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<document id="A8341D892956309642D6A8495D119108" ID-DOI="10.11646/zootaxa.5342.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="8323879" ID-ZooBank="A6D20DA1-1C12-454E-B422-D8457CF132FE" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1694069472299" checkinUser="plazi" docAuthor="Harvey, Mark S., Burger, Mieke A. A., Abrams, Kym M., Finston, Terrie L., Huey, Joel A. &amp; Perina, Giulia" docDate="2023" docId="010B87B4FFB00B63AFB0FF1EFB94F03B" docLanguage="en" docName="zootaxa.5342.1.1.pdf" docOrigin="Zootaxa 5342 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5342.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Indohya incomperta Harvey &amp; Burger 2023, n. sp." docType="treatment" docUuid="F216F6FE-397F-434D-97A2-FC254DE82C29" docUuidSource="ZooBank" docVersion="3" lastPageNumber="76" masterDocId="FD32FFCCFFFA0B28AF27FF89FFDAF444" masterDocTitle="The systematics of the pseudoscorpion genus Indohya (Pseudoscorpiones: Hyidae) in Australia" masterLastPageNumber="119" masterPageNumber="1" pageNumber="75" updateTime="1699635377361" updateUser="plazi">
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<mods:title id="8AB8A9410FC799CFA3680F58A5CDF62B">The systematics of the pseudoscorpion genus Indohya (Pseudoscorpiones: Hyidae) in Australia</mods:title>
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<mods:namePart id="1083F74263FE676B331557A284308A81">Harvey, Mark S.</mods:namePart>
<mods:affiliation id="D7573A91FC168AC5C7732551570D6BF3">Collections &amp; Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 &amp; Adjunct, School of Biological Sciences, University of Western Australia, Crawley, Western Australia 6009, Australia</mods:affiliation>
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<mods:namePart id="03D518EE72D39824839344362F927DBC">Burger, Mieke A. A.</mods:namePart>
<mods:affiliation id="1B98365B819FC00CDE35B0FA04C1D511">Collections &amp; Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238</mods:affiliation>
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<mods:namePart id="39DEA7904B7F6BC7B9202957DC56243E">Abrams, Kym M.</mods:namePart>
<mods:affiliation id="E2474EF4F7D0B2F6785DD96507FC726C">Collections &amp; Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 &amp; Present address: Department of Water and Environmental Regulation, Locked Bag 10, Joondalup DC, Western Australia 6919, Australia https: // orcid. org / 0000 - 0003 - 4543 - 9255</mods:affiliation>
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<mods:namePart id="DA2866C199307FA03DABC1034BD548EE">Finston, Terrie L.</mods:namePart>
<mods:affiliation id="00CDB507132471F75EC9638C8BF5E2DC">Adjunct, School of Biological Sciences, University of Western Australia, Crawley, Western Australia 6009, Australia &amp; Helix Molecular Solutions Pty Ltd, PO Box 155, Leederville, Western Australia 6153, Australia https: // orcid. org / 0000 - 0002 - 1750 - 4577</mods:affiliation>
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<mods:affiliation id="D697B5E677E04EB45BFC9919645CED0B">Collections &amp; Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 &amp; Adjunct, School of Biological Sciences, University of Western Australia, Crawley, Western Australia 6009, Australia &amp; Collections &amp; Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 &amp; Present address: Biologic Environmental Survey, East Perth, Western Australia 6004, Australia https: // orcid. org / 0000 - 0001 - 7108 - 0552</mods:affiliation>
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<mods:namePart id="C429451888618E98D3A70309359AC9DE">Perina, Giulia</mods:namePart>
<mods:affiliation id="D7C92CFE0A718862C6B9F98C921FC00F">Collections &amp; Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 &amp; Collections &amp; Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 &amp; Present address: Biologic Environmental Survey, East Perth, Western Australia 6004, Australia https: // orcid. org / 0000 - 0001 - 7108 - 0552 &amp; Collections &amp; Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 &amp; Subterranean Research and Groundwater Ecology (SuRGE) Group, Trace and Environmental DNA (TrEnD) Laboratory, School of Molecular and Life Sciences, Curtin University, Perth, Western Australia 6102, Australia https: // orcid. org / 0000 - 0002 - 0349 - 3803 Corresponding author</mods:affiliation>
<mods:nameIdentifier id="0F0027756321D27B3381380C8C78C527" type="email">mark.harvey@museum.wa.gov.au</mods:nameIdentifier>
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<mods:title id="9AF1FC64BFBC0BC262EAB0678146977E">Zootaxa</mods:title>
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<mods:date id="F275CFB69C5F2EE5B06B8FCFCC743AA6">2023</mods:date>
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<mods:number id="51096AA1FD14A544E75E871687D15C3F">2023-09-05</mods:number>
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<mods:number id="FE53D8C819E3FE07C49A1867876A946F">5342</mods:number>
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<treatment id="010B87B4FFB00B63AFB0FF1EFB94F03B" ID-DOI="http://doi.org/10.5281/zenodo.8323875" ID-Zenodo-Dep="8323875" LSID="urn:lsid:zoobank.org:act:F216F6FE-397F-434D-97A2-FC254DE82C29" httpUri="http://treatment.plazi.org/id/010B87B4FFB00B63AFB0FF1EFB94F03B" lastPageId="75" lastPageNumber="76" pageId="74" pageNumber="75">
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<taxonomicName id="4EA24D21FFB00B62AFB0FF1EFDABF4F5" authority="Harvey &amp; Burger" authorityName="Harvey &amp; Burger" authorityYear="2023" box="[151,625,151,178]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="incomperta" status="sp. nov.">
<emphasis id="BBD6EAB0FFB00B62AFB0FF1EFE57F4F5" bold="true" box="[151,397,151,178]" italics="true" pageId="74" pageNumber="75">Indohya incomperta</emphasis>
Harvey &amp; Burger
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,
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(
<figureCitation id="11992A27FFB00B62AFB8FF32FED1F491" box="[159,267,187,213]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="14.[151,250,1943,1967]" captionStartId-1="15.[151,250,1767,1791]" captionStartId-2="16.[151,250,1474,1498]" captionTargetBox-0="[212,1376,181,1919]" captionTargetBox-1="[151,1435,181,1744]" captionTargetBox-2="[151,1435,181,1449]" captionTargetId-0="figure-19@14.[212,1376,181,1919]" captionTargetId-1="figure-17@15.[151,1435,181,1744]" captionTargetId-2="figure-19@16.[151,1435,181,1449]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="16" captionText-0="FIGURE 1. Maximum likelihood phylogeny of Hyidae and other neobisioids, based on alignment of concatenated COI, 18S and 28S. Ultrafast bootstrap values are presented for nodes greater than 80%." captionText-1="FIGURE 2. Maximum likelihood phylogeny of Australian Indohya, using COI, 18S and 28S concatenated dataset. Bootstrap values are presented for all nodes. The genes amplified for each specimen are depicted with black squares in the gene matrix." captionText-2="FIGURE 3. Maximum likelihood phylogeny of Australian Indohya, using COI. Bootstrap values are presented for all nodes." figureDoi-0="http://doi.org/10.5281/zenodo.8323881" figureDoi-1="http://doi.org/10.5281/zenodo.8323885" figureDoi-2="http://doi.org/10.5281/zenodo.8323887" httpUri-0="https://zenodo.org/record/8323881/files/figure.png" httpUri-1="https://zenodo.org/record/8323885/files/figure.png" httpUri-2="https://zenodo.org/record/8323887/files/figure.png" pageId="74" pageNumber="75">Figs. 1–3</figureCitation>
,
<figureCitation id="11992A27FFB00B62AE30FF32FEFDF492" box="[279,295,187,214]" captionStart="FIGURE 5" captionStartId="21.[151,250,1919,1943]" captionTargetBox="[151,1436,181,1895]" captionTargetId="figure-17@21.[151,1436,181,1895]" captionTargetPageId="21" captionText="FIGURE 5. Maps of north-western Australia depicting recorded distributions of the subterranean species of Indohya from the Pilbara and Cape Range region." figureDoi="http://doi.org/10.5281/zenodo.8323893" httpUri="https://zenodo.org/record/8323893/files/figure.png" pageId="74" pageNumber="75">5</figureCitation>
,
<figureCitation id="11992A27FFB00B62AE14FF32FE8BF491" box="[307,337,187,213]" captionStart="FIGURE 37" captionStartId="74.[151,250,1944,1968]" captionTargetBox="[151,1436,1353,1921]" captionTargetId="figure-456@74.[151,1436,1353,1921]" captionTargetPageId="74" captionText="FIGURE 37. Indohya incomperta Harvey &amp; Burger, n. sp., holotype protonymph (WAM T149363): A, dorsal; B, ventral; C, cephalothorax, dorsal. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8323982" httpUri="https://zenodo.org/record/8323982/files/figure.png" pageId="74" pageNumber="75">37</figureCitation>
,
<figureCitation id="11992A27FFB00B62AE7AFF32FEA0F491" box="[349,378,187,213]" captionStart="FIGURE 38" captionStartId="75.[151,250,946,970]" captionTargetBox="[628,964,189,913]" captionTargetId="figure-17@75.[618,970,181,922]" captionTargetPageId="75" captionText="FIGURE 38. Indohya incomperta Harvey &amp; Burger, n. sp., holotype protonymph (WAM T149363): left chela, retrolateral. Scale line = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8323988" httpUri="https://zenodo.org/record/8323988/files/figure.png" pageId="74" pageNumber="75">38</figureCitation>
)
</paragraph>
<paragraph id="891D36A2FFB00B62AFB0FF56FC65F4BE" blockId="74.[151,959,151,250]" box="[151,959,223,250]" pageId="74" pageNumber="75">
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urn:lsid:zoobank.org:act:
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<emphasis id="BBD6EAB0FFB00B62AFB0FEAEFE5AF505" bold="true" box="[151,384,295,321]" pageId="74" pageNumber="75">Material examined.</emphasis>
<materialsCitation id="39CA3CFFFFB00B62AEA0FEAEFAFDF521" collectingDate="2019-05-06" collectionCode="WAM" collectorName="S. Callan" country="Australia" latitude="-23.121733" location="Newman" longLatPrecision="1" longitude="118.854034" municipality="West Angelas" pageId="74" pageNumber="75" specimenCode="T149363" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
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<typeStatus id="56198800FFB00B62AEA0FEAEFE34F505" box="[391,494,295,321]" pageId="74" pageNumber="75" type="holotype">Holotype</typeStatus>
protonymph
</emphasis>
.
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:
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<collectingRegion id="4B66F840FFB00B62AC14FEA1FC27F505" box="[819,1021,295,321]" country="Australia" name="Western Australia" pageId="74" pageNumber="75">Western Australia</collectingRegion>
:
</emphasis>
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, ca.
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WNW. of
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,
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, troglofauna scrape,
<collectorName id="24575374FFB00B62ACC8FEC2FB8FF521" box="[1007,1109,331,357]" pageId="74" pageNumber="75">S. Callan</collectorName>
(
<collectionCode id="EFB3AE67FFB00B62AB43FEC2FB71F521" box="[1124,1195,331,357]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="74" pageNumber="75" type="Museum">WAM</collectionCode>
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).
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</paragraph>
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<paragraph id="891D36A2FFB00B62AFE0FEE6FC14F5B1" blockId="74.[151,1437,295,1329]" pageId="74" pageNumber="75">
<emphasis id="BBD6EAB0FFB00B62AFE0FEE6FE98F5CD" bold="true" box="[199,322,367,393]" pageId="74" pageNumber="75">Diagnosis.</emphasis>
<taxonomicName id="4EA24D21FFB00B62AE6AFEE6FDEEF5CD" box="[333,564,367,393]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="incomperta">
<emphasis id="BBD6EAB0FFB00B62AE6AFEE6FDEEF5CD" box="[333,564,367,393]" italics="true" pageId="74" pageNumber="75">Indohya incomperta</emphasis>
</taxonomicName>
is a blind troglobite that differs from all other
<taxonomicName id="4EA24D21FFB00B62AB4EFEE6FAFBF5CD" box="[1129,1313,367,393]" pageId="74" pageNumber="75">
<emphasis id="BBD6EAB0FFB00B62AB4EFEE6FB1FF5CD" box="[1129,1221,367,393]" italics="true" pageId="74" pageNumber="75">Indohya</emphasis>
species
</taxonomicName>
for which sequence data are available by three synapomorphies in
<emphasis id="BBD6EAB0FFB00B62AC22FE1DFCEFF5EA" box="[773,821,404,430]" italics="true" pageId="74" pageNumber="75">COI</emphasis>
mtDNA: at base 375 there is a substitution to T; at base 377 there is a substitution to A; and at base 628 there is a substitution to C. The single sequenced specimen differs from all other sequenced specimens of
<taxonomicName id="4EA24D21FFB00B62AD69FE52FD70F5B1" box="[590,682,475,501]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="74" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis id="BBD6EAB0FFB00B62AD69FE52FD70F5B1" box="[590,682,475,501]" italics="true" pageId="74" pageNumber="75">Indohya</emphasis>
</taxonomicName>
by 16.3–25.2% (Table 2).
</paragraph>
</subSubSection>
<subSubSection id="C1B86529FFB00B62AFE0FE76FC00F7AA" pageId="74" pageNumber="75" type="description">
<paragraph id="891D36A2FFB00B62AFE0FE76FC33F65D" blockId="74.[151,1437,295,1329]" box="[199,1001,511,538]" pageId="74" pageNumber="75">
<emphasis id="BBD6EAB0FFB00B62AFE0FE76FDDDF65D" bold="true" box="[199,519,511,537]" pageId="74" pageNumber="75">Description (protonymph).</emphasis>
<emphasis id="BBD6EAB0FFB00B62AD29FE76FD0BF65D" box="[526,721,511,537]" italics="true" pageId="74" pageNumber="75">
Colour (
<figureCitation id="11992A27FFB00B62AD49FE76FD19F65D" box="[622,707,511,537]" captionStart="FIGURE 37" captionStartId="74.[151,250,1944,1968]" captionTargetBox="[151,1436,1353,1921]" captionTargetId="figure-456@74.[151,1436,1353,1921]" captionTargetPageId="74" captionText="FIGURE 37. Indohya incomperta Harvey &amp; Burger, n. sp., holotype protonymph (WAM T149363): A, dorsal; B, ventral; C, cephalothorax, dorsal. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8323982" httpUri="https://zenodo.org/record/8323982/files/figure.png" pageId="74" pageNumber="75">Fig. 37</figureCitation>
):
</emphasis>
very pale, yellow-white.
</paragraph>
<paragraph id="891D36A2FFB00B62AFE0FDAAFC82F67A" blockId="74.[151,1437,295,1329]" box="[199,856,547,574]" pageId="74" pageNumber="75">
<emphasis id="BBD6EAB0FFB00B62AFE0FDAAFEE4F679" box="[199,318,547,573]" italics="true" pageId="74" pageNumber="75">Chelicera:</emphasis>
hand with 4 setae, movable finger without seta.
</paragraph>
<paragraph id="891D36A2FFB00B62AFE0FDCEFB27F689" blockId="74.[151,1437,295,1329]" pageId="74" pageNumber="75">
<emphasis id="BBD6EAB0FFB00B62AFE0FDCEFEEFF625" box="[199,309,583,609]" italics="true" pageId="74" pageNumber="75">Pedipalp:</emphasis>
trochanter 2.06 ×, femur 5.41 ×, patella 3.11 ×, chela (with pedicel) 4.45 ×, chela (without pedicel) 4.23 ×, hand (without pedicel) 1.68 × longer than broad, movable finger 1.40 × longer than hand (without pedicel). Fixed finger with 3 trichobothria (
<figureCitation id="11992A27FFB00B62AD3DFD06FDB4F6ED" box="[538,622,655,681]" captionStart="FIGURE 38" captionStartId="75.[151,250,946,970]" captionTargetBox="[628,964,189,913]" captionTargetId="figure-17@75.[618,970,181,922]" captionTargetPageId="75" captionText="FIGURE 38. Indohya incomperta Harvey &amp; Burger, n. sp., holotype protonymph (WAM T149363): left chela, retrolateral. Scale line = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8323988" httpUri="https://zenodo.org/record/8323988/files/figure.png" pageId="74" pageNumber="75">Fig. 38</figureCitation>
),
<emphasis id="BBD6EAB0FFB00B62ADA4FD06FD47F6ED" box="[643,669,655,681]" italics="true" pageId="74" pageNumber="75">eb</emphasis>
,
<emphasis id="BBD6EAB0FFB00B62AD8BFD19FD1CF6ED" box="[684,710,656,681]" italics="true" pageId="74" pageNumber="75">ist</emphasis>
and
<emphasis id="BBD6EAB0FFB00B62ADD8FD19FCC9F6ED" box="[767,787,656,681]" italics="true" pageId="74" pageNumber="75">et</emphasis>
present; movable finger with 1 trichobothrium,
<emphasis id="BBD6EAB0FFB00B62AA13FD19FAE6F6ED" box="[1332,1340,656,681]" italics="true" pageId="74" pageNumber="75">t</emphasis>
present; fixed chelal finger with 34 low, juxtadentate teeth and quadrate; movable chelal finger with 11 teeth.
</paragraph>
<paragraph id="891D36A2FFB00B62AFE0FD5EFE17F751" blockId="74.[151,1437,295,1329]" pageId="74" pageNumber="75">
<emphasis id="BBD6EAB0FFB00B62AFE0FD5EFEA2F6B5" box="[199,376,727,753]" italics="true" pageId="74" pageNumber="75">Cephalothorax:</emphasis>
0.83 × longer than broad; with 12 setae arranged 4: 0: 4: 2: 2; without furrows; eyes absent. Coxal chaetotaxy 1: 1: 1: 1.
</paragraph>
<paragraph id="891D36A2FFB00B62AFE0FCA9FD8AF77D" blockId="74.[151,1437,295,1329]" box="[199,592,799,826]" pageId="74" pageNumber="75">
<emphasis id="BBD6EAB0FFB00B62AFE0FCA9FEDCF77D" box="[199,262,800,825]" italics="true" pageId="74" pageNumber="75">Legs:</emphasis>
metatarsi and tarsi not fused.
</paragraph>
<paragraph id="891D36A2FFB00B62AFE0FCCAFBC8F7C5" blockId="74.[151,1437,295,1329]" pageId="74" pageNumber="75">
<emphasis id="BBD6EAB0FFB00B62AFE0FCCAFEE0F719" box="[199,314,835,861]" italics="true" pageId="74" pageNumber="75">Abdomen:</emphasis>
tergal chaetotaxy 4: 4: 4: 4: 4: 4: 4: 4: 4: 4 (including 2 tactile setae): 4 (including 2 tactile setae): 2; sternal chaetotaxy 0: (0) 2 (0): (1) 2 (1): 4: 4: 4: 4: 4: 4: 2 (both tactile setae): 2.
</paragraph>
<paragraph id="891D36A2FFB00B62AFE0FC05FC00F7AA" blockId="74.[151,1437,295,1329]" pageId="74" pageNumber="75">
<emphasis id="BBD6EAB0FFB00B62AFE0FC05FE41F7E1" box="[199,411,907,933]" italics="true" pageId="74" pageNumber="75">Dimensions (mm):</emphasis>
<materialsCitation id="39CA3CFFFFB00B62AE80FC02FA93F7E2" box="[423,1353,907,934]" collectionCode="WAM" latitude="0.175" longLatPrecision="78" longitude="0.085" pageId="74" pageNumber="75" specimenCode="T149363" specimenCount="1" typeStatus="holotype">
<typeStatus id="56198800FFB00B62AE80FC02FDD3F7E1" box="[423,521,907,933]" pageId="74" pageNumber="75" type="holotype">holotype</typeStatus>
(
<collectionCode id="EFB3AE67FFB00B62AD3BFC02FDB9F7E1" box="[540,611,907,933]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="74" pageNumber="75" type="Museum">WAM</collectionCode>
<specimenCode id="D9049ED9FFB00B62AD49FC02FD02F7E1" box="[622,728,907,933]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="74" pageNumber="75">T149363</specimenCode>
). Body length 1.25. Pedipalp: trochanter
<geoCoordinate id="EC965065FFB00B62ABE6FC02FADBF7E2" box="[1217,1281,907,934]" degrees="0.175" direction="north" orientation="latitude" pageId="74" pageNumber="75" precision="55" value="0.175">0.175</geoCoordinate>
/
<geoCoordinate id="EC965065FFB00B62AA2EFC02FA93F7E2" box="[1289,1353,907,934]" degrees="0.085" direction="east" orientation="longitude" pageId="74" pageNumber="75" precision="55" value="0.085">0.085</geoCoordinate>
</materialsCitation>
, femur
<geoCoordinate id="EC965065FFB00B62AFB0FC26FF02F78E" box="[151,216,943,970]" degrees="0.460" direction="north" orientation="latitude" pageId="74" pageNumber="75" precision="55" value="0.46">0.460</geoCoordinate>
/
<geoCoordinate id="EC965065FFB00B62AFC7FC26FEC5F78E" box="[224,287,943,970]" degrees="0.085" direction="east" orientation="longitude" pageId="74" pageNumber="75" precision="55" value="0.085">0.085</geoCoordinate>
, patella
<geoCoordinate id="EC965065FFB00B62AEA5FC26FE18F78E" box="[386,450,943,970]" degrees="0.295" direction="north" orientation="latitude" pageId="74" pageNumber="75" precision="55" value="0.295">0.295</geoCoordinate>
/
<geoCoordinate id="EC965065FFB00B62AEECFC26FDD1F78E" box="[459,523,943,970]" degrees="0.095" direction="east" orientation="longitude" pageId="74" pageNumber="75" precision="55" value="0.095">0.095</geoCoordinate>
, chela (with pedicel)
<geoCoordinate id="EC965065FFB00B62AC21FC26FC9DF78E" box="[774,839,943,970]" degrees="0.690" direction="north" orientation="latitude" pageId="74" pageNumber="75" precision="55" value="0.69">0.690</geoCoordinate>
/
<geoCoordinate id="EC965065FFB00B62AC68FC26FC54F78E" box="[847,910,943,970]" degrees="0.155" direction="east" orientation="longitude" pageId="74" pageNumber="75" precision="55" value="0.155">0.155</geoCoordinate>
, chela (without pedicel) 0.655, hand (without pedicel) length 0.260, movable finger length 0.365. Carapace
<geoCoordinate id="EC965065FFB00B62AC6AFC5AFC57F7AA" box="[845,909,979,1006]" degrees="0.385" direction="north" orientation="latitude" pageId="74" pageNumber="75" precision="55" value="0.385">0.385</geoCoordinate>
/
<geoCoordinate id="EC965065FFB00B62ACB1FC5AFC0CF7AA" box="[918,982,979,1006]" degrees="0.465" direction="east" orientation="longitude" pageId="74" pageNumber="75" precision="55" value="0.465">0.465</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection id="C1B86529FFB00B62AFE0FC7EFF33F175" pageId="74" pageNumber="75" type="distribution">
<paragraph id="891D36A2FFB00B62AFE0FC7EFD9CF039" blockId="74.[151,1437,295,1329]" pageId="74" pageNumber="75">
<emphasis id="BBD6EAB0FFB00B62AFE0FC7EFDDCF055" bold="true" box="[199,518,1015,1041]" pageId="74" pageNumber="75">Distribution and remarks.</emphasis>
<taxonomicName id="4EA24D21FFB00B62AD37FC7EFD2FF055" box="[528,757,1015,1041]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="incomperta">
<emphasis id="BBD6EAB0FFB00B62AD37FC7EFD2FF055" box="[528,757,1015,1041]" italics="true" pageId="74" pageNumber="75">Indohya incomperta</emphasis>
</taxonomicName>
has been recorded in subterranean habitats within a single site at West Angelas mine site WNW. of Newman (
<figureCitation id="11992A27FFB00B62ADF2FB92FCC0F072" box="[725,794,1051,1078]" captionStart="FIGURE 5" captionStartId="21.[151,250,1919,1943]" captionTargetBox="[151,1436,181,1895]" captionTargetId="figure-17@21.[151,1436,181,1895]" captionTargetPageId="21" captionText="FIGURE 5. Maps of north-western Australia depicting recorded distributions of the subterranean species of Indohya from the Pilbara and Cape Range region." figureDoi="http://doi.org/10.5281/zenodo.8323893" httpUri="https://zenodo.org/record/8323893/files/figure.png" pageId="74" pageNumber="75">Fig. 5</figureCitation>
). The sequence data available for this nymphal specimen demonstrates that it belongs to a distinct species (
<figureCitation id="11992A27FFB00B62ADD2FBB6FCBDF01D" box="[757,871,1087,1113]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="14.[151,250,1943,1967]" captionStartId-1="15.[151,250,1767,1791]" captionStartId-2="16.[151,250,1474,1498]" captionTargetBox-0="[212,1376,181,1919]" captionTargetBox-1="[151,1435,181,1744]" captionTargetBox-2="[151,1435,181,1449]" captionTargetId-0="figure-19@14.[212,1376,181,1919]" captionTargetId-1="figure-17@15.[151,1435,181,1744]" captionTargetId-2="figure-19@16.[151,1435,181,1449]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="16" captionText-0="FIGURE 1. Maximum likelihood phylogeny of Hyidae and other neobisioids, based on alignment of concatenated COI, 18S and 28S. Ultrafast bootstrap values are presented for nodes greater than 80%." captionText-1="FIGURE 2. Maximum likelihood phylogeny of Australian Indohya, using COI, 18S and 28S concatenated dataset. Bootstrap values are presented for all nodes. The genes amplified for each specimen are depicted with black squares in the gene matrix." captionText-2="FIGURE 3. Maximum likelihood phylogeny of Australian Indohya, using COI. Bootstrap values are presented for all nodes." figureDoi-0="http://doi.org/10.5281/zenodo.8323881" figureDoi-1="http://doi.org/10.5281/zenodo.8323885" figureDoi-2="http://doi.org/10.5281/zenodo.8323887" httpUri-0="https://zenodo.org/record/8323881/files/figure.png" httpUri-1="https://zenodo.org/record/8323885/files/figure.png" httpUri-2="https://zenodo.org/record/8323887/files/figure.png" pageId="74" pageNumber="75">Figs. 1–3</figureCitation>
). The species was previously known by WAM identification code
<taxonomicName id="4EA24D21FFB00B62AE48FBEAFE11F039" authority="PSE" authorityName="PSE" box="[367,459,1123,1149]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="74" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis id="BBD6EAB0FFB00B62AE48FBEAFE11F039" box="[367,459,1123,1149]" italics="true" pageId="74" pageNumber="75">Indohya</emphasis>
</taxonomicName>
‘PSE147’.
</paragraph>
<paragraph id="891D36A2FFB00B62AFE0FB0EFF33F175" blockId="74.[151,1437,295,1329]" pageId="74" pageNumber="75">
<emphasis id="BBD6EAB0FFB00B62AFE0FB0EFE2AF0E5" bold="true" box="[199,496,1159,1185]" pageId="74" pageNumber="75">Conservation assessment</emphasis>
.
<taxonomicName id="4EA24D21FFB00B62AEDAFB0EFD05F0E5" box="[509,735,1159,1185]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="incomperta">
<emphasis id="BBD6EAB0FFB00B62AEDAFB0EFD05F0E5" box="[509,735,1159,1185]" italics="true" pageId="74" pageNumber="75">Indohya incomperta</emphasis>
</taxonomicName>
occurs in subterranean biotopes at West Angelas mine site. The total extent of the subterranean habitat is unknown, and the surface terrain is not heavily dissected or constrained by mesas. It is therefore likely that it’s distribution may extend further into the local area. The species does not appear to meet the status of threatened using the criteria developed by the International Union for Conservation of Nature (2012).
</paragraph>
</subSubSection>
<caption id="DDDD662AFFB00B62AFB0F811FD81F390" ID-DOI="http://doi.org/10.5281/zenodo.8323982" ID-Zenodo-Dep="8323982" httpUri="https://zenodo.org/record/8323982/files/figure.png" pageId="74" pageNumber="75" startId="74.[151,250,1944,1968]" targetBox="[151,1436,1353,1921]" targetPageId="74" targetType="figure">
<paragraph id="891D36A2FFB00B62AFB0F811FD81F390" blockId="74.[151,1436,1944,2005]" pageId="74" pageNumber="75">
<emphasis id="BBD6EAB0FFB00B62AFB0F811FEFBF3F4" bold="true" box="[151,289,1944,1968]" pageId="74" pageNumber="75">FIGURE 37.</emphasis>
<taxonomicName id="4EA24D21FFB00B62AE0FF810FD68F3F5" authority="Harvey &amp; Burger" authorityName="Harvey &amp; Burger" authorityYear="2023" box="[296,690,1945,1969]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="incomperta" status="sp. nov.">
<emphasis id="BBD6EAB0FFB00B62AE0FF810FE2FF3F4" box="[296,501,1945,1969]" italics="true" pageId="74" pageNumber="75">Indohya incomperta</emphasis>
Harvey &amp; Burger
</taxonomicName>
,
<emphasis id="BBD6EAB0FFB00B62AD9AF810FD2DF3F5" bold="true" box="[701,759,1945,1969]" pageId="74" pageNumber="75">
<taxonomicNameLabel id="A0E557CBFFB00B62AD9AF810FD2DF3F5" box="[701,759,1945,1969]" pageId="74" pageNumber="75" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype protonymph (WAM T149363): A, dorsal; B, ventral; C, cephalothorax, dorsal. Scale lines = 1.0 mm.
</paragraph>
</caption>
<caption id="DDDD662AFFB10B63AFB0FC3BFEB0F7AA" ID-DOI="http://doi.org/10.5281/zenodo.8323988" ID-Zenodo-Dep="8323988" httpUri="https://zenodo.org/record/8323988/files/figure.png" pageId="75" pageNumber="76" startId="75.[151,250,946,970]" targetBox="[628,964,189,913]" targetPageId="75" targetType="figure">
<paragraph id="891D36A2FFB10B63AFB0FC3BFEB0F7AA" blockId="75.[151,1437,946,1006]" pageId="75" pageNumber="76">
<emphasis id="BBD6EAB0FFB10B63AFB0FC3BFEF8F78E" bold="true" box="[151,290,946,970]" pageId="75" pageNumber="76">FIGURE 38.</emphasis>
<taxonomicName id="4EA24D21FFB10B63AE0CFC3BFD60F78F" authority="Harvey &amp; Burger" authorityName="Harvey &amp; Burger" authorityYear="2023" box="[299,698,946,971]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="incomperta" status="sp. nov.">
<emphasis id="BBD6EAB0FFB10B63AE0CFC3BFE23F78E" box="[299,505,946,970]" italics="true" pageId="75" pageNumber="76">Indohya incomperta</emphasis>
Harvey &amp; Burger
</taxonomicName>
,
<emphasis id="BBD6EAB0FFB10B63ADE1FC3BFCDBF78E" bold="true" box="[710,769,946,970]" pageId="75" pageNumber="76">
<taxonomicNameLabel id="A0E557CBFFB10B63ADE1FC3BFCDBF78E" box="[710,769,946,970]" pageId="75" pageNumber="76" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype protonymph (WAM T149363): left chela, retrolateral. Scale line = 0.2 mm.
</paragraph>
</caption>
<subSubSection id="C1B86529FFB10B63AFE0FBC8FB94F03B" pageId="75" pageNumber="76" type="etymology">
<paragraph id="891D36A2FFB10B63AFE0FBC8FB94F03B" blockId="75.[151,1436,1088,1151]" pageId="75" pageNumber="76">
<emphasis id="BBD6EAB0FFB10B63AFE0FBC8FE94F01F" bold="true" box="[199,334,1089,1115]" pageId="75" pageNumber="76">Etymology.</emphasis>
The specific epithet refers to the use of molecular sequence data to establish that this nymphal specimen represents a distinct species (
<emphasis id="BBD6EAB0FFB10B63AD69FBECFD02F03A" box="[590,728,1125,1150]" italics="true" pageId="75" pageNumber="76">incompertus</emphasis>
, Latin, unknown) (
<bibRefCitation id="ED334B53FFB10B63AC97FBECFB9BF03B" author="Brown, R. W." box="[944,1089,1124,1151]" pageId="75" pageNumber="76" refId="ref82031" refString="Brown, R. W. (1956) Composition of scientific words. Revised Edition. Smithsonian Institution Press, Washington, D. C., 882 pp." type="book" year="1956">Brown 1956</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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