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<mods:title id="5A299EFD75E2F7225B6186899DB3C870">The systematics of the pseudoscorpion genus Indohya (Pseudoscorpiones: Hyidae) in Australia</mods:title>
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<mods:affiliation id="3BD1785E1A57592C78BDE3C58B221494">Collections &amp; Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 &amp; Adjunct, School of Biological Sciences, University of Western Australia, Crawley, Western Australia 6009, Australia &amp; Collections &amp; Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 &amp; Present address: Biologic Environmental Survey, East Perth, Western Australia 6004, Australia https: // orcid. org / 0000 - 0001 - 7108 - 0552</mods:affiliation>
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<mods:affiliation id="B3F3D4BBEF4525C54FF3FD24D86CAD95">Collections &amp; Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 &amp; Collections &amp; Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 &amp; Present address: Biologic Environmental Survey, East Perth, Western Australia 6004, Australia https: // orcid. org / 0000 - 0001 - 7108 - 0552 &amp; Collections &amp; Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 &amp; Subterranean Research and Groundwater Ecology (SuRGE) Group, Trace and Environmental DNA (TrEnD) Laboratory, School of Molecular and Life Sciences, Curtin University, Perth, Western Australia 6102, Australia https: // orcid. org / 0000 - 0002 - 0349 - 3803 Corresponding author</mods:affiliation>
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<emphasis id="BBD6EAB0FF880B5AAFB0FF1EFEB3F4F6" bold="true" box="[151,361,151,178]" italics="true" pageId="114" pageNumber="115">Indohya typhlops</emphasis>
Harvey, 1993
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(
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, 
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)
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<emphasis id="BBD6EAB0FF880B5AAFB0FEAAFE9DF57F" box="[151,327,291,315]" italics="true" pageId="114" pageNumber="115">Indohya typhlops</emphasis>
Harvey 1993: 27
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, figs. 3, 81–88; 
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, figs. 5–9, 15.
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:
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WSW. of 
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staff (
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<emphasis id="BBD6EAB0FF880B5AAFE0FE5FFEB7F5B4" box="[199,365,470,496]" italics="true" pageId="114" pageNumber="115">Other material</emphasis>
. 
<materialsCitation id="39CA3CFFFF880B5AAE5FFE5FFD0EF67D" collectingDate="2022-07-27" collectionCode="WAM" collectorName="M. S. Harvey &amp; J. D. Wilson &amp; L. Kirkendale" country="Australia" latitude="-16.372223" location="Charnley River-Artesian Range Wildlife Sanctuary" longLatPrecision="21" longitude="125.208885" pageId="114" pageNumber="115" specimenCode="T158276, T158277, T158592, T158595, T158598" specimenCount="4" specimenCount-female="4" stateProvince="Western Australia">
<collectingCountry id="F1B57632FF880B5AAE5FFE5FFDC1F5B4" box="[376,539,470,496]" name="Australia" pageId="114" pageNumber="115">AUSTRALIA</collectingCountry>
: 
<emphasis id="BBD6EAB0FF880B5AAD04FE5EFD29F5B4" box="[547,755,470,496]" italics="true" pageId="114" pageNumber="115">
<collectingRegion id="4B66F840FF880B5AAD04FE5EFD36F5B4" box="[547,748,470,496]" country="Australia" name="Western Australia" pageId="114" pageNumber="115">Western Australia</collectingRegion>
:
</emphasis>
4 Ô, 
<specimenCount id="9FA4FD2BFF880B5AAC0AFE5FFC8CF5B4" box="[813,854,470,496]" pageId="114" pageNumber="115" type="female">4 ♀</specimenCount>
, 
<location id="8C7D6079FF880B5AAC45FE5FFA42F5B4" LSID="urn:lsid:plazi:treatment:010B87B4FF880B5AAFB0FF1EFADDF010:8C7D6079FF880B5AAC45FE5FFA42F5B4" box="[866,1432,470,496]" country="Australia" latitude="-16.372223" longLatPrecision="21" longitude="125.208885" name="Charnley River-Artesian Range Wildlife Sanctuary" pageId="114" pageNumber="115" stateProvince="Western Australia">Charnley River-Artesian Range Wildlife Sanctuary</location>
, 
<geoCoordinate id="EC965065FF880B5AAFB0FE72FEC1F652" box="[151,283,507,534]" degrees="16" direction="south" minutes="22" orientation="latitude" pageId="114" pageNumber="115" precision="15" seconds="20" value="-16.372223">16°22′20”S</geoCoordinate>
, 
<geoCoordinate id="EC965065FF880B5AAE00FE72FE60F652" box="[295,442,507,534]" degrees="125" direction="east" minutes="12" orientation="longitude" pageId="114" pageNumber="115" precision="15" seconds="32" value="125.208885">125°12′32”E</geoCoordinate>
, 
<date id="FD1C1062FF880B5AAEE1FE73FD80F650" box="[454,602,506,532]" pageId="114" pageNumber="115" value="2022-07-27">
<collectingDate id="ED58E98AFF880B5AAEE1FE73FD80F650" box="[454,602,506,532]" pageId="114" pageNumber="115" value="2022-07-27">27 July 2022</collectingDate>
</date>
, under rock, vine thicket, 
<collectorName id="24575374FF880B5AAC5AFE73FBD5F650" box="[893,1039,506,532]" pageId="114" pageNumber="115">M.S. Harvey</collectorName>
, 
<collectorName id="24575374FF880B5AAB3DFE73FB7BF650" box="[1050,1185,506,532]" pageId="114" pageNumber="115">J.D. Wilson</collectorName>
, 
<collectorName id="24575374FF880B5AAB8BFE73FA9DF650" box="[1196,1351,506,532]" pageId="114" pageNumber="115">L. Kirkendale</collectorName>
(
<collectionCode id="EFB3AE67FF880B5AAA71FE73FA47F650" box="[1366,1437,506,532]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="114" pageNumber="115" type="Museum">WAM</collectionCode>
<specimenCode id="D9049ED9FF880B5AAFB0FD97FF25F67D" box="[151,255,542,569]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="114" pageNumber="115">T158276</specimenCode>
, 
<specimenCode id="D9049ED9FF880B5AAE2DFD97FEABF67D" box="[266,369,542,569]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="114" pageNumber="115">T158277</specimenCode>
, 
<specimenCode id="D9049ED9FF880B5AAE5BFD97FE3FF67D" box="[380,485,542,569]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="114" pageNumber="115">T158592</specimenCode>
, 
<specimenCode id="D9049ED9FF880B5AAEC8FD97FD8DF67D" box="[495,599,542,569]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="114" pageNumber="115">T158595</specimenCode>
, 
<specimenCode id="D9049ED9FF880B5AAD45FD97FD11F67D" box="[610,715,542,569]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="114" pageNumber="115">T158598</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C1B86529FF880B5AAFE0FDC8FE48F738" pageId="114" pageNumber="115" type="diagnosis">
<paragraph id="891D36A2FF880B5AAFE0FDC8FD30F6A8" blockId="114.[151,1437,361,1108]" pageId="114" pageNumber="115">
<emphasis id="BBD6EAB0FF880B5AAFE0FDC8FE98F61F" bold="true" box="[199,322,577,603]" pageId="114" pageNumber="115">Diagnosis.</emphasis>
<taxonomicName id="4EA24D21FF880B5AAE6DFDCBFDD7F618" authorityName="Harvey WAMT" authorityYear="1993" box="[330,525,578,604]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="typhlops">
<emphasis id="BBD6EAB0FF880B5AAE6DFDCBFDD7F618" box="[330,525,578,604]" italics="true" pageId="114" pageNumber="115">Indohya typhlops</emphasis>
</taxonomicName>
has 16 setae on the carapace setae and lacks eyes, thus resembling 
<taxonomicName id="4EA24D21FF880B5AAA37FDCAFAA7F618" authorityName="Harvey &amp; Burger &amp; Abrams &amp; Finston &amp; Huey &amp; Perina" authorityYear="2023" box="[1296,1405,579,604]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="currani">
<emphasis id="BBD6EAB0FF880B5AAA37FDCAFAA7F618" box="[1296,1405,579,604]" italics="true" pageId="114" pageNumber="115">I. currani</emphasis>
</taxonomicName>
, 
<taxonomicName id="4EA24D21FF880B5AAAABFDCAFF3FF6C4" authorityName="Harvey and Volschenk" authorityYear="2007" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="gollum">
<emphasis id="BBD6EAB0FF880B5AAAABFDCAFF3FF6C4" italics="true" pageId="114" pageNumber="115">I. gollum</emphasis>
</taxonomicName>
and 
<taxonomicName id="4EA24D21FF880B5AAE3DFDEEFE6BF6C4" authorityName="Harvey and Volschenk" authorityYear="2007" box="[282,433,615,640]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="napierensis">
<emphasis id="BBD6EAB0FF880B5AAE3DFDEEFE6BF6C4" box="[282,433,615,640]" italics="true" pageId="114" pageNumber="115">I. napierensis</emphasis>
</taxonomicName>
. It differs from 
<taxonomicName id="4EA24D21FF880B5AAD46FDEEFD1FF6C4" authorityName="Harvey and Volschenk" authorityYear="2007" box="[609,709,614,640]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="gollum">
<emphasis id="BBD6EAB0FF880B5AAD46FDEEFD1FF6C4" box="[609,709,614,640]" italics="true" pageId="114" pageNumber="115">I. gollum</emphasis>
</taxonomicName>
and 
<taxonomicName id="4EA24D21FF880B5AADDDFDEEFC4BF6C4" authorityName="Harvey and Volschenk" authorityYear="2007" box="[762,913,615,640]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="napierensis">
<emphasis id="BBD6EAB0FF880B5AADDDFDEEFC4BF6C4" box="[762,913,615,640]" italics="true" pageId="114" pageNumber="115">I. napierensis</emphasis>
</taxonomicName>
in being smaller [e.g. chela (with pedicel) 0.77 (
<specimenCount id="9FA4FD2BFF880B5AAF85FD03FF6EF6E0" box="[162,180,650,676]" pageId="114" pageNumber="115" type="female">♀</specimenCount>
) mm vs. greater than 
<quantity id="4E5A9B47FF880B5AAE82FD03FE20F6E0" box="[421,506,650,676]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="114" pageNumber="115" unit="mm" value="1.8">1.8 mm</quantity>
in 
<taxonomicName id="4EA24D21FF880B5AAD3CFD02FDA5F6E0" authorityName="Harvey and Volschenk" authorityYear="2007" box="[539,639,650,676]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="gollum">
<emphasis id="BBD6EAB0FF880B5AAD3CFD02FDA5F6E0" box="[539,639,650,676]" italics="true" pageId="114" pageNumber="115">I. gollum</emphasis>
</taxonomicName>
and 
<taxonomicName id="4EA24D21FF880B5AAD95FD02FC93F6E0" authorityName="Harvey and Volschenk" authorityYear="2007" box="[690,841,651,676]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="napierensis">
<emphasis id="BBD6EAB0FF880B5AAD95FD02FC93F6E0" box="[690,841,651,676]" italics="true" pageId="114" pageNumber="115">I. napierensis</emphasis>
</taxonomicName>
], and from 
<taxonomicName id="4EA24D21FF880B5AACEEFD02FBE9F6E0" authorityName="Harvey &amp; Burger &amp; Abrams &amp; Finston &amp; Huey &amp; Perina" authorityYear="2023" box="[969,1075,651,676]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="currani">
<emphasis id="BBD6EAB0FF880B5AACEEFD02FBE9F6E0" box="[969,1075,651,676]" italics="true" pageId="114" pageNumber="115">I. currani</emphasis>
</taxonomicName>
by its broader chela [chela (with pedicel) 3.21 (
<specimenCount id="9FA4FD2BFF880B5AAE1AFD27FE95F68C" box="[317,335,686,712]" pageId="114" pageNumber="115" type="female">♀</specimenCount>
) × longer than broad vs. 3.98 (
<specimenCount id="9FA4FD2BFF880B5AAD94FD27FD1FF68C" box="[691,709,686,712]" pageId="114" pageNumber="115" type="female">♀</specimenCount>
) × longer than broad in 
<taxonomicName id="4EA24D21FF880B5AACFBFD26FB92F68C" authorityName="Harvey &amp; Burger &amp; Abrams &amp; Finston &amp; Huey &amp; Perina" authorityYear="2023" box="[988,1096,687,712]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="currani">
<emphasis id="BBD6EAB0FF880B5AACFBFD26FB92F68C" box="[988,1096,687,712]" italics="true" pageId="114" pageNumber="115">I. currani</emphasis>
</taxonomicName>
] and by having more teeth on the fixed chelal finger [41 teeth (
<specimenCount id="9FA4FD2BFF880B5AAD2EFD5BFDC6F6A8" box="[521,540,722,748]" pageId="114" pageNumber="115" type="female">♀</specimenCount>
), vs. 46 teeth (
<specimenCount id="9FA4FD2BFF880B5AADE0FD5BFD00F6A8" box="[711,730,722,748]" pageId="114" pageNumber="115" type="female">♀</specimenCount>
).
</paragraph>
<paragraph id="891D36A2FF880B5AAFE0FD7FFE48F738" blockId="114.[151,1437,361,1108]" pageId="114" pageNumber="115">
It also differs from all other 
<taxonomicName id="4EA24D21FF880B5AAD2AFD7FFD18F754" box="[525,706,758,784]" pageId="114" pageNumber="115">
<emphasis id="BBD6EAB0FF880B5AAD2AFD7FFDB3F754" box="[525,617,758,784]" italics="true" pageId="114" pageNumber="115">Indohya</emphasis>
species
</taxonomicName>
for which sequence data are available by three synapomorphies in 
<emphasis id="BBD6EAB0FF880B5AAF91FC93FF3CF770" box="[182,230,794,820]" italics="true" pageId="114" pageNumber="115">COI</emphasis>
mtDNA: at base 172 there is a substitution to G; at base 286 there is a substitution to T; and at base 492 there is a substitution to A. The two sequenced specimens differ from all other sequenced specimens of 
<taxonomicName id="4EA24D21FF880B5AAA39FCB7FAA0F71C" box="[1310,1402,830,856]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis id="BBD6EAB0FF880B5AAA39FCB7FAA0F71C" box="[1310,1402,830,856]" italics="true" pageId="114" pageNumber="115">Indohya</emphasis>
</taxonomicName>
by 18.8–26.4% (Table 2).
</paragraph>
</subSubSection>
<subSubSection id="C1B86529FF880B5AAFE0FC0CFD48F7E4" box="[199,658,901,928]" pageId="114" pageNumber="115" type="description">
<paragraph id="891D36A2FF880B5AAFE0FC0CFD48F7E4" blockId="114.[151,1437,361,1108]" box="[199,658,901,928]" pageId="114" pageNumber="115">
<emphasis id="BBD6EAB0FF880B5AAFE0FC0CFE6AF7DB" bold="true" box="[199,432,901,927]" pageId="114" pageNumber="115">Description (adult).</emphasis>
See Harvey (1993).
</paragraph>
</subSubSection>
<subSubSection id="C1B86529FF880B5AAFE0FC20FADDF010" pageId="114" pageNumber="115" type="distribution">
<paragraph id="891D36A2FF880B5AAFE0FC20FBDAF7AC" blockId="114.[151,1437,361,1108]" pageId="114" pageNumber="115">
<emphasis id="BBD6EAB0FF880B5AAFE0FC20FDD8F780" bold="true" box="[199,514,937,964]" pageId="114" pageNumber="115">Distribution and remarks.</emphasis>
<taxonomicName id="4EA24D21FF880B5AAD2DFC23FD17F780" authorityName="Harvey WAMT" authorityYear="1993" box="[522,717,938,964]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="typhlops">
<emphasis id="BBD6EAB0FF880B5AAD2DFC23FD17F780" box="[522,717,938,964]" italics="true" pageId="114" pageNumber="115">Indohya typhlops</emphasis>
</taxonomicName>
has been recorded from a single rainforest site on the Charnley River (
<figureCitation id="11992A27FF880B5AAFC2FC47FEE5F7AC" box="[229,319,974,1000]" captionStart="FIGURE 6" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[200,1392,188,1917]" captionTargetId="figure-19@22.[194,1394,181,1920]" captionTargetPageId="22" captionText="FIGURE 6. Maps of north-western Australia depicting recorded distributions of the three epigean species of Indohya from the Kimberley:A, epigean species; B, troglobitic species." figureDoi="http://doi.org/10.5281/zenodo.8323897" httpUri="https://zenodo.org/record/8323897/files/figure.png" pageId="114" pageNumber="115">Fig. 6A</figureCitation>
). The site is situated within the Northern Kimberley bioregion.
</paragraph>
<paragraph id="891D36A2FF880B5AAFE0FC78FADDF010" blockId="114.[151,1437,361,1108]" pageId="114" pageNumber="115">
<emphasis id="BBD6EAB0FF880B5AAFE0FC78FE2EF048" bold="true" box="[199,500,1009,1036]" pageId="114" pageNumber="115">Conservation assessment</emphasis>
. Although 
<taxonomicName id="4EA24D21FF880B5AAD51FC7BFCE1F048" authorityName="Harvey WAMT" authorityYear="1993" box="[630,827,1010,1036]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="typhlops">
<emphasis id="BBD6EAB0FF880B5AAD51FC7BFCE1F048" box="[630,827,1010,1036]" italics="true" pageId="114" pageNumber="115">Indohya typhlops</emphasis>
</taxonomicName>
has only been collected at a single location, there is no reason to believe that it is not more widely distributed in the region. The species does not meet the status of threatened using the criteria developed by the International Union for Conservation of Nature (2012).
</paragraph>
</subSubSection>
</treatment>
</document>