405 lines
47 KiB
XML
405 lines
47 KiB
XML
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<mods:title id="D86A99B070179766E0AE584E082BB8F1">The systematics of the pseudoscorpion genus Indohya (Pseudoscorpiones: Hyidae) in Australia</mods:title>
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<mods:namePart id="094E45448621ECCFC5FA9D57F3F70ED5">Harvey, Mark S.</mods:namePart>
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<mods:affiliation id="F10AF62E7729BE73BB7E27F2A0BF34EE">Collections & Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 & Adjunct, School of Biological Sciences, University of Western Australia, Crawley, Western Australia 6009, Australia</mods:affiliation>
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<mods:nameIdentifier id="6792609D3CFC0496CF7BD7ABF3443351" type="email">mark.harvey@museum.wa.gov.au</mods:nameIdentifier>
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<mods:namePart id="209B44518FE1703CD802364647342B6D">Burger, Mieke A. A.</mods:namePart>
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<mods:affiliation id="68A7BB05BFEAF5B670BF23E083A5073F">Collections & Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238</mods:affiliation>
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<mods:namePart id="04374049CDB0CBE0D915EE45806E70D9">Abrams, Kym M.</mods:namePart>
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<mods:affiliation id="5A9F195AA9233131450818FAE7533950">Collections & Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 & Present address: Department of Water and Environmental Regulation, Locked Bag 10, Joondalup DC, Western Australia 6919, Australia https: // orcid. org / 0000 - 0003 - 4543 - 9255</mods:affiliation>
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<mods:nameIdentifier id="F1C2A1A3280A49BC03A34D0D37D6DE4B" type="email">mark.harvey@museum.wa.gov.au</mods:nameIdentifier>
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<mods:affiliation id="987D4F7C2FE66FEFE662796994424B9D">Adjunct, School of Biological Sciences, University of Western Australia, Crawley, Western Australia 6009, Australia & Helix Molecular Solutions Pty Ltd, PO Box 155, Leederville, Western Australia 6153, Australia https: // orcid. org / 0000 - 0002 - 1750 - 4577</mods:affiliation>
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<mods:affiliation id="44225C768558DD1616F6E7633E84085B">Collections & Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 & Adjunct, School of Biological Sciences, University of Western Australia, Crawley, Western Australia 6009, Australia & Collections & Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 & Present address: Biologic Environmental Survey, East Perth, Western Australia 6004, Australia https: // orcid. org / 0000 - 0001 - 7108 - 0552</mods:affiliation>
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<mods:nameIdentifier id="06F94022CFA51D1F3E0C212910A7CD40" type="email">mark.harvey@museum.wa.gov.au</mods:nameIdentifier>
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<mods:namePart id="0594CD6FB0EB87D25A996F409F3E0160">Perina, Giulia</mods:namePart>
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<mods:affiliation id="E2474C7E690668913FA5808210D96496">Collections & Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 & Collections & Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 & Present address: Biologic Environmental Survey, East Perth, Western Australia 6004, Australia https: // orcid. org / 0000 - 0001 - 7108 - 0552 & Collections & Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 & Subterranean Research and Groundwater Ecology (SuRGE) Group, Trace and Environmental DNA (TrEnD) Laboratory, School of Molecular and Life Sciences, Curtin University, Perth, Western Australia 6102, Australia https: // orcid. org / 0000 - 0002 - 0349 - 3803 Corresponding author</mods:affiliation>
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<mods:nameIdentifier id="CBE0BFD272BAEEB045C9DA9516925F98" type="email">mark.harvey@museum.wa.gov.au</mods:nameIdentifier>
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<mods:title id="68A10818847ED82748F037815D73149B">Zootaxa</mods:title>
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<mods:date id="EB06BF01138A51C7B86B56F2A0F0D0EE">2023</mods:date>
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<mods:number id="B7566213A05227EF87EE8365EAB22338">2023-09-05</mods:number>
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<taxonomicName id="4EA24D21FFD00B02AFB0FD73FDEEF751" authority="Harvey & Burger" authorityName="Harvey & Burger" authorityYear="2023" box="[151,564,762,789]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="aquila" status="sp. nov.">
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<emphasis id="BBD6EAB0FFD00B02AFB0FD73FE8BF751" bold="true" box="[151,337,762,789]" italics="true" pageId="42" pageNumber="43">Indohya aquila</emphasis>
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Harvey & Burger
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,
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(
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,
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<figureCitation id="11992A27FFD00B02AFDCFC97FEC3F77D" box="[251,281,798,825]" captionStart="FIGURE 15" captionStartId="42.[151,250,1979,2003]" captionTargetBox="[151,1436,1389,1956]" captionTargetId="figure-469@42.[151,1436,1389,1956]" captionTargetPageId="42" captionText="FIGURE 15. Indohya aquila Harvey & Burger, n. sp., holotype male (WAM T130511):A, dorsal; B, ventral; C, cephalothorax, dorsal. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8323919" httpUri="https://zenodo.org/record/8323919/files/figure.png" pageId="42" pageNumber="43">15</figureCitation>
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,
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)
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<emphasis id="BBD6EAB0FFD00B02AFB0FC03FE5EF7E0" bold="true" box="[151,388,906,932]" pageId="42" pageNumber="43">Material examined.</emphasis>
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<materialsCitation id="39CA3CFFFFD00B02AEA8FC02FED6F7A9" collectingDate="2011-04-15" collectionCode="WAM" collectorName="D. C. Main & J. W. Quartermaine" country="Australia" latitude="-22.17886" location="Tom Price" longLatPrecision="1" longitude="117.47328" municipality="Eagle" pageId="42" pageNumber="43" specimenCode="T130511" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
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male
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.
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:
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:
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<collectingMunicipality id="6979ACD8FFD00B02ACFFFC02FBC1F7E1" box="[984,1051,907,933]" pageId="42" pageNumber="43">Eagle</collectingMunicipality>
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, ca.
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NNW. of
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,
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,
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,
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<collectingDate id="ED58E98AFFD00B02AD26FC27FD69F78D" box="[513,691,942,969]" pageId="42" pageNumber="43" value="2011-04-15">15 April 2011</collectingDate>
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, subterranean scrape,
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,
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<collectorName id="24575374FFD00B02AB46FC26FAE7F78D" box="[1121,1341,943,969]" pageId="42" pageNumber="43">J.W. Quartermaine</collectorName>
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(
|
||
<collectionCode id="EFB3AE67FFD00B02AA71FC26FA47F78D" box="[1366,1437,943,969]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="42" pageNumber="43" type="Museum">WAM</collectionCode>
|
||
<specimenCode id="D9049ED9FFD00B02AFB0FC5BFEDAF7A9" box="[151,256,978,1005]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="42" pageNumber="43">T130511</specimenCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C1B86529FFD00B02AFE0FC7FFB8CF039" pageId="42" pageNumber="43" type="diagnosis">
|
||
<paragraph id="891D36A2FFD00B02AFE0FC7FFB8CF039" blockId="42.[151,1437,906,1365]" pageId="42" pageNumber="43">
|
||
<emphasis id="BBD6EAB0FFD00B02AFE0FC7FFE98F054" bold="true" box="[199,322,1014,1040]" pageId="42" pageNumber="43">Diagnosis.</emphasis>
|
||
<taxonomicName id="4EA24D21FFD00B02AE6DFC7EFE2DF055" authorityName="Harvey & Burger" authorityYear="2023" box="[330,503,1015,1041]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="aquila">
|
||
<emphasis id="BBD6EAB0FFD00B02AE6DFC7EFE2DF055" box="[330,503,1015,1041]" italics="true" pageId="42" pageNumber="43">Indohya aquila</emphasis>
|
||
</taxonomicName>
|
||
belongs to a group of
|
||
<taxonomicName id="4EA24D21FFD00B02ADD9FC7EFC6EF055" box="[766,948,1015,1041]" pageId="42" pageNumber="43">
|
||
<emphasis id="BBD6EAB0FFD00B02ADD9FC7EFC80F055" box="[766,858,1015,1041]" italics="true" pageId="42" pageNumber="43">Indohya</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
that have 12 setae on the carapace and no eyes. It differs from all others except
|
||
<taxonomicName id="4EA24D21FFD00B02AD10FB92FD0EF070" authorityName="Harvey & Burger" authorityYear="2023" box="[567,724,1051,1076]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="anastomosa">
|
||
<emphasis id="BBD6EAB0FFD00B02AD10FB92FD0EF070" box="[567,724,1051,1076]" italics="true" pageId="42" pageNumber="43">I. anastomosa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4EA24D21FFD00B02AC2EFB92FC78F071" authorityName="WAMT" baseAuthorityName="Harvey" baseAuthorityYear="1993" box="[777,930,1051,1077]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="humphreysi">
|
||
<emphasis id="BBD6EAB0FFD00B02AC2EFB92FC78F071" box="[777,930,1051,1077]" italics="true" pageId="42" pageNumber="43">I. humphreysi</emphasis>
|
||
</taxonomicName>
|
||
by the presence of only 2 setae on tergite I. It differs from both of these species by its smaller size, e.g. pedipalpal chela (with pedicel)
|
||
<quantity id="4E5A9B47FFD00B02AB52FBB6FB3CF01D" box="[1141,1254,1087,1113]" metricMagnitude="-3" metricUnit="m" metricValue="1.12" pageId="42" pageNumber="43" unit="mm" value="1.12">1.120 mm</quantity>
|
||
(Ô) in
|
||
<taxonomicName id="4EA24D21FFD00B02AA10FBB6FA4CF01D" authorityName="Harvey & Burger" authorityYear="2023" box="[1335,1430,1087,1113]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="aquila">
|
||
<emphasis id="BBD6EAB0FFD00B02AA10FBB6FA4CF01D" box="[1335,1430,1087,1113]" italics="true" pageId="42" pageNumber="43">I. aquila</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="4E5A9B47FFD00B02AFB0FBEBFEE0F039" box="[151,314,1122,1149]" metricMagnitude="-3" metricUnit="m" metricValue="1.585" metricValueMax="1.65" metricValueMin="1.52" pageId="42" pageNumber="43" unit="mm" value="1.585" valueMax="1.65" valueMin="1.52">1.52–1.65 mm</quantity>
|
||
(Ô),
|
||
<quantity id="4E5A9B47FFD00B02AE51FBEAFE00F039" box="[374,474,1123,1149]" metricMagnitude="-3" metricUnit="m" metricValue="2.09" pageId="42" pageNumber="43" unit="mm" value="2.09">2.09 mm</quantity>
|
||
(
|
||
<specimenCount id="9FA4FD2BFFD00B02AECBFBEAFE24F039" box="[492,510,1123,1149]" pageId="42" pageNumber="43" type="female">♀</specimenCount>
|
||
) in
|
||
<taxonomicName id="4EA24D21FFD00B02AD0BFBEAFD1CF039" authorityName="WAMT" baseAuthorityName="Harvey" baseAuthorityYear="1993" box="[556,710,1123,1149]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="humphreysi">
|
||
<emphasis id="BBD6EAB0FFD00B02AD0BFBEAFD1CF039" box="[556,710,1123,1149]" italics="true" pageId="42" pageNumber="43">I. humphreysi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<quantity id="4E5A9B47FFD00B02ADDBFBEAFCBAF039" box="[764,864,1123,1149]" metricMagnitude="-3" metricUnit="m" metricValue="2.18" pageId="42" pageNumber="43" unit="mm" value="2.18">2.18 mm</quantity>
|
||
(
|
||
<specimenCount id="9FA4FD2BFFD00B02AC56FBEAFC59F039" box="[881,899,1123,1149]" pageId="42" pageNumber="43" type="female">♀</specimenCount>
|
||
) in
|
||
<taxonomicName id="4EA24D21FFD00B02AC95FBEAFB95F038" authorityName="Harvey & Burger" authorityYear="2023" box="[946,1103,1123,1148]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="anastomosa">
|
||
<emphasis id="BBD6EAB0FFD00B02AC95FBEAFB95F038" box="[946,1103,1123,1148]" italics="true" pageId="42" pageNumber="43">I. anastomosa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C1B86529FFD00B04AFE0FB0FFE89F0AD" lastPageId="44" lastPageNumber="45" pageId="42" pageNumber="43" type="description">
|
||
<paragraph id="891D36A2FFD00B02AFE0FB0FFB34F0E5" blockId="42.[151,1437,906,1365]" box="[199,1262,1158,1185]" pageId="42" pageNumber="43">
|
||
<emphasis id="BBD6EAB0FFD00B02AFE0FB0FFE6AF0E4" bold="true" box="[199,432,1158,1184]" pageId="42" pageNumber="43">Description (adult).</emphasis>
|
||
<emphasis id="BBD6EAB0FFD00B02AE9FFB0EFDA1F0E5" box="[440,635,1159,1185]" italics="true" pageId="42" pageNumber="43">
|
||
Colour (
|
||
<figureCitation id="11992A27FFD00B02AD30FB0EFDB1F0E5" box="[535,619,1159,1185]" captionStart="FIGURE 15" captionStartId="42.[151,250,1979,2003]" captionTargetBox="[151,1436,1389,1956]" captionTargetId="figure-469@42.[151,1436,1389,1956]" captionTargetPageId="42" captionText="FIGURE 15. Indohya aquila Harvey & Burger, n. sp., holotype male (WAM T130511):A, dorsal; B, ventral; C, cephalothorax, dorsal. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8323919" httpUri="https://zenodo.org/record/8323919/files/figure.png" pageId="42" pageNumber="43">Fig. 15</figureCitation>
|
||
):
|
||
</emphasis>
|
||
light yellow-brown, pedipalps and carapace red-brown.
|
||
</paragraph>
|
||
<paragraph id="891D36A2FFD00B02AFE0FB22FAE0F081" blockId="42.[151,1437,906,1365]" box="[199,1338,1195,1221]" pageId="42" pageNumber="43">
|
||
<emphasis id="BBD6EAB0FFD00B02AFE0FB22FE4AF081" box="[199,400,1195,1221]" italics="true" pageId="42" pageNumber="43">Setae and cuticle:</emphasis>
|
||
setae long, mostly straight and acicular; most cuticular surfaces smooth and glossy.
|
||
</paragraph>
|
||
<paragraph id="891D36A2FFD00B02AFE0FB46FB75F111" blockId="42.[151,1437,906,1365]" pageId="42" pageNumber="43">
|
||
<emphasis id="BBD6EAB0FFD00B02AFE0FB46FEE4F0AD" box="[199,318,1231,1257]" italics="true" pageId="42" pageNumber="43">Chelicera:</emphasis>
|
||
surface smooth; hand with 5 (Ô) setae, movable finger with 1 sub-medial seta, all setae acuminate; galea present, long, slender and slightly curved; fixed finger with 10 (Ô) small teeth, each approximately same size; movable finger with 6, distal teeth larger than basal teeth; with 2 dorsal and 1 ventral lyrifissures; lamina exterior absent; rallum with 6 blades, all blades with anterior spinules, basal blade shorter than others.
|
||
</paragraph>
|
||
<caption id="DDDD662AFFD00B02AFB0F832FE64F3B3" ID-DOI="http://doi.org/10.5281/zenodo.8323919" ID-Zenodo-Dep="8323919" httpUri="https://zenodo.org/record/8323919/files/figure.png" pageId="42" pageNumber="43" startId="42.[151,250,1979,2003]" targetBox="[151,1436,1389,1956]" targetPageId="42" targetType="figure">
|
||
<paragraph id="891D36A2FFD00B02AFB0F832FE64F3B3" blockId="42.[151,1437,1979,2040]" pageId="42" pageNumber="43">
|
||
<emphasis id="BBD6EAB0FFD00B02AFB0F832FEC5F397" bold="true" box="[151,287,1979,2003]" pageId="42" pageNumber="43">FIGURE 15.</emphasis>
|
||
<taxonomicName id="4EA24D21FFD00B02AE02F832FDAFF390" authority="Harvey & Burger" authorityName="Harvey & Burger" authorityYear="2023" box="[293,629,1979,2004]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="aquila" status="sp. nov.">
|
||
<emphasis id="BBD6EAB0FFD00B02AE02F832FE64F397" box="[293,446,1979,2003]" italics="true" pageId="42" pageNumber="43">Indohya aquila</emphasis>
|
||
Harvey & Burger
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="BBD6EAB0FFD00B02AD59F835FD6CF390" bold="true" box="[638,694,1980,2004]" pageId="42" pageNumber="43">
|
||
<taxonomicNameLabel id="A0E557CBFFD00B02AD59F835FD6CF390" box="[638,694,1980,2004]" pageId="42" pageNumber="43" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (WAM T130511): A, dorsal; B, ventral; C, cephalothorax, dorsal. Scale lines = 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DDDD662AFFD10B03AFB0F95FFCA0F373" ID-DOI="http://doi.org/10.5281/zenodo.8323921" ID-Zenodo-Dep="8323921" httpUri="https://zenodo.org/record/8323921/files/figure.png" pageId="43" pageNumber="44" startId="43.[151,250,1750,1774]" targetBox="[163,1425,196,1716]" targetPageId="43" targetType="figure">
|
||
<paragraph id="891D36A2FFD10B03AFB0F95FFCA0F373" blockId="43.[151,1437,1750,1847]" pageId="43" pageNumber="44">
|
||
<emphasis id="BBD6EAB0FFD10B03AFB0F95FFEFBF2AA" bold="true" box="[151,289,1750,1774]" pageId="43" pageNumber="44">FIGURE 16.</emphasis>
|
||
<taxonomicName id="4EA24D21FFD10B03AE0FF95FFDA4F2AB" authority="Harvey & Burger" authorityName="Harvey & Burger" authorityYear="2023" box="[296,638,1750,1775]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="aquila" status="sp. nov.">
|
||
<emphasis id="BBD6EAB0FFD10B03AE0FF95FFE19F2AA" box="[296,451,1750,1774]" italics="true" pageId="43" pageNumber="44">Indohya aquila</emphasis>
|
||
Harvey & Burger
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="BBD6EAB0FFD10B03ADAEF95EFD19F2AB" bold="true" box="[649,707,1751,1775]" pageId="43" pageNumber="44">
|
||
<taxonomicNameLabel id="A0E557CBFFD10B03ADAEF95EFD19F2AB" box="[649,707,1751,1775]" pageId="43" pageNumber="44" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (WAM T130511): A, carapace, dorsal; B, left pedipalp, dorsal; C, left chela, retrolateral; D, left movable finger, chelal teeth, retrolateral; E, left fixed finger, chelal teeth, retrolateral. Scale lines = 0.5 mm (Figs. A, C), 0.2 mm (Fig. A), 0.1 mm (Figs. D, E).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="891D36A2FFD10B04AFE0F800FB02F5B1" blockId="43.[151,1437,1928,2027]" lastBlockId="44.[151,1437,151,1653]" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">
|
||
<emphasis id="BBD6EAB0FFD10B03AFE0F800FE63F3E6" box="[199,441,1928,1955]" italics="true" pageId="43" pageNumber="44">
|
||
Pedipalp (
|
||
<figureCitation id="11992A27FFD10B03AE67F800FE71F3E6" box="[320,427,1928,1955]" captionStart="FIGURE 16" captionStartId="43.[151,250,1750,1774]" captionTargetBox="[163,1425,196,1716]" captionTargetId="figure-17@43.[151,1436,181,1726]" captionTargetPageId="43" captionText="FIGURE 16. Indohya aquila Harvey & Burger, n. sp., holotype male (WAM T130511): A, carapace, dorsal; B, left pedipalp, dorsal; C, left chela, retrolateral; D, left movable finger, chelal teeth, retrolateral; E, left fixed finger, chelal teeth, retrolateral. Scale lines = 0.5 mm (Figs. A, C), 0.2 mm (Fig. A), 0.1 mm (Figs. D, E)." figureDoi="http://doi.org/10.5281/zenodo.8323921" httpUri="https://zenodo.org/record/8323921/files/figure.png" pageId="43" pageNumber="44">Fig. 16B</figureCitation>
|
||
):
|
||
</emphasis>
|
||
long and slender; trochanter smooth, 2.04 (Ô) ×; femur slightly broadened medially, smooth, without dorsal tactile seta, with 3 stout retrolateral setae near base, 6.52 (Ô) ×; patella slightly broadened distally, smooth, pedicel not strongly pronounced but basal portion slimmer than distal portion, with several small lyrifissures situated basally on dorsal surface, 4.10 (Ô) ×; chela (
|
||
<figureCitation id="11992A27FFD60B04AC47FF1EFC1CF4F5" box="[864,966,151,177]" captionStart="FIGURE 16" captionStartId="43.[151,250,1750,1774]" captionTargetBox="[163,1425,196,1716]" captionTargetId="figure-17@43.[151,1436,181,1726]" captionTargetPageId="43" captionText="FIGURE 16. Indohya aquila Harvey & Burger, n. sp., holotype male (WAM T130511): A, carapace, dorsal; B, left pedipalp, dorsal; C, left chela, retrolateral; D, left movable finger, chelal teeth, retrolateral; E, left fixed finger, chelal teeth, retrolateral. Scale lines = 0.5 mm (Figs. A, C), 0.2 mm (Fig. A), 0.1 mm (Figs. D, E)." figureDoi="http://doi.org/10.5281/zenodo.8323921" httpUri="https://zenodo.org/record/8323921/files/figure.png" pageId="44" pageNumber="45">Fig. 16C</figureCitation>
|
||
): hand ovoid, smooth, chela (with pedicel) 4.39 (Ô) ×, chela (without pedicel) 4.14 (Ô) ×, hand (without pedicel) 1.67 (Ô) × longer than broad, movable finger 1.52 (Ô) × longer than hand (without pedicel). Fixed finger with 8 trichobothria,
|
||
<emphasis id="BBD6EAB0FFD60B04AB0AFF56FB9DF4BD" box="[1069,1095,223,249]" italics="true" pageId="44" pageNumber="45">eb</emphasis>
|
||
,
|
||
<emphasis id="BBD6EAB0FFD60B04AB71FF56FBA1F4BD" box="[1110,1147,223,249]" italics="true" pageId="44" pageNumber="45">esb</emphasis>
|
||
and
|
||
<emphasis id="BBD6EAB0FFD60B04AB94FF56FB0EF4BD" box="[1203,1236,223,249]" italics="true" pageId="44" pageNumber="45">isb</emphasis>
|
||
in straight row at base of finger,
|
||
<emphasis id="BBD6EAB0FFD60B04AE1BFE8AFE88F559" box="[316,338,259,285]" italics="true" pageId="44" pageNumber="45">ib</emphasis>
|
||
situated dorsally at the end of the chelal hand,
|
||
<emphasis id="BBD6EAB0FFD60B04AC7DFE8AFCA1F559" box="[858,891,259,285]" italics="true" pageId="44" pageNumber="45">isb</emphasis>
|
||
at the basal end of the fixed finger,
|
||
<emphasis id="BBD6EAB0FFD60B04AA20FE8AFAC7F559" box="[1287,1309,259,285]" italics="true" pageId="44" pageNumber="45">ib</emphasis>
|
||
situated on dorsum of chelal hand,
|
||
<emphasis id="BBD6EAB0FFD60B04AEBBFEAEFE76F504" box="[412,428,295,320]" italics="true" pageId="44" pageNumber="45">it</emphasis>
|
||
closer to
|
||
<emphasis id="BBD6EAB0FFD60B04AD3FFEAEFDF6F504" box="[536,556,295,320]" italics="true" pageId="44" pageNumber="45">et</emphasis>
|
||
than to
|
||
<emphasis id="BBD6EAB0FFD60B04ADA2FEAEFD7EF504" box="[645,676,295,320]" italics="true" pageId="44" pageNumber="45">est</emphasis>
|
||
,
|
||
<emphasis id="BBD6EAB0FFD60B04AD95FEAEFD1CF504" box="[690,710,295,320]" italics="true" pageId="44" pageNumber="45">et</emphasis>
|
||
slightly distal to
|
||
<emphasis id="BBD6EAB0FFD60B04ACA1FEAEFC4CF504" box="[902,918,295,320]" italics="true" pageId="44" pageNumber="45">it</emphasis>
|
||
; movable finger with 4 trichobothria,
|
||
<emphasis id="BBD6EAB0FFD60B04AA18FEAEFA82F505" box="[1343,1368,295,321]" italics="true" pageId="44" pageNumber="45">sb</emphasis>
|
||
much closer to
|
||
<emphasis id="BBD6EAB0FFD60B04AFD8FEC2FED7F521" box="[255,269,331,357]" italics="true" pageId="44" pageNumber="45">b</emphasis>
|
||
than to
|
||
<emphasis id="BBD6EAB0FFD60B04AE4CFEC2FEA4F520" box="[363,382,331,356]" italics="true" pageId="44" pageNumber="45">st</emphasis>
|
||
,
|
||
<emphasis id="BBD6EAB0FFD60B04AEABFEC2FE45F520" box="[396,415,331,356]" italics="true" pageId="44" pageNumber="45">st</emphasis>
|
||
situated much closer to
|
||
<emphasis id="BBD6EAB0FFD60B04AD94FEC2FD61F520" box="[691,699,331,356]" italics="true" pageId="44" pageNumber="45">t</emphasis>
|
||
than to
|
||
<emphasis id="BBD6EAB0FFD60B04AC3FFEC2FCEBF521" box="[792,817,331,357]" italics="true" pageId="44" pageNumber="45">sb</emphasis>
|
||
, ratio
|
||
<emphasis id="BBD6EAB0FFD60B04AC5CFEC2FC6FF520" box="[891,949,331,357]" italics="true" pageId="44" pageNumber="45">sb–st</emphasis>
|
||
/
|
||
<emphasis id="BBD6EAB0FFD60B04AC9AFEC2FC2BF521" box="[957,1009,331,357]" italics="true" pageId="44" pageNumber="45">sb–b</emphasis>
|
||
= 2.11 (Ô),
|
||
<emphasis id="BBD6EAB0FFD60B04AB58FEC2FB5DF520" box="[1151,1159,331,356]" italics="true" pageId="44" pageNumber="45">t</emphasis>
|
||
acuminate. Both fingers straight in lateral view. Fixed finger smooth, movable finger with several prolateral and retrolateral granulations basally. Chelal teeth (
|
||
<figureCitation id="11992A27FFD60B04AEAAFE1AFDC5F5E9" box="[397,543,403,429]" captionStart="FIGURE 16" captionStartId="43.[151,250,1750,1774]" captionTargetBox="[163,1425,196,1716]" captionTargetId="figure-17@43.[151,1436,181,1726]" captionTargetPageId="43" captionText="FIGURE 16. Indohya aquila Harvey & Burger, n. sp., holotype male (WAM T130511): A, carapace, dorsal; B, left pedipalp, dorsal; C, left chela, retrolateral; D, left movable finger, chelal teeth, retrolateral; E, left fixed finger, chelal teeth, retrolateral. Scale lines = 0.5 mm (Figs. A, C), 0.2 mm (Fig. A), 0.1 mm (Figs. D, E)." figureDoi="http://doi.org/10.5281/zenodo.8323921" httpUri="https://zenodo.org/record/8323921/files/figure.png" pageId="44" pageNumber="45">Figs. 16D, E</figureCitation>
|
||
) juxtadentate, fixed finger with 62 (Ô) teeth, teeth 1–6, 37, 41–43, 45–48, 50, 52–62 pointed, remainder rounded or truncate, movable finger with ca. 25 (Ô), low teeth. Venom apparatus present only in movable chelal finger, venom duct long, terminating in nodus ramosus between
|
||
<emphasis id="BBD6EAB0FFD60B04AB48FE52FB52F5B1" box="[1135,1160,475,501]" italics="true" pageId="44" pageNumber="45">sb</emphasis>
|
||
and
|
||
<emphasis id="BBD6EAB0FFD60B04AB98FE52FB08F5B0" box="[1215,1234,475,500]" italics="true" pageId="44" pageNumber="45">st</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="891D36A2FFD60B04AFE0FE76FAA8F6C1" blockId="44.[151,1437,151,1653]" pageId="44" pageNumber="45">
|
||
<emphasis id="BBD6EAB0FFD60B04AFE0FE76FEA2F65D" box="[199,376,511,537]" italics="true" pageId="44" pageNumber="45">Cephalothorax:</emphasis>
|
||
carapace (
|
||
<figureCitation id="11992A27FFD60B04AEC9FE76FD88F65D" box="[494,594,511,537]" captionStart="FIGURE 16" captionStartId="43.[151,250,1750,1774]" captionTargetBox="[163,1425,196,1716]" captionTargetId="figure-17@43.[151,1436,181,1726]" captionTargetPageId="43" captionText="FIGURE 16. Indohya aquila Harvey & Burger, n. sp., holotype male (WAM T130511): A, carapace, dorsal; B, left pedipalp, dorsal; C, left chela, retrolateral; D, left movable finger, chelal teeth, retrolateral; E, left fixed finger, chelal teeth, retrolateral. Scale lines = 0.5 mm (Figs. A, C), 0.2 mm (Fig. A), 0.1 mm (Figs. D, E)." figureDoi="http://doi.org/10.5281/zenodo.8323921" httpUri="https://zenodo.org/record/8323921/files/figure.png" pageId="44" pageNumber="45">Fig. 16A</figureCitation>
|
||
) sub-rectangular; smooth; anterior margin slightly convex; epistome absent; lateral margins slightly convex; posterior margin straight; 1.52 (Ô) × longer than broad; with 12 setae arranged 4: 0: 4: 2: 2 (Ô); without furrows; eyes absent. Manducatory process distally triangular, with 2 distal setae; pedipalpal coxa with 6 additional setae. Coxa I with anterior margin not modified; chaetotaxy of coxae I–IV: Ô, 4: 4: 4: 6.
|
||
</paragraph>
|
||
<paragraph id="891D36A2FFD60B04AFE0FD06FDD8F719" blockId="44.[151,1437,151,1653]" pageId="44" pageNumber="45">
|
||
<emphasis id="BBD6EAB0FFD60B04AFE0FD06FEDCF6EC" box="[199,262,655,680]" italics="true" pageId="44" pageNumber="45">Legs:</emphasis>
|
||
femora I and II much longer than patellae I and II, respectively; femora I and II with basal swelling; femora I and II each with 2 primary slit sensilla, 1 sub-basally directed longitudinally and 1 sub-distally directed transversely; junction between anterior femora and patellae perpendicular; junction between posterior femora and patellae slightly oblique; femora III and IV much smaller than patellae III and IV, respectively; femur + patella IV? (missing) (Ô); metatarsi shorter than tarsi; subterminal tarsal seta not known; claws smooth, arolium slightly shorter than claws; arolium not divided.
|
||
</paragraph>
|
||
<paragraph id="891D36A2FFD60B04AFE0FCEEFF2AF7A9" blockId="44.[151,1437,151,1653]" pageId="44" pageNumber="45">
|
||
<emphasis id="BBD6EAB0FFD60B04AFE0FCEEFEE0F7C5" box="[199,314,871,897]" italics="true" pageId="44" pageNumber="45">Abdomen:</emphasis>
|
||
tergites not divided; tergal chaetotaxy Ô, 2: 4: 4: 6: 6: 6: 6: 6: 6:?:?: 2; arranged in single rows; sternal chaetotaxy Ô, 18: (?) 22 (?): (?) 6 (?):?:?:?:?:?:?:?: 2; posterior tergites and sternites with several tactile setae; medial sternites without suture line; pleural membrane uniformly granulate; without setae; stigmatic helix present.
|
||
</paragraph>
|
||
<paragraph id="891D36A2FFD60B04AFE0FC7EFEE1F01D" blockId="44.[151,1437,151,1653]" pageId="44" pageNumber="45">
|
||
<emphasis id="BBD6EAB0FFD60B04AFE0FC7EFEE0F055" box="[199,314,1015,1041]" italics="true" pageId="44" pageNumber="45">Genitalia:</emphasis>
|
||
male: lateral genital sacs large and ovoid; median genital sac short and ovoid; ejaculatory duct atrium large and rounded; dorsal apodeme wing-like; without a pair of large dorsal genital sacs; with 2 pairs of setae within genital atrium.
|
||
</paragraph>
|
||
<paragraph id="891D36A2FFD60B04AFE0FBEAFE89F0AD" blockId="44.[151,1437,151,1653]" pageId="44" pageNumber="45">
|
||
<emphasis id="BBD6EAB0FFD60B04AFE0FBEAFE40F039" box="[199,410,1123,1149]" italics="true" pageId="44" pageNumber="45">Dimensions (mm):</emphasis>
|
||
<materialsCitation id="39CA3CFFFFD60B04AE83FBEAFE95F0AD" collectionCode="WAM" collectorName="Body & Pedipalp & Carapace & Leg IV" latitude="0.555" longLatPrecision="78" longitude="0.365" pageId="44" pageNumber="45" specimenCode="T130511" specimenCount="1" typeStatus="holotype">
|
||
male
|
||
<typeStatus id="56198800FFD60B04AEC2FBEAFD9DF039" box="[485,583,1123,1149]" pageId="44" pageNumber="45" type="holotype">holotype</typeStatus>
|
||
(
|
||
<collectionCode id="EFB3AE67FFD60B04AD7EFBEAFD7AF039" box="[601,672,1123,1149]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="44" pageNumber="45" type="Museum">WAM</collectionCode>
|
||
<specimenCode id="D9049ED9FFD60B04AD8EFBEBFCC8F039" box="[681,786,1122,1149]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="44" pageNumber="45">T130511</specimenCode>
|
||
).
|
||
<collectorName id="24575374FFD60B04AC0FFBEAFCBFF039" box="[808,869,1123,1149]" pageId="44" pageNumber="45">Body</collectorName>
|
||
length (excluding chelicerae) ca. 1.92.
|
||
<collectorName id="24575374FFD60B04AA15FBEAFA42F039" box="[1330,1432,1123,1149]" pageId="44" pageNumber="45">Pedipalp</collectorName>
|
||
: trochanter
|
||
<geoCoordinate id="EC965065FFD60B04AE35FB0FFE8BF0E5" box="[274,337,1158,1185]" degrees="0.265" direction="north" orientation="latitude" pageId="44" pageNumber="45" precision="55" value="0.265">0.265</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EC965065FFD60B04AE7DFB0FFE41F0E5" box="[346,411,1158,1185]" degrees="0.130" direction="east" orientation="longitude" pageId="44" pageNumber="45" precision="55" value="0.13">0.130</geoCoordinate>
|
||
, femur
|
||
<geoCoordinate id="EC965065FFD60B04AED2FB0FFDEFF0E5" box="[501,565,1158,1185]" degrees="0.815" direction="north" orientation="latitude" pageId="44" pageNumber="45" precision="55" value="0.815">0.815</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EC965065FFD60B04AD19FB0FFDA4F0E5" box="[574,638,1158,1185]" degrees="0.125" direction="east" orientation="longitude" pageId="44" pageNumber="45" precision="55" value="0.125">0.125</geoCoordinate>
|
||
, patella
|
||
<geoCoordinate id="EC965065FFD60B04ADF8FB0FFCC5F0E5" box="[735,799,1158,1185]" degrees="0.595" direction="north" orientation="latitude" pageId="44" pageNumber="45" precision="55" value="0.595">0.595</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EC965065FFD60B04AC0FFB0FFCB2F0E5" box="[808,872,1158,1185]" degrees="0.145" direction="east" orientation="longitude" pageId="44" pageNumber="45" precision="55" value="0.145">0.145</geoCoordinate>
|
||
, chela (with pedicel)
|
||
<geoCoordinate id="EC965065FFD60B04AB45FB0FFB7BF0E5" box="[1122,1185,1158,1185]" degrees="1.120" direction="north" orientation="latitude" pageId="44" pageNumber="45" precision="55" value="1.12">1.120</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EC965065FFD60B04AB8DFB0FFB31F0E5" box="[1194,1259,1158,1185]" degrees="0.255" direction="east" orientation="longitude" pageId="44" pageNumber="45" precision="55" value="0.255">0.255</geoCoordinate>
|
||
, chela (without pedicel) length 1.055, chelal hand (without pedicel) length 0.425, movable finger length 0.645.
|
||
<collectorName id="24575374FFD60B04AB84FB22FAD1F081" box="[1187,1291,1195,1221]" pageId="44" pageNumber="45">Carapace</collectorName>
|
||
<geoCoordinate id="EC965065FFD60B04AA28FB23FA95F081" box="[1295,1359,1194,1221]" degrees="0.555" direction="north" orientation="latitude" pageId="44" pageNumber="45" precision="55" value="0.555">0.555</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EC965065FFD60B04AA7EFB23FA43F081" box="[1369,1433,1194,1221]" degrees="0.365" direction="east" orientation="longitude" pageId="44" pageNumber="45" precision="55" value="0.365">0.365</geoCoordinate>
|
||
.
|
||
<collectorName id="24575374FFD60B04AFB0FB46FF30F0AD" box="[151,234,1231,1257]" pageId="44" pageNumber="45">Leg IV</collectorName>
|
||
: missing
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C1B86529FFD60B04AFE0FB7BFB1AF269" pageId="44" pageNumber="45" type="distribution">
|
||
<paragraph id="891D36A2FFD60B04AFE0FB7BFC34F111" blockId="44.[151,1437,151,1653]" pageId="44" pageNumber="45">
|
||
<emphasis id="BBD6EAB0FFD60B04AFE0FB7BFDD9F149" bold="true" box="[199,515,1266,1293]" pageId="44" pageNumber="45">Distribution and remarks.</emphasis>
|
||
The
|
||
<typeStatus id="56198800FFD60B04AD18FB7AFD7BF149" box="[575,673,1267,1293]" pageId="44" pageNumber="45" type="holotype">holotype</typeStatus>
|
||
is in poor condition and was most likely already dead when it was retrieved from the borehole, as evidenced by its macerated appearance (
|
||
<figureCitation id="11992A27FFD60B04ACE6FA9EFBCFF175" box="[961,1045,1302,1329]" captionStart="FIGURE 15" captionStartId="42.[151,250,1979,2003]" captionTargetBox="[151,1436,1389,1956]" captionTargetId="figure-469@42.[151,1436,1389,1956]" captionTargetPageId="42" captionText="FIGURE 15. Indohya aquila Harvey & Burger, n. sp., holotype male (WAM T130511):A, dorsal; B, ventral; C, cephalothorax, dorsal. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8323919" httpUri="https://zenodo.org/record/8323919/files/figure.png" pageId="44" pageNumber="45">Fig. 15</figureCitation>
|
||
). Indeed, the poor condition of the specimen precluded the successful amplification of DNA from a leg sample.
|
||
</paragraph>
|
||
<paragraph id="891D36A2FFD60B04AFE0FAD6FE70F185" blockId="44.[151,1437,151,1653]" pageId="44" pageNumber="45">
|
||
<taxonomicName id="4EA24D21FFD60B04AFE0FAD6FEAEF13D" authorityName="Harvey & Burger" authorityYear="2023" box="[199,372,1375,1401]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="aquila">
|
||
<emphasis id="BBD6EAB0FFD60B04AFE0FAD6FEAEF13D" box="[199,372,1375,1401]" italics="true" pageId="44" pageNumber="45">Indohya aquila</emphasis>
|
||
</taxonomicName>
|
||
is only known from subterranean habitats at the proposed Flinders Pilbara Iron Ore Project: Eagle deposit, which is situated northwest of Tom Price. The species was previously known by WAM identification code
|
||
<taxonomicName id="4EA24D21FFD60B04AFF4FA2EFEF5F185" authority="PSE" authorityName="PSE" box="[211,303,1447,1473]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BBD6EAB0FFD60B04AFF4FA2EFEF5F185" box="[211,303,1447,1473]" italics="true" pageId="44" pageNumber="45">Indohya</emphasis>
|
||
</taxonomicName>
|
||
‘PSE148’.
|
||
</paragraph>
|
||
<paragraph id="891D36A2FFD60B04AFE0FA43FB1AF269" blockId="44.[151,1437,151,1653]" pageId="44" pageNumber="45">
|
||
<emphasis id="BBD6EAB0FFD60B04AFE0FA43FE2BF1A1" bold="true" box="[199,497,1482,1509]" pageId="44" pageNumber="45">Conservation assessment</emphasis>
|
||
. The collecting site lies in a valley ca.
|
||
<quantity id="4E5A9B47FFD60B04ACBAFA43FC39F1A1" box="[925,995,1482,1509]" metricMagnitude="4" metricUnit="m" metricValue="5.4" pageId="44" pageNumber="45" unit="km" value="54.0">54 km</quantity>
|
||
NNW. of Tom Price, which is currently undeveloped. The presence of extensive similar surface habitat in the valley and adjacent areas suggests that
|
||
<taxonomicName id="4EA24D21FFD60B04AAABFA66FF05F269" authorityName="Harvey & Burger" authorityYear="2023" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="aquila">
|
||
<emphasis id="BBD6EAB0FFD60B04AAABFA66FF05F269" italics="true" pageId="44" pageNumber="45">I. aquila</emphasis>
|
||
</taxonomicName>
|
||
is likely to be located at other subterranean sites and therefore unlikely to be threatened.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C1B86529FFD60B04AFE0F9BEFE0BF231" pageId="44" pageNumber="45" type="etymology">
|
||
<paragraph id="891D36A2FFD60B04AFE0F9BEFE0BF231" blockId="44.[151,1437,151,1653]" pageId="44" pageNumber="45">
|
||
<emphasis id="BBD6EAB0FFD60B04AFE0F9BEFE94F215" bold="true" box="[199,334,1591,1617]" pageId="44" pageNumber="45">Etymology.</emphasis>
|
||
The specific epithet is a noun referring to the collection of this species at the Eagle deposit (
|
||
<taxonomicName id="4EA24D21FFD60B04AA69F9BEFF0FF231" authority=", Latin" authorityName="Latin" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="aquila">
|
||
<emphasis id="BBD6EAB0FFD60B04AA69F9BEFA4CF215" box="[1358,1430,1591,1617]" italics="true" pageId="44" pageNumber="45">aquila</emphasis>
|
||
, Latin
|
||
</taxonomicName>
|
||
, eagle) (
|
||
<bibRefCitation id="ED334B53FFD60B04AE14F9D2FE1EF231" author="Brown, R. W." box="[307,452,1626,1653]" pageId="44" pageNumber="45" refId="ref82031" refString="Brown, R. W. (1956) Composition of scientific words. Revised Edition. Smithsonian Institution Press, Washington, D. C., 882 pp." type="book" year="1956">Brown 1956</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |