, Christoph I.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5994-555X</mods:nameIdentifier>
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<mods:affiliation>Biencom Real Estate, Carretera Chapala-Jocotepec # 57 - 1, C. P. 45920, Ajijic, Jalisco, Mexico & Biodiversa A. C., Avenida de la Ribera # 203, C. P. 45900, Chapala, Jalisco, Mexico & Herp. mx A. C. C. P. 28989, Villa de Alvarez, Colima, Mexico</mods:affiliation>
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<mods:nameIdentifier type="email">cgruenwald@switaki.com</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Montano-Ruvalcaba, Carlos</mods:namePart>
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<mods:affiliation>Biodiversa A. C., Avenida de la Ribera # 203, C. P. 45900, Chapala, Jalisco, Mexico & Herp. mx A. C. C. P. 28989, Villa de Alvarez, Colima, Mexico</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jones, Jason M.</mods:namePart>
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<mods:affiliation>Biodiversa A. C., Avenida de la Ribera # 203, C. P. 45900, Chapala, Jalisco, Mexico & Herp. mx A. C. C. P. 28989, Villa de Alvarez, Colima, Mexico</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ahumada-Carrillo, Ivan</mods:namePart>
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<mods:affiliation>Biodiversa A. C., Avenida de la Ribera # 203, C. P. 45900, Chapala, Jalisco, Mexico & Herp. mx A. C. C. P. 28989, Villa de Alvarez, Colima, Mexico</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gruenwald, Andre J.</mods:namePart>
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<mods:affiliation>Herp. mx A. C. C. P. 28989, Villa de Alvarez, Colima, Mexico & Instituto Loyola C. P. 28989, San Antonio Tlayacapan, Jalisco, Mexico</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zheng, Jiacheng</mods:namePart>
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<mods:affiliation>Department of Biology, University of South Alabama, 5871 USA Dr. N., Mobile, P. C. 36688, Alabama, USA</mods:affiliation>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Strickland, Jason L.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1927-7259</mods:nameIdentifier>
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<mods:affiliation>Department of Biology, University of South Alabama, 5871 USA Dr. N., Mobile, P. C. 36688, Alabama, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Reyes-Velasco, Jacobo</mods:namePart>
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<mods:affiliation>Herp. mx A. C. C. P. 28989, Villa de Alvarez, Colima, Mexico & Department of Biology, Zoology and Evolutionary Biology, Universitaet Konstanz, 78457 Konstanz, Germany</mods:affiliation>
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<mods:title>Herpetozoa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-04-24</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>36</mods:number>
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</mods:detail>
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<mods:start>95</mods:start>
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<mods:url>http://dx.doi.org/10.3897/herpetozoa.36.e104707</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/herpetozoa.36.e104707</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2682-955X-36-95</mods:identifier>
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<mods:identifier type="ZooBank">C17C61D315384F32B934B28A37E1A376</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:18CE837E-C4B3-4468-BA37-AFB9213A9509" httpUri="http://treatment.plazi.org/id/01355F73A0DA52D5B33EDBE47A4A9AA0" lastPageNumber="95" pageId="0" pageNumber="95">
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<taxonomicName LSID="https://zoobank.org/18CE837E-C4B3-4468-BA37-AFB9213A9509" authority="sp. nov." authorityName="sp. nov." class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus franzi" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="species" species="franzi">Eleutherodactylus franzi sp. nov.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="95" type="description">
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<paragraph pageId="0" pageNumber="95">
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||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype of Eleutherodactylus franzi, sp. nov., INIRENA 2900 (CIG 01725), El Cucharillo, Municipio de Ixcateopan de Cuauhtemoc, Guerrero, Mexico in life (A-C) and in preservative (D, E)." figureDoi="10.3897/herpetozoa.36.e104707.figure2" httpUri="https://binary.pensoft.net/fig/841843" pageId="0" pageNumber="95">Figs 2</figureCitation>
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Some of the paratypes of Eleutherodactylus franzi sp. nov., in life. A-C. INIRENA 2902 (CIG 01717) El Cucharillo, Municipio de Ixcateopan de Cuauhtemoc, Guerrero, Mexico; D-F. INIRENA 2910 (CIG 01726) El Cucharillo, Municipio de Ixcateopan de Cuauhtemoc, Guerrero, Mexico; G-I. INIRENA 2895 (CIG 01727), 3 km E of Ixcateopan de Cuauhtemoc, Municipio de Ixcateopan de Cuauhtemoc, Guerrero, Mexico; J-L. INIRENA 2896 (CIG 01728), 3 km E of Ixcateopan de Cuauhtemoc, Municipio de Ixcateopan de Cuauhtemoc, Guerrero, Mexico; M-O. INIRENA 2898 (CIG 01731), between El Cucharillo and Chichila, Municipio de Taxco de Alarcon, Guerrero, Mexico." figureDoi="10.3897/herpetozoa.36.e104707.figure3" httpUri="https://binary.pensoft.net/fig/841844" pageId="0" pageNumber="95">, 3</figureCitation>
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Spectrograms (top) and oscillograms (bottom) of the advertisement calls of Eleutherodactylus franzi sp. nov., recorded at El Cucharillo, Municipio de Ixcateopan de Cuauhtemoc, Guerrero, Mexico (A, B) and at 3 km E of Ixcateopan de Cuauhtemoc, Guerrero, Mexico (C, D)." figureDoi="10.3897/herpetozoa.36.e104707.figure4" httpUri="https://binary.pensoft.net/fig/841845" pageId="0" pageNumber="95">, 4</figureCitation>
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A. Ventral aspect of the hand of the holotype of Eleutherodactylus franzi sp. nov., INIRENA 2900 (CIG 01725) from El Cucharillo, Municipio de Ixcateopan de Cuauhtemoc, Guerrero, Mexico; B. Ventral aspect of the hand of a Eleutherodactylus humboldti, INIRENA 2911 (CIG 01703) from 9.4 km N of Valle de Bravo junction on Valle de Bravo - Toluca toll road, near San Bartolo, Municipio de Amanalco, Estado de Mexico, Mexico." figureDoi="10.3897/herpetozoa.36.e104707.figure5" httpUri="https://binary.pensoft.net/fig/841846" pageId="0" pageNumber="95">
|
||
, 5A
|
||
<normalizedToken originalValue="Franz’s">Franz's</normalizedToken>
|
||
Piping Frog / Rana gaitera de Franz
|
||
</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="95" type="type material">
|
||
<paragraph pageId="0" pageNumber="95">Type material.</paragraph>
|
||
<paragraph continue="true" pageId="0" pageNumber="95">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Holotype</emphasis>
|
||
.
|
||
</emphasis>
|
||
INIRENA 2900 (CIG 01725). Adult male (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype of Eleutherodactylus franzi, sp. nov., INIRENA 2900 (CIG 01725), El Cucharillo, Municipio de Ixcateopan de Cuauhtemoc, Guerrero, Mexico in life (A-C) and in preservative (D, E)." figureDoi="10.3897/herpetozoa.36.e104707.figure2" httpUri="https://binary.pensoft.net/fig/841843" pageId="0" pageNumber="95">2</figureCitation>
|
||
), El Cucharillo, Municipio de Ixcateopan de
|
||
<normalizedToken originalValue="Cuauhtémoc">Cuauhtemoc</normalizedToken>
|
||
(
|
||
<geoCoordinate degrees="18.5331" direction="north" orientation="latitude" precision="5" value="18.5331">18.5331</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="99.7335" direction="west" orientation="longitude" precision="5" value="-99.7335">-99.7335</geoCoordinate>
|
||
, 2,315 m a.s.l.; datum = WGS84), Guerrero, Mexico, collected on 27 June 2020 by
|
||
<normalizedToken originalValue="Héctor">Hector</normalizedToken>
|
||
<normalizedToken originalValue="Franz-Chávez">Franz-Chavez</normalizedToken>
|
||
, Christoph I.
|
||
<normalizedToken originalValue="Grünwald">Gruenwald</normalizedToken>
|
||
,
|
||
<normalizedToken originalValue="André">Andre</normalizedToken>
|
||
J.
|
||
<normalizedToken originalValue="Grünwald">Gruenwald</normalizedToken>
|
||
and Kimberly
|
||
<normalizedToken originalValue="Montelongo-Chávez">Montelongo-Chavez</normalizedToken>
|
||
.
|
||
<emphasis bold="true" pageId="0" pageNumber="95">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Paratypes</emphasis>
|
||
(n = 14; Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Some of the paratypes of Eleutherodactylus franzi sp. nov., in life. A-C. INIRENA 2902 (CIG 01717) El Cucharillo, Municipio de Ixcateopan de Cuauhtemoc, Guerrero, Mexico; D-F. INIRENA 2910 (CIG 01726) El Cucharillo, Municipio de Ixcateopan de Cuauhtemoc, Guerrero, Mexico; G-I. INIRENA 2895 (CIG 01727), 3 km E of Ixcateopan de Cuauhtemoc, Municipio de Ixcateopan de Cuauhtemoc, Guerrero, Mexico; J-L. INIRENA 2896 (CIG 01728), 3 km E of Ixcateopan de Cuauhtemoc, Municipio de Ixcateopan de Cuauhtemoc, Guerrero, Mexico; M-O. INIRENA 2898 (CIG 01731), between El Cucharillo and Chichila, Municipio de Taxco de Alarcon, Guerrero, Mexico." figureDoi="10.3897/herpetozoa.36.e104707.figure3" httpUri="https://binary.pensoft.net/fig/841844" pageId="0" pageNumber="95">3</figureCitation>
|
||
).
|
||
</emphasis>
|
||
INIRENA 2901--10 (CIG 01716-24, 01726), 10 adult males, collected at same locality and on same date as holotype; INIRENA 2895--97 (CIG 01727-29), 3 adult males, 3 km E of Ixcateopan de
|
||
<normalizedToken originalValue="Cuauhtémoc">Cuauhtemoc</normalizedToken>
|
||
, Municipio de Ixcateopan de
|
||
<normalizedToken originalValue="Cuauhtémoc">Cuauhtemoc</normalizedToken>
|
||
(
|
||
<geoCoordinate degrees="18.5076" direction="north" orientation="latitude" precision="5" value="18.5076">18.5076</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="99.7329" direction="west" orientation="longitude" precision="5" value="-99.7329">-99.7329</geoCoordinate>
|
||
, 2,060 m a.s.l.; datum = WGS84), Guerrero collected on 27 June 2020 by
|
||
<normalizedToken originalValue="Héctor">Hector</normalizedToken>
|
||
<normalizedToken originalValue="Franz-Chávez">Franz-Chavez</normalizedToken>
|
||
, Christoph I.
|
||
<normalizedToken originalValue="Grünwald">Gruenwald</normalizedToken>
|
||
,
|
||
<normalizedToken originalValue="André">Andre</normalizedToken>
|
||
J.
|
||
<normalizedToken originalValue="Grünwald">Gruenwald</normalizedToken>
|
||
and Kimberly
|
||
<normalizedToken originalValue="Montelongo-Chávez">Montelongo-Chavez</normalizedToken>
|
||
; INIRENA 2898 (CIG 01731), 1 adult male, between El Cucharillo and Chichila, Municipio de Taxco de
|
||
<normalizedToken originalValue="Alarcón">Alarcon</normalizedToken>
|
||
, (
|
||
<geoCoordinate degrees="18.5335" direction="north" orientation="latitude" precision="5" value="18.5335">18.5335</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="99.7117" direction="west" orientation="longitude" precision="5" value="-99.7117">-99.7117</geoCoordinate>
|
||
, 2,260 m a.s.l.; datum = WGS84), Guerrero, Mexico, collected on 27 June 2020 by
|
||
<normalizedToken originalValue="Héctor">Hector</normalizedToken>
|
||
<normalizedToken originalValue="Franz-Chávez">Franz-Chavez</normalizedToken>
|
||
, Christoph I.
|
||
<normalizedToken originalValue="Grünwald">Gruenwald</normalizedToken>
|
||
,
|
||
<normalizedToken originalValue="André">Andre</normalizedToken>
|
||
J.
|
||
<normalizedToken originalValue="Grünwald">Gruenwald</normalizedToken>
|
||
and Kimberly
|
||
<normalizedToken originalValue="Montelongo-Chávez">Montelongo-Chavez</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
<caption doi="10.3897/herpetozoa.36.e104707.figure2" httpUri="https://binary.pensoft.net/fig/841843" pageId="0" pageNumber="95" start="Figure 2" startId="F2">
|
||
<paragraph pageId="0" pageNumber="95">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Figure 2.</emphasis>
|
||
Holotype of
|
||
<taxonomicName class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus franzi" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Eleutherodactylus franzi</emphasis>
|
||
</taxonomicName>
|
||
, sp. nov., INIRENA 2900 (CIG 01725), El Cucharillo, Municipio de Ixcateopan de
|
||
<normalizedToken originalValue="Cuauhtémoc">Cuauhtemoc</normalizedToken>
|
||
, Guerrero, Mexico in life (
|
||
<emphasis bold="true" pageId="0" pageNumber="95">A-C</emphasis>
|
||
) and in preservative (
|
||
<emphasis bold="true" pageId="0" pageNumber="95">D, E</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/herpetozoa.36.e104707.figure3" httpUri="https://binary.pensoft.net/fig/841844" pageId="0" pageNumber="95" start="Figure 3" startId="F3">
|
||
<paragraph pageId="0" pageNumber="95">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Figure 3.</emphasis>
|
||
Some of the paratypes of
|
||
<taxonomicName class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus franzi" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Eleutherodactylus franzi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., in life.
|
||
<emphasis bold="true" pageId="0" pageNumber="95">A-C.</emphasis>
|
||
INIRENA 2902 (CIG 01717) El Cucharillo, Municipio de Ixcateopan de
|
||
<normalizedToken originalValue="Cuauhtémoc">Cuauhtemoc</normalizedToken>
|
||
, Guerrero, Mexico;
|
||
<emphasis bold="true" pageId="0" pageNumber="95">D-F.</emphasis>
|
||
INIRENA 2910 (CIG 01726) El Cucharillo, Municipio de Ixcateopan de
|
||
<normalizedToken originalValue="Cuauhtémoc">Cuauhtemoc</normalizedToken>
|
||
, Guerrero, Mexico;
|
||
<emphasis bold="true" pageId="0" pageNumber="95">G-I.</emphasis>
|
||
INIRENA 2895 (CIG 01727), 3 km E of Ixcateopan de
|
||
<normalizedToken originalValue="Cuauhtémoc">Cuauhtemoc</normalizedToken>
|
||
, Municipio de Ixcateopan de
|
||
<normalizedToken originalValue="Cuauhtémoc">Cuauhtemoc</normalizedToken>
|
||
, Guerrero, Mexico;
|
||
<emphasis bold="true" pageId="0" pageNumber="95">J-L.</emphasis>
|
||
INIRENA 2896 (CIG 01728), 3 km E of Ixcateopan de
|
||
<normalizedToken originalValue="Cuauhtémoc">Cuauhtemoc</normalizedToken>
|
||
, Municipio de Ixcateopan de
|
||
<normalizedToken originalValue="Cuauhtémoc">Cuauhtemoc</normalizedToken>
|
||
, Guerrero, Mexico;
|
||
<emphasis bold="true" pageId="0" pageNumber="95">M-O.</emphasis>
|
||
INIRENA 2898 (CIG 01731), between El Cucharillo and Chichila, Municipio de Taxco de
|
||
<normalizedToken originalValue="Alarcón">Alarcon</normalizedToken>
|
||
, Guerrero, Mexico.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="95" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="95">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="95">
|
||
Based on our phylogenetic analysis, this is a member of the genus
|
||
<taxonomicName authorityName="Dumeril & Bibron" authorityYear="1841" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Eleutherodactylus</emphasis>
|
||
</taxonomicName>
|
||
, subgenus
|
||
<taxonomicName class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus (Syrrhophus)" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="subGenus" subGenus="Syrrhophus">Eleutherodactylus Syrrhophus</taxonomicName>
|
||
, as defined by
|
||
<bibRefCitation author="Hedges, SB" journalOrPublisher="Zootaxa" pageId="0" pageNumber="95" pagination="1 - 182" refId="B16" refString="Hedges, SB, Duellman, WE, Heinicke, MP, 2008. New World direct-developing frogs (Anura: Terrarana); molecular phylogeny, classification, biogeography, and conservation. Zootaxa 1737: 1 - 182" title="New World direct-developing frogs (Anura: Terrarana); molecular phylogeny, classification, biogeography, and conservation." volume="1737" year="2008">Hedges et al. (2008)</bibRefCitation>
|
||
. In the
|
||
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1870" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus (Syrrhophus) nitidus" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="species" species="nitidus" subGenus="Syrrhophus">Eleutherodactylus (Syrrhophus) nitidus</taxonomicName>
|
||
species series and the
|
||
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1870" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus (Syrrhophus) nitidus" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="species" species="nitidus" subGenus="Syrrhophus">Eleutherodactylus (Syrrhophus) nitidus</taxonomicName>
|
||
species group as defined by
|
||
<bibRefCitation author="Gruenwald, CI" journalOrPublisher="Mesoamerican Herpetology" pageId="0" pageNumber="95" pagination="6 - 83" refId="B14" refString="Gruenwald, CI, Reyes-Velasco, J, Franz-Chavez, H, Morales-Flores, KI, Ahumada-Carrillo, IT, Jones, JM, Boissinot, S, 2018. Six new species of Eleutherodactylus (Anura: systematic relationships and the validity of related species. Mesoamerican Herpetology 5 (1): 6 - 83" title="Six new species of Eleutherodactylus (Anura: systematic relationships and the validity of related species." volume="5" year="2018">
|
||
<normalizedToken originalValue="Grünwald">Gruenwald</normalizedToken>
|
||
et al. (2018)
|
||
</bibRefCitation>
|
||
, based on the condition of the tympanic annuli, ventral epidermis and visceral peritoneum. A small frog, but relatively large
|
||
<taxonomicName class="Amphibia" family="Eleutherodactylidae" genus="Syrrhophus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Syrrhophus" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Syrrhophus</emphasis>
|
||
</taxonomicName>
|
||
, adult males measure 25.6-29.5 mm SVL; vocal slits are present in males, readily visible under partially translucent ventral epidermis; digital tips are widely expanded, 1.8-2.6 times the width of the narrowest part of the finger on the third and fourth fingers; fingers moderately long, finger lengths are I-II-IV-III with third finger length ranging from 15-18% of SVL; compact lumbo-inguinal gland above the inguinal region present, raised, indistinct, but visible in live specimen; ventral epidermis is partially translucent and visceral peritoneum is clear, not white, thus abdominal vein is not clearly visible against a white background on the venter of live specimens and viscera is partially visible through translucent dark grey ventral epidermis; limbs moderate, TL/SVL ratio is 0.40-0.50, FeL/SVL ratio is 0.31-0.38 and TotFL/ SVL ratio is 0.50-0.60; snout relatively short, END/ SVL ratio is 0.08-0.11; tympanum small, indistinct and round, tympanic annuli not visible in live specimen; TW/ED ratio is 0.40-0.60. The dorsal and lateral skin is slightly shagreened, whereas the ventral skin is smooth. Dorsal colouration cream or tan, with darker brown reticulations on dorsal and lateral portions of head and back; loreal and region dark brown, from snout to behind tympanum, with some pale spots and speckling present on labial region same colour as pale dorsal colouration; pale interorbital bar variable, may be present or absent, when present same colour as pale dorsal colouration of head and body; distinct irregular-edged dark brown transverse bands present on legs; upper arms same colouration and pattern as forearms; no pale mid-dorsal stripe; upper flanks same colour as dorsum, lower flanks whitish with some grey marbling; venter pale lavender with some sparse white spotting and pale grey on throat. No inguinal flash colours present on thighs and groin. The mating call of adult males is a short low-pitched pipe (see below; Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Spectrograms (top) and oscillograms (bottom) of the advertisement calls of Eleutherodactylus franzi sp. nov., recorded at El Cucharillo, Municipio de Ixcateopan de Cuauhtemoc, Guerrero, Mexico (A, B) and at 3 km E of Ixcateopan de Cuauhtemoc, Guerrero, Mexico (C, D)." figureDoi="10.3897/herpetozoa.36.e104707.figure4" httpUri="https://binary.pensoft.net/fig/841845" pageId="0" pageNumber="95">4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/herpetozoa.36.e104707.figure4" httpUri="https://binary.pensoft.net/fig/841845" pageId="0" pageNumber="95" start="Figure 4" startId="F4">
|
||
<paragraph pageId="0" pageNumber="95">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Figure 4.</emphasis>
|
||
Spectrograms (top) and oscillograms (bottom) of the advertisement calls of
|
||
<taxonomicName class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus franzi" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Eleutherodactylus franzi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., recorded at El Cucharillo, Municipio de Ixcateopan de
|
||
<normalizedToken originalValue="Cuauhtémoc">Cuauhtemoc</normalizedToken>
|
||
, Guerrero, Mexico (
|
||
<emphasis bold="true" pageId="0" pageNumber="95">A, B</emphasis>
|
||
) and at 3 km E of Ixcateopan de
|
||
<normalizedToken originalValue="Cuauhtémoc">Cuauhtemoc</normalizedToken>
|
||
, Guerrero, Mexico (
|
||
<emphasis bold="true" pageId="0" pageNumber="95">C, D</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="95" type="comparisons">
|
||
<paragraph pageId="0" pageNumber="95">Comparisons.</paragraph>
|
||
<paragraph pageId="0" pageNumber="95">
|
||
<taxonomicName class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus franzi" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Eleutherodactylus franzi</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from all species in the
|
||
<taxonomicName baseAuthorityName="Baird" baseAuthorityYear="1859" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus (Syrrhophus) longipes" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="species" species="longipes" subGenus="Syrrhophus">Eleutherodactylus (Syrrhophus) longipes</taxonomicName>
|
||
species series by: possessing a small, indistinct tympanum with no tympanic annulus visible and with a diameter less than 50% of the diameter of the eye; by possessing a ventral epidermis which is semi-translucent and combined with a visceral peritoneum which is not white, an abdominal vein on the venter is not clearly evident against a white background in life; by possessing indistinct, but visible raised lumbo-inguinal gland above the inguinal region.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="95">
|
||
<taxonomicName class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus franzi" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Eleutherodactylus franzi</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from most species of the
|
||
<taxonomicName baseAuthorityName="Taylor" baseAuthorityYear="1942" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus (Syrrhophus) modestus" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="species" species="modestus" subGenus="Syrrhophus">Eleutherodactylus (Syrrhophus) modestus</taxonomicName>
|
||
species group by the combination of possessing a compact, protruding lumbo-inguinal gland above the inguinal region, digital tips which are expanded more than 1.8 times the width of the narrowest part the finger on the third and fourth fingers and the lack of a distinct interorbital bar a colour distinct from the dorsal ground colouration. It can further be distinguished from the superficially similar
|
||
<taxonomicName lsidName="E. grunwaldi" pageId="0" pageNumber="95" rank="species" species="grunwaldi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. grunwaldi</emphasis>
|
||
</taxonomicName>
|
||
by its smaller body size, 25.6-29.5 mm (vs. 28.4-32.4 mm), less expanded fingertips, 1.8-2.5 times the width of the narrowest part of the finger on fingers three and four (vs. 2.8-3.2) and the presence of a visible raised lumbo-inguinal gland. From the superficially very similar
|
||
<taxonomicName lsidName="E. saxatilis" pageId="0" pageNumber="95" rank="species" species="saxatilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. saxatilis</emphasis>
|
||
</taxonomicName>
|
||
, it can be distinguished by snout shape, eye size and head colouration. These two frogs, although not closely related, are very similar in appearance, but come from two widely-separated mountain ranges in central Mexico and are genetically distinct.
|
||
<taxonomicName class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus franzi" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Eleutherodactylus franzi</emphasis>
|
||
</taxonomicName>
|
||
has an angular canthus rostralis, with a shorter snout that is distinctly truncated from a lateral profile. It has a larger eye, with a larger ETD and, generally, there is a pale interorbital region that lacks dark markings. In
|
||
<taxonomicName lsidName="E. saxatilis" pageId="0" pageNumber="95" rank="species" species="saxatilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. saxatilis</emphasis>
|
||
</taxonomicName>
|
||
, the canthus rostralis is noticeably rounded and the snout is acuminate from a lateral profile. The eyes are smaller and located closer to the tympanum and there is no noticeable lack of dark dorsal markings in the interorbital area.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="95">
|
||
Within its own species group,
|
||
<taxonomicName lsidName="E. franzi" pageId="0" pageNumber="95" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. franzi</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from most species by possessing a compact inguinal gland that is indistinct, but visible in live specimens. This character may or may not be visible in preserved specimens depending on how they were preserved. This species differs from
|
||
<taxonomicName lsidName="E. pipilans" pageId="0" pageNumber="95" rank="species" species="pipilans">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. pipilans</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="E. erythrochomus" pageId="0" pageNumber="95" rank="species" species="erythrochomus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. erythrochomus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="E. nebulosus" pageId="0" pageNumber="95" rank="species" species="nebulosus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. nebulosus</emphasis>
|
||
</taxonomicName>
|
||
, which lack visible compact lumbo-inguinal glands altogether. All other known species in the
|
||
<taxonomicName lsidName="E. (Syrrhophus) nitidus" pageId="0" pageNumber="95" rank="species" species="nitidus" subGenus="Syrrhophus">E. (Syrrhophus) nitidus</taxonomicName>
|
||
species group have readily visible compact lumbo-inguinal glands above the inguinal region, except
|
||
<taxonomicName lsidName="E. maculabialis" pageId="0" pageNumber="95" rank="species" species="maculabialis">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. maculabialis</emphasis>
|
||
</taxonomicName>
|
||
, which has similarly visible, but indistinct lumbo-inguinal glands.
|
||
<taxonomicName class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus franzi" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Eleutherodactylus franzi</emphasis>
|
||
</taxonomicName>
|
||
can be further distinguished from
|
||
<taxonomicName lsidName="E. pipilans" pageId="0" pageNumber="95" rank="species" species="pipilans">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. pipilans</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="E. nebulosus" pageId="0" pageNumber="95" rank="species" species="nebulosus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. nebulosus</emphasis>
|
||
</taxonomicName>
|
||
by possessing digital tips which are expanded more than 1.8 times the width of the narrowest part of the finger and from
|
||
<taxonomicName lsidName="E. erythrochomus" pageId="0" pageNumber="95" rank="species" species="erythrochomus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. erythrochomus</emphasis>
|
||
</taxonomicName>
|
||
by possessing digital tips which are more than 1.5 times, but less than 3.0 times, the width of the narrowest part of the finger. It is distinguished from
|
||
<taxonomicName lsidName="E. albolabris" pageId="0" pageNumber="95" rank="species" species="albolabris">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. albolabris</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="E. nitidus" pageId="0" pageNumber="95" rank="species" species="nitidus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. nitidus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="E. petersi" pageId="0" pageNumber="95" rank="species" species="petersi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. petersi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="E. jamesdixoni" pageId="0" pageNumber="95" rank="species" species="jamesdixoni">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. jamesdixoni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="E. orarius" pageId="0" pageNumber="95" rank="species" species="orarius">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. orarius</emphasis>
|
||
</taxonomicName>
|
||
by the combination of larger size, smoother skin, longer limbs and tips of digits which are expanded more than 1.8 times the narrowest part of the finger on the third and fourth fingers. This species is distinguished from
|
||
<taxonomicName lsidName="E. dilatus" pageId="0" pageNumber="95" rank="species" species="dilatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. dilatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="E. humboldti" pageId="0" pageNumber="95" rank="species" species="humboldti">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. humboldti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="E. maurus" pageId="0" pageNumber="95" rank="species" species="maurus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. maurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="E. sentinelus" pageId="0" pageNumber="95" rank="species" species="sentinelus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. sentinelus</emphasis>
|
||
</taxonomicName>
|
||
by its larger size, smoother skin, pale dorsal colouration with dark reticulations and lack of a pale interorbital bar which is paler than the pale dorsal colouration. Furthermore, all these species, except
|
||
<taxonomicName lsidName="E. humboldti" pageId="0" pageNumber="95" rank="species" species="humboldti">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. humboldti</emphasis>
|
||
</taxonomicName>
|
||
, present multi-note whistles, while
|
||
<taxonomicName lsidName="E. franzi" pageId="0" pageNumber="95" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. franzi</emphasis>
|
||
</taxonomicName>
|
||
has a call that consists of a single note, low-pitched pipe.
|
||
<taxonomicName class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus franzi" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Eleutherodactylus franzi</emphasis>
|
||
</taxonomicName>
|
||
may be distinguished from
|
||
<taxonomicName lsidName="E. maculabialis" pageId="0" pageNumber="95" rank="species" species="maculabialis">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. maculabialis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="E. syristes" pageId="0" pageNumber="95" rank="species" species="syristes">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. syristes</emphasis>
|
||
</taxonomicName>
|
||
by its larger size, more expanded fingertips, lack of inguinal flash colouration, as well as those
|
||
<normalizedToken originalValue="species’">species'</normalizedToken>
|
||
unique advertisement call which consists of a trill rather than a short pipe. In Guerrero, two other species of saxicolous
|
||
<taxonomicName authorityName="Dumeril & Bibron" authorityYear="1841" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Eleutherodactylus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName lsidName="E. pipilans" pageId="0" pageNumber="95" rank="species" species="pipilans">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. pipilans</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="E. erythrochomus" pageId="0" pageNumber="95" rank="species" species="erythrochomus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. erythrochomus</emphasis>
|
||
</taxonomicName>
|
||
) have similar colouration, similar smooth skin and widely expanded digital pads. Both are readily distinguishable from
|
||
<taxonomicName lsidName="E. franzi" pageId="0" pageNumber="95" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. franzi</emphasis>
|
||
</taxonomicName>
|
||
by possessing an indistinct lumbo-inguinal gland which is barely visible in life and by a male advertisement call which is a peep instead of a pipe. Furthermore,
|
||
<taxonomicName lsidName="E. franzi" pageId="0" pageNumber="95" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. franzi</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName lsidName="E. pipilans" pageId="0" pageNumber="95" rank="species" species="pipilans">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. pipilans</emphasis>
|
||
</taxonomicName>
|
||
by its more expanded digital tips on the third and fourth fingers, 1.8-2.5 times the width of the narrowest part of the finger (vs. 1.3-1.9) and distinct lack of dark markings in the interorbital region (vs. no difference from rest of dorsum). It can further be distinguished from
|
||
<taxonomicName lsidName="E. erythrochomus" pageId="0" pageNumber="95" rank="species" species="erythrochomus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. erythrochomus</emphasis>
|
||
</taxonomicName>
|
||
by the conspicuous dark pattern on a pale dorsal colouration, lesser expanded digital tips no more than 2.5 times the width of the narrowest part of the digit on the third and fourth fingers (vs. 2.3-3.8) and a less distinct tympanum. General characteristics for the
|
||
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1870" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus nitidus" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="species" species="nitidus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Eleutherodactylus nitidus</emphasis>
|
||
</taxonomicName>
|
||
species group are given in Table
|
||
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Key Comparative Characters of the Eleutherodactylus nitidus species group." httpUri="http://table.plazi.org/id/D1565CB02F53EA0F9DE95ECCD9FAC6F5" pageId="0" pageNumber="95" tableUuid="D1565CB02F53EA0F9DE95ECCD9FAC6F5">1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption ID-Table-UUID="D1565CB02F53EA0F9DE95ECCD9FAC6F5" httpUri="http://table.plazi.org/id/D1565CB02F53EA0F9DE95ECCD9FAC6F5" pageId="0" pageNumber="95" start="Table 1" startId="T1">
|
||
<paragraph pageId="0" pageNumber="95">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Table 1.</emphasis>
|
||
Key Comparative Characters of the
|
||
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1870" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus nitidus" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="species" species="nitidus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Eleutherodactylus nitidus</emphasis>
|
||
</taxonomicName>
|
||
species group.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="95">
|
||
<table pageId="0" pageNumber="95">
|
||
<tr pageId="0" pageNumber="95">
|
||
<th colspan="1" isEmpty="true" pageId="0" pageNumber="95" rowspan="1">-</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<taxonomicName lsidName="E. albolabris" pageId="0" pageNumber="95" rank="species" species="albolabris">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. albolabris</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<taxonomicName lsidName="E. dilatus" pageId="0" pageNumber="95" rank="species" species="dilatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. dilatus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<taxonomicName lsidName="E. erythrochomus" pageId="0" pageNumber="95" rank="species" species="erythrochomus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. erythrochomus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<taxonomicName lsidName="E. franzi" pageId="0" pageNumber="95" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. franzi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<taxonomicName lsidName="E. humboldti" pageId="0" pageNumber="95" rank="species" species="humboldti">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. humboldti</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<taxonomicName lsidName="E. jamesdixoni" pageId="0" pageNumber="95" rank="species" species="jamesdixoni">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. jamesdixoni</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<taxonomicName lsidName="E. maculabialis" pageId="0" pageNumber="95" rank="species" species="maculabialis">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. maculabialis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<taxonomicName lsidName="E. maurus" pageId="0" pageNumber="95" rank="species" species="maurus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. maurus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<taxonomicName lsidName="E. nebulosus" pageId="0" pageNumber="95" rank="species" species="nebulosus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. nebulosus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<taxonomicName lsidName="E. nitidus" pageId="0" pageNumber="95" rank="species" species="nitidus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. nitidus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<taxonomicName lsidName="E. orarius" pageId="0" pageNumber="95" rank="species" species="orarius">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. orarius</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<taxonomicName lsidName="E. petersi" pageId="0" pageNumber="95" rank="species" species="petersi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. petersi</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<taxonomicName lsidName="E. pipilans" pageId="0" pageNumber="95" rank="species" species="pipilans">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. pipilans</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<taxonomicName lsidName="E. rubrimaculatus" pageId="0" pageNumber="95" rank="species" species="rubrimaculatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. rubrimaculatus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<taxonomicName lsidName="E. sentinelus" pageId="0" pageNumber="95" rank="species" species="sentinelus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. sentinelus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<taxonomicName lsidName="E. syristes" pageId="0" pageNumber="95" rank="species" species="syristes">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. syristes</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Size</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Medium</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Medium</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Large</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Large</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Medium</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Medium</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Small</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Small</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Medium</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Medium</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Medium</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Medium</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Large</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Small</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Medium</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Small</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">SVL adult males (range) in mm</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">23.0-26.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">23.8-25.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">24.9-30.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">25.6-29.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">23.4-25.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">24.3-26.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">17.9-24.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">20.7-24.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">22.9-28.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">24.3-26.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">24.6-28.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">23.9-26.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">25.5-29.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">18.2-23.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">23.3-25.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">21.7-24.6</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Condition of Interorbital Bar</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Indistinct, Pale</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Present, Pale</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Usually Present, Pale</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Indistinct, Pale</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Present, Pale</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Present, Pale</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Indistinct, Pale</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Indistinct, Pale</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Indistinct, Pale</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Present, Pale</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Present, Pale</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Pale Mid-dorsal Blotch</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Sometime Present</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Present</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Present</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Present</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Sometimes Present</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Present</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Present</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Absent</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Colouration of Lip</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">White</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Dark with pale flecking</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">As head</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Grey, with flecking of dorsal colouration</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Pale, with white flecking,</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Mottled</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Dark with pale spots</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Dark with pale flecking</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Dark with pale spots</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Mottled</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">White, mottled with dark</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Mottled</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">As head</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Dark with white or pale spots</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Dark with pale flecking</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Dark, variedly spotted</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Inguinal Flash Colouration</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Orange</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Yellow</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Yellow or orange</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Faint, yellow-orange</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Varied, Yellow / Orange</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Varied, usually Absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Faint, yellow-orange</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Faint, yellow-orange</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Faint, yellow-orange</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Yellow</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Varied, Yellow / Orange</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Ventral Colouration</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">White with black spots</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Grey with white and black</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Transparent</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">White or pinkish</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Grey, with white and black</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">White with dark mottling</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Grey with white and black</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Grey with white and black</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Transparent</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">White with dark mottling</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">White with dark mottling</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">White with dark mottling</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Transparent</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Pale grey with white spotting</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Transparent with white and black</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Transparent and White, with black spots</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Dorsal Skin Texture</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Smooth</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Not Smooth</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Smooth</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Smooth</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Not Smooth</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Not Smooth</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Smooth</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Not Smooth</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Smooth</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Not Smooth</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Not Smooth</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Not Smooth</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Smooth</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Smooth</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Smooth</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Smooth</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Ventral Skin Texture</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Slightly Rugose</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Rugose</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Smooth</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Smooth</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Rugose</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Slightly Rugose</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Smooth</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Rugose</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Smooth</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Slightly Rugose</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Slightly Rugose</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Slightly Rugose</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Smooth</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Smooth</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Smooth</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Smooth</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Condition of Lumbo-Inguinal Gland</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Very distinct</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Very distinct</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Indistinct</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Distinct</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Very distinct</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Very distinct</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Distinct</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Very distinct</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Indistinct</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Very distinct</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Very distinct</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Very distinct</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Indistinct</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Indistinct</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Very distinct</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Distinct</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">3FPW/3FW</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.3-1.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.5-1.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.3-3.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.8-2.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.9-2.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.1-1.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.4-2.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.3-1.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.1-1.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.1-1.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.2-1.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.1-1.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.5-1.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.5-1.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.7-2.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.1-1.9</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">4FPW/4FW</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.3-1.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.5-1.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.3-3.5-</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.8-2.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.3-2.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.1-1.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.4-2.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.3-1.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.1-1.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.1-1.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.2-1.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.1-1.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.5-2.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.5-1.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.5-2.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.2-1.8</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">TW/ED</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.27-0.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.25-0.35</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.33-0.51</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.40-0.61</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.37-0.52</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.25-0.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.25-0.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.28-0.31</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.35-0.38</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.25-0.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.25-0.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.25-0.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.30-0.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.25-0.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.26-0.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.25-0.29</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Call</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Whistle</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Peep</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Peep</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Pipe</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Pipe</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Whistle</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Trill</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Pipe</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Peep</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Whistle</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Whistle</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Whistle</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Peep</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Peep</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Peep</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Trill</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="95" type="description">
|
||
<paragraph pageId="0" pageNumber="95">Description of the holotype.</paragraph>
|
||
<paragraph pageId="0" pageNumber="95">
|
||
Adult male, relatively large (26.2 mm SVL); head as wide (9.5 mm) as long (9.5 mm), head wider than body; snout rounded from a dorsal view and rounded to slightly truncate from a lateral profile; tympanum indistinct, rounded with no supra-tympanic fold present; tympanum small, oval, greatest width of tympanum 1.4 mm; greatest diameter of eye 2.8 mm; tympanum width to eye-diameter 0.51; eyelid width 1.6 mm, approximately 38% of the IOD; first finger shorter than second finger; finger lengths from shortest to longest I-II-IV-III; digital pads on fingers two, three and four expanded, 2.1 times the narrowest point of the digit on fingers three and four; expanded finger pads widely expanded, truncate, three palmar tubercles; inner palmar tubercle 70% of middle palmar tubercle and outer palmar tubercles about 60% as large as middle palmar tubercle, (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A. Ventral aspect of the hand of the holotype of Eleutherodactylus franzi sp. nov., INIRENA 2900 (CIG 01725) from El Cucharillo, Municipio de Ixcateopan de Cuauhtemoc, Guerrero, Mexico; B. Ventral aspect of the hand of a Eleutherodactylus humboldti, INIRENA 2911 (CIG 01703) from 9.4 km N of Valle de Bravo junction on Valle de Bravo - Toluca toll road, near San Bartolo, Municipio de Amanalco, Estado de Mexico, Mexico." figureDoi="10.3897/herpetozoa.36.e104707.figure5" httpUri="https://binary.pensoft.net/fig/841846" pageId="0" pageNumber="95">5A</figureCitation>
|
||
); toe lengths from shortest to longest I-V-II-III-IV, TL1 and TL5 very similar; outer metatarsal conical with a round base moderate, approximately 56% of inner metatarsal tubercle; inner metatarsal tubercle spherical shape with oval base, large, approximately 1.1 mm in length. Dorsal skin smooth, lateral skin slightly shagreened with some low tubercles; ventral skin smooth. Vocal slits present.
|
||
</paragraph>
|
||
<caption doi="10.3897/herpetozoa.36.e104707.figure5" httpUri="https://binary.pensoft.net/fig/841846" pageId="0" pageNumber="95" start="Figure 5" startId="F5">
|
||
<paragraph pageId="0" pageNumber="95">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Figure 5.</emphasis>
|
||
<emphasis bold="true" pageId="0" pageNumber="95">A.</emphasis>
|
||
Ventral aspect of the hand of the holotype of
|
||
<taxonomicName class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus franzi" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Eleutherodactylus franzi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., INIRENA 2900 (CIG 01725) from El Cucharillo, Municipio de Ixcateopan de
|
||
<normalizedToken originalValue="Cuauhtémoc">Cuauhtemoc</normalizedToken>
|
||
, Guerrero, Mexico;
|
||
<emphasis bold="true" pageId="0" pageNumber="95">B.</emphasis>
|
||
Ventral aspect of the hand of a
|
||
<taxonomicName class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus humboldti" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="species" species="humboldti">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Eleutherodactylus humboldti</emphasis>
|
||
</taxonomicName>
|
||
, INIRENA 2911 (CIG 01703) from 9.4 km N of Valle de Bravo junction on Valle de Bravo - Toluca toll road, near San Bartolo, Municipio de Amanalco, Estado de
|
||
<normalizedToken originalValue="México">Mexico</normalizedToken>
|
||
, Mexico.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="95">In life, the holotype had a yellowish-tan dorsal colouration on the back, with darker brown blotches on the back and flanks. Head yellowish-tan with some dark brown speckling, no pale interorbital bar; however, the interorbital region mostly lacks darker brown speckling giving it the resemblance of a pale interorbital bar. Labial region pale grey with some tan and some white speckling. Three and four white-tipped tubercles present at the rictal. A dark brown stripe present from the tip of the snout posteriorly through loreal region, eye and tympanum to right above rictal tubercles. Forearms, thighs, femur and tarsus tan with indistinct pale brown banding. The upper arms were unmarked, tan to slightly orange. No inguinal flash colouration was present on groin or thighs. Ventral colouration was lavender with some sparse white and on sides, ventral colouration on throat grey. Ventral skin was slightly translucent and visceral peritoneum clear, so no visible red abdominal vein and viscera were visible in life.</paragraph>
|
||
<paragraph pageId="0" pageNumber="95">
|
||
Colouration in preservative is pale tan on dorsum, with darker brown reticulations. Pale tan interorbital bar present. Dark canthal bar is dark brown. Unmarked upper arms are cream to white. Limbs cream with dark brown cross-banding. The dorsal surfaces of the legs are light brown and the groin and posterior surfaces of the thighs are brown. Ventral surfaces yellowish-cream, unmarked, slightly darker brownish pigmentation on throat and chin. Ventral surfaces of hands and feet brown, with dark brown spots. (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype of Eleutherodactylus franzi, sp. nov., INIRENA 2900 (CIG 01725), El Cucharillo, Municipio de Ixcateopan de Cuauhtemoc, Guerrero, Mexico in life (A-C) and in preservative (D, E)." figureDoi="10.3897/herpetozoa.36.e104707.figure2" httpUri="https://binary.pensoft.net/fig/841843" pageId="0" pageNumber="95">2D, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="95" type="description">
|
||
<paragraph pageId="0" pageNumber="95">Measurements of the holotype (in millimetres).</paragraph>
|
||
<paragraph pageId="0" pageNumber="95">IND 2.2, IOD 4.2, END 2.5, ETD 0.9, UpL 6.4, FoL 7.5, PaL 2.3, HaL 6.7, F1L 2.3, F1PW 0.5, F1W 0.4, F2L 2.8, F2PW 0.8, F2W 0.4, F3L 4.7, F3PW 1.1, F3W 0.5, F4L 3.5, F4PW 1.1, F4W 0.5, IPTL 0.7, MPTL 1.0, OPTL 0.6, FeL 10.0, TL 11.0, TaL 7.1, TotFL 11.4, T1L 2.4, T1PW 0.62, T1W 0.5, T2L 3.6, T2PW 0.7, T2W 0.5, T3L 4.4, T3PW 0.7, T3W 0.5, T4L 6.0, T4PW 0.7, T4W 0.5, T5L 2.5, T5PW 0.6, T5W 0.4, IMTL 1.1, OMTL 0.6, FeL/SVL 38%, TL/SVL 42%, HaL/SVL 26%, TotFL/SVL 56%, HL/SVL 35%, HW/SVL 36%.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="95" type="variation">
|
||
<paragraph pageId="0" pageNumber="95">Variation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="95">
|
||
SVL from 25.6-29.5 mm (27.15
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
1.77). Expanded finger pads vary from 1.8-2.5 times the narrowest part of the digit on the third finger and from 1.8-2.6 times the narrowest part of the digit on the fourth finger, with average 2.0
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.21 on the third finger and average 2.2
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.21 on the fourth finger. Dorsal ground colouration cream or tan, but varied with some greenish, reddish or yellowish tinge, always with darker brown blotches or reticulations. The extent of the darker brown blotches or reticulation varied greatly. The condition of the interorbital area ranged from unmarked and same colour as ground colouration to heavily marked by dark speckling same colour as dark dorsal blotches or reticulation. Venter always lavender, but with varying amounts of white spots. Morphological variation of
|
||
<taxonomicName lsidName="E. franzi" pageId="0" pageNumber="95" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. franzi</emphasis>
|
||
</taxonomicName>
|
||
is presented in Table
|
||
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Eleutherodactylus franzi sp. nov. Morphological Measurements (in millimetres)." httpUri="http://table.plazi.org/id/9937AC210025DF1B590D37F47B1EFA22" pageId="0" pageNumber="95" tableUuid="9937AC210025DF1B590D37F47B1EFA22">2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption ID-Table-UUID="9937AC210025DF1B590D37F47B1EFA22" httpUri="http://table.plazi.org/id/9937AC210025DF1B590D37F47B1EFA22" pageId="0" pageNumber="95" start="Table 2" startId="T2">
|
||
<paragraph pageId="0" pageNumber="95">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Table 2.</emphasis>
|
||
<taxonomicName class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus franzi" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Eleutherodactylus franzi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Morphological Measurements (in millimetres).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="95">
|
||
<table pageId="0" pageNumber="95">
|
||
<tr pageId="0" pageNumber="95">
|
||
<th colspan="1" isEmpty="true" pageId="0" pageNumber="95" rowspan="1">-</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">INIRENA 2901</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">INIRENA 2902</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">INIRENA 2903</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">INIRENA 2904</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">INIRENA 2905</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">INIRENA 2906</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">INIRENA 2907</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">INIRENA 2908</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">INIRENA 2909</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">INIRENA 2900</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">INIRENA 2910</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">INIRENA 2895</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">INIRENA 2896</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">INIRENA 2897</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">INIRENA 2898</th>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">SVL</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">27.72</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">25.58</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">27.72</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">26.86</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">26.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">26.04</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">28.62</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">26.68</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">25.55</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">26.24</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">27.97</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">28.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">27.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">29.51</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">26.56</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">HL</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">8.76</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.35</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">8.61</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">8.33</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">8.76</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">8.73</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">8.68</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.38</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">8.79</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.04</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">8.61</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">HW</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.78</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.03</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.81</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.78</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.82</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.07</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.41</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.99</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.68</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.67</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.66</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.61</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">TW</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.53</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.52</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.88</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.74</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.38</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.39</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.39</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.34</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.42</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">ED</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">3.74</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.62</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.79</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">3.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.63</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.74</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.78</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.81</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.76</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.77</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.78</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.38</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.94</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">ElW</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.39</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.52</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.68</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.19</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.49</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.67</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.69</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.59</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.35</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.78</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.51</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.62</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.9</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">IOD</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">5.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">5.02</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">5.19</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">5.09</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">5.04</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">4.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">5.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">5.09</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">5.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">4.24</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">4.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">5.09</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">5.14</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">5.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">5.37</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">IND</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.25</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.31</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.16</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.38</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.17</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.53</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.37</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.21</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.3</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">END</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">3.15</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.61</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.51</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.53</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.77</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.78</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.96</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.79</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.61</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.69</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.33</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.55</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.37</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">ETD</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.96</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.92</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.98</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.86</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.02</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.82</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.89</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.89</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.76</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.84</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.76</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.95</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.98</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">UpL</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.19</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.31</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.02</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.62</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.31</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.43</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.39</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.31</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.39</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.88</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.58</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Fol</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.86</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.39</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.78</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.49</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.58</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.56</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.51</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.82</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.51</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.58</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.92</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.62</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">HaL</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.57</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.82</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.41</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.23</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.72</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.46</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.86</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.93</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.58</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.24</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.53</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">F3PW/F3W</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.78</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.94</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.33</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.23</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.77</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.97</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.09</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.75</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.43</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.87</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.53</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">F4PW/F4W</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.77</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.34</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.15</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.38</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.84</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.94</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.17</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.46</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.97</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">2.64</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">FeL</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">10</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.11</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.24</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.19</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.87</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.57</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.95</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.81</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.82</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.34</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.22</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">9.9</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">TL</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">12.78</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">11.79</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">11.88</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">11.82</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">11.46</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">11.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">11.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">12.41</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">12.69</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">11.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">11.81</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">11.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">11.21</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">12.39</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">11.34</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">TaL</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.41</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.51</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.31</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.93</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">6.84</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.92</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.14</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.59</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">7.83</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">FL</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">4.49</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">4.33</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">4.97</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">4.31</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">3.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">4.22</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">4.49</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">4.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">4.21</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">4.58</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">4.82</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">4.35</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">4.25</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">4.58</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">4.75</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">3TL</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">3.72</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">3.74</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">3.64</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">3.59</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">3.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">3.16</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">3.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">3.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">3.17</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">3.53</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">3.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">3.34</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">3.48</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">3.38</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">3.71</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">TotFL</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">11.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">11.37</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">11.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">11.41</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">11.15</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">10.82</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">12.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">11.16</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">11.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">11.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">11.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">11.31</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">11.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">11.96</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">12.46</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">IPT</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.68</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.89</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.86</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.88</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.86</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.64</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.92</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.91</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.86</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.68</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.62</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.94</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.78</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">MPT</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.09</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.19</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.16</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.17</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.14</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.02</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.26</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">OPT</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.52</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.56</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.58</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.54</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.62</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.58</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.76</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.66</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.68</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.58</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.52</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.71</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.76</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.64</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">IMTL</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.02</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.14</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.01</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.26</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.16</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.15</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.16</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.1</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">OMTL</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.72</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.77</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.78</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.74</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.92</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.75</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.68</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.74</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.62</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.78</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.65</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.91</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.81</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.98</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">TW/ED</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.41</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.52</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.54</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.61</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.55</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.53</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.51</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.58</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.48</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.48</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">F3PW</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.14</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.26</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.26</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.25</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.62</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">F3W</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.64</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.68</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.66</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.54</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.61</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.67</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.72</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.62</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.54</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.63</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.56</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.66</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.64</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">F4PW</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.33</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.31</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.14</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.41</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.26</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.34</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.23</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.38</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">1.48</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">F4W</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.62</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.58</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.62</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.55</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.62</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.62</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.72</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.68</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.65</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.53</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.52</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.55</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">0.56</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="95" type="advertisement call">
|
||
<paragraph pageId="0" pageNumber="95">Advertisement call.</paragraph>
|
||
<paragraph pageId="0" pageNumber="95">
|
||
The advertisement call of the males of this species consists of a single, short, low-pitched pipe that lasts about 106 ms and has a dominant frequency of 2612.7
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
40.6 Hz (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Spectrograms (top) and oscillograms (bottom) of the advertisement calls of Eleutherodactylus franzi sp. nov., recorded at El Cucharillo, Municipio de Ixcateopan de Cuauhtemoc, Guerrero, Mexico (A, B) and at 3 km E of Ixcateopan de Cuauhtemoc, Guerrero, Mexico (C, D)." figureDoi="10.3897/herpetozoa.36.e104707.figure4" httpUri="https://binary.pensoft.net/fig/841845" pageId="0" pageNumber="95">4A-D</figureCitation>
|
||
). The note has limited amplitude modulation and the highest energy is displayed at the end of the note. The call is similar to that of
|
||
<taxonomicName lsidName="E. pipilans" pageId="0" pageNumber="95" rank="species" species="pipilans">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. pipilans</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="E. erythrochomus" pageId="0" pageNumber="95" rank="species" species="erythrochomus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. erythrochomus</emphasis>
|
||
</taxonomicName>
|
||
. The call differs from the nearby
|
||
<taxonomicName lsidName="E. maurus" pageId="0" pageNumber="95" rank="species" species="maurus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. maurus</emphasis>
|
||
</taxonomicName>
|
||
and the sympatric
|
||
<taxonomicName lsidName="E. humboldti" pageId="0" pageNumber="95" rank="species" species="humboldti">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. humboldti</emphasis>
|
||
</taxonomicName>
|
||
, which also have a pipe, by being shorter and lower pitched. We present call data for both
|
||
<taxonomicName lsidName="E. franzi" pageId="0" pageNumber="95" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. franzi</emphasis>
|
||
</taxonomicName>
|
||
and the sympatric
|
||
<taxonomicName lsidName="E. humboldti" pageId="0" pageNumber="95" rank="species" species="humboldti">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. humboldti</emphasis>
|
||
</taxonomicName>
|
||
in Table
|
||
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Advertisement call data of Eleutherodactylus franzi sp. nov. and the sympatric Eleutherodactylus humboldti." httpUri="http://table.plazi.org/id/C54456A9B27E391FCDF0519119FFB2C0" pageId="0" pageNumber="95" tableUuid="C54456A9B27E391FCDF0519119FFB2C0">3</tableCitation>
|
||
. The advertisement call also differs from the sympatric or near sympatric
|
||
<taxonomicName lsidName="E. petersi" pageId="0" pageNumber="95" rank="species" species="petersi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. petersi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="E. nitidus" pageId="0" pageNumber="95" rank="species" species="nitidus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. nitidus</emphasis>
|
||
</taxonomicName>
|
||
as these two species have a call that consists of a multi-note whistle rather than a single-note pipe.
|
||
</paragraph>
|
||
<caption ID-Table-UUID="C54456A9B27E391FCDF0519119FFB2C0" httpUri="http://table.plazi.org/id/C54456A9B27E391FCDF0519119FFB2C0" pageId="0" pageNumber="95" start="Table 3" startId="T3">
|
||
<paragraph pageId="0" pageNumber="95">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Table 3.</emphasis>
|
||
Advertisement call data of
|
||
<taxonomicName class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus franzi" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Eleutherodactylus franzi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and the sympatric
|
||
<taxonomicName class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus humboldti" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="species" species="humboldti">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Eleutherodactylus humboldti</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="95">
|
||
<table pageId="0" pageNumber="95">
|
||
<tr pageId="0" pageNumber="95">
|
||
<th colspan="1" isEmpty="true" pageId="0" pageNumber="95" rowspan="1">-</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<taxonomicName class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus franzi" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Eleutherodactylus franzi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<taxonomicName class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus humboldti" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="species" species="humboldti">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Eleutherodactylus humboldti</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Individuals</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">4</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">3</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Call type</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Pipe</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">Pipe</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Dominant frequency (kHz)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
2.61
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.04
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
2.89
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.23
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Call length (ms)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
106.0
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
5.2
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
256.8
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
12.8
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Call rate (/m)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
6.88
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
2.16
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
1.73
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Call rise time (ms)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
63.64
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
4.06
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
90.82
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
45.37
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Pulse Rate</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
2.05
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.62
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="95">
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Call Interval</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
13.03
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
3.35
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="95" rowspan="1">
|
||
39.75
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
14.77
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="95" type="distribution">
|
||
<paragraph pageId="0" pageNumber="95">Distribution and ecology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="95">
|
||
<taxonomicName class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus franzi" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Eleutherodactylus franzi</emphasis>
|
||
</taxonomicName>
|
||
appears to be endemic to the Sierra de Taxco Region of northern Guerrero (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Map showing the type localities and distribution of Eleutherodactylus which are either closely related or superficially similar to the species described herein from southern Mexico. White diamond represents the type locality of E. franzi sp. nov. and white circle represents additional locality." figureDoi="10.3897/herpetozoa.36.e104707.figure6" httpUri="https://binary.pensoft.net/fig/841847" pageId="0" pageNumber="95">6</figureCitation>
|
||
). This is an extension of the Mexican Transverse Volcanic Belt which extends south into the Balsas Basin. This frog occurs at high elevations between 2,000-2,400 m a.s.l. and has been collected in tropical deciduous forest, oak woodland and pine-oak forest, as well as mixtures of these assemblages. It is restricted to areas of karstic rock outcroppings and the associated sinkholes and caves systems. This species may also occur in nearby Estado de
|
||
<normalizedToken originalValue="México">Mexico</normalizedToken>
|
||
, as habitat is continuous and the State line is near the type locality. Old reports of
|
||
<taxonomicName lsidName="E. pipilans" pageId="0" pageNumber="95" rank="species" species="pipilans">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. pipilans</emphasis>
|
||
</taxonomicName>
|
||
from that State may be referable to this species; however, we have made attempts to examine these specimens at MZFC and were not able to find the specimens (C.
|
||
<normalizedToken originalValue="Grünwald">Gruenwald</normalizedToken>
|
||
, pers. obs.)
|
||
</paragraph>
|
||
<caption doi="10.3897/herpetozoa.36.e104707.figure6" httpUri="https://binary.pensoft.net/fig/841847" pageId="0" pageNumber="95" start="Figure 6" startId="F6">
|
||
<paragraph pageId="0" pageNumber="95">
|
||
<emphasis bold="true" pageId="0" pageNumber="95">Figure 6.</emphasis>
|
||
Map showing the type localities and distribution of
|
||
<taxonomicName authorityName="Dumeril & Bibron" authorityYear="1841" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Eleutherodactylus</emphasis>
|
||
</taxonomicName>
|
||
which are either closely related or superficially similar to the species described herein from southern Mexico. White diamond represents the type locality of
|
||
<taxonomicName lsidName="E. franzi" pageId="0" pageNumber="95" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">E. franzi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and white circle represents additional locality.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="95" type="etymology">
|
||
<paragraph pageId="0" pageNumber="95">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="95">
|
||
This species is named after
|
||
<normalizedToken originalValue="Héctor">Hector</normalizedToken>
|
||
<normalizedToken originalValue="Franz-Chávez">Franz-Chavez</normalizedToken>
|
||
, Mexican herpetologist and avid field collector who collected the type material and who helped collect an extensive sampling of the
|
||
<taxonomicName authorityName="Dumeril & Bibron" authorityYear="1841" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eleutherodactylus" order="Anura" pageId="0" pageNumber="95" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="95">Eleutherodactylus</emphasis>
|
||
</taxonomicName>
|
||
specimens to be used in our succession of studies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |